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<document id="8B4101DA5F4FBCC1AB7DF23EBAA3366E" ID-CLB-Dataset="9764" ID-DOI="10.5852/ejt.2020.720.1109" ID-GBIF-Dataset="545aa4ea-1874-4f74-a4c4-a2aeb97ed657" ID-ISSN="2118-9773" ID-Zenodo-Dep="4068355" ID-ZooBank="0CF83E89-2532-407B-921F-C05F9A29C68D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1601993099685" checkinUser="valdenar" docAuthor="Bergsten, Johannes, Ranarilalatiana, Tolotra &amp; Biström, Olof" docDate="2020" docId="03828798D3726460FDC3FE68AFD3F80A" docLanguage="en" docName="ejt-720_begsten_ranarilalatinana_bistroem.pdf.imf" docOrigin="European Journal of Taxonomy 720" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Bidessus anjozorobe Bergsten &amp; Ranarilalatiana &amp; Biström 2020, sp. nov." docType="treatment" docUuid="C6F11BD1-C6DC-41B6-8994-35C58E062999" docUuidSource="ZooBank" docVersion="10" lastPageNumber="9" masterDocId="FFBBFFE0D3756468FFAEFFBDAD21FFBB" masterDocTitle="A new species of Bidessus from Anjozorobe-Angavo and a review of Malagasy Bidessus (Coleoptera, Dytiscidae)" masterLastPageNumber="18" masterPageNumber="1" pageNumber="8" updateTime="1698792528866" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="F78A99F6A07FD357EE9A5B621A63D450" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="0061666199A64ED20516E8290D510079">
<mods:title id="E75DD147BE5072CFDD3A7ACF73FFAF23">A new species of Bidessus from Anjozorobe-Angavo and a review of Malagasy Bidessus (Coleoptera, Dytiscidae)</mods:title>
</mods:titleInfo>
<mods:name id="D1D4F18BDBF1CC241D8D32E9A223176D" type="personal">
<mods:role id="FE7E5AE8B5918655E59E116C78425923">
<mods:roleTerm id="263A5FB2DBC37D0FC30D3122EBD2A9C9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6791D4BEFB0089925A24DB2A708A64CD">Bergsten, Johannes</mods:namePart>
<mods:nameIdentifier id="B5DF285435D2E62C2E69437584FB193E" type="LSID">64120AE6-1D4B-4C29-AA37-46DA7C7A26A8</mods:nameIdentifier>
<mods:affiliation id="4622252FA38AE4214CF1857D1BADF577">Department of Zoology, Swedish Museum of Natural History, Box 50007, SE- 10405 Stockholm, Sweden. Department of Entomology, Faculty of Sciences, Box 906, University of Antananarivo, 101 Antananarivo, Madagascar. Finnish Museum of Natural History, Zoology Unit, P. O. Box 17, FI- 00014 University of Helsinki, Finland.</mods:affiliation>
<mods:nameIdentifier id="DCE90AE1B74B9AAD57628DA1B956BF89" type="email">johannes.bergsten@nrm.se</mods:nameIdentifier>
</mods:name>
<mods:name id="31DEBD3526B8A7F693E8F7DBE7EEDF60" type="personal">
<mods:role id="D6DBAC4FE2979718338C90B60DA6DDF0">
<mods:roleTerm id="319EBFB2D71300ABF5B511D2CE2122DC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="206BFE6C749C0567ACBFE741DB54A867">Ranarilalatiana, Tolotra</mods:namePart>
<mods:nameIdentifier id="D34CE706757454BBECEA47978306D3AA" type="LSID">4E61E817-7646-4993-A25A-78E2B160F5B8</mods:nameIdentifier>
<mods:affiliation id="3CAE69FFEF688B925C1DAC7F54C79D51">Department of Entomology, Faculty of Sciences, Box 906, University of Antananarivo, 101 Antananarivo, Madagascar.</mods:affiliation>
<mods:nameIdentifier id="3D9441EF84515990AE2FC61AF3D8BD69" type="email">t.ranarilalatiana@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="5A96F4A06F54857456B35795876A9AF0" type="personal">
<mods:role id="586E20A83FFA9B845E8B07A2430F563B">
<mods:roleTerm id="97C5DB624F9ECB1998A513CF306A636D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="697CCB4E60C83ABEA6BC01E8BA57D07D">Biström, Olof</mods:namePart>
<mods:nameIdentifier id="DDB45B326371E470DBF3DED979288090" type="LSID">2F25C707-9602-438B-BCCE-D7A9B7103BC1</mods:nameIdentifier>
<mods:affiliation id="C13ADA860C47DDF510A556BC66B75CB0">Finnish Museum of Natural History, Zoology Unit, P. O. Box 17, FI- 00014 University of Helsinki, Finland</mods:affiliation>
<mods:nameIdentifier id="962EA006C86ACB5DE7CEF60AF058B812" type="email">olof.bistrom@helsinki.fi</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="56004B6E9A491C18DBD0302553F5EC7F">text</mods:typeOfResource>
<mods:relatedItem id="B41D13CA97829D39A58A5BA454DF8197" type="host">
<mods:titleInfo id="69E1A7CB12F8DDD83603B1139C2207EE">
<mods:title id="CDB18615F980DCA588187D26075A2F50">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="9851DB7D8AD8781704EC2383C06585A5">
<mods:date id="44D1E59B63CC24D4131C43983765BA38">2020</mods:date>
<mods:detail id="E8AB9038C1B7803CADDD4D38FFF53768" type="pubDate">
<mods:number id="BAEC846D02C36F2D8E9616E0218762FE">2020-10-05</mods:number>
</mods:detail>
<mods:detail id="C2842884A99C9B6B889C93EDA243DBB0" type="volume">
<mods:number id="4D131FF296784C50341B9D19F2E2988D">720</mods:number>
</mods:detail>
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<mods:start id="556D25AE46ECB18B62A73EED278BD53B">1</mods:start>
<mods:end id="A0EB6EBD38C5ED9EF56E3CFAA19AEAB2">18</mods:end>
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<mods:classification id="AF00EDC834BA3BA3A602B9F8789A6399">journal article</mods:classification>
<mods:identifier id="4A58BA66D0B192F89BD71DBF8332CC2B" type="CLB-Dataset">9764</mods:identifier>
<mods:identifier id="B8C9E450CB3408119EB34A29AEF33BED" type="DOI">10.5852/ejt.2020.720.1109</mods:identifier>
<mods:identifier id="2C7385E6307532B47FEB1D63DB71E4CF" type="GBIF-Dataset">545aa4ea-1874-4f74-a4c4-a2aeb97ed657</mods:identifier>
<mods:identifier id="952EDA5418FC85921790332BEFB76B48" type="ISSN">2118-9773</mods:identifier>
<mods:identifier id="AEE6A0F3812D7803A1CAF5AB31CDD46B" type="Zenodo-Dep">4068355</mods:identifier>
<mods:identifier id="62BBF7B0B402DD2CD5DEB2D05776B547" type="ZooBank">0CF83E89-2532-407B-921F-C05F9A29C68D</mods:identifier>
</mods:mods>
<treatment id="03828798D3726460FDC3FE68AFD3F80A" ID-DOI="http://doi.org/10.5281/zenodo.4329747" ID-GBIF-Taxon="168237712" ID-Zenodo-Dep="4329747" LSID="urn:lsid:zoobank.org:act:C6F11BD1-C6DC-41B6-8994-35C58E062999" httpUri="http://treatment.plazi.org/id/03828798D3726460FDC3FE68AFD3F80A" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
