354 lines
52 KiB
XML
354 lines
52 KiB
XML
<document id="159F6C503CFDAE96E93A9E07FFF214EE" ID-DOI="10.11646/zootaxa.4759.2.7" ID-GBIF-Dataset="a1f8fff2-71f1-4cda-9b90-484ece852445" ID-ISSN="1175-5326" ID-Zenodo-Dep="3737489" ID-ZooBank="EE7F7824-3672-4B28-BABC-F87600417A70" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1585803901140" checkinUser="plazi" docAuthor="Lien, Nguyen Thi Phuong" docDate="2020" docId="038287A80C791E3DFF40FC42927D1720" docLanguage="en" docName="zootaxa.4759.2.7.pdf" docOrigin="Zootaxa 4759 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Polistes (Polistella) caobangensis Lien 2020, sp. nov." docType="treatment" docVersion="5" lastPageNumber="255" masterDocId="FFBBFFD00C781E3EFFD7FFC590471543" masterDocTitle="Contribution to the taxonomy of the paper wasp genus Polistes (Polistella) (Hymenoptera: Vespidae: Polistinae) from Vietnam, with description of a new species" masterLastPageNumber="260" masterPageNumber="252" pageNumber="253" updateTime="1698734784073" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="268AEBC57B50F0E4E8322A6E2AE53F60">Contribution to the taxonomy of the paper wasp genus Polistes (Polistella) (Hymenoptera: Vespidae: Polistinae) from Vietnam, with description of a new species</mods:title>
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<mods:namePart id="435424A1CF068C10471555E8A2B9B91D">Lien, Nguyen Thi Phuong</mods:namePart>
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<mods:title id="F767202A1C07512DA299F9A8E31EAE1F">Zootaxa</mods:title>
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<mods:part id="ED95E240853CEABCBD45B87552410628">
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<mods:date id="9301163028CC1763BB75E36F1FC187A4">2020</mods:date>
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<mods:number id="0CF0FBBAB24A38A12E5FA41F0A829C97">2020-04-02</mods:number>
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<mods:number id="F1D648909EAECD3CA427681740926890">4759</mods:number>
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<treatment id="038287A80C791E3DFF40FC42927D1720" ID-DOI="http://doi.org/10.5281/zenodo.3811437" ID-GBIF-Taxon="163420901" ID-Zenodo-Dep="3811437" LSID="urn:lsid:plazi:treatment:038287A80C791E3DFF40FC42927D1720" httpUri="http://treatment.plazi.org/id/038287A80C791E3DFF40FC42927D1720" lastPageId="3" lastPageNumber="255" pageId="1" pageNumber="253">
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<subSubSection id="C33165350C791E3FFF40FC4292D116E1" box="[151,662,903,930]" pageId="1" pageNumber="253" type="nomenclature">
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<paragraph id="8B9436BE0C791E3FFF40FC4292D116E1" blockId="1.[151,662,903,966]" box="[151,662,903,930]" pageId="1" pageNumber="253">
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<heading id="D0DC81D20C791E3FFF40FC4292D116E1" bold="true" box="[151,662,903,930]" fontSize="11" level="1" pageId="1" pageNumber="253" reason="1">
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<emphasis id="B95FEAAC0C791E3FFF40FC4292D116E1" bold="true" box="[151,662,903,930]" pageId="1" pageNumber="253">
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<taxonomicName id="4C2B4D3D0C791E3FFF40FC42926D16E1" authority="Lien, 2020" authorityName="Lien" authorityYear="2020" box="[151,554,903,930]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="253" phylum="Arthropoda" rank="species" species="caobangensis" status="sp. nov." subGenus="Polistella">
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<emphasis id="B95FEAAC0C791E3FFF40FC4290B316E1" bold="true" box="[151,244,903,930]" italics="true" pageId="1" pageNumber="253">Polistes</emphasis>
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(
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<emphasis id="B95FEAAC0C791E3FFED2FC42913216E1" bold="true" box="[261,373,903,930]" italics="true" pageId="1" pageNumber="253">Polistella</emphasis>
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)
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<emphasis id="B95FEAAC0C791E3FFE50FC42926D16E1" bold="true" box="[391,554,903,930]" italics="true" pageId="1" pageNumber="253">caobangensis</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A26C57D70C791E3FFDEFFC4D92D116E1" box="[568,662,904,930]" pageId="1" pageNumber="253" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33165350C791E3FFF40FC6E91671685" box="[151,288,939,966]" pageId="1" pageNumber="253" type="description">
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<paragraph id="8B9436BE0C791E3FFF40FC6E91671685" blockId="1.[151,662,903,966]" box="[151,288,939,966]" pageId="1" pageNumber="253">
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(
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<figureCitation id="13102A3B0C791E3FFF48FC6E915F1685" box="[159,280,939,966]" captionStart-0="FIGURES 1–5" captionStart-1="FIGURES 6–10" captionStartId-0="3.[151,264,1948,1973]" captionStartId-1="4.[151,264,1943,1968]" captionTargetBox-0="[148,1439,633,1923]" captionTargetBox-1="[148,1432,181,1918]" captionTargetId-0="figure@3.[148,1440,633,1923]" captionTargetId-1="figure@4.[148,1439,181,1918]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURES 1–5. Polistes caobangensis, new species, female (holotype): 1. Head, frontal view. 2. Head, dorsal view. 3. Scutellum, metanotum and propodeum, dorsal view. 4. Metasomal segments 1–6, lateral view. 5. Habitus." captionText-1="FIGURES 6–10. Polistes caobangensis, new species, male: 6. Head, frontal view. 7. Part of head with left antenna. Genitalia. 8. Inner aspect of paramere with digitus and volsella. 9, 10. Aedeagus. (9. Ventral and 10. Lateral view). Scale 1mm." figureDoi-0="http://doi.org/10.5281/zenodo.3737491" figureDoi-1="http://doi.org/10.5281/zenodo.3737493" httpUri-0="https://zenodo.org/record/3737491/files/figure.png" httpUri-1="https://zenodo.org/record/3737493/files/figure.png" pageId="1" pageNumber="253">Figs. 1–10</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C33165350C791E3FFF40FC359332114B" box="[151,885,1007,1032]" pageId="1" pageNumber="253" type="reference_group">
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<paragraph id="8B9436BE0C791E3FFF40FC359332114B" blockId="1.[151,885,1007,1032]" box="[151,885,1007,1032]" pageId="1" pageNumber="253">
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<taxonomicName id="4C2B4D3D0C791E3FFF40FC3591AF114B" ID-CoL="4KYF9" authority="Perez, 1905" authorityName="Perez" authorityYear="1905" box="[151,488,1007,1032]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="253" phylum="Arthropoda" rank="species" species="nipponensis">
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<emphasis id="B95FEAAC0C791E3FFF40FC359120114B" box="[151,359,1008,1032]" italics="true" pageId="1" pageNumber="253">Polistes nipponensis</emphasis>
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<bibRefCitation id="EFBA4B4F0C791E3FFEBAFC3591AF114B" author="Perez, J." box="[365,488,1007,1032]" pageId="1" pageNumber="253" pagination="23 - 41" refId="ref5585" refString="Perez, J. (1905) Hymenopteres recueillis dans le Japon central par M. Harmand, Ministre Plenipotentiaire de France a Tokio. Bulletin du Museum National d'Histoire Naturelle, Paris, 1905, 23 - 41 + 79 - 87 + 148 - 158." type="journal article" year="1905">Pérez, 1905</bibRefCitation>
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</taxonomicName>
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: Nguyen, 2017: 225, misidentification.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33165350C791E3FFF40FBF0914011FF" pageId="1" pageNumber="253" type="materials_examined">
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<paragraph id="8B9436BE0C791E3FFF40FBF0914011FF" blockId="1.[151,1437,1077,2041]" pageId="1" pageNumber="253">
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<emphasis id="B95FEAAC0C791E3FFF40FBF091C71113" bold="true" box="[151,384,1077,1104]" pageId="1" pageNumber="253">Material examined.</emphasis>
