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<document id="EA375C52AD49E2BE7146C8C0ED590C07" ID-DOI="10.11646/zootaxa.5266.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="7840582" ID-ZooBank="079536BC-B8C4-4974-90EA-BF600D990D14" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1681817008050" checkinUser="plazi" docAuthor="Wood, Thomas James" docDate="2023" docId="03828781F83D702ED6C7FA0C8C29A170" docLanguage="en" docName="zootaxa.5266.1.1.pdf" docOrigin="Zootaxa 5266 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5266.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Andrena (Leimelissa) claves Wood 2023, spec. nov." docType="treatment" docVersion="4" lastPageNumber="30" masterDocId="FFBBFFF9F8267033D650FFC68E49A015" masterDocTitle="New Old World Andrena species, with a focus on Turkey (Hymenoptera: Andrenidae)" masterLastPageNumber="72" masterPageNumber="1" pageNumber="28" updateTime="1699487009086" updateUser="plazi">
<mods:mods id="F301BB8B22C28FA3B936F391D54D35CD" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="89623BEE65C4412D8D8231B05026874E">
<mods:title id="47B8DF2E71C3382BE586068A50A4FE41">New Old World Andrena species, with a focus on Turkey (Hymenoptera: Andrenidae)</mods:title>
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<mods:namePart id="AEB92779A4739673C4D817D7068416A3">Wood, Thomas James</mods:namePart>
<mods:nameIdentifier id="368759090EB214180365A5E409FB690A" type="ORCID">0000-0001-5653-224X</mods:nameIdentifier>
<mods:nameIdentifier id="E87EF25685CBCBEEB6511D3B7D5F5E59" type="email">thomasjames.wood@umons.ac.be</mods:nameIdentifier>
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<mods:typeOfResource id="33272DFCF8E098DFF44EE0A6ABE24E73">text</mods:typeOfResource>
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<mods:title id="D2CC9D9A7FEFB2E71C1DDFD440D40A78">Zootaxa</mods:title>
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<mods:part id="C336BF7A35E4822254C630CD6095CCB0">
<mods:date id="9B24187D7369B88F27C0796EC0190B1C">2023</mods:date>
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<mods:number id="AC2CE306E992408AC2DC9A1EE9A4BC67">2023-04-18</mods:number>
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<mods:number id="1AEC00483BC1069AC6FE4176F3FE40B0">5266</mods:number>
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<mods:number id="4D4F41BF0B5501F2058575750AE966C3">1</mods:number>
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<mods:start id="4804E4A531B33D2093E3EE4FB893550A">1</mods:start>
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<mods:url id="BBBACEE3193F7E53821303136872A56C">http://dx.doi.org/10.11646/zootaxa.5266.1.1</mods:url>
</mods:location>
<mods:classification id="EEEA2323C33C8D22F7C1270844DD2B90">journal article</mods:classification>
<mods:identifier id="A0E7B15918CB84EAC7CEFB976481CCA5" type="DOI">10.11646/zootaxa.5266.1.1</mods:identifier>
<mods:identifier id="3069B97852E2826306D8FBF09012CC43" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="05A64FDA5C74E8C6B2050899BE8C7E29" type="Zenodo-Dep">7840582</mods:identifier>
<mods:identifier id="21221E2A8D1C0FD39DA254F39A3A32E0" type="ZooBank">079536BC-B8C4-4974-90EA-BF600D990D14</mods:identifier>
</mods:mods>
<treatment id="03828781F83D702ED6C7FA0C8C29A170" ID-DOI="http://doi.org/10.5281/zenodo.7840604" ID-Zenodo-Dep="7840604" LSID="urn:lsid:plazi:treatment:03828781F83D702ED6C7FA0C8C29A170" httpUri="http://treatment.plazi.org/id/03828781F83D702ED6C7FA0C8C29A170" lastPageId="29" lastPageNumber="30" pageId="27" pageNumber="28">
<subSubSection id="C331651CF83D7028D6C7FA0C8C25A5F0" box="[151,620,1482,1509]" pageId="27" pageNumber="28" type="nomenclature">
<paragraph id="8B943697F83D7028D6C7FA0C8C25A5F0" blockId="27.[151,620,1482,1509]" box="[151,620,1482,1509]" pageId="27" pageNumber="28">
<heading id="D0DC81FBF83D7028D6C7FA0C8C25A5F0" bold="true" box="[151,620,1482,1509]" fontSize="11" level="2" pageId="27" pageNumber="28" reason="4">
<emphasis id="B95FEA85F83D7028D6C7FA0C8C25A5F0" bold="true" box="[151,620,1482,1509]" pageId="27" pageNumber="28">
<taxonomicName id="4C2B4D14F83D7028D6C7FA0C8FA4A5F0" authority="Wood, 2023" authorityName="Wood" authorityYear="2023" box="[151,493,1482,1509]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="claves" status="sp. nov." subGenus="Leimelissa">
<emphasis id="B95FEA85F83D7028D6C7FA0C8F48A5F0" bold="true" box="[151,257,1482,1509]" italics="true" pageId="27" pageNumber="28">Andrena</emphasis>
(
<emphasis id="B95FEA85F83D7028D742FA0C8FDAA5F0" bold="true" box="[274,403,1482,1509]" italics="true" pageId="27" pageNumber="28">Leimelissa</emphasis>
)
<emphasis id="B95FEA85F83D7028D7F4FA0C8FA4A5F0" bold="true" box="[420,493,1482,1509]" italics="true" pageId="27" pageNumber="28">claves</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26C57FEF83D7028D7A5FA0D8C25A5F0" box="[501,620,1483,1509]" pageId="27" pageNumber="28" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C331651CF83D7028D6C7F9D48AC5A662" pageId="27" pageNumber="28" type="materials_examined">
<paragraph id="8B943697F83D7028D6C7F9D48A0CA646" blockId="27.[151,1436,1554,2013]" pageId="27" pageNumber="28">
<materialsCitation id="3B433CCAF83D7028D6C7F9D48A0CA646" collectingDate="1980-06-03" collectionCode="OOLM" collectorName="Z. Padr" country="Kazakhstan" county="Ujuk" elevation="1000" location="Dzambul" municipality="Kara Tau" pageId="27" pageNumber="28" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<emphasis id="B95FEA85F83D7028D6C7F9D48C5DA639" bold="true" box="[151,532,1554,1580]" pageId="27" pageNumber="28">
<typeStatus id="54908835F83D7028D6C7F9D48F73A639" box="[151,314,1554,1580]" pageId="27" pageNumber="28" type="holotype">HOLOTYPE</typeStatus>
:
<collectingCountry id="F33C7607F83D7028D716F9D48C59A639" box="[326,528,1554,1580]" name="Kazakhstan" pageId="27" pageNumber="28">KAZAKHSTAN</collectingCountry>
:
</emphasis>
<location id="8EF4604CF83D7028D44BF9D58CCBA638" LSID="urn:lsid:plazi:treatment:03828781F83D702ED6C7FA0C8C29A170:8EF4604CF83D7028D44BF9D58CCBA638" box="[539,642,1555,1581]" country="Kazakhstan" county="Ujuk" municipality="Kara Tau" name="Dzambul" pageId="27" pageNumber="28">Džambul</location>
env. [Russian name, Dzhambul, now Taraz],
<collectingMunicipality id="6BF0ACEDF83D7028D22DF9D58AACA638" box="[1149,1253,1555,1581]" pageId="27" pageNumber="28">Kara Tau</collectingMunicipality>
, průsmyk [pass]
<collectingCounty id="62F54E1BF83D7028D6C7F9F18E86A644" box="[151,207,1591,1617]" pageId="27" pageNumber="28">Ujuk</collectingCounty>
[inferred 43.1597
<superScript id="7C5E9BDFF83D7028D7CAF9F18FF0A644" attach="right" box="[410,441,1591,1617]" fontSize="6" pageId="27" pageNumber="28">oN</superScript>
, 70.2016
<superScript id="7C5E9BDFF83D7028D44FF9F18C74A644" attach="right" box="[543,573,1591,1617]" fontSize="6" pageId="27" pageNumber="28">oE</superScript>
],
<quantity id="4CD39B72F83D7028D400F9F18CEEA644" box="[592,679,1591,1617]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="27" pageNumber="28" unit="m" value="1000.0">
<elevation id="0006D1A4F83D7028D400F9F18CEEA644" box="[592,679,1591,1617]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="27" pageNumber="28" unit="m" value="1000.0">1000 m</elevation>
</quantity>
,
<date id="FF951057F83D7028D4E3F9F18D56A644" box="[691,799,1591,1617]" pageId="27" pageNumber="28" value="1980-06-03">
<collectingDate id="EFD1E9BFF83D7028D4E3F9F18D56A644" box="[691,799,1591,1617]" pageId="27" pageNumber="28" value="1980-06-03">3.vi.1980</collectingDate>
</date>
,
<specimenCount id="9D2DFD1EF83D7028D57BF9F18D07A644" box="[811,846,1591,1617]" pageId="27" pageNumber="28" type="female">1♀</specimenCount>
, leg.