<subSubSection id="C3316505D372646FFDC3FE68AEE7FE4B" box="[621,966,469,496]" pageId="7" pageNumber="8" type="nomenclature">
<paragraph id="8B94368ED372646FFDC3FE68AEE7FE4B" blockId="7.[370,1217,469,566]" box="[621,966,469,496]" pageId="7" pageNumber="8">
<heading id="D0DC81E2D372646FFDC3FE68AEE7FE4B" box="[621,966,469,496]" centered="true" fontSize="11" level="2" pageId="7" pageNumber="8" reason="2">
<taxonomicName id="4C2B4D0DD372646FFDC3FE68AE44FE4B" ID-CoL="8LZJJ" authority="Bergsten &amp; Ranarilalatiana &amp; Biström, 2020" authorityName="Bergsten &amp; Ranarilalatiana &amp; Biström" authorityYear="2020" box="[621,869,469,496]" class="Insecta" family="Dytiscidae" genus="Bidessus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="anjozorobe" status="sp. nov.">
<emphasis id="B95FEA9CD372646FFDC3FE68AE44FE4B" bold="true" box="[621,869,469,496]" italics="true" pageId="7" pageNumber="8">Bidessus anjozorobe</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26C57E7D372646FFCC2FE6BAEE7FE4B" box="[876,966,470,496]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3316505D372646FFEDCFE44AE41FD8D" pageId="7" pageNumber="8" type="description">
<paragraph id="8B94368ED372646FFEDCFE44A9E0FDAF" blockId="7.[370,1217,469,566]" box="[370,1217,505,532]" pageId="7" pageNumber="8">
<uri id="FFBA3A8CD372646FFEDCFE44A9E0FDAF" box="[370,1217,505,532]" pageId="7" pageNumber="8">
urn:lsid:zoobank.org:act:
<uuid id="FF8D0C5BD372646FFD3BFE44A9E0FDAF" box="[661,1217,505,532]" pageId="7" pageNumber="8">
C6F11BD1-C6DC-41B6-8994-35C58E
<date id="FF95104ED372646FFBCCFE44A9E0FDAF" box="[1122,1217,505,532]" pageId="7" pageNumber="8" value="99-06-29">062999</date>
</uuid>
</uri>
</paragraph>
<paragraph id="8B94368ED372646FFD7DFDA1AE41FD8D" blockId="7.[370,1217,469,566]" box="[723,864,540,566]" pageId="7" pageNumber="8">
<figureCitation id="13102A0BD372646FFD7DFDA1AE10FD8D" box="[723,817,540,566]" captionStart="Fig" captionStartId="3.[189,232,1887,1913]" captionTargetBox="[233,1355,265,1851]" captionTargetId="figure-11@3.[233,1355,265,1851]" captionTargetPageId="3" captionText="Fig. 1. Habitus photographs of Malagasy species of Bidessus Sharp, 1882. A. B. apicidens Bistr̂m &amp; Sanfilippo, 1986, ♀, paratype, http://id.luomus.fi/GL.8408 (FMNH). B. B. cf. ceratus Guignot, 1941, http://id.luomus.fi/GC.2001 (FMNH). C. B. anjozorobe sp. nov., paratype (NHRS-JLKB000025034). D. B. cf. nero Gschwendtner, 1933 (NHRS-JLKB000025032)." figureDoi="http://doi.org/10.5281/zenodo.4068357" httpUri="https://zenodo.org/record/4068357/files/figure.png" pageId="7" pageNumber="8">Figs 1C</figureCitation>
,
<figureCitation id="13102A0BD372646FFC90FDA1AE41FD8D" box="[830,864,540,566]" captionStart="Fig" captionStartId="9.[189,232,1888,1914]" captionTargetBox="[366,1209,282,1833]" captionTargetId="figure-11@9.[308,1280,265,1849]" captionTargetPageId="9" captionText="Fig. 3. Male genitalia of the Malagasy species of Bidessus Sharp, 1882. For each subpanel from left to right penis in dorsal view, penis in lateral view and paramere in lateral view. A. B. apicidens Bistr̂m &amp; Sanfilippo, 1986. B. B. cf. ceratus Guignot, 1941. C. B. anjozorobe sp. nov. D. B. cf. nero Gschwendtner, 1933. E. B. nesioticus Guignot, 1956. F. B. perexiguus Kolbe, 1883. G. B. longistriga Régimbart, 1895." figureDoi="http://doi.org/10.5281/zenodo.4068361" httpUri="https://zenodo.org/record/4068361/files/figure.png" pageId="7" pageNumber="8">3C</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3316505D372646FFF13FDE0ACB6FBF6" pageId="7" pageNumber="8" type="diagnosis">
<paragraph id="8B94368ED372646FFF13FDE0ACEDFDCC" blockId="7.[189,460,605,631]" box="[189,460,605,631]" pageId="7" pageNumber="8">
<heading id="D0DC81E2D372646FFF13FDE0ACEDFDCC" bold="true" box="[189,460,605,631]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="B95FEA9CD372646FFF13FDE0ACEDFDCC" bold="true" box="[189,460,605,631]" pageId="7" pageNumber="8">Diagnostic characters</emphasis>
</heading>
</paragraph>
<paragraph id="8B94368ED372646FFF13FD31ACB6FBF6" blockId="7.[189,1399,652,1101]" pageId="7" pageNumber="8">
Head slightly posterior to frontal edge with a shallow, transverse depression, which medially is vague. Pronotum discally with broad area with fine, sparse and irregular punctures. Pronotal striae at base strongly impressed; angled (not smoothly curved). Basal striae of elytra medium long, straight (close to base, strongly curved outwards); at base quite strongly impressed. Sutural lines close to frontal edge of elytra distinct; posteriorly lines gradually fade away. Female with outline of elytra posteriorly smooth (minor process absent). Penis in lateral view curved, medially slightly straightened. Penis apex with a long and slender extension (
<figureCitation id="13102A0BD372646FFDA2FCE2AF48FCC2" box="[524,617,863,889]" captionStart="Fig" captionStartId="9.[189,232,1888,1914]" captionTargetBox="[366,1209,282,1833]" captionTargetId="figure-11@9.[308,1280,265,1849]" captionTargetPageId="9" captionText="Fig. 3. Male genitalia of the Malagasy species of Bidessus Sharp, 1882. For each subpanel from left to right penis in dorsal view, penis in lateral view and paramere in lateral view. A. B. apicidens Bistr̂m &amp; Sanfilippo, 1986. B. B. cf. ceratus Guignot, 1941. C. B. anjozorobe sp. nov. D. B. cf. nero Gschwendtner, 1933. E. B. nesioticus Guignot, 1956. F. B. perexiguus Kolbe, 1883. G. B. longistriga Régimbart, 1895." figureDoi="http://doi.org/10.5281/zenodo.4068361" httpUri="https://zenodo.org/record/4068361/files/figure.png" pageId="7" pageNumber="8">Fig. 3C</figureCitation>
), a feature which separates
<taxonomicName id="4C2B4D0DD372646FFC1BFCDDA97BFCC2" authorityName="Bergsten &amp; Ranarilalatiana &amp; Biström" authorityYear="2020" box="[949,1114,863,890]" class="Insecta" family="Dytiscidae" genus="Bidessus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="anjozorobe">
<emphasis id="B95FEA9CD372646FFC1BFCDDAEE6FCC2" box="[949,967,864,889]" italics="true" pageId="7" pageNumber="8">B</emphasis>
.
