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<document id="9687824D5A74141D3ADD78CD4A5748CE" ID-CLB-Dataset="51608" ID-DOI="10.11646/zootaxa.4091.1.1" ID-GBIF-Dataset="bdd057d5-b4d6-4b57-940a-d7839f483e25" ID-ISSN="1175-5326" ID-Zenodo-Dep="265332" ID-ZooBank="urn:lsid:zoobank.org:pub:76D13D36-682E-4E91-AC91-693CA9D3D465" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1457950950343" checkinUser="plazi" docAuthor="Chris J. Hodgson &amp; Douglas J. Williams" docDate="2016" docId="03F2FF48815E0D4924B6ABE9FDC9FEEA" docLanguage="en" docName="15613-5135-1-PB.pdf.imf" docOrigin="Zootaxa 4091 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Antecerococcus steppicus Balachowsky, comb. nov." docType="treatment" docVersion="16" lastPageNumber="120" masterDocId="FFCB8730812B0D3E2421AB7EFF8EFF8E" masterDocTitle="(Hemiptera: Sternorrhyncha, Coccomorpha) with particular reference to species from the Afrotropical, western Palaearctic and western Oriental Regions, with the revival of Antecerococcus Green and description of a new genus and fifteen new species, and with ten new synonomies" masterLastPageNumber="175" masterPageNumber="1" pageNumber="118" updateTime="1698185172771" updateUser="ExternalLinkService">
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<mods:title id="40151C4D5AB4AECF182C318B98847649">(Hemiptera: Sternorrhyncha, Coccomorpha) with particular reference to species from the Afrotropical, western Palaearctic and western Oriental Regions, with the revival of Antecerococcus Green and description of a new genus and fifteen new species, and with ten new synonomies</mods:title>
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<mods:namePart id="485F62486AA694EEC25EE993E1DB35F3">Chris J. Hodgson</mods:namePart>
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<mods:namePart id="D2B3DA91239890C11ACFB8045B68E4AC">Douglas J. Williams</mods:namePart>
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<mods:title id="5D06ECE5A32A1209BB5EF89402588EB4">Zootaxa</mods:title>
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<mods:date id="252501825613745FA5F2CC62E0BA2210">2016</mods:date>
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<treatment id="03F2FF48815E0D4924B6ABE9FDC9FEEA" ID-DOI="http://doi.org/10.5281/zenodo.6081619" ID-GBIF-Taxon="117911318" ID-Zenodo-Dep="6081619" LSID="urn:lsid:plazi:treatment:03F2FF48815E0D4924B6ABE9FDC9FEEA" httpUri="http://treatment.plazi.org/id/03F2FF48815E0D4924B6ABE9FDC9FEEA" lastPageId="119" lastPageNumber="120" pageId="117" pageNumber="118">
<subSubSection id="C3411DD5815E0D4B24B6ABE9FD03FE96" pageId="117" pageNumber="118" type="nomenclature">
<paragraph id="8BE44E5E815E0D4B24B6ABE9FCAEFF3C" blockId="117.[151,800,151,211]" box="[151,800,151,178]" pageId="117" pageNumber="118">
<heading id="D0ACF932815E0D4B24B6ABE9FCAEFF3C" bold="true" box="[151,800,151,178]" fontSize="11" level="1" pageId="117" pageNumber="118" reason="1">
<emphasis id="B92F924C815E0D4B24B6ABE9FCAEFF3C" bold="true" box="[151,800,151,178]" pageId="117" pageNumber="118">
<taxonomicName id="4C5B35DD815E0D4B24B6ABE9FD03FF3C" authority="Balachowsky" authorityName="Balachowsky" box="[151,653,151,178]" class="Insecta" family="Cerococcidae" genus="Antecerococcus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="117" pageNumber="118" phylum="Arthropoda" rank="species" species="steppicus" status="comb. nov.">