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<materialsCitation id="3B433CE30C791E3FFE50FBF395DE1136" ID-GBIF-Occurrence="2594759899" collectionCode="IEBR, T, V" country="Vietnam" county="Phia Oac" elevation="1000" latitude="22.585472" location="Nguyen Binh" longLatPrecision="1" longitude="105.86661" municipality="Pha Den" pageId="1" pageNumber="253" specimenCount="1" stateProvince="Cao Bang" typeStatus="holotype">
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<typeStatus id="5490881C0C791E3FFE50FBF391B51113" box="[391,498,1078,1104]" pageId="1" pageNumber="253" type="holotype">Holotype</typeStatus>
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, female [
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], labeled “
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<collectingCountry id="F33C762E0C791E3FFCC6FBF393DF1113" box="[785,920,1078,1104]" name="Vietnam" pageId="1" pageNumber="253">VIETNAM</collectingCountry>
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,
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,
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<collectingMunicipality id="6BF0ACC40C791E3FFBC1FBF394301112" box="[1046,1143,1078,1105]" pageId="1" pageNumber="253">Pha Den</collectingMunicipality>
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,
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,
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,
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, alt.
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</quantity>
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, Nest#VN-CB-2014-P-01, 25–28 May 2014, coll.
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. Hoang
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</materialsCitation>
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.
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<materialsCitation id="3B433CE30C791E3FFF40FBBA944911DB" ID-GBIF-Occurrence="2594759902" box="[151,1038,1149,1177]" collectionCode="IEBR" country="Vietnam" county="Phia Oac" elevation="1000" latitude="22.585472" location="Nguyen Binh" longLatPrecision="1" longitude="105.86661" municipality="Pha Den" pageId="1" pageNumber="253" specimenCount="40" specimenCount-female="15" specimenCount-male="25" stateProvince="Cao Bang" typeStatus="paratype">
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<typeStatus id="5490881C0C791E3FFF40FBBA914F11DA" box="[151,264,1151,1177]" pageId="1" pageNumber="253" type="paratype">Paratypes</typeStatus>
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:
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:
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,
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, same data as holotype (
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)
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</materialsCitation>
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,
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<materialsCitation id="3B433CE30C791E3FFBCBFBBB914411FF" ID-GBIF-Occurrence="2594759903" collectionCode="VNMN" country="Vietnam" county="Phia Oac" elevation="1000" latitude="22.585472" location="Nguyen Binh" longLatPrecision="1" longitude="105.86661" municipality="Pha Den" pageId="1" pageNumber="253" specimenCount="3" specimenCount-female="3" stateProvince="Cao Bang" typeStatus="paratype">
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<specimenCount id="9D2DFD370C791E3FFBCBFBBB94C911DB" box="[1052,1166,1150,1176]" pageId="1" pageNumber="253" type="female">3 females</specimenCount>
|
||
, same data as holotype (
|
||
<collectionCode id="ED3AAE7B0C791E3FFF48FB6790BC11FF" box="[159,251,1186,1212]" country="Viet Nam" httpUri="http://grbio.org/cool/65ma-2bzq" name="Vietnam National Museum of Nature" pageId="1" pageNumber="253">VNMN</collectionCode>
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)
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</materialsCitation>
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.
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||
</paragraph>
|
||
</subSubSection>
|
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<subSubSection id="C33165350C791E3FFF10FB00940D109F" pageId="1" pageNumber="253" type="diagnosis">
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<paragraph id="8B9436BE0C791E3FFF10FB00940D109F" blockId="1.[151,1437,1077,2041]" pageId="1" pageNumber="253">
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||
<emphasis id="B95FEAAC0C791E3FFF10FB00910511A3" bold="true" box="[199,322,1221,1248]" pageId="1" pageNumber="253">Diagnosis.</emphasis>
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||
The new species can be distinguished from all other species of the subgenus
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||
<taxonomicName id="4C2B4D3D0C791E3FFB45FB0394BA11A3" authorityName="Ashmead" authorityYear="1904" box="[1170,1277,1222,1248]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="253" phylum="Arthropoda" rank="subGenus" subGenus="Polistella">
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||
<emphasis id="B95FEAAC0C791E3FFB45FB0394BA11A3" box="[1170,1277,1222,1248]" italics="true" pageId="1" pageNumber="253">Polistella</emphasis>
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||
</taxonomicName>
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||
by the combination of the following characters: female clypeus in frontal view 1.1 times as wide as high, with tomentum on clypeus extended to its dorsal three fifth to two third from its base; vertex strongly sloping down behind posterior ocelli towards occipital carina; pronotal carina sharply raised; S
|
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<quantity id="4CD39B5B0C791E3FFCC5FAF7937A100F" box="[786,829,1330,1356]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="1" pageNumber="253" unit="in" value="2.0">2 in</quantity>
|
||
lateral view strongly swollen ventrally in anterior half, then ventral margin bluntly angled into nearly straight line parallel to dorsal margin of the tergum; male clypeus in frontal view 1.1 times as wide as high; S7 with large and round tubercle; genitalia with penis valves 0.8 times longer than basal apodeme, in lateral view with proximoventral corner produced into nearly right angle; ventral margin of penis valves finely serrated along entire the length; jugal lobe of hind wing rounded.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33165350C791E3CFF10FA20951613FC" lastPageId="2" lastPageNumber="254" pageId="1" pageNumber="253" type="description">
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|
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<emphasis id="B95FEAAC0C791E3FFF10FA20911E1343" bold="true" box="[199,345,1509,1536]" pageId="1" pageNumber="253">Description.</emphasis>
|
||
<emphasis id="B95FEAAC0C791E3FFEB7FA2391F31343" box="[352,436,1510,1536]" italics="true" pageId="1" pageNumber="253">Female</emphasis>
|
||
. Body length
|
||
<quantity id="4CD39B5B0C791E3FFD83FA2392B01342" box="[596,759,1510,1537]" metricMagnitude="-2" metricUnit="m" metricValue="1.1099999999999999" metricValueMax="1.16" metricValueMin="1.06" pageId="1" pageNumber="253" unit="mm" value="11.1" valueMax="11.6" valueMin="10.6">10.6–11.6 mm</quantity>
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(
|
||
<typeStatus id="5490881C0C791E3FFCD2FA23932E1343" box="[773,873,1510,1536]" pageId="1" pageNumber="253" type="holotype">holotype</typeStatus>
|
||
11.0 mm); fore wing length 10.0–
|
||
<quantity id="4CD39B5B0C791E3FFB3AFA2395161342" box="[1261,1361,1510,1537]" metricMagnitude="-2" metricUnit="m" metricValue="1.09" pageId="1" pageNumber="253" unit="mm" value="10.9">10.9 mm</quantity>
|
||
(
|
||
<typeStatus id="5490881C0C791E3FFAB7FA2390801366" pageId="1" pageNumber="253" type="holotype">holotype</typeStatus>
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<quantity id="4CD39B5B0C791E3FFF19F9CC91711366" box="[206,310,1545,1573]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="1" pageNumber="253" unit="mm" value="10.5">10.5 mm</quantity>
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).