<emphasis id="B95FEA9CD372646FFC79FCE2A97BFCC2" box="[983,1114,863,889]" italics="true" pageId="7" pageNumber="8">anjozorobe</emphasis>
</taxonomicName>
from all other
<taxonomicName id="4C2B4D0DD372646FFABEFCE2A856FCC2" authorityName="Sharp" authorityYear="1882" box="[1296,1399,863,889]" class="Insecta" family="Dytiscidae" genus="Bidessus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B95FEA9CD372646FFABEFCE2A856FCC2" box="[1296,1399,863,889]" italics="true" pageId="7" pageNumber="8">Bidessus</emphasis>
</taxonomicName>
species known from
<collectingCountry id="F33C761ED372646FFE1BFC3FAF64FC27" box="[437,581,898,924]" name="Madagascar" pageId="7" pageNumber="8">Madagascar</collectingCountry>
. Considering morphology
<taxonomicName id="4C2B4D0DD372646FFC2CFC3EA906FC27" authorityName="Bergsten &amp; Ranarilalatiana &amp; Biström" authorityYear="2020" box="[898,1063,898,925]" class="Insecta" family="Dytiscidae" genus="Bidessus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="anjozorobe">
<emphasis id="B95FEA9CD372646FFC2CFC3EAEB5FC27" box="[898,916,899,924]" italics="true" pageId="7" pageNumber="8">B</emphasis>
.
<emphasis id="B95FEA9CD372646FFC0AFC3FA906FC27" box="[932,1063,898,924]" italics="true" pageId="7" pageNumber="8">anjozorobe</emphasis>
</taxonomicName>
most resembles
<taxonomicName id="4C2B4D0DD372646FFB41FC3EADDAFC04" authority="Bistr" authorityName="Bistr" class="Insecta" family="Dytiscidae" genus="Bidessus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="perrinae">
<emphasis id="B95FEA9CD372646FFB41FC3EA857FC27" box="[1263,1398,899,924]" italics="true" pageId="7" pageNumber="8">B. perrinae</emphasis>
Bistr
</taxonomicName>
̂m, 1985 thus far known from
<collectingCountry id="F33C761ED372646FFDDDFC18AFECFC04" box="[627,717,933,959]" name="Angola" pageId="7" pageNumber="8">Angola</collectingCountry>
. The two species are separated as follows: elytra almost unicolored dark, black to dark piceous in
<taxonomicName id="4C2B4D0DD372646FFD0AFC74AE66FC58" authorityName="Bergsten &amp; Ranarilalatiana &amp; Biström" authorityYear="2020" box="[676,839,969,995]" class="Insecta" family="Dytiscidae" genus="Bidessus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="anjozorobe">
<emphasis id="B95FEA9CD372646FFD0AFC74AF97FC59" box="[676,694,969,994]" italics="true" pageId="7" pageNumber="8">B</emphasis>
.
<emphasis id="B95FEA9CD372646FFD6AFC74AE66FC58" box="[708,839,969,995]" italics="true" pageId="7" pageNumber="8">anjozorobe</emphasis>
</taxonomicName>
while distinctly bicolored in
<taxonomicName id="4C2B4D0DD372646FFB31FC74A804FC59" box="[1183,1317,969,994]" class="Insecta" family="Dytiscidae" genus="Bidessus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="perrinae">
<emphasis id="B95FEA9CD372646FFB31FC74A804FC59" box="[1183,1317,969,994]" italics="true" pageId="7" pageNumber="8">B. perrinae</emphasis>
</taxonomicName>
, elytra being brownish to dark ferrugineous with distinct testaceous markings. Furthermore, body slightly larger and more robust in
<taxonomicName id="4C2B4D0DD372646FFE0FFBADAF64FB92" authorityName="Bergsten &amp; Ranarilalatiana &amp; Biström" authorityYear="2020" box="[417,581,1039,1066]" class="Insecta" family="Dytiscidae" genus="Bidessus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="anjozorobe">
<emphasis id="B95FEA9CD372646FFE0FFBADAC92FB92" box="[417,435,1040,1065]" italics="true" pageId="7" pageNumber="8">B</emphasis>
.
<emphasis id="B95FEA9CD372646FFE6CFBB2AF64FB92" box="[450,581,1039,1065]" italics="true" pageId="7" pageNumber="8">anjozorobe</emphasis>
</taxonomicName>
than in
<taxonomicName id="4C2B4D0DD372646FFD0AFBADAE0BFB92" box="[676,810,1040,1065]" class="Insecta" family="Dytiscidae" genus="Bidessus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="perrinae">
<emphasis id="B95FEA9CD372646FFD0AFBADAE0BFB92" box="[676,810,1040,1065]" italics="true" pageId="7" pageNumber="8">B. perrinae</emphasis>
</taxonomicName>
(length of body
<quantity id="4CD39B6BD372646FFC40FBB2A96CFB91" box="[1006,1101,1039,1066]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="7" pageNumber="8" unit="mm" value="1.6">1.6 mm</quantity>
). See original description in Bistr̂m (1985).
</paragraph>
</subSubSection>
<subSubSection id="C3316505D372646FFF13FBC9AFCBFB5B" pageId="7" pageNumber="8" type="etymology">
<paragraph id="8B94368ED372646FFF13FBC9AC65FB35" blockId="7.[189,324,1140,1166]" box="[189,324,1140,1166]" pageId="7" pageNumber="8">
<heading id="D0DC81E2D372646FFF13FBC9AC65FB35" bold="true" box="[189,324,1140,1166]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="B95FEA9CD372646FFF13FBC9AC65FB35" bold="true" box="[189,324,1140,1166]" pageId="7" pageNumber="8">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8B94368ED372646FFF13FB1FAFCBFB5B" blockId="7.[189,1399,1186,1248]" pageId="7" pageNumber="8">
Named after the
<typeStatus id="5490882CD372646FFE27FB1EAC9DFB06" box="[393,444,1187,1213]" pageId="7" pageNumber="8">type</typeStatus>
locality Anjozorobe forest, part of the Anjozorobe-Angavo protected area. The epithet is a noun in apposition (ICZN 11.9.1.2).