<emphasis id="B92F924C815E0D4B24B6ABE9FE42FF3F" bold="true" box="[151,460,151,177]" italics="true" pageId="117" pageNumber="118">Antecerococcus steppicus</emphasis>
(Balachowsky)
</taxonomicName>
,
<taxonomicNameLabel id="A21C2F37815E0D4B26B8ABE6FCAEFF3C" box="[665,800,152,178]" pageId="117" pageNumber="118">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE44E5E815E0D4B24B6ABC4FF76FF5D" blockId="117.[151,800,151,211]" box="[151,248,186,211]" pageId="117" pageNumber="118">
(
<figureCitation id="136052DB815E0D4B24BEABC4FF7FFF5D" box="[159,241,186,211]" captionStart="FIGURE 45" captionStartId="118.[151,250,1823,1845]" captionTargetBox="[210,1392,195,1776]" captionTargetId="figure@118.[190,1397,195,1802]" captionTargetPageId="118" captionText="FIGURE 45. Antecerococcus steppicus (Balachowsky). Adult female." httpUri="https://zenodo.org/record/265378/files/figure.png" pageId="117" pageNumber="118">Fig. 45</figureCitation>
)
</paragraph>
<paragraph id="8BE44E5E815E0D4B24B6AA7CFD03FE96" blockId="117.[151,653,258,280]" box="[151,653,258,280]" pageId="117" pageNumber="118">
<taxonomicName id="4C5B35DD815E0D4B24B6AA7CFDEEFE96" authority="Balachowsky 1941: 87" authorityName="Balachowsky" authorityPageNumber="87" authorityYear="1941" box="[151,608,258,280]" class="Insecta" family="Cerococcidae" genus="Cerococcus" kingdom="Animalia" order="Hemiptera" pageId="117" pageNumber="118" phylum="Arthropoda" rank="species" species="steppicus">
<emphasis id="B92F924C815E0D4B24B6AA7CFEFFFE96" box="[151,369,258,280]" italics="true" pageId="117" pageNumber="118">Cerococcus steppicus</emphasis>
Balachowsky 1941: 87
</taxonomicName>
90.
</paragraph>
</subSubSection>
<subSubSection id="C3411DD5815E0D4B24B6AA38FAB2FBDC" pageId="117" pageNumber="118" type="materials_examined">
<paragraph id="8BE44E5E815E0D4B24B6AA38FE69FE44" blockId="117.[151,1437,325,1934]" pageId="117" pageNumber="118">
<emphasis id="B92F924C815E0D4B24B6AA38FE36FED0" bold="true" box="[151,440,325,350]" pageId="117" pageNumber="118">
<typeStatus id="54E0F0FC815E0D4B24B6AA38FF5CFED3" box="[151,210,326,349]" pageId="117" pageNumber="118">Type</typeStatus>
details.
<collectingCountry id="F34C0ECE815E0D4B2514AA3BFE36FED0" box="[309,440,325,350]" name="Algeria" pageId="117" pageNumber="118">ALGERIA</collectingCountry>
</emphasis>
, Ben-Toumi, Sud Beghani, on
<taxonomicName id="4C5B35DD815E0D4B2737AA3BFC7DFED3" box="[790,1011,325,350]" class="Magnoliopsida" family="Asteraceae" genus="Atractylis" kingdom="Animalia" order="Asterales" pageId="117" pageNumber="118" phylum="Tracheophyta" rank="species" species="cespitosa">
<emphasis id="B92F924C815E0D4B2737AA3BFC7DFED3" box="[790,1011,325,350]" italics="true" pageId="117" pageNumber="118">Atractylis cespitosa</emphasis>
</taxonomicName>
[misspelling of
<taxonomicName id="4C5B35DD815E0D4B208AAA38FAAFFED3" box="[1195,1313,326,349]" class="Magnoliopsida" family="Asteraceae" genus="Atractylis" kingdom="Animalia" order="Asterales" pageId="117" pageNumber="118" phylum="Tracheophyta" rank="species" species="caespitosa">
<emphasis id="B92F924C815E0D4B208AAA38FAAFFED3" box="[1195,1313,326,349]" italics="true" pageId="117" pageNumber="118">caespitosa</emphasis>
</taxonomicName>
], -.
<date id="FFE5689E815E0D4B2164AA38FA19FED0" box="[1349,1431,326,350]" pageId="117" pageNumber="118" value="1939-05">v.1939</date>
, A. Pasquier.
