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<document id="E74E82D86D8811DC578E134CAEA3A335" ID-CLB-Dataset="5429" ID-DOI="10.11646/zootaxa.4981.3.3" ID-GBIF-Dataset="bd4313a1-f17a-4745-ba2d-655124108edc" ID-ISSN="1175-5326" ID-Zenodo-Dep="4921137" ID-ZooBank="CB36C384-5538-494A-8802-4F821C7E25FE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623280791010" checkinUser="plazi" docAuthor="Chernyshev, Alexei V. &amp; Goedert, James L." docDate="2021" docId="038287AEC1401749FF7F5E7F0B69082C" docLanguage="en" docName="zootaxa.4981.3.3.pdf" docOrigin="Zootaxa 4981 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Orbitestella palaiopacifica Squires &amp; Goedert 1996" docType="treatment" docVersion="8" lastPageNumber="471" masterDocId="FFBBFFD6C142174BFFE85A1D090F0F6B" masterDocTitle="New species of latest Eocene / earliest Oligocene microgastropods (Heterobranchia Orbitestellidae and Omalogyridae) from the Gries Ranch Formation, Lewis County, Washington State, USA" masterLastPageNumber="480" masterPageNumber="469" pageNumber="471" updateTime="1699009098020" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="65A83D1F82A7E674CA39A31CBFB33D4E">New species of latest Eocene / earliest Oligocene microgastropods (Heterobranchia Orbitestellidae and Omalogyridae) from the Gries Ranch Formation, Lewis County, Washington State, USA</mods:title>
</mods:titleInfo>
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<mods:namePart id="930D748D3913BAEA1167E75BF500F306">Chernyshev, Alexei V.</mods:namePart>
<mods:affiliation id="BF78A646CD7C0D2F16B8A4CBC3988952">A. V. Zhirmunsky National Scientific Center of Marine Biology, Far Eastern Branch, Russian Academy of Sciences, Palchevskogo Street 17, Vladivostok 690041, Russia.</mods:affiliation>
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<mods:namePart id="79A81CBCA07878F8D7CEC364A0CDC959">Goedert, James L.</mods:namePart>
<mods:affiliation id="C3FF4FF77490E5EF59C0C93C2D1DEE3E">Burke Museum of Natural History and Culture, University of Washington, Seattle, WA 98195, USA.</mods:affiliation>
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<mods:title id="B0EFEBBE54872B4727309AAA158A9CC2">Zootaxa</mods:title>
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<mods:date id="D041430D29C7E1F5924CB1ADB0271176">2021</mods:date>
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<mods:number id="54C1E842A9D8BC5F204E27F0862E3AEE">2021-06-08</mods:number>
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<mods:number id="951866E8D9B754BCC4CC5BE910E5ED45">4981</mods:number>
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<mods:identifier id="3F4C7330EE6B8F10299065309D11EF81" type="CLB-Dataset">5429</mods:identifier>
<mods:identifier id="A30838F44BFF2796D861B41319EE9C4B" type="DOI">10.11646/zootaxa.4981.3.3</mods:identifier>
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<treatment id="038287AEC1401749FF7F5E7F0B69082C" ID-DOI="http://doi.org/10.5281/zenodo.5109761" ID-GBIF-Taxon="183548939" ID-Zenodo-Dep="5109761" LSID="urn:lsid:plazi:treatment:038287AEC1401749FF7F5E7F0B69082C" httpUri="http://treatment.plazi.org/id/038287AEC1401749FF7F5E7F0B69082C" lastPageNumber="471" pageId="2" pageNumber="471">