</paragraph>
</subSubSection>
<subSubSection id="C3316505D372646FFF13FABAAC93F870" pageId="7" pageNumber="8" type="materials_examined">
<paragraph id="8B94368ED372646FFF13FABAAC4DFA9A" blockId="7.[189,364,1287,1313]" box="[189,364,1287,1313]" pageId="7" pageNumber="8">
<heading id="D0DC81E2D372646FFF13FABAAC4DFA9A" bold="true" box="[189,364,1287,1313]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="B95FEA9CD372646FFF13FABAAC4DFA9A" bold="true" box="[189,364,1287,1313]" pageId="7" pageNumber="8">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="8B94368ED372646FFF44FA8BAC7DFAEB" blockId="7.[189,1398,1334,1501]" box="[234,348,1334,1360]" pageId="7" pageNumber="8">
<emphasis id="B95FEA9CD372646FFF44FA8BAC7DFAEB" bold="true" box="[234,348,1334,1360]" pageId="7" pageNumber="8">
<typeStatus id="5490882CD372646FFF44FA8BAC7DFAEB" box="[234,348,1334,1360]" pageId="7" pageNumber="8" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8B94368ED372646FFF13FAE4A9D4FA66" blockId="7.[189,1398,1334,1501]" pageId="7" pageNumber="8">
<materialsCitation id="3B433CD3D372646FFF13FAE4A9D4FA66" ID-GBIF-Occurrence="2871419309" collectingDate="2014-11-23" collectorName="J. Bergsten &amp; R. Bukontaite &amp; S. Holmgren &amp; J. H. Randriamihaja &amp; T. Ranarilalatiana" country="Madagascar" county="Analamanga region" elevation="1320" latitude="-18.4128" location="Saha forest" longLatPrecision="7" longitude="47.9439" municipality="Anjozorobe-Angavo" pageId="7" pageNumber="8" specimenCount="1" stateProvince="Antananarivo" typeStatus="holotype">
<collectingCountry id="F33C761ED372646FFF13FAE4ACAFFAC8" box="[189,398,1369,1395]" name="Madagascar" pageId="7" pageNumber="8">MADAGASCAR</collectingCountry>
• 1 ♂;
<collectingRegion id="49EFF86CD372646FFE49FAE4AFA6FAC8" box="[487,647,1369,1395]" country="Madagascar" name="Antananarivo" pageId="7" pageNumber="8">Antananarivo</collectingRegion>
former province,
<collectingCounty id="62F54E02D372646FFCF3FAE4A969FAC8" box="[861,1096,1369,1395]" pageId="7" pageNumber="8">Analamanga region</collectingCounty>
,
<locationDeviation id="9A262FACD372646FFBFBFAE4A852FAC8" box="[1109,1395,1369,1395]" location="Anjozorobe" pageId="7" pageNumber="8">
<quantity id="4CD39B6BD372646FFBFBFAE4A980FAC8" box="[1109,1185,1369,1395]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="8" unit="km" value="10.0">10 km</quantity>
E of Anjozorobe
</locationDeviation>
,
<collectingMunicipality id="6BF0ACF4D372646FFF13FAC1AC8FFA2D" box="[189,430,1404,1430]" pageId="7" pageNumber="8">Anjozorobe-Angavo</collectingMunicipality>
protected area, marsh next to the stream in
<location id="8EF46055D372646FFC10FAC1A966FA2D" LSID="urn:lsid:plazi:treatment:03828798D3726460FDC3FE68AFD3F80A:8EF46055D372646FFC10FAC1A966FA2D" box="[958,1095,1404,1430]" country="Madagascar" county="Analamanga region" latitude="-18.4128" longLatPrecision="7" longitude="47.9439" municipality="Anjozorobe-Angavo" name="Saha forest" pageId="7" pageNumber="8" stateProvince="Antananarivo">Saha forest</location>
;
<geoCoordinate id="EE1F5049D372646FFBFAFAC1A9FBFA2D" box="[1108,1242,1404,1430]" degrees="18.4128" direction="south" orientation="latitude" pageId="7" pageNumber="8" precision="5" value="-18.4128">18.4128° S</geoCoordinate>
,
<geoCoordinate id="EE1F5049D372646FFB46FAC1A850FA2D" box="[1256,1393,1404,1430]" degrees="47.9439" direction="east" orientation="longitude" pageId="7" pageNumber="8" precision="5" value="47.9439">47.9439° E</geoCoordinate>
;
<elevation id="0006D1BDD372646FFF13FA22AC44FA02" box="[189,357,1439,1466]" metricMagnitude="3" metricUnit="m" metricValue="1.3199999999999998" pageId="7" pageNumber="8" unit="m" value="1320.0">
<quantity id="4CD39B6BD372646FFF13FA22AC3FFA01" box="[189,286,1439,1466]" metricMagnitude="3" metricUnit="m" metricValue="1.3199999999999998" pageId="7" pageNumber="8" unit="m" value="1320.0">1320 m</quantity>
a.s.l.
</elevation>
;
<date id="FF95104ED372646FFED4FA22AF0AFA02" box="[378,555,1439,1466]" pageId="7" pageNumber="8" value="2014-11-23">
<collectingDate id="EFD1E9A6D372646FFED4FA22AF0AFA02" box="[378,555,1439,1466]" pageId="7" pageNumber="8" value="2014-11-23">23 Nov. 2014</collectingDate>
</date>
;
<collectorName id="26DE5358D372646FFD91FA1DAFEAFA01" box="[575,715,1440,1466]" pageId="7" pageNumber="8">J. Bergsten</collectorName>
,
<collectorName id="26DE5358D372646FFD70FA1DAEADFA02" box="[734,908,1439,1466]" pageId="7" pageNumber="8">R. Bukontaite</collectorName>
,
<collectorName id="26DE5358D372646FFC31FA22A91EFA02" box="[927,1087,1439,1465]" pageId="7" pageNumber="8">S. Holmgren</collectorName>
,
<collectorName id="26DE5358D372646FFBFCFA1DA81FFA02" box="[1106,1342,1439,1466]" pageId="7" pageNumber="8">J.H. Randriamihaja</collectorName>
and
<collectorName id="26DE5358D372646FFF13FA7EACACFA67" box="[189,397,1474,1501]" pageId="7" pageNumber="8">T. Ranarilalatiana</collectorName>
leg.; collecting event MAD14-78; forest marsh; NHRS-JLKB000025033.