<emphasis id="B92F924C815E0D4B2506AA17FD9AFE0F" bold="true" box="[295,532,361,386]" pageId="117" pageNumber="118">Depository: MNHN</emphasis>
:
<typeStatus id="54E0F0FC815E0D4B2602AA14FD00FE0F" box="[547,654,362,385]" pageId="117" pageNumber="118" type="lectotype">
<emphasis id="B92F924C815E0D4B2602AA14FD00FE0F" bold="true" box="[547,654,362,385]" pageId="117" pageNumber="118">lectotype</emphasis>
</typeStatus>
adf (designated by Lambdin &amp; Kosztarab 1977: 216) (MNHN 4869- 2);
<typeStatus id="54E0F0FC815E0D4B249DAAF0FED1FE2B" box="[188,351,398,421]" pageId="117" pageNumber="118" type="paralectotype">
<emphasis id="B92F924C815E0D4B249DAAF0FED1FE2B" bold="true" box="[188,351,398,421]" pageId="117" pageNumber="118">paralectotype</emphasis>
</typeStatus>
ff: data as for
<typeStatus id="54E0F0FC815E0D4B2621AAF3FDE4FE28" box="[512,618,397,422]" pageId="117" pageNumber="118" type="lectotype">lectotype</typeStatus>
, 4/8 adff (MNHN 4869-1: 2adff); MNHN 4869-3: 2adff; MNHN 4869-4: 2adff; MNHN 4869-5: 2adff).
</paragraph>
<paragraph id="8BE44E5E815E0D4B24E6AAABFDA4FDB8" blockId="117.[151,1437,325,1934]" pageId="117" pageNumber="118">
<emphasis id="B92F924C815E0D4B24E6AAABFE10FE60" bold="true" box="[199,414,469,494]" pageId="117" pageNumber="118">Material studied.</emphasis>
<typeStatus id="54E0F0FC815E0D4B258EAAA8FD91FE63" box="[431,543,470,493]" pageId="117" pageNumber="118" type="lectotype">Lectotype</typeStatus>
f:
<collectingCountry id="F34C0ECE815E0D4B2670AAA8FD41FE60" box="[593,719,470,494]" name="Algeria" pageId="117" pageNumber="118">ALGERIA</collectingCountry>
, Ben-Toumi, Sud Beghani, on
<taxonomicName id="4C5B35DD815E0D4B2078AAABFAC3FE63" box="[1113,1357,469,494]" class="Magnoliopsida" family="Asteraceae" genus="Atractylis" kingdom="Animalia" order="Asterales" pageId="117" pageNumber="118" phylum="Tracheophyta" rank="species" species="caespitosa">
<emphasis id="B92F924C815E0D4B2078AAABFAC3FE63" box="[1113,1357,469,494]" italics="true" pageId="117" pageNumber="118">Atractylis caespitosa</emphasis>
</taxonomicName>
(spelt
<taxonomicName id="4C5B35DD815E0D4B24B6AA84FF71FD9F" box="[151,255,506,529]" class="Magnoliopsida" family="Asteraceae" genus="Atractylis" kingdom="Animalia" order="Asterales" pageId="117" pageNumber="118" phylum="Tracheophyta" rank="species" species="cespitosa">
<emphasis id="B92F924C815E0D4B24B6AA84FF71FD9F" box="[151,255,506,529]" italics="true" pageId="117" pageNumber="118">cespitosa</emphasis>
</taxonomicName>
) (
<taxonomicName id="4C5B35DD815E0D4B2500AA87FE2FFD9C" box="[289,417,505,530]" class="Magnoliopsida" family="Asteraceae" kingdom="Plantae" order="Asterales" pageId="117" pageNumber="118" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
), -.