|
||
</paragraph>
|
||
<paragraph id="8B9436BE0C791E3FFF10F9EB93771297" blockId="1.[151,1437,1077,2041]" pageId="1" pageNumber="253">
|
||
<typeStatus id="5490881C0C791E3FFF10F9EB9168130B" box="[199,303,1582,1608]" pageId="1" pageNumber="253" type="holotype">Holotype</typeStatus>
|
||
(
|
||
<figureCitation id="13102A3B0C791E3FFEE9F9EB91C5130B" box="[318,386,1581,1608]" captionStart="FIGURES 1–5" captionStartId="3.[151,264,1948,1973]" captionTargetBox="[148,1439,633,1923]" captionTargetId="figure@3.[148,1440,633,1923]" captionTargetPageId="3" captionText="FIGURES 1–5. Polistes caobangensis, new species, female (holotype): 1. Head, frontal view. 2. Head, dorsal view. 3. Scutellum, metanotum and propodeum, dorsal view. 4. Metasomal segments 1–6, lateral view. 5. Habitus." figureDoi="http://doi.org/10.5281/zenodo.3737491" httpUri="https://zenodo.org/record/3737491/files/figure.png" pageId="1" pageNumber="253">Fig. 5</figureCitation>
|
||
). Head in frontal view nearly 1.2 × as wide as high (
|
||
<figureCitation id="13102A3B0C791E3FFC11F9EB9441130B" box="[966,1030,1582,1608]" captionStart="FIGURES 1–5" captionStartId="3.[151,264,1948,1973]" captionTargetBox="[148,1439,633,1923]" captionTargetId="figure@3.[148,1440,633,1923]" captionTargetPageId="3" captionText="FIGURES 1–5. Polistes caobangensis, new species, female (holotype): 1. Head, frontal view. 2. Head, dorsal view. 3. Scutellum, metanotum and propodeum, dorsal view. 4. Metasomal segments 1–6, lateral view. 5. Habitus." figureDoi="http://doi.org/10.5281/zenodo.3737491" httpUri="https://zenodo.org/record/3737491/files/figure.png" pageId="1" pageNumber="253">Fig. 1</figureCitation>
|
||
); in dorsal view weakly swollen laterally just behind eyes, and then narrowed posteriorly. Vertex with slightly raised area inside ocelli, greatly sloping down behind posterior ocelli towards occipital carina; POD: OOD = 1:1.7; POD about two times Od (
|
||
<figureCitation id="13102A3B0C791E3FFADFF9B3950A13D3" box="[1288,1357,1654,1680]" captionStart="FIGURES 1–5" captionStartId="3.[151,264,1948,1973]" captionTargetBox="[148,1439,633,1923]" captionTargetId="figure@3.[148,1440,633,1923]" captionTargetPageId="3" captionText="FIGURES 1–5. Polistes caobangensis, new species, female (holotype): 1. Head, frontal view. 2. Head, dorsal view. 3. Scutellum, metanotum and propodeum, dorsal view. 4. Metasomal segments 1–6, lateral view. 5. Habitus." figureDoi="http://doi.org/10.5281/zenodo.3737491" httpUri="https://zenodo.org/record/3737491/files/figure.png" pageId="1" pageNumber="253">Fig. 2</figureCitation>
|
||
). Gena in lateral view 0.9 × as wide as eye; occipital carina fine, evanescent in ventral one-third of gena. Inner eye margins in frontal view slightly further apart from each other at vertex than at clypeus (
|
||
<figureCitation id="13102A3B0C791E3FFBD5F97B9404139B" box="[1026,1091,1726,1752]" captionStart="FIGURES 1–5" captionStartId="3.[151,264,1948,1973]" captionTargetBox="[148,1439,633,1923]" captionTargetId="figure@3.[148,1440,633,1923]" captionTargetPageId="3" captionText="FIGURES 1–5. Polistes caobangensis, new species, female (holotype): 1. Head, frontal view. 2. Head, dorsal view. 3. Scutellum, metanotum and propodeum, dorsal view. 4. Metasomal segments 1–6, lateral view. 5. Habitus." figureDoi="http://doi.org/10.5281/zenodo.3737491" httpUri="https://zenodo.org/record/3737491/files/figure.png" pageId="1" pageNumber="253">Fig. 1</figureCitation>
|
||
). Distance between inner margins of antennal sockets about 1.7 × of distance from outer margin of antennal socket to inner eye margin; anterior tentorial pit as distant from antennal socket as from inner eye margin; interantennal space weakly raised. Clypeus in frontal view 1.1 × as wide as high (
|
||
<figureCitation id="13102A3B0C791E3FFDE9F8EF92381207" box="[574,639,1834,1860]" captionStart="FIGURES 1–5" captionStartId="3.[151,264,1948,1973]" captionTargetBox="[148,1439,633,1923]" captionTargetId="figure@3.[148,1440,633,1923]" captionTargetPageId="3" captionText="FIGURES 1–5. Polistes caobangensis, new species, female (holotype): 1. Head, frontal view. 2. Head, dorsal view. 3. Scutellum, metanotum and propodeum, dorsal view. 4. Metasomal segments 1–6, lateral view. 5. Habitus." figureDoi="http://doi.org/10.5281/zenodo.3737491" httpUri="https://zenodo.org/record/3737491/files/figure.png" pageId="1" pageNumber="253">Fig. 1</figureCitation>
|
||
), produced ventrally into blunt angle; in lateral view convex from base to apical margin; lateral margin of clypeus lying along inner eye margin as long as diameter of antennal socket and length of malar space. Antennal scape 3 × as long as its maximum width; F1 2.9 × as long as its maximum width, 1.2 × as long as F2 and 3 combined; F2 longer than wide, F3 & 4 as long as wide; F5–9 wider than long; terminal flagellomere bullet-shaped, 1.2 × as long as its basal width.