</materialsCitation>
</paragraph>
<paragraph id="8B94368ED372646FFF44F9B9AC46F9A5" blockId="7.[189,1399,1540,1848]" box="[234,359,1540,1566]" pageId="7" pageNumber="8">
<emphasis id="B95FEA9CD372646FFF44F9B9AC46F9A5" bold="true" box="[234,359,1540,1566]" pageId="7" pageNumber="8">
<typeStatus id="5490882CD372646FFF44F9B9AC46F9A5" box="[234,359,1540,1566]" pageId="7" pageNumber="8" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8B94368ED372646FFF13F99AAEC7F883" blockId="7.[189,1399,1540,1848]" pageId="7" pageNumber="8">
<materialsCitation id="3B433CD3D372646FFF13F99AA9D2F9F9" ID-GBIF-Occurrence="2871419310" box="[189,1267,1575,1602]" collectingDate="2014-11-23" collectorName="J. Bergsten &amp; R. Bukontaite &amp; S. Holmgren &amp; J. H. Randriamihaja &amp; T. Ranarilalatiana" country="Madagascar" county="Analamanga region" elevation="1320" latitude="-18.4128" location="Saha forest" longLatPrecision="7" longitude="47.9439" municipality="Anjozorobe-Angavo" pageId="7" pageNumber="8" specimenCount="1" stateProvince="Antananarivo" typeStatus="holotype">
<collectingCountry id="F33C761ED372646FFF13F99AACAFF9FA" box="[189,398,1575,1601]" name="Madagascar" pageId="7" pageNumber="8">MADAGASCAR</collectingCountry>
•
<specimenCount id="9D2DFD07D372646FFE04F99AAF20F9FA" box="[426,513,1575,1601]" pageId="7" pageNumber="8" type="generic">1 spec.</specimenCount>
; same collecting data as for
<typeStatus id="5490882CD372646FFCF7F99AAEE2F9FA" box="[857,963,1575,1601]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
; NHRS-JLKB000025034
</materialsCitation>
•
<materialsCitation id="3B433CD3D372646FFABEF99AA997F9DE" ID-GBIF-Occurrence="2871419312" collectingDate="2014-11-23" collectorName="J. Bergsten &amp; R. Bukontaite &amp; S. Holmgren &amp; J. H. Randriamihaja &amp; T. Ranarilalatiana" country="Madagascar" county="Analamanga region" elevation="1320" latitude="-18.4128" location="Saha forest" longLatPrecision="7" longitude="47.9439" municipality="Anjozorobe-Angavo" pageId="7" pageNumber="8" specimenCount="1" stateProvince="Antananarivo" typeStatus="holotype">
27 specs (preserved in ethanol); same collecting data as for
<typeStatus id="5490882CD372646FFCB5F9F7AEA4F9DF" box="[795,901,1610,1636]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
; NHRS-JLKB000025027
</materialsCitation>
•
<materialsCitation id="3B433CD3D372646FFB7CF9F7AFA1F933" ID-GBIF-Occurrence="2871419319" collectingDate="2014-11-23" collectionCode="FMNH" collectorName="J. Bergsten &amp; R. Bukontaite &amp; S. Holmgren &amp; J. H. Randriamihaja &amp; T. Ranarilalatiana" country="Madagascar" county="Analamanga region" elevation="1320" latitude="-18.4128" location="Saha forest" longLatPrecision="7" longitude="47.9439" municipality="Anjozorobe-Angavo" pageId="7" pageNumber="8" specimenCount="1" stateProvince="Antananarivo" typeStatus="holotype">
5 specs; same collecting data as for
<typeStatus id="5490882CD372646FFE18F9D0AF01F93C" box="[438,544,1645,1671]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
;
<collectionCode id="ED3AAE4BD372646FFD85F9D3AFA1F933" box="[555,640,1646,1672]" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="7" pageNumber="8" type="Museum">FMNH</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B433CD3D372646FFD39F9D0A822F933" ID-GBIF-Occurrence="2871419317" box="[663,1283,1645,1672]" collectingDate="2014-11-23" collectionCode="BMNH" collectorName="J. Bergsten &amp; R. Bukontaite &amp; S. Holmgren &amp; J. H. Randriamihaja &amp; T. Ranarilalatiana" country="Madagascar" county="Analamanga region" elevation="1320" latitude="-18.4128" location="Saha forest" longLatPrecision="7" longitude="47.9439" municipality="Anjozorobe-Angavo" pageId="7" pageNumber="8" specimenCount="1" stateProvince="Antananarivo" typeStatus="holotype">
5 specs; same collecting data as for
<typeStatus id="5490882CD372646FFB9BF9D0A9BEF93C" box="[1077,1183,1645,1671]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
;
<collectionCode id="ED3AAE4BD372646FFB05F9D3A822F933" box="[1195,1283,1646,1672]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="7" pageNumber="8">BMNH</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B433CD3D372646FFAB4F9D0AE01F910" ID-GBIF-Occurrence="2871419307" collectingDate="2014-11-23" collectionCode="PBZT, MBC" collectorName="J. Bergsten &amp; R. Bukontaite &amp; S. Holmgren &amp; J. H. Randriamihaja &amp; T. Ranarilalatiana" country="Madagascar" county="Analamanga region" elevation="1320" latitude="-18.4128" location="Saha forest" longLatPrecision="7" longitude="47.9439" municipality="Anjozorobe-Angavo" pageId="7" pageNumber="8" specimenCount="1" stateProvince="Antananarivo" typeStatus="holotype">
5 specs; same collecting data as for
<typeStatus id="5490882CD372646FFDBAF92DAF5FF911" box="[532,638,1680,1706]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
;
<collectionCode id="ED3AAE4BD372646FFD21F92CAFF6F910" box="[655,727,1681,1707]" collectionName="Madagascar, Antananarivo, Parc Botanique et Zoologique de Tsimbazaza" country="Madagascar" httpUri="http://grbio.org/cool/a6tz-5rcd" name="Parc Botanique et Zoologique de Tsimbazaza" pageId="7" pageNumber="8">PBZT</collectionCode>