<date id="FFE5689E815E0D4B25EBAA87FD92FD9F" box="[458,540,505,529]" pageId="117" pageNumber="118" value="1939-05">v.1939</date>
, A. Pasquier (MNHN): 1/1 adf (g, labelled co-type) and (MNHN): 1/2
<typeStatus id="54E0F0FC815E0D4B24B6A963FEA0FDB8" box="[151,302,541,566]" pageId="117" pageNumber="118" type="paralectotype">paralectotype</typeStatus>
adff (f, labelled
<typeStatus id="54E0F0FC815E0D4B25C9A963FD93FDB8" box="[488,541,541,566]" pageId="117" pageNumber="118">type</typeStatus>
).
</paragraph>
<paragraph id="8BE44E5E815E0D4B24E6A93CFBF3FDD4" blockId="117.[151,1437,325,1934]" box="[199,1149,577,602]" pageId="117" pageNumber="118">
<emphasis id="B92F924C815E0D4B24E6A93CFE29FDD4" bold="true" box="[199,423,577,602]" pageId="117" pageNumber="118">Mounted material.</emphasis>
Body roundly pear-shaped,
<quantity id="4CA3E3BB815E0D4B26C5A93FFCE5FDD7" box="[740,875,577,601]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.4" metricValueMin="1.6" pageId="117" pageNumber="118" unit="mm" value="2.0" valueMax="2.4" valueMin="1.6">1.62.4 mm</quantity>
long,
<quantity id="4CA3E3BB815E0D4B2793A93FFBB7FDD7" box="[946,1081,577,601]" metricMagnitude="-3" metricUnit="m" metricValue="1.8499999999999999" metricValueMax="2.3" metricValueMin="1.4" pageId="117" pageNumber="118" unit="mm" value="1.8499999999999999" valueMax="2.3" valueMin="1.4">1.42.3 mm</quantity>
wide.
</paragraph>
<paragraph id="8BE44E5E815E0D4B24E6A918FAB2FBDC" blockId="117.[151,1437,325,1934]" pageId="117" pageNumber="118">
<emphasis id="B92F924C815E0D4B24E6A918FEA0FDF3" bold="true" box="[199,302,614,637]" pageId="117" pageNumber="118">Dorsum.</emphasis>
Eight-shaped pores all small, of 3 sizes: (i) largest each
<date id="FFE5689E815E0D4B2788A918FB8AFDF3" box="[937,1028,614,638]" pageId="117" pageNumber="118" value="2011-10-12">1112 x</date>
6.57.0 µm, restricted to close to or within each stigmatic pore band, with 05 near apex of each band, plus a few (3 or 4) along margins of posteriormost abdominal segments; (ii) intermediate-sized pores slightly smaller, each 9.510.0 x 6.06.5 µm, abundant throughout most of dorsum, randomly distributed but absent medially posterior to abdominal segment IV; and (iii) a small pore, each 6.5
<date id="FFE5689E815E0D4B25BDA988FE4FFC80" box="[412,449,758,782]" pageId="117" pageNumber="118">x 3</date>
µm, sparsely distributed medially posterior to cribriform plates; smallest pores absent from stigmatic pore bands. Simple pores, each 2.02.5 µm wide, rare. Cribriform plates roundish, small, each mainly 1620 µm wide, in 2 groups of 3 or 4 submedially on abdominal segment IV; each plate with a narrow border and moderate-sized micropores. Dorsal setae showing nothing distinctive. Tubular ducts with each outer duct 2325 µm long and about 2.5 µm wide, slightly broader than those on venter; abundant throughout. Anal lobes mainly membranous, but with sclerotized inner margins; each lobe with a long apical seta, all broken but at least 180 µm long; fleshy setae on dorsal surface near apex each 2022 µm long; more anterior basal fleshy setae each 2531 µm long; ventral setose seta near apical setae, each about 35 µm long; medioventral setae possibly absent; outer margin setae each 15 µm long; each lobe with two or three 8-shaped pores. Median anal plate roundly pointed, 3340 µm long and 3340 µm wide at base. Anal ring with 4 pairs of setae, each 7075 µm long.