|
||
</paragraph>
|
||
<paragraph id="8B9436BE0C791E3CFF10F81B956B1400" blockId="1.[151,1437,1077,2041]" lastBlockId="2.[151,1437,153,2015]" lastPageId="2" lastPageNumber="254" pageId="1" pageNumber="253">
|
||
Pronotal carina sharply raised, produced dorsally into thin lamella, slightly sinuate backward on lateral sides, reaching ventral corner of pronotum. Mesoscutum strongly convex, slightly longer than wide between tegulae; anterior margin broadly rounded. Scutellum convex, disc slightly concave at middle. Disc of metanotum almost flat. Propodeum (
|
||
<figureCitation id="13102A3B0C7A1E3CFEF1FF24912F15B8" box="[294,360,225,251]" captionStart="FIGURES 1–5" captionStartId="3.[151,264,1948,1973]" captionTargetBox="[148,1439,633,1923]" captionTargetId="figure@3.[148,1440,633,1923]" captionTargetPageId="3" captionText="FIGURES 1–5. Polistes caobangensis, new species, female (holotype): 1. Head, frontal view. 2. Head, dorsal view. 3. Scutellum, metanotum and propodeum, dorsal view. 4. Metasomal segments 1–6, lateral view. 5. Habitus." figureDoi="http://doi.org/10.5281/zenodo.3737491" httpUri="https://zenodo.org/record/3737491/files/figure.png" pageId="2" pageNumber="254">Fig. 3</figureCitation>
|
||
) short; posterior face widely (nearly half of maximum propodeal width) and shallowly excavated medially, more or less smoothly passing into lateral face; propodeal orifice elongate, 2.0 × as long as wide (measured at widest part), somewhat narrowed in dorsal half. Fore wing brown, transparent, vein dark brown.
|
||
</paragraph>
|
||
<paragraph id="8B9436BE0C7A1E3CFF10FE8891CC1490" blockId="2.[151,1437,153,2015]" pageId="2" pageNumber="254">
|
||
T1 short and thick, about 0.4 × as long as its apical width, in lateral view abruptly swollen dorsally just behind propodeal orifice; corner between anterior and dorsal faces bluntly angled. S
|
||
<quantity id="4CD39B5B0C7A1E3CFBC4FEB4940414C8" box="[1043,1091,369,395]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="2" pageNumber="254" unit="in" value="2.0">2 in</quantity>
|
||
lateral view (
|
||
<figureCitation id="13102A3B0C7A1E3CFB30FEB4956914C8" box="[1255,1326,369,395]" captionStart="FIGURES 1–5" captionStartId="3.[151,264,1948,1973]" captionTargetBox="[148,1439,633,1923]" captionTargetId="figure@3.[148,1440,633,1923]" captionTargetPageId="3" captionText="FIGURES 1–5. Polistes caobangensis, new species, female (holotype): 1. Head, frontal view. 2. Head, dorsal view. 3. Scutellum, metanotum and propodeum, dorsal view. 4. Metasomal segments 1–6, lateral view. 5. Habitus." figureDoi="http://doi.org/10.5281/zenodo.3737491" httpUri="https://zenodo.org/record/3737491/files/figure.png" pageId="2" pageNumber="254">Fig. 4</figureCitation>
|
||
) strongly swollen ventrally in anterior one half, then ventral margin bluntly angled into nearly straight line parallel to dorsal margin of the tergum.
|
||
</paragraph>
|
||
<paragraph id="8B9436BE0C7A1E3CFF10FE18937F16C0" blockId="2.[151,1437,153,2015]" pageId="2" pageNumber="254">Body with short, golden setae except posterior part of propodeum with long silver setae. Clypeus with dense large punctures at basal half and scattered large punctures at apical half, each bearing a sharply pointed golden bristle, tomentum on clypeus restricted to its dorsal one half. Mandible with scattered medium punctures. Frons with deep definite punctures. Vertex and gena with sparse and strong punctures; area around ocelli smooth; ventral one third of gena with scattered strong punctures. Pronotum with dense deep punctures; space between punctures very narrow, slightly raised to form reticulation. Mesoscutum with coarse dense flat-bottom punctures; punctures on scutellum and metanotum coarse and dense, smaller than those on mesoscutum. Mesepisternum with punctures similar to those on pronotum, densely with punctures in posterodorsal part and scattered punctures in anteroventral part; border between them indistinct. Dorsal metapleuron with large smooth area, with some striae and sparse punctures, ventral metapleuron with sparse punctures in between smooth areas. Propodeum with strong sharply-raised transverse striae. Metasomal segments densely covered with minute punctures in addition to sparse small punctures (ill-defined on terga; more or less well-defined on sternum).</paragraph>
|
||
<paragraph id="8B9436BE0C7A1E3CFF10FC4895651174" blockId="2.[151,1437,153,2015]" pageId="2" pageNumber="254">
|
||
<emphasis id="B95FEAAC0C7A1E3CFF10FC48914F16E4" box="[199,264,909,935]" italics="true" pageId="2" pageNumber="254">Color</emphasis>
|
||
. Black, following parts dark brown: clypeus except basal margin, gena, mandible, antennal scape and all flagellomeres beneath, pronotum entirely, tegulae, scutellum, metanotum; a spot on upper part of mesepisternum; half of T1 and 2 at apical margin, T3–6 entirely, S2–6 except black band at basal margin of S2; apical half of fore femur and tibia; upper part of middle and hind femur and middle and hind tibia entirely, tarsi of fore legs except a brighter brown longitudinal band on tarsus 5. Two spots on each side of propodeum and valvula yellow.