<subSubSection id="C3316533C1401749FF7F5E7F0A1D0B17" box="[151,786,1122,1149]" pageId="2" pageNumber="471" type="nomenclature">
<paragraph id="8B9436B8C1401749FF7F5E7F0A1D0B17" blockId="2.[151,786,1122,1185]" box="[151,786,1122,1149]" pageId="2" pageNumber="471">
<heading id="D0DC81D4C1401749FF7F5E7F0A1D0B17" bold="true" box="[151,786,1122,1149]" fontSize="11" level="1" pageId="2" pageNumber="471" reason="1">
<taxonomicName id="4C2B4D3BC1401749FF7F5E7F0A1D0B17" ID-CoL="8V8X6" authority="Squires &amp; Goedert, 1996" authorityName="Squires &amp; Goedert" authorityYear="1996" box="[151,786,1122,1149]" class="Gastropoda" family="Orbitestellidae" genus="Orbitestella" kingdom="Animalia" order="Heterostropha" pageId="2" pageNumber="471" phylum="Mollusca" rank="species" species="palaiopacifica">
<emphasis id="B95FEAAAC1401749FF7F5E7F0A1D0B17" bold="true" box="[151,786,1122,1149]" pageId="2" pageNumber="471">
<emphasis id="B95FEAAAC1401749FF7F5E7F08D90B16" bold="true" box="[151,470,1122,1149]" italics="true" pageId="2" pageNumber="471">Orbitestella palaiopacifica</emphasis>
Squires &amp; Goedert, 1996
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3316533C1401749FF7F5E9A08250BCA" box="[151,298,1159,1185]" pageId="2" pageNumber="471" type="description">
<paragraph id="8B9436B8C1401749FF7F5E9A08250BCA" blockId="2.[151,786,1122,1185]" box="[151,298,1159,1185]" pageId="2" pageNumber="471">
<figureCitation id="13102A3DC1401749FF7F5E9A08250BCA" box="[151,298,1159,1185]" captionStart="FIGURE 1" captionStartId="3.[151,250,1017,1041]" captionTargetBox="[151,1435,179,991]" captionTargetId="figure-19@3.[151,1435,179,991]" captionTargetPageId="3" captionText="FIGURE 1. Orbitestella palaiopacifica Squires &amp; Goedert (SEM). A, specimen MIMB 41133–1, apical view; B, specimen MIMB 41133–2, umbilical view; C, specimen MIMB 41133–3, apertural view. Scales: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4921141" httpUri="https://zenodo.org/record/4921141/files/figure.png" pageId="2" pageNumber="471">Figure 1A–C</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3316533C1401749FF7F5ED60A710B88" box="[151,894,1227,1251]" pageId="2" pageNumber="471" type="reference_group">
<paragraph id="8B9436B8C1401749FF7F5ED60A710B88" blockId="2.[151,894,1227,1251]" box="[151,894,1227,1251]" pageId="2" pageNumber="471">
<taxonomicName id="4C2B4D3BC1401749FF7F5ED60BF40B88" ID-CoL="8V8X6" authority="Squires &amp; Goedert, 1996, p. 233" authorityName="Squires &amp; Goedert" authorityPageNumber="233" authorityYear="1996" box="[151,763,1227,1251]" class="Gastropoda" family="Orbitestellidae" genus="Orbitestella" kingdom="Animalia" order="Heterostropha" pageId="2" pageNumber="471" phylum="Mollusca" rank="species" species="palaiopacifica">
<emphasis id="B95FEAAAC1401749FF7F5ED608A60B88" box="[151,425,1227,1251]" italics="true" pageId="2" pageNumber="471">Orbitestella palaiopacifica</emphasis>
Squires &amp; Goedert, 1996, p. 233
</taxonomicName>
, figs. 22–26.