<collectorName id="26DE5341F83D7028D5DCF9F18DA9A644" box="[908,992,1591,1617]" pageId="27" pageNumber="28">Z. Pádr</collectorName>
,
<collectionCode id="ED3AAE52F83D7028D5BBF9FE8A08A646" box="[1003,1089,1592,1619]" pageId="27" pageNumber="28">OÖLM</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8B943697F83D7028D697F99C8AC5A662" blockId="27.[151,1436,1554,2013]" box="[199,1164,1626,1655]" pageId="27" pageNumber="28">
<materialsCitation id="3B433CCAF83D7028D697F99C8AC1A662" box="[199,1160,1626,1655]" collectionCode="OOLM, TJWC" country="Kazakhstan" location="Same" pageId="27" pageNumber="28" specimenCount="3" specimenCount-female="3" typeStatus="paratype">
<emphasis id="B95FEA85F83D7028D697F99C8C03A661" bold="true" box="[199,586,1626,1652]" pageId="27" pageNumber="28">
<typeStatus id="54908835F83D7028D697F99C8F26A661" box="[199,367,1626,1652]" pageId="27" pageNumber="28" type="paratype">PARATYPES</typeStatus>
:
<collectingCountry id="F33C7607F83D7028D72BF99C8C0CA661" box="[379,581,1626,1652]" name="Kazakhstan" pageId="27" pageNumber="28">KAZAKHSTAN</collectingCountry>
:
</emphasis>
<location id="8EF4604CF83D7028D401F99D8CC6A660" LSID="urn:lsid:plazi:treatment:03828781F83D702ED6C7FA0C8C29A170:8EF4604CF83D7028D401F99D8CC6A660" box="[593,655,1627,1653]" country="Kazakhstan" name="Same" pageId="27" pageNumber="28">Same</location>
information as
<typeStatus id="54908835F83D7028D510F99D8DECA660" box="[832,933,1627,1653]" pageId="27" pageNumber="28" type="holotype">holotype</typeStatus>
,
<specimenCount id="9D2DFD1EF83D7028D5E0F99D8D9DA660" box="[944,980,1627,1653]" pageId="27" pageNumber="28" type="female">3♀</specimenCount>
,
<collectionCode id="ED3AAE52F83D7028D5B1F99A8A7CA662" box="[993,1077,1628,1655]" pageId="27" pageNumber="28">OÖLM</collectionCode>
/
<collectionCode id="ED3AAE52F83D7028D26DF99A8AC1A662" box="[1085,1160,1628,1655]" pageId="27" pageNumber="28">TJWC</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C331651CF83D702FD697F9B88FC3A2D8" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28" type="description">
<paragraph id="8B943697F83D702FD697F9B88AB5A2BC" blockId="27.[151,1436,1554,2013]" lastBlockId="28.[151,1437,151,933]" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28">
<emphasis id="B95FEA85F83D7028D697F9B88F8AA68C" bold="true" box="[199,451,1662,1689]" pageId="27" pageNumber="28">Description: Female:</emphasis>
Body length:
<quantity id="4CD39B72F83D7028D436F9B98C96A68C" box="[614,735,1663,1689]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="27" pageNumber="28" unit="mm" value="11.5" valueMax="12.0" valueMin="11.0">11–12 mm</quantity>
(
<figureCitation id="13102A12F83D7028D4BFF9B98D3FA68C" box="[751,886,1662,1689]" captionStart="FIGURE 15" captionStartId="29.[152,255,1846,1870]" captionTargetBox="[151,1436,381,1823]" captionTargetId="figure-105@29.[151,1436,381,1823]" captionTargetPageId="29" captionText="FIGURE 15. Andrena (Leimelissa) claves spec. nov. female. A. Habitus, lateral view, B. Face, frontal view, C. Head, dorsal view, D. Dorsum, E. Propodeum, posterior view, F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7840636" httpUri="https://zenodo.org/record/7840636/files/figure.png" pageId="27" pageNumber="28">Figure 15A</figureCitation>
).
<emphasis id="B95FEA85F83D7028D5DAF9B98D99A68C" box="[906,976,1663,1689]" italics="true" pageId="27" pageNumber="28">Head:</emphasis>
Dark, 1.3 times wider than long (
<figureCitation id="13102A12F83D7028D301F9B98E80A6A8" captionStart="FIGURE 15" captionStartId="29.[152,255,1846,1870]" captionTargetBox="[151,1436,381,1823]" captionTargetId="figure-105@29.[151,1436,381,1823]" captionTargetPageId="29" captionText="FIGURE 15. Andrena (Leimelissa) claves spec. nov. female. A. Habitus, lateral view, B. Face, frontal view, C. Head, dorsal view, D. Dorsum, E. Propodeum, posterior view, F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7840636" httpUri="https://zenodo.org/record/7840636/files/figure.png" pageId="27" pageNumber="28">Figure 15B</figureCitation>
). Clypeus strongly flattened, medially slightly concave, with two weak and subtle longitudinal impressions; clypeus shallowly and irregularly punctate, punctures separated by 1–3 puncture diameters, broad impunctate longitudinal midline present; underlying surface strongly shagreened, dull. Process of labrum extremely broad, 5 times wider than long, basally with short latitudinal impression, medially with narrow emargination. Gena equalling width of compound eye, densely but shallowly and obscurely punctate with small punctures, punctures separated by 0.5 puncture diameters; ocelloccipital distance slightly exceeding 1 diameter of lateral ocellus. Foveae broad, occupying three quarters distance between lateral ocellus and compound eye, filled with white hairs (
<figureCitation id="13102A12F83D7028D35BF8BD8BC6A780" box="[1291,1423,1914,1941]" captionStart="FIGURE 15" captionStartId="29.[152,255,1846,1870]" captionTargetBox="[151,1436,381,1823]" captionTargetId="figure-105@29.[151,1436,381,1823]" captionTargetPageId="29" captionText="FIGURE 15. Andrena (Leimelissa) claves spec. nov. female. A. Habitus, lateral view, B. Face, frontal view, C. Head, dorsal view, D. Dorsum, E. Propodeum, posterior view, F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7840636" httpUri="https://zenodo.org/record/7840636/files/figure.png" pageId="27" pageNumber="28">Figure 15C</figureCitation>
). Gena, vertex, face, and scape with short white hairs. Antennae dark, A4–5 and A6 basally orange-marked ventrally; A3 =A4+5+6.