|
||
</paragraph>
|
||
<paragraph id="8B9436BE0C7A1E3CFF10FB8495151070" blockId="2.[151,1437,153,2015]" pageId="2" pageNumber="254">
|
||
<emphasis id="B95FEAAC0C7A1E3CFF10FB84914F1118" box="[199,264,1089,1115]" italics="true" pageId="2" pageNumber="254">Male.</emphasis>
|
||
Body length
|
||
<quantity id="4CD39B5B0C7A1E3CFE75FB8492001118" box="[418,583,1089,1115]" metricMagnitude="-2" metricUnit="m" metricValue="1.08" metricValueMax="1.14" metricValueMin="1.02" pageId="2" pageNumber="254" unit="mm" value="10.8" valueMax="11.4" valueMin="10.2">10.2–11.4 mm</quantity>
|
||
; fore wing length 9.0–10.0 mm. Structure as in female, but differing as follows: head 1.3 × as wide as high in frontal view (
|
||
<figureCitation id="13102A3B0C7A1E3CFD17FBA0934311C3" box="[704,772,1125,1152]" captionStart="FIGURES 6–10" captionStartId="4.[151,264,1943,1968]" captionTargetBox="[148,1432,181,1918]" captionTargetId="figure@4.[148,1439,181,1918]" captionTargetPageId="4" captionText="FIGURES 6–10. Polistes caobangensis, new species, male: 6. Head, frontal view. 7. Part of head with left antenna. Genitalia. 8. Inner aspect of paramere with digitus and volsella. 9, 10. Aedeagus. (9. Ventral and 10. Lateral view). Scale 1mm." figureDoi="http://doi.org/10.5281/zenodo.3737493" httpUri="https://zenodo.org/record/3737493/files/figure.png" pageId="2" pageNumber="254">Fig. 6</figureCitation>
|
||
); eye strongly swollen laterally; inner eye margins 1.2 × as distant from each other at vertex as at clypeus; gena narrow, in lateral view 0.4 × as wide as eye; clypeus in frontal view 1.1 × as wide as high (
|
||
<figureCitation id="13102A3B0C7A1E3CFE18FB6892551184" box="[463,530,1196,1223]" captionStart="FIGURES 1–5" captionStartId="3.[151,264,1948,1973]" captionTargetBox="[148,1439,633,1923]" captionTargetId="figure@3.[148,1440,633,1923]" captionTargetPageId="3" captionText="FIGURES 1–5. Polistes caobangensis, new species, female (holotype): 1. Head, frontal view. 2. Head, dorsal view. 3. Scutellum, metanotum and propodeum, dorsal view. 4. Metasomal segments 1–6, lateral view. 5. Habitus." figureDoi="http://doi.org/10.5281/zenodo.3737491" httpUri="https://zenodo.org/record/3737491/files/figure.png" pageId="2" pageNumber="254">Fig. 5</figureCitation>
|
||
), slightly produced ventrally, evenly and very weakly rounded apically, in lateral view weakly convex dorsally. Antenna slenderer than in female (
|
||
<figureCitation id="13102A3B0C7A1E3CFCBAFB1493F611A8" box="[877,945,1233,1259]" captionStart="FIGURES 6–10" captionStartId="4.[151,264,1943,1968]" captionTargetBox="[148,1432,181,1918]" captionTargetId="figure@4.[148,1439,181,1918]" captionTargetPageId="4" captionText="FIGURES 6–10. Polistes caobangensis, new species, male: 6. Head, frontal view. 7. Part of head with left antenna. Genitalia. 8. Inner aspect of paramere with digitus and volsella. 9, 10. Aedeagus. (9. Ventral and 10. Lateral view). Scale 1mm." figureDoi="http://doi.org/10.5281/zenodo.3737493" httpUri="https://zenodo.org/record/3737493/files/figure.png" pageId="2" pageNumber="254">Fig. 7</figureCitation>
|
||
); scape short, 2.3 × as long as its maximum width; F1 about 0.8 × as long as the length of the length of F2 and 3 combined; F1–11each longer than wide; terminal flagellomere elongate, slightly curved, 3.3 × as long as its basal width. S7 with large and round tubercle.
|
||
</paragraph>
|
||
<paragraph id="8B9436BE0C7A1E3CFF10FAF894F11014" blockId="2.[151,1437,153,2015]" box="[199,1206,1341,1367]" pageId="2" pageNumber="254">Body surface sculpture as in female except clypeus with tomentum entirely on its surface.</paragraph>
|
||
<paragraph id="8B9436BE0C7A1E3CFF10FAA493F81080" blockId="2.[151,1437,153,2015]" pageId="2" pageNumber="254">
|
||
<emphasis id="B95FEAAC0C7A1E3CFF10FAA4914F1038" box="[199,264,1377,1403]" italics="true" pageId="2" pageNumber="254">Color</emphasis>
|
||
. Similar to female except with more yellow parts: clypeus to ocular sinus, antennal beneath, a thin band along gena just behind eye, margins of pronotum, spots on lateral sides of scutellum, spots on lateral sides at basal margin, large spots on each side of propodeum, and underneath of body.
|
||
</paragraph>
|
||
<paragraph id="8B9436BE0C7A1E3CFF10FA08951613FC" blockId="2.[151,1437,153,2015]" pageId="2" pageNumber="254">
|
||
<emphasis id="B95FEAAC0C7A1E3CFF10FA08912C10A4" box="[199,363,1485,1511]" italics="true" pageId="2" pageNumber="254">Male genitalia</emphasis>
|
||
. Generally similar to that of
|
||
<taxonomicName id="4C2B4D3D0C7A1E3CFD70FA0B93A710A4" authority="Gusenleitner, 2006" authorityName="Gusenleitner" authorityYear="2006" box="[679,992,1485,1511]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="254" phylum="Arthropoda" rank="species" species="affinis">
|
||
<emphasis id="B95FEAAC0C7A1E3CFD70FA0B934010A4" box="[679,775,1485,1511]" italics="true" pageId="2" pageNumber="254">P. affinis</emphasis>
|
||
<bibRefCitation id="EFBA4B4F0C7A1E3CFCDAFA0893A710A4" author="Gusenleitner, J." box="[781,992,1485,1511]" pageId="2" pageNumber="254" pagination="671 - 675" refId="ref5007" refString="Gusenleitner, J. (2006) Drei neue Polistes-Arten aus Laos und Vietnam (Hymenoptera: Vespidae). Linzer Biologische Beitrage, 38 (1), 671 - 675." type="journal article" year="2006">Gusenleitner, 2006</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2B4D3D0C7A1E3CFBC3FA0B950710A4" authority="Perez, 1905" authorityName="Perez" authorityYear="1905" box="[1044,1344,1484,1511]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="254" phylum="Arthropoda" rank="species" species="nipponensis">
|
||
<emphasis id="B95FEAAC0C7A1E3CFBC3FA0B94F310A4" box="[1044,1204,1486,1511]" italics="true" pageId="2" pageNumber="254">P. nipponensis</emphasis>
|
||