</paragraph>
</subSubSection>
<subSubSection id="C3411DD5815E0D4B24E6AF20FE07F938" pageId="117" pageNumber="118" type="description">
<paragraph id="8BE44E5E815E0D4B24E6AF20FE07F938" blockId="117.[151,1437,325,1934]" pageId="117" pageNumber="118">
<emphasis id="B92F924C815E0D4B24E6AF20FE96FBFB" bold="true" box="[199,280,1118,1141]" pageId="117" pageNumber="118">Venter.</emphasis>
Eight-shaped pores similar to intermediate-sized pores on dorsum, in a sparse, fairly broad submarginal band and also in broad transverse bands anterior to each band of multilocular disc-pores on abdomen. Simple pores rare. Small bilocular pores, each about
<date id="FFE5689E815E0D4B26C5AFD8FC88FB33" box="[740,774,1190,1213]" pageId="117" pageNumber="118">5 x</date>
4 µm, frequent medially on head and thorax, not extending onto anterior abdominal segments. Spiracular disc-pores small, each 45 µm wide (smallest near spiracles), mainly with 5 loculi, each band mainly 25 pores wide but widening slightly at apex on dorsum; posterior bands bifurcated; each band with 4560 pores + 05 largest 8-shaped pores; also with small groups between each spiracle and anterior and posterior leg stubs, plus 2 quinquelocular disc-pores near each antenna and 2 near labium. Small convex closed pores absent. Multilocular disc-pores, each 7.58.0 µm wide with mainly 10 loculi, in transverse bands 24 pores wide across most abdominal segments, as follows:
<date id="FFE5689E815E0D4B2781AE00FB8DFA18" box="[928,1027,1406,1430]" pageId="117" pageNumber="118">VIII 46</date>
submarginally and 0 or 1 medially; VII with 1 or 2 on each side of vulva, none medially; VI with 14 submarginally + 1417 medially; V with 36 submarginally + 3341 medially; IV with 612 submarginally + 4869 medially;
<date id="FFE5689E815E0D4B2012AEB8FB01FA50" box="[1075,1167,1478,1502]" pageId="117" pageNumber="118" value="2012-03-06">III 612</date>
submarginally + 4658 medially;
<date id="FFE5689E815E0D4B252BAE94FED0F98C" box="[266,350,1514,1538]" pageId="117" pageNumber="118" value="2015-02-08">II 815</date>
submarginally + 3335 medially; also with 1 or 2 laterad to each metathoracic leg stub; none medially on metathorax. Tubular ducts slightly narrower than those on dorsum; present throughout. Ventral setae showing nothing significant; preanal setae each 90100 µm long; companion setae short. Leg stubs present. Antennae unsegmented, each 3340 µm long, 3340 µm wide, without either a sclerotized conical point on apex or a distinct setal cavity; with 7 or 8 setose and fleshy setae. Clypeolabral shield 165 µm long. Spiracular peritremes each 3840 µm wide.
</paragraph>
</subSubSection>
<subSubSection id="C3411DD5815E0D4924E6ADBFFDC9FEEA" lastPageId="119" lastPageNumber="120" pageId="117" pageNumber="118" type="discussion">
<paragraph id="8BE44E5E815E0D4924E6ADBFFB1EFE93" blockId="117.[151,1437,325,1934]" lastBlockId="119.[151,1436,151,357]" lastPageId="119" lastPageNumber="120" pageId="117" pageNumber="118">
<emphasis id="B92F924C815E0D4B24E6ADBFFEB3F954" bold="true" box="[199,317,1729,1754]" pageId="117" pageNumber="118">Comment</emphasis>
. The above description is similar to that of Lambdin and Kosztarab (1977) but some of the measurements are slightly larger and they did not note the smaller 8-shaped pores medially on the posterior abdominal segments. The adult female of
<taxonomicName id="4C5B35DD815E0D4B2652AC74FD75F8AF" box="[627,763,1802,1826]" class="Insecta" family="Cerococcidae" genus="Cerococcus" kingdom="Animalia" order="Hemiptera" pageId="117" pageNumber="118" phylum="Arthropoda" rank="species" species="steppicus">
<emphasis id="B92F924C815E0D4B2652AC74FD75F8AF" box="[627,763,1802,1826]" italics="true" pageId="117" pageNumber="118">A. steppicus</emphasis>
</taxonomicName>
is distinctive, with the following combination of characterstates: (i) dorsum with three sizes of 8-shaped pores, all small; (ii) largest 8-shaped pores very close to or restricted to within each stigmatic pore band; (iii) smallest 8-shaped pores restricted to medially on posterior abdominal segments; (iv) intermediate-sized 8-shaped pore randomly distributed over rest of dorsum; (v) cribriform plates in a submedial group of three or four on each side of abdominal segment IV; (vi) leg stubs present; (vii) posterior discpore bands bifurcated; (viii) multilocular disc-pores fairly abundant in transverse bands across all abdominal segments and submarginally on metathorax; (ix) a few quinquelocular disc-pores also present between each spiracle and associated leg stubs; and (x) antennae without a cone-like apex or setal cavity.
</paragraph>
<caption id="DF241ED6815D0D4824B6AC61FCECF8BB" box="[151,866,1823,1845]" httpUri="https://zenodo.org/record/265378/files/figure.png" pageId="118" pageNumber="119" targetBox="[210,1392,195,1776]" targetPageId="118">
<paragraph id="8BE44E5E815D0D4824B6AC61FCECF8BB" blockId="118.[151,866,1823,1845]" box="[151,866,1823,1845]" pageId="118" pageNumber="119">
<emphasis id="B92F924C815D0D4824B6AC61FEAEF8BB" bold="true" box="[151,288,1823,1845]" pageId="118" pageNumber="119">FIGURE 45.</emphasis>
<taxonomicName id="4C5B35DD815D0D482504AC61FD42F8BB" authority="Balachowsky" authorityName="Balachowsky" box="[293,716,1823,1845]" class="Insecta" family="Cerococcidae" genus="Antecerococcus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="118" pageNumber="119" phylum="Arthropoda" rank="species" species="steppicus">
<emphasis id="B92F924C815D0D482504AC61FDA7F8BB" box="[293,553,1823,1845]" italics="true" pageId="118" pageNumber="119">Antecerococcus steppicus</emphasis>
(Balachowsky)
</taxonomicName>
. Adult female.
</paragraph>
</caption>
<paragraph id="8BE44E5E815C0D4924E6AA59FDC9FEEA" blockId="119.[151,1436,151,357]" pageId="119" pageNumber="120">
The adult female of
<taxonomicName id="4C5B35DD815C0D492593AA56FDB2FEB1" box="[434,572,296,320]" class="Insecta" family="Cerococcidae" genus="Antecerococcus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="119" pageNumber="120" phylum="Arthropoda" rank="species" species="steppicus">
<emphasis id="B92F924C815C0D492593AA56FDB2FEB1" box="[434,572,296,320]" italics="true" pageId="119" pageNumber="120">A. steppicus</emphasis>
</taxonomicName>
falls within Group B in the key to species of
<taxonomicName id="4C5B35DD815C0D492070AA56FA8FFECE" box="[1105,1281,296,320]" class="Insecta" family="Cerococcidae" genus="Antecerococcus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="119" pageNumber="120" phylum="Arthropoda" rank="genus">
<emphasis id="B92F924C815C0D492070AA56FA8FFECE" box="[1105,1281,296,320]" italics="true" pageId="119" pageNumber="120">Antecerococcus</emphasis>
</taxonomicName>
and keys out close to
<taxonomicName id="4C5B35DD815C0D4924D5AA32FE1FFEEB" box="[244,401,332,357]" class="Insecta" family="Cerococcidae" genus="Antecerococcus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="119" pageNumber="120" phylum="Arthropoda" rank="species" species="kakamegae">
<emphasis id="B92F924C815C0D4924D5AA32FE1FFEEB" box="[244,401,332,357]" italics="true" pageId="119" pageNumber="120">A. kakamegae</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5B35DD815C0D4925E6AA32FDB1FEEA" box="[455,575,332,356]" class="Insecta" family="Cerococcidae" genus="Antecerococcus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="119" pageNumber="120" phylum="Arthropoda" rank="species" species="royenae">
<emphasis id="B92F924C815C0D4925E6AA32FDB1FEEA" box="[455,575,332,356]" italics="true" pageId="119" pageNumber="120">A. royenae</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>