/
<collectionCode id="ED3AAE4BD372646FFD4EF92CAE01F910" box="[736,800,1681,1707]" country="USA" name="Montgomery Botanical Center" pageId="7" pageNumber="8">MBC</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B433CD3D372646FFCECF92DAC2CF975" ID-GBIF-Occurrence="2871419315" collectingDate="2014-11-23" collectionCode="DEUA" collectorName="J. Bergsten &amp; R. Bukontaite &amp; S. Holmgren &amp; J. H. Randriamihaja &amp; T. Ranarilalatiana" country="Madagascar" county="Analamanga region" elevation="1320" latitude="-18.4128" location="Saha forest" longLatPrecision="7" longitude="47.9439" municipality="Anjozorobe-Angavo" pageId="7" pageNumber="8" specimenCount="1" stateProvince="Antananarivo" typeStatus="holotype">
5 specs; same collecting data as for
<typeStatus id="5490882CD372646FFAA6F92DA853F911" box="[1288,1394,1680,1706]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
;
<collectionCode id="ED3AAE4BD372646FFF13F909AC2CF975" box="[189,269,1716,1742]" pageId="7" pageNumber="8">DEUA</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B433CD3D372646FFE88F909A844F8AE" ID-GBIF-Occurrence="2871419320" collectingDate="2016-04-05" collectorName="T. Ranarilalatiana" country="Madagascar" county="Analamanga region" elevation="1341" latitude="-18.41266" location="Antsahabe" longLatPrecision="5" longitude="47.9438" municipality="Anjozorobe Angavo" pageId="7" pageNumber="8" specimenCount="1" stateProvince="Antananarivo" typeStatus="paratype">
<specimenCount id="9D2DFD07D372646FFE88F909AC75F975" box="[294,340,1714,1742]" pageId="7" pageNumber="8" type="male">1 ♂</specimenCount>
;
<collectingRegion id="49EFF86CD372646FFECFF909AF20F975" box="[353,513,1716,1742]" country="Madagascar" name="Antananarivo" pageId="7" pageNumber="8">Antananarivo</collectingRegion>
former province,
<collectingCounty id="62F54E02D372646FFD7AF909AE9FF975" box="[724,958,1716,1742]" pageId="7" pageNumber="8">Analamanga region</collectingCounty>
,
<locationDeviation id="9A262FACD372646FFC64F909A9C3F975" box="[970,1250,1716,1742]" location="Anjozorobe" pageId="7" pageNumber="8">
<quantity id="4CD39B6BD372646FFC64F909A935F975" box="[970,1044,1716,1742]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="8" unit="km" value="10.0">10 km</quantity>
E of Anjozorobe
</locationDeviation>
,
<collectingMunicipality id="6BF0ACF4D372646FFB42F909AC3BF94A" pageId="7" pageNumber="8">Anjozorobe Angavo</collectingMunicipality>
protected area,
<location id="8EF46055D372646FFE7EF96AAF6EF94A" LSID="urn:lsid:plazi:treatment:03828798D3726460FDC3FE68AFD3F80A:8EF46055D372646FFE7EF96AAF6EF94A" box="[464,591,1751,1777]" country="Madagascar" county="Analamanga region" latitude="-18.41266" longLatPrecision="5" longitude="47.9438" municipality="Anjozorobe Angavo" name="Antsahabe" pageId="7" pageNumber="8" stateProvince="Antananarivo">Antsahabe</location>
, circuit bamboo;
<geoCoordinate id="EE1F5049D372646FFCB8F96AAE89F94A" box="[790,936,1751,1777]" degrees="18.41266" direction="south" orientation="latitude" pageId="7" pageNumber="8" precision="1" value="-18.41266">18.41266° S</geoCoordinate>
,
<geoCoordinate id="EE1F5049D372646FFC1AF96AA91BF94A" box="[948,1082,1751,1777]" degrees="47.9438" direction="east" orientation="longitude" pageId="7" pageNumber="8" precision="5" value="47.9438">47.9438° E</geoCoordinate>
;
<elevation id="0006D1BDD372646FFBEBF96AA9FAF94A" box="[1093,1243,1751,1777]" metricMagnitude="3" metricUnit="m" metricValue="1.341" pageId="7" pageNumber="8" unit="m" value="1341.0">
<quantity id="4CD39B6BD372646FFBEBF96AA9BCF94A" box="[1093,1181,1751,1777]" metricMagnitude="3" metricUnit="m" metricValue="1.341" pageId="7" pageNumber="8" unit="m" value="1341.0">1341 m</quantity>
a.s.l.
</elevation>
;
<date id="FF95104ED372646FFB49F96AA853F94A" box="[1255,1394,1751,1778]" pageId="7" pageNumber="8" value="2016-04-05">
<collectingDate id="EFD1E9A6D372646FFB49F96AA853F94A" box="[1255,1394,1751,1778]" pageId="7" pageNumber="8" value="2016-04-05">5 Apr. 2016</collectingDate>
</date>
;
<collectorName id="26DE5358D372646FFF13F946ACACF8AF" box="[189,397,1786,1813]" pageId="7" pageNumber="8">T. Ranarilalatiana</collectorName>
leg.; collecting event MAD16-40; marsh and stream pool; NHRS-JLKB000065489
</materialsCitation>
•
<materialsCitation id="3B433CD3D372646FFF13F8A0AEC3F883" ID-GBIF-Occurrence="2871419306" box="[189,994,1821,1848]" collectingDate="2016-04-05" collectorName="T. Ranarilalatiana" country="Madagascar" county="Analamanga region" elevation="1341" latitude="-18.41266" location="Antsahabe" longLatPrecision="5" longitude="47.9438" municipality="Anjozorobe Angavo" pageId="7" pageNumber="8" specimenCount="1" specimenCount-female="1" stateProvince="Antananarivo" typeStatus="paratype">
<specimenCount id="9D2DFD07D372646FFF13F8A0ADCBF883" box="[189,234,1821,1848]" pageId="7" pageNumber="8" type="female">1 ♀</specimenCount>
; same collecting data as for preceding; NHRS-JLKB000065490
</materialsCitation>
.