<bibRefCitation id="EFBA4B4F0C7A1E3CFB6DFA08950710A4" author="Perez, J." box="[1210,1344,1484,1511]" pageId="2" pageNumber="254" pagination="23 - 41" refId="ref5585" refString="Perez, J. (1905) Hymenopteres recueillis dans le Japon central par M. Harmand, Ministre Plenipotentiaire de France a Tokio. Bulletin du Museum National d'Histoire Naturelle, Paris, 1905, 23 - 41 + 79 - 87 + 148 - 158." type="journal article" year="1905">Pérez, 1905</bibRefCitation>
|
||
</taxonomicName>
|
||
. Digitus in inner aspect of paramere (
|
||
<figureCitation id="13102A3B0C7A1E3CFE0AFA3492661348" box="[477,545,1521,1547]" captionStart="FIGURES 6–10" captionStartId="4.[151,264,1943,1968]" captionTargetBox="[148,1432,181,1918]" captionTargetId="figure@4.[148,1439,181,1918]" captionTargetPageId="4" captionText="FIGURES 6–10. Polistes caobangensis, new species, male: 6. Head, frontal view. 7. Part of head with left antenna. Genitalia. 8. Inner aspect of paramere with digitus and volsella. 9, 10. Aedeagus. (9. Ventral and 10. Lateral view). Scale 1mm." figureDoi="http://doi.org/10.5281/zenodo.3737493" httpUri="https://zenodo.org/record/3737493/files/figure.png" pageId="2" pageNumber="254">Fig. 8</figureCitation>
|
||
) 3.2 × as long as wide (measured at widest part), distinctly swollen near base, gradually narrowed apically to mid-length, then slightly swollen towards the rounded apex. Aedeagus (
|
||
<figureCitation id="13102A3B0C7A1E3CFAC2F9D095C9136C" box="[1301,1422,1557,1583]" captionStart="FIGURES 6–10" captionStartId="4.[151,264,1943,1968]" captionTargetBox="[148,1432,181,1918]" captionTargetId="figure@4.[148,1439,181,1918]" captionTargetPageId="4" captionText="FIGURES 6–10. Polistes caobangensis, new species, male: 6. Head, frontal view. 7. Part of head with left antenna. Genitalia. 8. Inner aspect of paramere with digitus and volsella. 9, 10. Aedeagus. (9. Ventral and 10. Lateral view). Scale 1mm." figureDoi="http://doi.org/10.5281/zenodo.3737493" httpUri="https://zenodo.org/record/3737493/files/figure.png" pageId="2" pageNumber="254">Figs. 9, 10</figureCitation>
|
||
), penis valves 0.8 × longer than basal apodeme, in ventral view narrowest at two fifth from base, nearly straight from two fifth then strongly diversion and distinctly produced laterally near proximal margins, in lateral view slightly thickened in proximal one fourth and with dorsal margin weakly and smoothly sinuate, with proximoventral corner produced into nearly right angle (
|
||
<figureCitation id="13102A3B0C7A1E3CFDD9F960922613FC" box="[526,609,1701,1727]" captionStart="FIGURES 6–10" captionStartId="4.[151,264,1943,1968]" captionTargetBox="[148,1432,181,1918]" captionTargetId="figure@4.[148,1439,181,1918]" captionTargetPageId="4" captionText="FIGURES 6–10. Polistes caobangensis, new species, male: 6. Head, frontal view. 7. Part of head with left antenna. Genitalia. 8. Inner aspect of paramere with digitus and volsella. 9, 10. Aedeagus. (9. Ventral and 10. Lateral view). Scale 1mm." figureDoi="http://doi.org/10.5281/zenodo.3737493" httpUri="https://zenodo.org/record/3737493/files/figure.png" pageId="2" pageNumber="254">Fig. 10</figureCitation>
|
||
); ventral margin of penis valves finely serrated along entire length.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33165350C7A1E3DFF10F90C95D71758" lastPageId="3" lastPageNumber="255" pageId="2" pageNumber="254" type="discussion">
|
||
<paragraph id="8B9436BE0C7A1E3DFF10F90C91B11594" blockId="2.[151,1437,153,2015]" lastBlockId="3.[151,1436,153,611]" lastPageId="3" lastPageNumber="255" pageId="2" pageNumber="254">
|
||
<emphasis id="B95FEAAC0C7A1E3CFF10F90C917313A0" bold="true" box="[199,308,1737,1763]" pageId="2" pageNumber="254">Remarks</emphasis>
|
||
. This species is similar to
|
||
<taxonomicName id="4C2B4D3D0C7A1E3CFD89F90F93CC13A0" authority="Perez, 1905" authorityName="Perez" authorityYear="1905" box="[606,907,1736,1763]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="254" phylum="Arthropoda" rank="species" species="nipponensis">
|
||
<emphasis id="B95FEAAC0C7A1E3CFD89F90F92B813A0" box="[606,767,1738,1763]" italics="true" pageId="2" pageNumber="254">P. nipponensis</emphasis>
|
||
<bibRefCitation id="EFBA4B4F0C7A1E3CFCD1F90C93CC13A0" author="Perez, J." box="[774,907,1736,1763]" pageId="2" pageNumber="254" pagination="23 - 41" refId="ref5585" refString="Perez, J. (1905) Hymenopteres recueillis dans le Japon central par M. Harmand, Ministre Plenipotentiaire de France a Tokio. Bulletin du Museum National d'Histoire Naturelle, Paris, 1905, 23 - 41 + 79 - 87 + 148 - 158." type="journal article" year="1905">Pérez, 1905</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2B4D3D0C7A1E3CFC16F90F954613A0" authority="Gusenleitner, 2006" authorityName="Gusenleitner" authorityYear="2006" box="[961,1281,1737,1763]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="254" phylum="Arthropoda" rank="species" species="affinis">
|
||
<emphasis id="B95FEAAC0C7A1E3CFC16F90F946413A0" box="[961,1059,1737,1763]" italics="true" pageId="2" pageNumber="254">P. affinis</emphasis>
|
||
<bibRefCitation id="EFBA4B4F0C7A1E3CFBFDF90C954613A0" author="Gusenleitner, J." box="[1066,1281,1737,1763]" pageId="2" pageNumber="254" pagination="671 - 675" refId="ref5007" refString="Gusenleitner, J. (2006) Drei neue Polistes-Arten aus Laos und Vietnam (Hymenoptera: Vespidae). Linzer Biologische Beitrage, 38 (1), 671 - 675." type="journal article" year="2006">Gusenleitner, 2006</bibRefCitation>
|
||
</taxonomicName>
|
||
, but differs in the following characters: female and male clypeus in frontal view 1.1 × as wide as high (as wide as high in
|
||
<taxonomicName id="4C2B4D3D0C7A1E3CFAEFF92B909A1268" authorityName="Perez" authorityYear="1905" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="254" phylum="Arthropoda" rank="species" species="nipponensis">
|
||
<emphasis id="B95FEAAC0C7A1E3CFAEFF92B909A1268" italics="true" pageId="2" pageNumber="254">P. nipponensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2B4D3D0C7A1E3CFEC5F8D791341268" authorityName="Gusenleitner" authorityYear="2006" box="[274,371,1809,1835]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="254" phylum="Arthropoda" rank="species" species="affinis">