</paragraph>
</subSubSection>
<subSubSection id="C3316533C1401749FF7F5F0C088A0AD7" pageId="2" pageNumber="471" type="materials_examined">
<paragraph id="8B9436B8C1401749FF7F5F0C09FE0A24" blockId="2.[151,1436,1297,1863]" pageId="2" pageNumber="471">
<emphasis id="B95FEAAAC1401749FF7F5F0C08880A40" bold="true" box="[151,391,1297,1323]" pageId="2" pageNumber="471">Examined material.</emphasis>
<materialsCitation id="3B433CE5C1401749FE7A5F0F0A280A47" ID-GBIF-Occurrence="3323273310" box="[402,807,1297,1324]" collectionCode="MIMB" pageId="2" pageNumber="471" specimenCode="MIMB 41133" specimenCount="6">
<specimenCode id="DB8D9EC3C1401749FE7A5F0F0B200A40" box="[402,559,1297,1324]" collectionCode="MIMB" country="Russia" httpUri="http://grbio.org/cool/063z-3mpp" name="Museum of the Institute of Marine Biology" pageId="2" pageNumber="471">MIMB 41133</specimenCode>
(lot of
<specimenCount id="9D2DFD31C1401749FD655F0C0A100A47" box="[653,799,1297,1324]" pageId="2" pageNumber="471" type="generic">6 specimens</specimenCount>
)
</materialsCitation>
,
<materialsCitation id="3B433CE5C1401749FCDE5F0C09E20A24" ID-GBIF-Occurrence="3323273309" collectionCode="SBMNH" pageId="2" pageNumber="471" specimenCode="SBMNH 467055" specimenCount="2">
<specimenCode id="DB8D9EC3C1401749FCDE5F0C0AF70A47" box="[822,1016,1297,1324]" collectionCode="SBMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34834" lsid="urn:lsid:biocol.org:col:34834" name="Santa Barbara Museum of Natural History" pageId="2" pageNumber="471" type="Museum">SBMNH 467055</specimenCode>
(lot of
<specimenCount id="9D2DFD31C1401749FBBE5F0C0DE70A47" box="[1110,1256,1297,1324]" pageId="2" pageNumber="471" type="generic">2 specimens</specimenCount>
), from the
<typeStatus id="5490881AC1401749FA845F0F0C930A47" box="[1388,1436,1298,1324]" pageId="2" pageNumber="471">type</typeStatus>
locality
</materialsCitation>
.
</paragraph>
<paragraph id="8B9436B8C1401749FF2F5F440D4E0A18" blockId="2.[151,1436,1297,1863]" box="[199,1089,1369,1396]" pageId="2" pageNumber="471">
<emphasis id="B95FEAAAC1401749FF2F5F44087A0A18" bold="true" box="[199,373,1369,1395]" pageId="2" pageNumber="471">Type material.</emphasis>
<materialsCitation id="3B433CE5C1401749FE945F440BA40A1F" ID-GBIF-Occurrence="3323273316" box="[380,683,1369,1396]" collectionCode="LACMIP" pageId="2" pageNumber="471" specimenCode="LACMIP 11365" specimenCount="1" typeStatus="holotype">
<typeStatus id="5490881AC1401749FE945F4408E60A18" box="[380,489,1369,1395]" pageId="2" pageNumber="471" type="holotype">Holotype</typeStatus>
:
<specimenCode id="DB8D9EC3C1401749FE1B5F440BA40A1F" box="[499,683,1369,1396]" collectionCode="LACMIP" pageId="2" pageNumber="471">LACMIP 11365</specimenCode>
</materialsCitation>
;
<materialsCitation id="3B433CE5C1401749FD5E5F470D320A18" ID-GBIF-Occurrence="3323273304" box="[694,1085,1369,1396]" collectionCode="LACMIP" pageId="2" pageNumber="471" specimenCode="LACMIP 11366-11367" specimenCount="1" typeStatus="paratype">
<typeStatus id="5490881AC1401749FD5E5F470A290A1F" box="[694,806,1370,1396]" pageId="2" pageNumber="471" type="paratype">Paratypes</typeStatus>
,
<specimenCode id="DB8D9EC3C1401749FCD95F440D320A18" box="[817,1085,1369,1395]" collectionCode="LACMIP" pageId="2" pageNumber="471">LACMIP 11366–11367</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8B9436B8C1401749FF2F5F60088A0AD7" blockId="2.[151,1436,1297,1863]" pageId="2" pageNumber="471">
<materialsCitation id="3B433CE5C1401749FF2F5F60088E0AD7" ID-GBIF-Occurrence="3323273303" collectionCode="LACMIP" country="United States of America" county="Eocene" location="Larch Mountain" municipality="Crescent Formation" pageId="2" pageNumber="471" specimenCount="1" stateProvince="Washington" typeStatus="holotype">
<emphasis id="B95FEAAAC1401749FF2F5F60086B0AFC" bold="true" box="[199,356,1405,1431]" pageId="2" pageNumber="471">
<typeStatus id="5490881AC1401749FF2F5F60080D0AFC" box="[199,258,1405,1431]" pageId="2" pageNumber="471">Type</typeStatus>
locality.