</paragraph>
<paragraph id="8B94368ED372646FFF13F8E2AC7CF8C3" blockId="7.[189,349,1886,1913]" box="[189,349,1886,1913]" pageId="7" pageNumber="8">
<heading id="D0DC81E2D372646FFF13F8E2AC7CF8C3" bold="true" box="[189,349,1886,1913]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="B95FEA9CD372646FFF13F8E2AC7CF8C3" bold="true" box="[189,349,1886,1913]" pageId="7" pageNumber="8">Type locality</emphasis>
</heading>
</paragraph>
<paragraph id="8B94368ED372646FFF13F830AC93F870" blockId="7.[189,1399,1933,1995]" pageId="7" pageNumber="8">
<collectingCountry id="F33C761ED372646FFF13F830AC6CF81C" box="[189,333,1933,1959]" name="Madagascar" pageId="7" pageNumber="8">Madagascar</collectingCountry>
,
<collectingRegion id="49EFF86CD372646FFEF8F830ACD8F81C" box="[342,505,1933,1959]" country="Madagascar" name="Antananarivo" pageId="7" pageNumber="8">Antananarivo</collectingRegion>
, Analamanga, Anjozorobe-Angavo Protected Area, Saha forest (
<geoCoordinate id="EE1F5049D372646FFB45F830A850F81C" box="[1259,1393,1933,1959]" degrees="18.4128" direction="south" orientation="latitude" pageId="7" pageNumber="8" precision="5" value="-18.4128">18.4128° S</geoCoordinate>
,
<geoCoordinate id="EE1F5049D372646FFF13F80CAC63F870" box="[189,322,1969,1995]" degrees="47.9439" direction="east" orientation="longitude" pageId="7" pageNumber="8" precision="5" value="47.9439">47.9439° E</geoCoordinate>
) (
<figureCitation id="13102A0BD372646FFEF5F80DAC82F870" box="[347,419,1968,1995]" captionStart="Fig" captionStartId="12.[189,232,1888,1914]" captionTargetBox="[189,1398,942,1849]" captionTargetId="figure-106@12.[189,1398,942,1849]" captionTargetPageId="12" captionText="Fig. 5. Type locality [MAD14-78] for Bidessus anjozorobe sp. nov. in Anjozorobe forest in Anjozorobe- Angavo protected area on the Central Highlands. A glade in the rainforest with marsh vegetation. Multiple individuals of Bidessus anjozorobe sp. nov. were found by tramping in the vegetation at the edges of the marsh." figureDoi="http://doi.org/10.5281/zenodo.4068365" httpUri="https://zenodo.org/record/4068365/files/figure.png" pageId="7" pageNumber="8">Fig. 5</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3316505D37D6460FF13FEB5AE90F99A" pageId="8" pageNumber="9" type="description">
<paragraph id="8B94368ED37D6460FF13FEB5AC6FFE99" blockId="8.[189,334,264,290]" box="[189,334,264,290]" pageId="8" pageNumber="9">
<heading id="D0DC81E2D37D6460FF13FEB5AC6FFE99" bold="true" box="[189,334,264,290]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="B95FEA9CD37D6460FF13FEB5AC6FFE99" bold="true" box="[189,334,264,290]" pageId="8" pageNumber="9">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8B94368ED37D6460FF44FE8AAC0BFEEA" blockId="8.[189,1389,311,373]" box="[234,298,311,337]" pageId="8" pageNumber="9">
<emphasis id="B95FEA9CD37D6460FF44FE8AAC0BFEEA" bold="true" box="[234,298,311,337]" pageId="8" pageNumber="9">Male</emphasis>
</paragraph>
<paragraph id="8B94368ED37D6460FF13FEE6A84CFECF" blockId="8.[189,1389,311,373]" box="[189,1389,346,373]" pageId="8" pageNumber="9">
BODY. Length
<quantity id="4CD39B6BD37D6460FEC8FEE7ACD7FECE" box="[358,502,346,373]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="1.8" metricValueMin="1.7" pageId="8" pageNumber="9" unit="mm" value="1.75" valueMax="1.8" valueMin="1.7">1.7–1.8 mm</quantity>
. Dorsal aspect as in
<figureCitation id="13102A0BD37D6460FD48FEE7AE61FECF" box="[742,832,346,372]" captionStart="Fig" captionStartId="3.[189,232,1887,1913]" captionTargetBox="[233,1355,265,1851]" captionTargetId="figure-11@3.[233,1355,265,1851]" captionTargetPageId="3" captionText="Fig. 1. Habitus photographs of Malagasy species of Bidessus Sharp, 1882. A. B. apicidens Bistr̂m &amp; Sanfilippo, 1986, ♀, paratype, http://id.luomus.fi/GL.8408 (FMNH). B. B. cf. ceratus Guignot, 1941, http://id.luomus.fi/GC.2001 (FMNH). C. B. anjozorobe sp. nov., paratype (NHRS-JLKB000025034). D. B. cf. nero Gschwendtner, 1933 (NHRS-JLKB000025032)." figureDoi="http://doi.org/10.5281/zenodo.4068357" httpUri="https://zenodo.org/record/4068357/files/figure.png" pageId="8" pageNumber="9">Fig. 1C</figureCitation>
. Angle between pronotum and elytra indistinct.
</paragraph>
<paragraph id="8B94368ED37D6460FF13FE21ACC8FDDD" blockId="8.[189,1399,412,614]" pageId="8" pageNumber="9">HEAD. Dark brown to dark piceous. Frontally between eyes not margined. Slightly posterior to frontal edge with a shallow, transverse depression, which medially is vague. Shiny although covered with fine, slightly indistinct reticulation; meshes when distinguished are isodiametric. Posterior to cervical line reticulation indistinct; surface is shagreened. With fine, sparse and somewhat irregular punctures. Posterior to cervical line impunctate. Antenna with basal segments 1–4 pale, testaceous; segments 5–11 darker; brown to piceous.</paragraph>
<paragraph id="8B94368ED37D6460FF13FD33A957FCAA" blockId="8.[189,1399,653,785]" pageId="8" pageNumber="9">PRONOTUM. Broadest at posterior corners. Sides posteriorly straight; anteriorly clearly curved inwards. Bright testaceous; anteriorly and between striae with narrow blackish to dark brown, transverse area. At posterior corners with vague, darker spots. Discally with broad area with fine, sparse and irregular punctures. At pronotal edges punctures coarser and denser. Shiny, not reticulated.</paragraph>
<paragraph id="8B94368ED37D6460FF13FC84AEE6FBF3" blockId="8.[189,1399,824,1097]" pageId="8" pageNumber="9">ELYTRA. Black to dark piceous; apically with a vague, paler area. Basal striae medium long and straight (close to frontal edge strongly curved outwards for a short distance). Sutural lines close to frontal edge of elytra, distinct; posteriorly lines gradually fade away. Punctures fine to rather fine, somewhat sparse and with slightly irregular distribution. On each side of elytral stria, punctures of almost equal size and density. Towards apical region punctures become slightly finer. In frontal half of elytra with a lateral row of slightly irregular punctures. Row of punctures form a shallow but clearly discernible furrow which in posterior half disappears. Narrow area between furrow and epipleuron pale brownish and with finer and sparser punctures. Shiny, not reticulated. Epipleura pale brownish.</paragraph>
<paragraph id="8B94368ED37D6460FF13FBCCA9BCFA81" blockId="8.[189,1399,1136,1339]" pageId="8" pageNumber="9">VENTRAL ASPECT. Pronotum testaceous, with lateral edges narrowly blackish. Metathorax, metacoxal plates and abdomen blackish to dark piceous. Rather shiny, reticulation absent. Punctures distributed as follows: metathorax medially with two rows of fine punctures and at frontal and lateral edge with a single row. Metacoxal plates medially with a few scattered punctures and at inner margin of metacoxal lines with a row of punctures. Abdomen at base with irregular coarse punctures and ventrite in apical half with dense, quite coarse punctures. Otherwise ventral aspect almost impunctate.</paragraph>
<paragraph id="8B94368ED37D6460FF13FADFA90FFAC0" blockId="8.[189,1070,1377,1404]" box="[189,1070,1377,1404]" pageId="8" pageNumber="9">LEGS. Testaceous to brownish. Protarsi slightly enlarged; mesotarsi slender.</paragraph>
<paragraph id="8B94368ED37D6460FF13FA1EACC0FA06" blockId="8.[189,481,1443,1469]" box="[189,481,1443,1469]" pageId="8" pageNumber="9">
MALE GENITALIA.
<figureCitation id="13102A0BD37D6460FE2CFA1EACFCFA06" box="[386,477,1443,1469]" captionStart="Fig" captionStartId="9.[189,232,1888,1914]" captionTargetBox="[366,1209,282,1833]" captionTargetId="figure-11@9.[308,1280,265,1849]" captionTargetPageId="9" captionText="Fig. 3. Male genitalia of the Malagasy species of Bidessus Sharp, 1882. For each subpanel from left to right penis in dorsal view, penis in lateral view and paramere in lateral view. A. B. apicidens Bistr̂m &amp; Sanfilippo, 1986. B. B. cf. ceratus Guignot, 1941. C. B. anjozorobe sp. nov. D. B. cf. nero Gschwendtner, 1933. E. B. nesioticus Guignot, 1956. F. B. perexiguus Kolbe, 1883. G. B. longistriga Régimbart, 1895." figureDoi="http://doi.org/10.5281/zenodo.4068361" httpUri="https://zenodo.org/record/4068361/files/figure.png" pageId="8" pageNumber="9">Fig. 3C</figureCitation>
.
</paragraph>
<paragraph id="8B94368ED37D6460FF44FA59AC64FA45" blockId="8.[189,945,1508,1570]" box="[234,325,1508,1534]" pageId="8" pageNumber="9">
<emphasis id="B95FEA9CD37D6460FF44FA59AC64FA45" bold="true" box="[234,325,1508,1534]" pageId="8" pageNumber="9">Female</emphasis>
</paragraph>
<paragraph id="8B94368ED37D6460FF13F9BAAE90F99A" blockId="8.[189,945,1508,1570]" box="[189,945,1543,1570]" pageId="8" pageNumber="9">Similar to male but protarsi not enlarged, narrower than in male.</paragraph>
</subSubSection>
<subSubSection id="C3316505D37D6460FF13F9F4AC11F90E" pageId="8" pageNumber="9" type="distribution">
<paragraph id="8B94368ED37D6460FF13F9F4AC77F9D8" blockId="8.[189,342,1609,1635]" box="[189,342,1609,1635]" pageId="8" pageNumber="9">
<heading id="D0DC81E2D37D6460FF13F9F4AC77F9D8" bold="true" box="[189,342,1609,1635]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="B95FEA9CD37D6460FF13F9F4AC77F9D8" bold="true" box="[189,342,1609,1635]" pageId="8" pageNumber="9">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8B94368ED37D6460FF13F9CAAC11F90E" blockId="8.[189,1399,1655,1717]" pageId="8" pageNumber="9">
Endemic to
<collectingCountry id="F33C761ED37D6460FEE4F9CAACF9F92A" box="[330,472,1655,1681]" name="Madagascar" pageId="8" pageNumber="9">Madagascar</collectingCountry>
and only known from the
<typeStatus id="5490882CD37D6460FCA5F9C5AE1FF929" box="[779,830,1656,1682]" pageId="8" pageNumber="9">type</typeStatus>
locality in Anjozorobe forest, Central Highlands (
<figureCitation id="13102A0BD37D6460FF68F926AC02F90E" box="[198,291,1691,1717]" captionStart="Fig" captionStartId="11.[189,232,1888,1914]" captionTargetBox="[270,1323,299,1826]" captionTargetId="figure-12@11.[232,1355,265,1852]" captionTargetPageId="11" captionText="Fig. 4. Distribution maps of the Malagasy species of Bidessus Sharp, 1882. A. B. longistriga Régimbart, 1895. B. B. perexiguus Kolbe, 1883. C. B. cf. nero Gschwendtner, 1933 (circle), B. nesioticus Guignot, 1956 (square), B. anjozorobe sp. nov. (triangle). D. B. apicidens Biström &amp; Sanfilippo, 1986 (square), B. cf. ceratus Guignot, 1941 (circle)." figureDoi="http://doi.org/10.5281/zenodo.4068363" httpUri="https://zenodo.org/record/4068363/files/figure.png" pageId="8" pageNumber="9">Fig. 4C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3316505D37D6460FF13F961AFD3F80A" pageId="8" pageNumber="9" type="description">
<paragraph id="8B94368ED37D6460FF13F961ACD5F94D" blockId="8.[189,500,1756,1782]" box="[189,500,1756,1782]" pageId="8" pageNumber="9">
<heading id="D0DC81E2D37D6460FF13F961ACD5F94D" bold="true" box="[189,500,1756,1782]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="B95FEA9CD37D6460FF13F961ACD5F94D" bold="true" box="[189,500,1756,1782]" pageId="8" pageNumber="9">Collecting circumstances</emphasis>
</heading>
</paragraph>
<paragraph id="8B94368ED37D6460FF13F8B6AFD3F80A" blockId="8.[189,1399,1802,1969]" pageId="8" pageNumber="9">
The type locality is a vegetation rich marsh in a forest glade at an elevation of
<quantity id="4CD39B6BD37D6460FBDCF8B6A9ECF89E" box="[1138,1229,1803,1829]" metricMagnitude="3" metricUnit="m" metricValue="1.3199999999999998" pageId="8" pageNumber="9" unit="m" value="1320.0">1320 m</quantity>
a.s.l. (
<figureCitation id="13102A0BD37D6460FAB0F8B6A849F89E" box="[1310,1384,1802,1829]" captionStart="Fig" captionStartId="12.[189,232,1888,1914]" captionTargetBox="[189,1398,942,1849]" captionTargetId="figure-106@12.[189,1398,942,1849]" captionTargetPageId="12" captionText="Fig. 5. Type locality [MAD14-78] for Bidessus anjozorobe sp. nov. in Anjozorobe forest in Anjozorobe- Angavo protected area on the Central Highlands. A glade in the rainforest with marsh vegetation. Multiple individuals of Bidessus anjozorobe sp. nov. were found by tramping in the vegetation at the edges of the marsh." figureDoi="http://doi.org/10.5281/zenodo.4068365" httpUri="https://zenodo.org/record/4068365/files/figure.png" pageId="8" pageNumber="9">Fig. 5</figureCitation>
). Small water pools were visible at the edges of the marsh during the visit at the end of
<date id="FF95104ED37D6460FB16F893A856F8F3" box="[1208,1399,1838,1864]" pageId="8" pageNumber="9" value="2014-11">November 2014</date>
but mostly derived through stamping in the vegetation. Anjozorobe-Angavo forest corridor with its westerly location is a higher elevation forest and represents a transition
<typeStatus id="5490882CD37D6460FB8CF8C8A974F834" box="[1058,1109,1909,1935]" pageId="8" pageNumber="9">type</typeStatus>
between typical eastern humid forest and sub-humid high plateau forest.
</paragraph>
</subSubSection>
</treatment>
</document>
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