|
||
<emphasis id="B95FEAAC0C7A1E3CFEC5F8D791341268" box="[274,371,1809,1835]" italics="true" pageId="2" pageNumber="254">P. affinis</emphasis>
|
||
</taxonomicName>
|
||
); vertex strongly sloping down behind posterior ocelli towards occipital carina (slightly sloping down behind posterior ocelli towards occipical carina in
|
||
<taxonomicName id="4C2B4D3D0C7A1E3CFCF6F8F39382120C" authorityName="Perez" authorityYear="1905" box="[801,965,1846,1871]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="254" phylum="Arthropoda" rank="species" species="nipponensis">
|
||
<emphasis id="B95FEAAC0C7A1E3CFCF6F8F39382120C" box="[801,965,1846,1871]" italics="true" pageId="2" pageNumber="254">P. nipponensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2B4D3D0C7A1E3CFC29F8F39425120C" authorityName="Gusenleitner" authorityYear="2006" box="[1022,1122,1845,1871]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="254" phylum="Arthropoda" rank="species" species="affinis">
|
||
<emphasis id="B95FEAAC0C7A1E3CFC29F8F39425120C" box="[1022,1122,1845,1871]" italics="true" pageId="2" pageNumber="254">P. affinis</emphasis>
|
||
</taxonomicName>
|
||
); male gena in lateral view 0.4 × as wide as eye (0.3 × as wide as eye in
|
||
<taxonomicName id="4C2B4D3D0C7A1E3CFD41F89F92BF1230" authorityName="Gusenleitner" authorityYear="2006" box="[662,760,1881,1907]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="254" phylum="Arthropoda" rank="species" species="affinis">
|
||
<emphasis id="B95FEAAC0C7A1E3CFD41F89F92BF1230" box="[662,760,1881,1907]" italics="true" pageId="2" pageNumber="254">P. affinis</emphasis>
|
||
</taxonomicName>
|
||
); female F5–6 wider than long (as wide as long in
|
||
<taxonomicName id="4C2B4D3D0C7A1E3CFAEDF89F95DB1230" authorityName="Gusenleitner" authorityYear="2006" box="[1338,1436,1881,1907]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="254" phylum="Arthropoda" rank="species" species="affinis">
|
||
<emphasis id="B95FEAAC0C7A1E3CFAEDF89F95DB1230" box="[1338,1436,1881,1907]" italics="true" pageId="2" pageNumber="254">P. affinis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2B4D3D0C7A1E3CFF1FF8BB912D12D4" authorityName="Perez" authorityYear="1905" box="[200,362,1918,1943]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="254" phylum="Arthropoda" rank="species" species="nipponensis">
|
||
<emphasis id="B95FEAAC0C7A1E3CFF1FF8BB912D12D4" box="[200,362,1918,1943]" italics="true" pageId="2" pageNumber="254">P. nipponensis</emphasis>
|
||
</taxonomicName>
|
||
); S
|
||
<quantity id="4CD39B5B0C7A1E3CFE45F8B891F812D4" box="[402,447,1917,1943]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="2" pageNumber="254" unit="in" value="2.0">2 in</quantity>
|
||
lateral view swollen ventrally in anterior one half (S
|
||
<quantity id="4CD39B5B0C7A1E3CFBCFF8B8940212D4" box="[1048,1093,1917,1943]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="2" pageNumber="254" unit="in" value="2.0">2 in</quantity>
|
||
lateral view swollen ventrally in anterior one-third in
|
||
<taxonomicName id="4C2B4D3D0C7A1E3CFE4AF867927812F8" authorityName="Perez" authorityYear="1905" box="[413,575,1954,1979]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="254" phylum="Arthropoda" rank="species" species="nipponensis">
|
||
<emphasis id="B95FEAAC0C7A1E3CFE4AF867927812F8" box="[413,575,1954,1979]" italics="true" pageId="2" pageNumber="254">P. nipponensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2B4D3D0C7A1E3CFDA1F867929012F8" authorityName="Gusenleitner" authorityYear="2006" box="[630,727,1953,1979]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="254" phylum="Arthropoda" rank="species" species="affinis">
|
||
<emphasis id="B95FEAAC0C7A1E3CFDA1F867929012F8" box="[630,727,1953,1979]" italics="true" pageId="2" pageNumber="254">P. affinis</emphasis>
|
||
</taxonomicName>
|
||
); male S7 with large and round tubercle (without tubercle in
|
||
<taxonomicName id="4C2B4D3D0C7A1E3CFA5FF867915A129C" authorityName="Perez" authorityYear="1905" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="254" phylum="Arthropoda" rank="species" species="nipponensis">
|
||
<emphasis id="B95FEAAC0C7A1E3CFA5FF867915A129C" italics="true" pageId="2" pageNumber="254">P. nipponensis</emphasis>
|
||
</taxonomicName>
|
||
); body color black and brown with some yellow mark, metasomal terga without yellow band at apical margin (body color black with extensive yellow mark, metasomal terga with thick yellow band at apical margin of each tergum in
|
||
<taxonomicName id="4C2B4D3D0C7B1E3DFE93FF7B91A11594" authorityName="Perez" authorityYear="1905" box="[324,486,190,215]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="255" phylum="Arthropoda" rank="species" species="nipponensis">
|
||
<emphasis id="B95FEAAC0C7B1E3DFE93FF7B91A11594" box="[324,486,190,215]" italics="true" pageId="3" pageNumber="255">P. nipponensis</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9436BE0C7B1E3DFF10FF2490AF1490" blockId="3.[151,1436,153,611]" pageId="3" pageNumber="255">
|
||
This species is also similar to
|
||
<taxonomicName id="4C2B4D3D0C7B1E3DFDCCFF2793D715B8" authority="de Saussure, 1853" authorityName="de Saussure" authorityYear="1853" box="[539,912,224,251]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="255" phylum="Arthropoda" rank="species" species="mandarinus">
|
||
<emphasis id="B95FEAAC0C7B1E3DFDCCFF2792FA15B8" box="[539,701,225,251]" italics="true" pageId="3" pageNumber="255">P. mandarinus</emphasis>
|
||
de Saussure, 1853
|
||
</taxonomicName>
|
||
but differs in the following characters: female head in frontal view 1.2 times as wide as high (1.1 times as wide as high in
|
||
<taxonomicName id="4C2B4D3D0C7B1E3DFC35FEC394C4145C" authorityName="de Saussure" authorityYear="1853" box="[994,1155,261,287]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="255" phylum="Arthropoda" rank="species" species="mandarinus">
|
||