</emphasis>
<collectionCode id="ED3AAE7DC1401749FE825F6008DC0AFC" box="[362,467,1405,1431]" pageId="2" pageNumber="471">LACMIP</collectionCode>
loc. 16655,
<location id="8EF46063C1401749FDB25F600A1E0AFC" LSID="urn:lsid:plazi:treatment:038287AEC1401749FF7F5E7F0B69082C:8EF46063C1401749FDB25F600A1E0AFC" box="[602,785,1405,1431]" country="United States of America" county="Eocene" municipality="Crescent Formation" name="Larch Mountain" pageId="2" pageNumber="471" stateProvince="Washington">Larch Mountain</location>
,
<location id="8EF46063C1401749FCF25F600A950AFC" LSID="urn:lsid:plazi:treatment:038287AEC1401749FF7F5E7F0B69082C:8EF46063C1401749FCF25F600A950AFC" box="[794,922,1405,1431]" country="United States of America" county="Eocene" municipality="Crescent Formation" name="Black Hills" pageId="2" pageNumber="471" stateProvince="Washington">Black Hills</location>
,
<collectingRegion id="49EFF85AC1401749FC4C5F600D650AFC" box="[932,1130,1405,1431]" country="United States of America" name="Washington" pageId="2" pageNumber="471">Washington State</collectingRegion>
,
<collectingCountry id="F33C7628C1401749FB9B5F600DA00AFC" box="[1139,1199,1405,1431]" name="United States of America" pageId="2" pageNumber="471">USA</collectingCountry>
;
<collectingMunicipality id="6BF0ACC2C1401749FB515F600C960AFC" box="[1209,1433,1405,1431]" pageId="2" pageNumber="471">Crescent Formation</collectingMunicipality>
, middle early
<collectingCounty id="62F54E34C1401749FEC35FBF088E0AD7" box="[299,385,1442,1468]" pageId="2" pageNumber="471">Eocene</collectingCounty>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3316533C1401749FF2F5FD80C5E0943" pageId="2" pageNumber="471" type="diagnosis">
<paragraph id="8B9436B8C1401749FF2F5FD80C5E0943" blockId="2.[151,1436,1297,1863]" pageId="2" pageNumber="471">
<emphasis id="B95FEAAAC1401749FF2F5FD8084D0AB4" bold="true" box="[199,322,1477,1503]" pageId="2" pageNumber="471">Diagnosis.</emphasis>
“An
<taxonomicName id="4C2B4D3BC1401749FE955FD80B0C0AB4" authorityName="Iredale" authorityYear="1917" box="[381,515,1477,1503]" class="Gastropoda" family="Orbitestellidae" genus="Orbitestella" kingdom="Animalia" order="Heterostropha" pageId="2" pageNumber="471" phylum="Mollusca" rank="genus">
<emphasis id="B95FEAAAC1401749FE955FD80B0C0AB4" box="[381,515,1477,1503]" italics="true" pageId="2" pageNumber="471">Orbitestella</emphasis>
</taxonomicName>
with an upper whorl surface crossed by narrow axial ribs, a keel-like carina on the shoulder, axial ribs in the interspace between the carina on the basal margin and the carina on the edge of the umbilicus, and a wide but deep umbilicus showing overlapping whorls in its interior.” (Squires &amp; Goedert 1996).
</paragraph>
</subSubSection>
<subSubSection id="C3316533C1401749FF2F5C2C0B69082C" pageId="2" pageNumber="471" type="discussion">
<paragraph id="8B9436B8C1401749FF2F5C2C0B69082C" blockId="2.[151,1436,1297,1863]" pageId="2" pageNumber="471">
<emphasis id="B95FEAAAC1401749FF2F5C2C08340920" bold="true" box="[199,315,1585,1611]" pageId="2" pageNumber="471">Remarks.</emphasis>
Shells of
<taxonomicName id="4C2B4D3BC1401749FE475C2F0B7A0920" authorityName="Squires &amp; Goedert" authorityYear="1996" box="[431,629,1585,1612]" class="Gastropoda" family="Orbitestellidae" genus="Orbitestella" kingdom="Animalia" order="Heterostropha" pageId="2" pageNumber="471" phylum="Mollusca" rank="species" species="palaiopacifica">
<emphasis id="B95FEAAAC1401749FE475C2F0B7A0920" box="[431,629,1585,1612]" italics="true" pageId="2" pageNumber="471">O. palaiopacifica</emphasis>
</taxonomicName>
were reported to be up to
<quantity id="4CD39B5DC1401749FC4B5C2C0AF50927" box="[931,1018,1585,1612]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="471" unit="mm" value="0.3">0.3 mm</quantity>
high with a diameter of
<quantity id="4CD39B5DC1401749FAFB5C2C0C760927" box="[1299,1401,1585,1612]" metricMagnitude="-4" metricUnit="m" metricValue="5.3" pageId="2" pageNumber="471" unit="mm" value="0.53">0.53 mm</quantity>
by Squires &amp; Goedert (1996); a new specimen (MIMB 41133–2;
<figureCitation id="13102A3DC1401749FCB25C480AA00904" box="[858,943,1621,1647]" captionStart="FIGURE 1" captionStartId="3.[151,250,1017,1041]" captionTargetBox="[151,1435,179,991]" captionTargetId="figure-19@3.[151,1435,179,991]" captionTargetPageId="3" captionText="FIGURE 1. Orbitestella palaiopacifica Squires &amp; Goedert (SEM). A, specimen MIMB 41133–1, apical view; B, specimen MIMB 41133–2, umbilical view; C, specimen MIMB 41133–3, apertural view. Scales: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4921141" httpUri="https://zenodo.org/record/4921141/files/figure.png" pageId="2" pageNumber="471">Fig. 1B</figureCitation>
) is now the largest known specimen with a diameter of
<quantity id="4CD39B5DC1401749FEF65C64087809FF" box="[286,375,1657,1684]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="2" pageNumber="471" unit="mm" value="0.9">0.9 mm</quantity>
. New specimens of
<taxonomicName id="4C2B4D3BC1401749FDB05C670A1309F8" authorityName="Squires &amp; Goedert" authorityYear="1996" box="[600,796,1657,1684]" class="Gastropoda" family="Orbitestellidae" genus="Orbitestella" kingdom="Animalia" order="Heterostropha" pageId="2" pageNumber="471" phylum="Mollusca" rank="species" species="palaiopacifica">
<emphasis id="B95FEAAAC1401749FDB05C670A1309F8" box="[600,796,1657,1684]" italics="true" pageId="2" pageNumber="471">O. palaiopacifica</emphasis>
</taxonomicName>
show that the axial ribs cross the prominent keel-like carina on the shoulder, and spiral threads are present on most surfaces, including the apical and basal carinae. Squires &amp; Goedert (1996) stated that the protoconch of
<taxonomicName id="4C2B4D3BC1401749FD5C5CDF0A7609B0" authorityName="Squires &amp; Goedert" authorityYear="1996" box="[692,889,1729,1756]" class="Gastropoda" family="Orbitestellidae" genus="Orbitestella" kingdom="Animalia" order="Heterostropha" pageId="2" pageNumber="471" phylum="Mollusca" rank="species" species="palaiopacifica">
<emphasis id="B95FEAAAC1401749FD5C5CDF0A7609B0" box="[692,889,1729,1756]" italics="true" pageId="2" pageNumber="471">O. palaiopacifica</emphasis>
</taxonomicName>
was apparently smooth, but we think this is due to poor preservation of the protoconch. The protoconch surface of the newly available specimens appears also to be corroded.
<taxonomicName id="4C2B4D3BC1401749FEE05D140B380848" authorityName="Squires &amp; Goedert" authorityYear="1996" box="[264,567,1801,1827]" class="Gastropoda" family="Orbitestellidae" genus="Orbitestella" kingdom="Animalia" order="Heterostropha" pageId="2" pageNumber="471" phylum="Mollusca" rank="species" species="palaiopacifica">
<emphasis id="B95FEAAAC1401749FEE05D140B380848" box="[264,567,1801,1827]" italics="true" pageId="2" pageNumber="471">Orbitestella palaiopacifica</emphasis>
</taxonomicName>
is still known only from two localities of the Crescent Formation in the Black Hills southwest of Olympia,
<collectingRegion id="49EFF85AC1401749FE315D300B6D082C" box="[473,610,1837,1863]" country="United States of America" name="Washington" pageId="2" pageNumber="471">Washington</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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