<taxonomicName id="4C2B4D14F83D7028D76AF8058C5FA7C8" authority=": Scutum" authorityName="Scutum" box="[314,534,1987,2013]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="B95FEA85F83D7028D76AF8058FF2A7C9" box="[314,443,1987,2012]" italics="true" pageId="27" pageNumber="28">Mesosoma:</emphasis>
Scutum
</taxonomicName>
irregularly punctate, punctures separated by 0.5–2 puncture diameters, underlying surface finely shagreened, weakly shining. Scutellum smooth and shining, sparsely and irregularly punctate (
<figureCitation id="13102A12F83A702FD301FF518E82A0C0" captionStart="FIGURE 15" captionStartId="29.[152,255,1846,1870]" captionTargetBox="[151,1436,381,1823]" captionTargetId="figure-105@29.[151,1436,381,1823]" captionTargetPageId="29" captionText="FIGURE 15. Andrena (Leimelissa) claves spec. nov. female. A. Habitus, lateral view, B. Face, frontal view, C. Head, dorsal view, D. Dorsum, E. Propodeum, posterior view, F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7840636" httpUri="https://zenodo.org/record/7840636/files/figure.png" pageId="28" pageNumber="29">Figure 15D</figureCitation>
). Pronotum without humeral angle, surface with shallow punctures. Mesepisternum and dorsolateral parts of propodeum shagreened, weakly shining, densely and regularly punctate with shallow punctures, punctures separated by 0.5–1 puncture diameter; propodeal triangle impunctate, internal surface finely shagreened, more strongly shining; propodeal triangle with short inconspicuous longitudinal rugae basally (
<figureCitation id="13102A12F83A702FD276FEE18AE3A154" box="[1062,1194,294,321]" captionStart="FIGURE 15" captionStartId="29.[152,255,1846,1870]" captionTargetBox="[151,1436,381,1823]" captionTargetId="figure-105@29.[151,1436,381,1823]" captionTargetPageId="29" captionText="FIGURE 15. Andrena (Leimelissa) claves spec. nov. female. A. Habitus, lateral view, B. Face, frontal view, C. Head, dorsal view, D. Dorsum, E. Propodeum, posterior view, F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7840636" httpUri="https://zenodo.org/record/7840636/files/figure.png" pageId="28" pageNumber="29">Figure 15E</figureCitation>
). Mesepisternum and propodeum with long white hairs, not exceeding length of scape, propodeal corbicula dorsally with dense white simple hairs, internal surface with sparse, long white hairs. Scutum and scutellum with sparse, short, simple white hairs. Legs dark, apical tarsal segments dark brown. Flocculus and femoral scopa snow-white; tibial scopa white ventrally, dark brown dorsally. Remaining pubescence of legs dark brown. Hind tarsal claws with strong inner tooth. Wings hyaline, stigma and venation dark brown, nervulus postfurcal.
<emphasis id="B95FEA85F83A702FD5FBFE1D8A60A1E1" box="[939,1065,475,500]" italics="true" pageId="28" pageNumber="29">Metasoma:</emphasis>
Terga dark, smooth and shining, disc of T1 clearly punctate, punctures separated by 1–3 puncture diameters, discs of T2–4 more densely punctate with finer punctures, punctures separated by 0.5–1 puncture diameters; marginal areas of T2–4 with smaller punctures, separated by 1 puncture diameter (
<figureCitation id="13102A12F83A702FD44CFD818CD4A274" box="[540,669,582,609]" captionStart="FIGURE 15" captionStartId="29.[152,255,1846,1870]" captionTargetBox="[151,1436,381,1823]" captionTargetId="figure-105@29.[151,1436,381,1823]" captionTargetPageId="29" captionText="FIGURE 15. Andrena (Leimelissa) claves spec. nov. female. A. Habitus, lateral view, B. Face, frontal view, C. Head, dorsal view, D. Dorsum, E. Propodeum, posterior view, F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7840636" httpUri="https://zenodo.org/record/7840636/files/figure.png" pageId="28" pageNumber="29">Figure 15F</figureCitation>
). T1–4 laterally with pairs of large white hair patches overlying and obscuring underlying surface, clearly separated medially. Apical fringe of T5 and hairs flanking pygidial plate dark chocolate brown, pygidial plate narrowly triangular, with obscure longitudinal raised ridge medially.
</paragraph>
<paragraph id="8B943697F83A702FD697FD758FC3A2D8" blockId="28.[151,1437,151,933]" box="[199,394,691,717]" pageId="28" pageNumber="29">
<emphasis id="B95FEA85F83A702FD697FD758F44A2D8" bold="true" box="[199,269,691,717]" pageId="28" pageNumber="29">Male:</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C331651CF83A702ED697FD108AF8A0EC" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29" type="diagnosis">
<paragraph id="8B943697F83A702FD697FD108BCEA3B0" blockId="28.[151,1437,151,933]" pageId="28" pageNumber="29">
<emphasis id="B95FEA85F83A702FD697FD108F0DA2E5" bold="true" box="[199,324,726,752]" pageId="28" pageNumber="29">Diagnosis:</emphasis>
This species can be placed into the subgenus
<taxonomicName id="4C2B4D14F83A702FD53DFD118DACA2E4" box="[877,997,727,753]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="subGenus" subGenus="Leimelissa">
<emphasis id="B95FEA85F83A702FD53DFD118DACA2E4" box="[877,997,727,753]" italics="true" pageId="28" pageNumber="29">Leimelissa</emphasis>
</taxonomicName>
due to the strongly flattened clypeus (
<figureCitation id="13102A12F83A702FD6CFFD3D8F6AA300" box="[159,291,762,789]" captionStart="FIGURE 15" captionStartId="29.[152,255,1846,1870]" captionTargetBox="[151,1436,381,1823]" captionTargetId="figure-105@29.[151,1436,381,1823]" captionTargetPageId="29" captionText="FIGURE 15. Andrena (Leimelissa) claves spec. nov. female. A. Habitus, lateral view, B. Face, frontal view, C. Head, dorsal view, D. Dorsum, E. Propodeum, posterior view, F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7840636" httpUri="https://zenodo.org/record/7840636/files/figure.png" pageId="28" pageNumber="29">Figure 15B</figureCitation>
), the short and broad process of the labrum with a shallow emargination medially (
<figureCitation id="13102A12F83A702FD28EFD3D8B2AA300" box="[1246,1379,762,789]" captionStart="FIGURE 15" captionStartId="29.[152,255,1846,1870]" captionTargetBox="[151,1436,381,1823]" captionTargetId="figure-105@29.[151,1436,381,1823]" captionTargetPageId="29" captionText="FIGURE 15. Andrena (Leimelissa) claves spec. nov. female. A. Habitus, lateral view, B. Face, frontal view, C. Head, dorsal view, D. Dorsum, E. Propodeum, posterior view, F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7840636" httpUri="https://zenodo.org/record/7840636/files/figure.png" pageId="28" pageNumber="29">Figure 15B</figureCitation>
), the densely punctate mesepisternum and dorsolateral part of the propodeum (
<figureCitation id="13102A12F83A702FD58EFCD98A2BA32C" box="[990,1122,798,825]" captionStart="FIGURE 15" captionStartId="29.[152,255,1846,1870]" captionTargetBox="[151,1436,381,1823]" captionTargetId="figure-105@29.[151,1436,381,1823]" captionTargetPageId="29" captionText="FIGURE 15. Andrena (Leimelissa) claves spec. nov. female. A. Habitus, lateral view, B. Face, frontal view, C. Head, dorsal view, D. Dorsum, E. Propodeum, posterior view, F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7840636" httpUri="https://zenodo.org/record/7840636/files/figure.png" pageId="28" pageNumber="29">Figure 15E</figureCitation>
), and the finely shagreened and impunctate propodeal triangle which lacks rugae (
<figureCitation id="13102A12F83A702FD573FC858DE5A348" box="[803,940,834,861]" captionStart="FIGURE 15" captionStartId="29.[152,255,1846,1870]" captionTargetBox="[151,1436,381,1823]" captionTargetId="figure-105@29.[151,1436,381,1823]" captionTargetPageId="29" captionText="FIGURE 15. Andrena (Leimelissa) claves spec. nov. female. A. Habitus, lateral view, B. Face, frontal view, C. Head, dorsal view, D. Dorsum, E. Propodeum, posterior view, F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7840636" httpUri="https://zenodo.org/record/7840636/files/figure.png" pageId="28" pageNumber="29">Figure 15E</figureCitation>
). Eight species of
<taxonomicName id="4C2B4D14F83A702FD2C4FC858B45A348" box="[1172,1292,835,861]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="54" phylum="Arthropoda" rank="subGenus" subGenus="Leimelissa">
<emphasis id="B95FEA85F83A702FD2C4FC858B45A348" box="[1172,1292,835,861]" italics="true" pageId="28" pageNumber="29">Leimelissa</emphasis>
</taxonomicName>
are known, predominantly from central Asia and
<collectingCountry id="F33C7607F83A702FD415FCA18CC1A394" box="[581,648,871,897]" name="China" pageId="28" pageNumber="29">China</collectingCountry>
(
<tableCitation id="C6A9032CF83A702FD4C8FCA18CB8A394" box="[664,753,871,897]" captionStart="TABLE 2" captionStartId="28.[152,244,980,1004]" captionText="TABLE 2. Species currently included in the subgenus Leimelissa and their known distributions." pageId="28" pageNumber="29">Table 2</tableCitation>
), with the new transfer of
<taxonomicName id="4C2B4D14F83A702FD276FCA18B78A394" authority="Nurse, 1904" authorityName="Nurse" authorityYear="1904" box="[1062,1329,871,897]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="flagella">
<emphasis id="B95FEA85F83A702FD276FCA18AD4A394" box="[1062,1181,871,897]" italics="true" pageId="28" pageNumber="29">A. flagella</emphasis>
<bibRefCitation id="EFBA4B66F83A702FD2F6FCA18B78A394" author="Nurse, C. G." box="[1190,1329,871,897]" pageId="28" pageNumber="29" pagination="557 - 585" refId="ref43614" refString="Nurse, C. G. (1904) New species of Indian Hymenoptera. Apidae. Journal of Bombay Natural History Society, 15, 557 - 585." type="journal article" year="1904">Nurse, 1904</bibRefCitation>
</taxonomicName>
from the subgenus
<taxonomicName id="4C2B4D14F83A702FD756FC4D8FEBA3B0" box="[262,418,907,933]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="subGenus" subGenus="Lepidandrena">
<emphasis id="B95FEA85F83A702FD756FC4D8FEBA3B0" box="[262,418,907,933]" italics="true" pageId="28" pageNumber="29">Lepidandrena</emphasis>
</taxonomicName>
(see placement of
<bibRefCitation id="EFBA4B66F83A702FD426FC4D8D83A3B0" author="Gusenleitner, F. &amp; Schwarz, M." box="[630,970,907,933]" pageId="28" pageNumber="29" pagination="1 - 1280" refId="ref43489" refString="Gusenleitner, F. &amp; Schwarz, M. (2002) Weltweite Checkliste der Bienengattung Andrena mit Bemerkungen und Erganzungen zu palaarktischen Arten (Hymenoptera, Apidae, Andreninae, Andrena). Entomofauna, Supplement 10, 1 - 1280." type="journal article" year="2002">Gusenleitner &amp; Schwarz 2002</bibRefCitation>
) following
<typeStatus id="54908835F83A702FD21CFC4D8A35A3B0" box="[1100,1148,907,933]" pageId="28" pageNumber="29">type</typeStatus>
inspection (see below).
</paragraph>
<caption id="DF54661FF83A702FD6C8FC128AD4A3F9" box="[152,1181,980,1004]" pageId="28" pageNumber="29" startId="28.[152,244,980,1004]">
<paragraph id="8B943697F83A702FD6C8FC128AD4A3F9" blockId="28.[152,1181,980,1004]" box="[152,1181,980,1004]" pageId="28" pageNumber="29">
<emphasis id="B95FEA85F83A702FD6C8FC128F46A3F9" bold="true" box="[152,271,980,1004]" pageId="28" pageNumber="29">TABLE 2.</emphasis>
Species currently included in the subgenus
<taxonomicName id="4C2B4D14F83A702FD4B4FC128D1FA3F9" box="[740,854,980,1004]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="subGenus" subGenus="Leimelissa">
<emphasis id="B95FEA85F83A702FD4B4FC128D1FA3F9" box="[740,854,980,1004]" italics="true" pageId="28" pageNumber="29">Leimelissa</emphasis>
</taxonomicName>
and their known distributions.
</paragraph>
</caption>
<paragraph id="8B943697F83A702FD6F6FC3A8B01A5D7" pageId="28" pageNumber="29">
<table id="F92BC437F83A8FCCD6F6FC3A8BC2A5F3" box="[166,1419,1020,1510]" colsContinueFrom="11.[166,1410,191,376]" colsContinueIn="48.[166,1413,588,1186]" gridcols="3" gridrows="10" pageId="28" pageNumber="29">
<tr id="351B34D5F83A8FCCD6F6FC3A8BC2A401" box="[166,1419,1020,1044]" gridrow="0" pageId="28" pageNumber="29">
<th id="76CA5DA9F83A8FCCD6F6FC3A8C2CA401" box="[166,613,1020,1044]" gridcol="0" gridrow="0" pageId="28" pageNumber="29">
<emphasis id="B95FEA85F83A702FD6F6FC3A8EBCA401" bold="true" box="[166,245,1020,1044]" pageId="28" pageNumber="29">Species</emphasis>
</th>
<th id="76CA5DA9F83A8FCCD4D7FC3A8DA8A401" box="[647,993,1020,1044]" gridcol="1" gridrow="0" pageId="28" pageNumber="29">
<emphasis id="B95FEA85F83A702FD4D7FC3A8D42A401" bold="true" box="[647,779,1020,1044]" pageId="28" pageNumber="29">Distribution</emphasis>
</th>
<th id="76CA5DA9F83A8FCCD255FC3A8BC2A401" box="[1029,1419,1020,1044]" gridcol="2" gridrow="0" pageId="28" pageNumber="29">
<emphasis id="B95FEA85F83A702FD255FC3A8A06A401" bold="true" box="[1029,1103,1020,1044]" pageId="28" pageNumber="29">Source</emphasis>
</th>
</tr>
<tr id="351B34D5F83A8FCCD6F6FBE28BC2A428" box="[166,1419,1060,1085]" gridrow="1" pageId="28" pageNumber="29">
<th id="76CA5DA9F83A8FCCD6F6FBE28C2CA428" box="[166,613,1060,1085]" gridcol="0" gridrow="1" pageId="28" pageNumber="29">
<taxonomicName id="4C2B4D14F83A702FD6F6FBE28C77A429" authority="Morawitz, 1876" authorityName="Morawitz" authorityYear="1876" box="[166,574,1060,1084]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="bairacumensis">
<emphasis id="B95FEA85F83A702FD6F6FBE28FDFA429" box="[166,406,1060,1084]" italics="true" pageId="28" pageNumber="29">Andrena bairacumensis</emphasis>
Morawitz, 1876
</taxonomicName>
</th>
<td id="76CA5DA9F83A8FCCD4D7FBE28DA8A428" box="[647,993,1060,1085]" gridcol="1" gridrow="1" pageId="28" pageNumber="29">Kazakhstan</td>
<td id="76CA5DA9F83A8FCCD255FBE28BC2A428" box="[1029,1419,1060,1085]" gridcol="2" gridrow="1" pageId="28" pageNumber="29">
<bibRefCitation id="EFBA4B66F83A702FD255FBE28B4DA428" author="Astafurova, Y. A. &amp; Proshchalykin, M. Y. &amp; Sidorov, D. A." box="[1029,1284,1060,1085]" pageId="28" pageNumber="29" pagination="105 - 176" refId="ref42923" refString="Astafurova, Y. A., Proshchalykin, M. Y. &amp; Sidorov, D. A. (2022 a) The bees of the genus Andrena Fabricius, 1775 (Hymenoptera, Andrenidae) described by Ferdinand Morawitz from the collection of Aleksey Fedtschenko. Zookeys, 1120, 105 - 176. https: // doi. org / 10.3897 / zookeys. 1120.90206" type="journal article" year="2022">
Astafurova
<emphasis id="B95FEA85F83A702FD22CFBE38AE6A429" box="[1148,1199,1060,1084]" italics="true" pageId="28" pageNumber="29">et al.</emphasis>
(2022a)
</bibRefCitation>
</td>
</tr>
<tr id="351B34D5F83A8FCCD6F6FB8B8BC2A470" box="[166,1419,1101,1125]" gridrow="2" pageId="28" pageNumber="29">
<th id="76CA5DA9F83A8FCCD6F6FB8B8C2CA470" box="[166,613,1101,1125]" gridcol="0" gridrow="2" pageId="28" pageNumber="29">
<taxonomicName id="4C2B4D14F83A702FD6F6FB8B8FA9A470" authority="Xu, 1994" authorityName="Xu" authorityYear="1994" box="[166,480,1101,1125]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="beijingensis">
<emphasis id="B95FEA85F83A702FD6F6FB8B8F32A470" box="[166,379,1101,1125]" italics="true" pageId="28" pageNumber="29">Andrena beijingensis</emphasis>
<bibRefCitation id="EFBA4B66F83A702FD7D2FB8B8FA9A470" author="Xu, H. - L." box="[386,480,1101,1125]" pageId="28" pageNumber="29" pagination="197 - 204" refId="ref45800" refString="Xu, H. - L. (1994) Descriptions of new species of genus Andrena from China (Apoidea, Andrenidae). Sinozoologia, 11, 197 - 204. [in Chinese]" type="journal article" year="1994">Xu, 1994</bibRefCitation>
</taxonomicName>
</th>
<td id="76CA5DA9F83A8FCCD4D7FB8B8DA8A470" box="[647,993,1101,1125]" gridcol="1" gridrow="2" pageId="28" pageNumber="29">China (Beijing)</td>
<td id="76CA5DA9F83A8FCCD255FB8B8BC2A470" box="[1029,1419,1101,1125]" gridcol="2" gridrow="2" pageId="28" pageNumber="29">
<bibRefCitation id="EFBA4B66F83A702FD255FB8B8AA3A470" author="Xu, H. - L. &amp; Tadauchi, O." box="[1029,1258,1101,1125]" pageId="28" pageNumber="29" pagination="63 - 66" refId="ref45916" refString="Xu, H. - L. &amp; Tadauchi, O. (2008) The Subgenera Margandrena, Leimelissa, Stenomelissa and Zonandrena of the Genus Andrena of Eastern Asia (Hymenoptera, Andrenidae). Journal of the Faculty of Agriculture, Kyushu University, 53 (1), 63 - 66." type="journal article" year="2008">Xu &amp; Tadauchi (2008)</bibRefCitation>
</td>
</tr>
<tr id="351B34D5F83A8FCCD6F6FBB38BC2A498" box="[166,1419,1141,1165]" gridrow="3" pageId="28" pageNumber="29">
<th id="76CA5DA9F83A8FCCD6F6FBB38C2CA498" box="[166,613,1141,1165]" gridcol="0" gridrow="3" pageId="28" pageNumber="29">
<taxonomicName id="4C2B4D14F83A702FD6F6FBB38C67A498" authority="Osytshnjuk, 1984" authorityName="Osytshnjuk" authorityYear="1984" box="[166,558,1141,1165]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="ermolenkoi">
<emphasis id="B95FEA85F83A702FD6F6FBB38F3DA498" box="[166,372,1141,1165]" italics="true" pageId="28" pageNumber="29">Andrena ermolenkoi</emphasis>
Osytshnjuk, 1984
</taxonomicName>
</th>
<td id="76CA5DA9F83A8FCCD4D7FBB38DA8A498" box="[647,993,1141,1165]" gridcol="1" gridrow="3" pageId="28" pageNumber="29">Turkey*, Armenia</td>
<td id="76CA5DA9F83A8FCCD255FBB38BC2A498" box="[1029,1419,1141,1165]" gridcol="2" gridrow="3" pageId="28" pageNumber="29">
<bibRefCitation id="EFBA4B66F83A702FD255FBB38AB7A498" author="Osytshnjuk, A. Z. &amp; Romasenko, L. &amp; Banaszak, J. &amp; Motyka, E." box="[1029,1278,1141,1165]" pageId="28" pageNumber="29" refId="ref43693" refString="Osytshnjuk, A. Z., Romasenko, L., Banaszak, J. &amp; Motyka, E. (2008). Andreninae of the Central and Eastern Palearctic. Part 2. Polish Entomological Monographs V. Polish Entomological Society, Poznan, 233 pp." type="book" year="2008">
Osytshnjuk
<emphasis id="B95FEA85F83A702FD22FFBB08AFBA498" box="[1151,1202,1141,1165]" italics="true" pageId="28" pageNumber="29">et al.</emphasis>
(2008)
</bibRefCitation>
; current work
</td>
</tr>
<tr id="351B34D5F83A8FCCD6F6FB5B8BC2A4CC" box="[166,1419,1181,1241]" gridrow="4" pageId="28" pageNumber="29">
<th id="76CA5DA9F83A8FCCD6F6FB5B8C2CA4CC" box="[166,613,1181,1241]" gridcol="0" gridrow="4" pageId="28" pageNumber="29">
<taxonomicName id="4C2B4D14F83A702FD6F6FB5B8FBDA4A0" authority="Eversmann, 1852" authorityName="Eversmann" authorityYear="1852" box="[166,500,1181,1205]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="fallax">
<emphasis id="B95FEA85F83A702FD6F6FB5B8F75A4A0" box="[166,316,1181,1205]" italics="true" pageId="28" pageNumber="29">Andrena fallax</emphasis>
Eversmann, 1852
</taxonomicName>
(=
<taxonomicName id="4C2B4D14F83A702FD6ECFB048FFEA4CC" authority="Warncke, 1965" authorityName="Warncke" authorityYear="1965" box="[188,439,1217,1241]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="ispida">
<emphasis id="B95FEA85F83A702FD6ECFB048F5FA4CC" box="[188,278,1217,1241]" italics="true" pageId="28" pageNumber="29">A. ispida</emphasis>
<bibRefCitation id="EFBA4B66F83A702FD74DFB078FFEA4CC" author="Warncke, K." box="[285,439,1217,1241]" pageId="28" pageNumber="29" pagination="27 - 76" refId="ref45141" refString="Warncke, K. (1965) Beitrag zur Kenntnis der Bienengattung Andrena Fabricius in Griechenland. Beitrage zur Entomologie, 15, 27 - 76." type="journal article" year="1965">Warncke, 1965</bibRefCitation>
</taxonomicName>
)
</th>
<td id="76CA5DA9F83A8FCCD4D7FB5B8DA8A4CC" box="[647,993,1181,1241]" gridcol="1" gridrow="4" pageId="28" pageNumber="29">Balkans, Turkey, Syria, Caucasus, Iran</td>
<td id="76CA5DA9F83A8FCCD255FB5B8BC2A4CC" box="[1029,1419,1181,1241]" gridcol="2" gridrow="4" pageId="28" pageNumber="29">
<bibRefCitation id="EFBA4B66F83A702FD255FB5B8B41A4A0" author="Astafurova, Y. A. &amp; Proshchalykin, M. Y. &amp; Sidorov, D. A." box="[1029,1288,1181,1205]" pageId="28" pageNumber="29" pagination="393 - 418" refId="ref42992" refString="Astafurova, Y. A., Proshchalykin, M. Y. &amp; Sidorov, D. A. (2022 b) The type specimens of bees (Hymenoptera, Apoidea) deposited in the Zoological Institute of the Russian Academy of Sciences, St. Petersburg. Contribution V. Family Andrenidae, genus Andrena Fabricius, 1775, species described by E. Eversmann. Zootaxa, 5190 (3), 393 - 418. https: // doi. org / 10.11646 / zootaxa. 5190.3.4" type="journal article" year="2022">
Astafurova
<emphasis id="B95FEA85F83A702FD22CFB588AE6A4A0" box="[1148,1199,1181,1205]" italics="true" pageId="28" pageNumber="29">et al.</emphasis>
(2022b)
</bibRefCitation>
;
<bibRefCitation id="EFBA4B66F83A702FD342FB5B8AFAA4CC" author="Wood, T. J. &amp; Monfared, A." pageId="28" pageNumber="29" pagination="1 - 136" refId="ref45741" refString="Wood, T. J. &amp; Monfared, A. (2022) A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy, 843, 1 - 136. https: // doi. org / 10.5852 / ejt. 2022.843.1947" type="journal article" year="2022">Wood &amp; Monfared (2022)</bibRefCitation>
</td>
</tr>
<tr id="351B34D5F83A8FCCD6F6FB2C8BC2A517" box="[166,1419,1258,1282]" gridrow="5" pageId="28" pageNumber="29">
<th id="76CA5DA9F83A8FCCD6F6FB2C8C2CA517" box="[166,613,1258,1282]" gridcol="0" gridrow="5" pageId="28" pageNumber="29">
<taxonomicName id="4C2B4D14F83A702FD6F6FB2C8F9BA517" authority="Nurse, 1904" authorityName="Nurse" authorityYear="1904" box="[166,466,1258,1282]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="flagella">
<emphasis id="B95FEA85F83A702FD6F6FB2C8F06A517" box="[166,335,1258,1282]" italics="true" pageId="28" pageNumber="29">Andrena flagella</emphasis>
<bibRefCitation id="EFBA4B66F83A702FD706FB2C8F9BA517" author="Nurse, C. G." box="[342,466,1258,1282]" pageId="28" pageNumber="29" pagination="557 - 585" refId="ref43614" refString="Nurse, C. G. (1904) New species of Indian Hymenoptera. Apidae. Journal of Bombay Natural History Society, 15, 557 - 585." type="journal article" year="1904">Nurse, 1904</bibRefCitation>
</taxonomicName>
</th>
<td id="76CA5DA9F83A8FCCD4D7FB2C8DA8A517" box="[647,993,1258,1282]" gridcol="1" gridrow="5" pageId="28" pageNumber="29">Pakistan</td>
<td id="76CA5DA9F83A8FCCD255FB2C8BC2A517" box="[1029,1419,1258,1282]" gridcol="2" gridrow="5" pageId="28" pageNumber="29">
<bibRefCitation id="EFBA4B66F83A702FD255FB2C8AC2A517" author="Nurse, C. G." box="[1029,1163,1258,1282]" pageId="28" pageNumber="29" pagination="557 - 585" refId="ref43614" refString="Nurse, C. G. (1904) New species of Indian Hymenoptera. Apidae. Journal of Bombay Natural History Society, 15, 557 - 585." type="journal article" year="1904">Nurse (1904)</bibRefCitation>
</td>
</tr>
<tr id="351B34D5F83A8FCCD6F6FAD48BC2A55B" box="[166,1419,1298,1358]" gridrow="6" pageId="28" pageNumber="29">
<th id="76CA5DA9F83A8FCCD6F6FAD48C2CA55B" box="[166,613,1298,1358]" gridcol="0" gridrow="6" pageId="28" pageNumber="29">
<taxonomicName id="4C2B4D14F83A702FD6F6FAD48FB7A53F" authority="Cockerell, 1929" authorityName="Cockerell" authorityYear="1929" box="[166,510,1298,1322]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="mimetes">
<emphasis id="B95FEA85F83A702FD6F6FAD48F1CA53F" box="[166,341,1298,1322]" italics="true" pageId="28" pageNumber="29">Andrena mimetes</emphasis>
<bibRefCitation id="EFBA4B66F83A702FD70CFAD48FB7A53F" author="Cockerell, T. D. A." box="[348,510,1298,1322]" pageId="28" pageNumber="29" pagination="205 - 207" refId="ref43242" refString="Cockerell, T. D. A. (1929) Redhaired bees from China. Entomologist, 62, 205 - 207." type="journal article" year="1929">Cockerell, 1929</bibRefCitation>
</taxonomicName>
</th>
<td id="76CA5DA9F83A8FCCD4D7FAD48DA8A55B" box="[647,993,1298,1358]" gridcol="1" gridrow="6" pageId="28" pageNumber="29">China (Beijing, Inner Mongolia, Shandong)</td>
<td id="76CA5DA9F83A8FCCD255FAD48BC2A55B" box="[1029,1419,1298,1358]" gridcol="2" gridrow="6" pageId="28" pageNumber="29">
<bibRefCitation id="EFBA4B66F83A702FD255FAD48AA3A53F" author="Xu, H. - L. &amp; Tadauchi, O." box="[1029,1258,1298,1322]" pageId="28" pageNumber="29" pagination="63 - 66" refId="ref45916" refString="Xu, H. - L. &amp; Tadauchi, O. (2008) The Subgenera Margandrena, Leimelissa, Stenomelissa and Zonandrena of the Genus Andrena of Eastern Asia (Hymenoptera, Andrenidae). Journal of the Faculty of Agriculture, Kyushu University, 53 (1), 63 - 66." type="journal article" year="2008">Xu &amp; Tadauchi (2008)</bibRefCitation>
</td>
</tr>
<tr id="351B34D5F83A8FCCD6F6FA988BC2A562" box="[166,1419,1374,1399]" gridrow="7" pageId="28" pageNumber="29">
<th id="76CA5DA9F83A8FCCD6F6FA988C2CA562" box="[166,613,1374,1399]" gridcol="0" gridrow="7" pageId="28" pageNumber="29">
<taxonomicName id="4C2B4D14F83A702FD6F6FA988C0CA563" authority="Osytshnjuk, 1983" authorityName="Osytshnjuk" authorityYear="1983" box="[166,581,1374,1398]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="ponomarevae">
<emphasis id="B95FEA85F83A702FD6F6FA988FC2A563" box="[166,395,1374,1398]" italics="true" pageId="28" pageNumber="29">Andrena ponomarevae</emphasis>
Osytshnjuk, 1983
</taxonomicName>
</th>
<td id="76CA5DA9F83A8FCCD4D7FA988DA8A562" box="[647,993,1374,1399]" gridcol="1" gridrow="7" pageId="28" pageNumber="29">Uzbekistan, Kazakhstan, Kyr-</td>
<td id="76CA5DA9F83A8FCCD255FA988BC2A562" box="[1029,1419,1374,1399]" gridcol="2" gridrow="7" pageId="28" pageNumber="29">
<bibRefCitation id="EFBA4B66F83A702FD255FA988AB7A563" author="Osytshnjuk, A. Z. &amp; Romasenko, L. &amp; Banaszak, J. &amp; Motyka, E." box="[1029,1278,1374,1398]" pageId="28" pageNumber="29" refId="ref43693" refString="Osytshnjuk, A. Z., Romasenko, L., Banaszak, J. &amp; Motyka, E. (2008). Andreninae of the Central and Eastern Palearctic. Part 2. Polish Entomological Monographs V. Polish Entomological Society, Poznan, 233 pp." type="book" year="2008">
Osytshnjuk
<emphasis id="B95FEA85F83A702FD22FFA998AFBA563" box="[1151,1202,1374,1398]" italics="true" pageId="28" pageNumber="29">et al.</emphasis>
(2008)
</bibRefCitation>
; Shebl &amp;
</td>
</tr>
<tr id="351B34D5F83A8FCCD6F6FA448BC2A58E" box="[166,1419,1410,1435]" gridrow="8" pageId="28" pageNumber="29" rowspan-0="1">
<td id="76CA5DA9F83A8FCCD4D7FA448DA8A58E" box="[647,993,1410,1435]" gridcol="1" gridrow="8" pageId="28" pageNumber="29">gyzstan</td>
<td id="76CA5DA9F83A8FCCD255FA448BC2A58E" box="[1029,1419,1410,1435]" gridcol="2" gridrow="8" pageId="28" pageNumber="29">Tadauchi (2009)</td>
</tr>
<tr id="351B34D5F83A8FCCD6F6FA6C8BC2A5F3" box="[166,1419,1450,1510]" gridrow="9" pageId="28" pageNumber="29">
<th id="76CA5DA9F83A8FCCD6F6FA6C8C2CA5F3" box="[166,613,1450,1510]" gridcol="0" gridrow="9" pageId="28" pageNumber="29">
<taxonomicName id="4C2B4D14F83A702FD6F6FA6C8E90A5F3" authority="Gusenleitner &amp; Schwarz, 2000" authorityName="Gusenleitner &amp; Schwarz" authorityYear="2000" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="westrichi">
<emphasis id="B95FEA85F83A702FD6F6FA6C8F16A5D7" box="[166,351,1450,1474]" italics="true" pageId="28" pageNumber="29">Andrena westrichi</emphasis>
Gusenleitner &amp; Schwarz, 2000
</taxonomicName>
</th>
<td id="76CA5DA9F83A8FCCD4D7FA6C8DA8A5F3" box="[647,993,1450,1510]" gridcol="1" gridrow="9" pageId="28" pageNumber="29">China (Inner Mongolia)</td>
<td id="76CA5DA9F83A8FCCD255FA6C8BC2A5F3" box="[1029,1419,1450,1510]" gridcol="2" gridrow="9" pageId="28" pageNumber="29">
<bibRefCitation id="EFBA4B66F83A702FD255FA6C8B01A5D7" author="Gusenleitner, F. &amp; Schwarz, M." box="[1029,1352,1450,1474]" pageId="28" pageNumber="29" pagination="1 - 1280" refId="ref43489" refString="Gusenleitner, F. &amp; Schwarz, M. (2002) Weltweite Checkliste der Bienengattung Andrena mit Bemerkungen und Erganzungen zu palaarktischen Arten (Hymenoptera, Apidae, Andreninae, Andrena). Entomofauna, Supplement 10, 1 - 1280." type="journal article" year="2002">Gusenleitner &amp; Schwarz (2002)</bibRefCitation>
</td>
</tr>
</table>
</paragraph>
<paragraph id="8B943697F83A702FD697F9D08CD6A745" blockId="28.[151,1437,1558,2016]" pageId="28" pageNumber="29">
<taxonomicName id="4C2B4D14F83A702FD697F9D08F3AA625" authorityName="Wood" authorityYear="2023" box="[199,371,1558,1584]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="claves">
<emphasis id="B95FEA85F83A702FD697F9D08F3AA625" box="[199,371,1558,1584]" italics="true" pageId="28" pageNumber="29">Andrena claves</emphasis>
</taxonomicName>
can be diagnosed by the simple scutal hairs (
<figureCitation id="13102A12F83A702FD536F9D08DA3A624" box="[870,1002,1558,1585]" captionStart="FIGURE 15" captionStartId="29.[152,255,1846,1870]" captionTargetBox="[151,1436,381,1823]" captionTargetId="figure-105@29.[151,1436,381,1823]" captionTargetPageId="29" captionText="FIGURE 15. Andrena (Leimelissa) claves spec. nov. female. A. Habitus, lateral view, B. Face, frontal view, C. Head, dorsal view, D. Dorsum, E. Propodeum, posterior view, F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7840636" httpUri="https://zenodo.org/record/7840636/files/figure.png" pageId="28" pageNumber="29">Figure 15A</figureCitation>
; with dense, short, and semi-squamous hairs in
<taxonomicName id="4C2B4D14F83A702FD6A1F9FC8C38A641" authority="Morawitz, 1876" authorityName="Morawitz" authorityYear="1876" box="[241,625,1594,1620]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="bairacumensis">
<emphasis id="B95FEA85F83A702FD6A1F9FC8FFAA641" box="[241,435,1594,1620]" italics="true" pageId="28" pageNumber="29">A. bairacumensis</emphasis>
Morawitz, 1876
</taxonomicName>
), by the grey and white vestiture and dark metasomal terga (
<figureCitation id="13102A12F83A702FD342F9FC8BDEA640" box="[1298,1431,1594,1621]" captionStart="FIGURE 15" captionStartId="29.[152,255,1846,1870]" captionTargetBox="[151,1436,381,1823]" captionTargetId="figure-105@29.[151,1436,381,1823]" captionTargetPageId="29" captionText="FIGURE 15. Andrena (Leimelissa) claves spec. nov. female. A. Habitus, lateral view, B. Face, frontal view, C. Head, dorsal view, D. Dorsum, E. Propodeum, posterior view, F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7840636" httpUri="https://zenodo.org/record/7840636/files/figure.png" pageId="28" pageNumber="29">Figure 15D</figureCitation>
; body with extensive red vestiture, terga also yellow-red in
<taxonomicName id="4C2B4D14F83A702FD571F9988B4EA66D" authority="Gusenleitner &amp; Schwarz, 2000" authorityName="Gusenleitner &amp; Schwarz" authorityYear="2000" box="[801,1287,1630,1656]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="westrichi">
<emphasis id="B95FEA85F83A702FD571F9988DEFA66D" box="[801,934,1630,1656]" italics="true" pageId="28" pageNumber="29">A. westrichi</emphasis>
Gusenleitner &amp; Schwarz, 2000
</taxonomicName>
), by the short clypeus with the lower margin only slightly projecting beyond the lower margin of the compound eyes (
<figureCitation id="13102A12F83A702FD301F9448E81A6D4" captionStart="FIGURE 15" captionStartId="29.[152,255,1846,1870]" captionTargetBox="[151,1436,381,1823]" captionTargetId="figure-105@29.[151,1436,381,1823]" captionTargetPageId="29" captionText="FIGURE 15. Andrena (Leimelissa) claves spec. nov. female. A. Habitus, lateral view, B. Face, frontal view, C. Head, dorsal view, D. Dorsum, E. Propodeum, posterior view, F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7840636" httpUri="https://zenodo.org/record/7840636/files/figure.png" pageId="28" pageNumber="29">Figure 15B</figureCitation>
; strongly and clearly projecting beyond the lower margin of the compound eyes in
<taxonomicName id="4C2B4D14F83A702FD22DF9608BD1A6D5" authority="Xu, 1994" authorityName="Xu" authorityYear="1994" box="[1149,1432,1702,1728]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="beijingensis">
<emphasis id="B95FEA85F83A702FD22DF9608B6DA6D5" box="[1149,1316,1702,1728]" italics="true" pageId="28" pageNumber="29">A. beijingensis</emphasis>
<bibRefCitation id="EFBA4B66F83A702FD37CF9608BD1A6D5" author="Xu, H. - L." box="[1324,1432,1702,1728]" pageId="28" pageNumber="29" pagination="197 - 204" refId="ref45800" refString="Xu, H. - L. (1994) Descriptions of new species of genus Andrena from China (Apoidea, Andrenidae). Sinozoologia, 11, 197 - 204. [in Chinese]" type="journal article" year="1994">Xu, 1994</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C2B4D14F83A702FD6C7F90C8C44A6F1" authority="Osytshnjuk, 1984" authorityName="Osytshnjuk" authorityYear="1984" box="[151,525,1738,1764]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="ermolenkoi">
<emphasis id="B95FEA85F83A702FD6C7F90C8F7EA6F1" box="[151,311,1738,1764]" italics="true" pageId="28" pageNumber="29">A. ermolenkoi</emphasis>
Osytshnjuk, 1984
</taxonomicName>
, and
<taxonomicName id="4C2B4D14F83A702FD41DF90C8D94A6F1" authority="Osytshnjuk, 1983" authorityName="Osytshnjuk" authorityYear="1983" box="[589,989,1738,1764]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="ponomarevae">
<emphasis id="B95FEA85F83A702FD41DF90C8D4EA6F1" box="[589,775,1738,1764]" italics="true" pageId="28" pageNumber="29">A. ponomarevae</emphasis>
Osytshnjuk, 1983
</taxonomicName>
), by the short glossa and labial palpus (distinctly elongate in
<taxonomicName id="4C2B4D14F83A702FD7CCF9288C0DA71D" authorityName="Xu" authorityYear="1994" box="[412,580,1774,1800]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="beijingensis">
<emphasis id="B95FEA85F83A702FD7CCF9288C0DA71D" box="[412,580,1774,1800]" italics="true" pageId="28" pageNumber="29">A. beijingensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2B4D14F83A702FD4D0F9288D8BA71D" authority="Cockerell, 1929" authorityName="Cockerell" authorityYear="1929" box="[640,962,1774,1800]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="mimetes">
<emphasis id="B95FEA85F83A702FD4D0F9288CB6A71D" box="[640,767,1774,1800]" italics="true" pageId="28" pageNumber="29">A. mimetes</emphasis>
<bibRefCitation id="EFBA4B66F83A702FD559F9288D8BA71D" author="Cockerell, T. D. A." box="[777,962,1774,1800]" pageId="28" pageNumber="29" pagination="205 - 207" refId="ref43242" refString="Cockerell, T. D. A. (1929) Redhaired bees from China. Entomologist, 62, 205 - 207." type="journal article" year="1929">Cockerell, 1929</bibRefCitation>
</taxonomicName>
,
<typeStatus id="54908835F83A702FD580F9288A7EA71D" box="[976,1079,1774,1800]" pageId="28" pageNumber="29" type="lectotype">lectotype</typeStatus>
designated below), and by the short ocelloccipital distance slightly exceeding the diameter of a lateral ocellus (
<figureCitation id="13102A12F83A702FD247F8D48AD3A738" box="[1047,1178,1810,1837]" captionStart="FIGURE 15" captionStartId="29.[152,255,1846,1870]" captionTargetBox="[151,1436,381,1823]" captionTargetId="figure-105@29.[151,1436,381,1823]" captionTargetPageId="29" captionText="FIGURE 15. Andrena (Leimelissa) claves spec. nov. female. A. Habitus, lateral view, B. Face, frontal view, C. Head, dorsal view, D. Dorsum, E. Propodeum, posterior view, F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7840636" httpUri="https://zenodo.org/record/7840636/files/figure.png" pageId="28" pageNumber="29">Figure 15C</figureCitation>
; equal to two times the diameter of a lateral ocellus in
<taxonomicName id="4C2B4D14F83A702FD7A2F8F08CC6A745" authorityName="Osytshnjuk" authorityYear="1984" box="[498,655,1846,1872]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="ermolenkoi">
<emphasis id="B95FEA85F83A702FD7A2F8F08CC6A745" box="[498,655,1846,1872]" italics="true" pageId="28" pageNumber="29">A. ermolenkoi</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8B943697F83A702ED697F89C8AF8A0EC" blockId="28.[151,1437,1558,2016]" lastBlockId="29.[151,1437,151,357]" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29">
<taxonomicName id="4C2B4D14F83A702FD697F89C8F3DA761" authorityName="Wood" authorityYear="2023" box="[199,372,1882,1908]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="claves">
<emphasis id="B95FEA85F83A702FD697F89C8F3DA761" box="[199,372,1882,1908]" italics="true" pageId="28" pageNumber="29">Andrena claves</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="4C2B4D14F83A702FD417F89C8D3BA760" authority="Eversmann, 1852" authorityName="Eversmann" authorityYear="1852" box="[583,882,1882,1909]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="fallax">
<emphasis id="B95FEA85F83A702FD417F89C8CEFA761" box="[583,678,1882,1908]" italics="true" pageId="28" pageNumber="29">A. fallax</emphasis>
Eversmann, 1852
</taxonomicName>
(=
<taxonomicName id="4C2B4D14F83A702FD5C3F89C8AE3A760" authority="Warncke, 1965" authorityName="Warncke" authorityYear="1965" box="[915,1194,1882,1909]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="ispida">
<emphasis id="B95FEA85F83A702FD5C3F89C8DBFA761" box="[915,1014,1882,1908]" italics="true" pageId="28" pageNumber="29">A. ispida</emphasis>
<bibRefCitation id="EFBA4B66F83A702FD5AEF89C8AE3A760" author="Warncke, K." box="[1022,1194,1882,1909]" pageId="28" pageNumber="29" pagination="27 - 76" refId="ref45141" refString="Warncke, K. (1965) Beitrag zur Kenntnis der Bienengattung Andrena Fabricius in Griechenland. Beitrage zur Entomologie, 15, 27 - 76." type="journal article" year="1965">Warncke, 1965</bibRefCitation>
</taxonomicName>
, see
<bibRefCitation id="EFBA4B66F83A702FD2B0F89C8E96A78D" author="Astafurova, Y. A. &amp; Proshchalykin, M. Y. &amp; Sidorov, D. A." pageId="28" pageNumber="29" pagination="393 - 418" refId="ref42992" refString="Astafurova, Y. A., Proshchalykin, M. Y. &amp; Sidorov, D. A. (2022 b) The type specimens of bees (Hymenoptera, Apoidea) deposited in the Zoological Institute of the Russian Academy of Sciences, St. Petersburg. Contribution V. Family Andrenidae, genus Andrena Fabricius, 1775, species described by E. Eversmann. Zootaxa, 5190 (3), 393 - 418. https: // doi. org / 10.11646 / zootaxa. 5190.3.4" type="journal article" year="2022">
Astafurova
<emphasis id="B95FEA85F83A702FD334F89D8BD4A761" box="[1380,1437,1882,1908]" italics="true" pageId="28" pageNumber="29">et al.</emphasis>
2022b
</bibRefCitation>
) due to the short clypeus. It can be separated by the slightly narrower ocelloccipital distance that is as long as 1 diameter of a lateral ocellus (slightly longer than the diameter of a lateral ocellus), by the broader foveae that occupy two thirds of the distance between the lateral ocellus and the compound eye, the foveae being filled with white hairs (foveae occupying half of this distance, filled with light brown hairs), by the strongly separated hairbands on T2–4 (tergal hairbands complete at least on T3–4), and by the completely flattened and sparsely punctate clypeus (clypeus weakly domed, very densely punctate either side of an impunctate longitudinal line).
</paragraph>
</subSubSection>
<subSubSection id="C331651CF83B702ED697FEC58B72A154" pageId="29" pageNumber="30" type="etymology">
<paragraph id="8B943697F83B702ED697FEC58B72A154" blockId="29.[151,1437,151,357]" pageId="29" pageNumber="30">
<emphasis id="B95FEA85F83B702ED697FEC58F1BA108" bold="true" box="[199,338,259,285]" pageId="29" pageNumber="30">Etymology:</emphasis>
The name is the nominative plural of the Latin noun
<emphasis id="B95FEA85F83B702ED5F9FEC58DA3A108" box="[937,1002,259,285]" italics="true" pageId="29" pageNumber="30">clavis</emphasis>
meaning ‘key’, as the pattern of white metasoma hairbands on the contrasting black terga is reminiscent of piano keys. It is a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C331651CF83B702ED697FE8C8C29A170" box="[199,608,330,357]" pageId="29" pageNumber="30" type="distribution">
<paragraph id="8B943697F83B702ED697FE8C8C29A170" blockId="29.[151,1437,151,357]" box="[199,608,330,357]" pageId="29" pageNumber="30">
<emphasis id="B95FEA85F83B702ED697FE8C8F2AA171" bold="true" box="[199,355,330,356]" pageId="29" pageNumber="30">Distribution:</emphasis>
Southern
<collectingCountry id="F33C7607F83B702ED786FE8D8C15A170" box="[470,604,331,357]" name="Kazakhstan" pageId="29" pageNumber="30">Kazakhstan</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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