<emphasis id="B95FEAAC0C7B1E3DFC35FEC394C4145C" box="[994,1155,261,287]" italics="true" pageId="3" pageNumber="255">P. mandarinus</emphasis>
|
||
</taxonomicName>
|
||
); vertex strongly sloping down behind posterior ocelli towards occipital carina (slightly sloping down behind posterior ocelli towards occipical carina in
|
||
<taxonomicName id="4C2B4D3D0C7B1E3DFE85FE8B91B31424" authorityName="de Saussure" authorityYear="1853" box="[338,500,333,359]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="255" phylum="Arthropoda" rank="species" species="mandarinus">
|
||
<emphasis id="B95FEAAC0C7B1E3DFE85FE8B91B31424" box="[338,500,333,359]" italics="true" pageId="3" pageNumber="255">P. mandarinus</emphasis>
|
||
</taxonomicName>
|
||
); male S7 with large and round tubercle (without tubercle in
|
||
<taxonomicName id="4C2B4D3D0C7B1E3DFB70FE8B950E1424" authorityName="de Saussure" authorityYear="1853" box="[1191,1353,333,359]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="255" phylum="Arthropoda" rank="species" species="mandarinus">
|
||
<emphasis id="B95FEAAC0C7B1E3DFB70FE8B950E1424" box="[1191,1353,333,359]" italics="true" pageId="3" pageNumber="255">P. mandarinus</emphasis>
|
||
</taxonomicName>
|
||
); penis valves in lateral view with proximoventral corner produced into nearly right angle (penis valves in lateral view with proximoventral corner not produced into right angle in
|
||
<taxonomicName id="4C2B4D3D0C7B1E3DFCE8FE5393A614EC" authorityName="de Saussure" authorityYear="1853" box="[831,993,405,431]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="255" phylum="Arthropoda" rank="species" species="mandarinus">
|
||
<emphasis id="B95FEAAC0C7B1E3DFCE8FE5393A614EC" box="[831,993,405,431]" italics="true" pageId="3" pageNumber="255">P. mandarinus</emphasis>
|
||
</taxonomicName>
|
||
, only slightly produced (
|
||
<bibRefCitation id="EFBA4B4F0C7B1E3DFB2CFE5090951490" author="Nguyen, L. T. P. & Kojima, J. & Saito, F." pageId="3" pageNumber="255" pagination="1 - 30" refId="ref5276" refString="Nguyen, L. T. P., Kojima, J. & Saito, F. (2011) Polistes (Polistella) wasps (Hymenoptera: Vespidae: Polistinae) from mountainous areas of northern Vietnam, with description of five new species. Zootaxa, 3060 (1), 1 - 30. https: // doi. org / 10.11646 / zootaxa. 3060.1.1" type="journal article" year="2011">
|
||
Nguyen
|
||
<emphasis id="B95FEAAC0C7B1E3DFA8AFE5395C914EC" box="[1373,1422,405,431]" italics="true" pageId="3" pageNumber="255">et al</emphasis>
|
||
., 2011
|
||
</bibRefCitation>
|
||
)).
|
||
</paragraph>
|
||
<paragraph id="8B9436BE0C7B1E3DFF10FE1895D71758" blockId="3.[151,1436,153,611]" pageId="3" pageNumber="255">
|
||
In the previous study,
|
||
<taxonomicName id="4C2B4D3D0C7B1E3DFE10FE1B922C14B4" authorityName="de Saussure" authorityYear="1853" box="[455,619,477,503]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="255" phylum="Arthropoda" rank="species" species="mandarinus">
|
||
<emphasis id="B95FEAAC0C7B1E3DFE10FE1B922C14B4" box="[455,619,477,503]" italics="true" pageId="3" pageNumber="255">P. mandarinus</emphasis>
|
||
</taxonomicName>
|
||
was also recorded from Phia Oac NR,
|
||
<collectingRegion id="49EFF85C0C7B1E3DFBE6FE1894E414B4" box="[1073,1187,477,503]" country="Vietnam" name="Cao Bang" pageId="3" pageNumber="255">Cao Bang</collectingRegion>
|
||
together with its nest (
|
||
<bibRefCitation id="EFBA4B4F0C7B1E3DFF48FDC491FC1758" author="Nguyen, L. T. P. & Kojima, J." box="[159,443,513,539]" pageId="3" pageNumber="255" pagination="45 - 63" refId="ref5211" refString="Nguyen, L. T. P. & Kojima, J. (2014) Distribution and nests of paper wasps of Polistes (Polistella) in northeastern Vietnam, with description of a new species (Hymenoptera, Vespidae, Polistinae). Zookeys, 368, 45 - 63. https: // doi. org / 10.3897 / zookeys. 368.6426" type="journal article" year="2014">Nguyen & Kojima, 2014</bibRefCitation>
|
||
), but this nest has white cocoons while the nest of
|
||
<taxonomicName id="4C2B4D3D0C7B1E3DFC25FDC794E21758" authorityName="Lien" authorityYear="2020" box="[1010,1189,513,539]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="255" phylum="Arthropoda" rank="species" species="caobangensis">
|
||
<emphasis id="B95FEAAC0C7B1E3DFC25FDC794E21758" box="[1010,1189,513,539]" italics="true" pageId="3" pageNumber="255">P. caobangensis</emphasis>
|
||
</taxonomicName>
|
||
has yellow cocoons.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33165350C7B1E3DFF10FDE095D7177C" box="[199,1424,549,575]" pageId="3" pageNumber="255" type="etymology">
|
||
<paragraph id="8B9436BE0C7B1E3DFF10FDE095D7177C" blockId="3.[151,1436,153,611]" box="[199,1424,549,575]" pageId="3" pageNumber="255">
|
||
<emphasis id="B95FEAAC0C7B1E3DFF10FDE0910F177C" bold="true" box="[199,328,549,575]" pageId="3" pageNumber="255">Etymology</emphasis>
|
||
. The specific name refers to the
|
||
<typeStatus id="5490881C0C7B1E3DFD60FDE092A0177C" box="[695,743,549,575]" pageId="3" pageNumber="255">type</typeStatus>
|
||
locality,
|
||
<collectingRegion id="49EFF85C0C7B1E3DFC99FDE09461177C" box="[846,1062,549,575]" country="Vietnam" name="Cao Bang" pageId="3" pageNumber="255">Cao Bang province</collectingRegion>
|
||
in the northern part of
|
||
<collectingCountry id="F33C762E0C7B1E3DFAFEFDE095CB177C" box="[1321,1420,549,575]" name="Vietnam" pageId="3" pageNumber="255">Vietnam</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33165350C7B1E3DFF10FD8D927D1720" box="[199,570,584,611]" pageId="3" pageNumber="255" type="distribution">
|
||
<paragraph id="8B9436BE0C7B1E3DFF10FD8D927D1720" blockId="3.[151,1436,153,611]" box="[199,570,584,611]" pageId="3" pageNumber="255">
|
||
<emphasis id="B95FEAAC0C7B1E3DFF10FD8D91261720" bold="true" box="[199,353,584,611]" pageId="3" pageNumber="255">Distribution.</emphasis>
|
||
Northern
|
||
<collectingCountry id="F33C762E0C7B1E3DFE04FD8C92711720" box="[467,566,585,611]" name="Vietnam" pageId="3" pageNumber="255">Vietnam</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |