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<document id="C8F21765387C38DCA4C35E7841CC7DD6" ID-DOI="10.11646/zootaxa.4748.2.7" ID-GBIF-Dataset="ae7f2b81-fd03-4ded-b33d-9f086dd3a0c8" ID-ISSN="1175-5326" ID-Zenodo-Dep="3698881" ID-ZooBank="1110012B-A68A-449D-BA03-AEB0DE911BFB" IM.bibliography_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1583482775224" checkinUser="plazi" docAuthor="Wang, Feng-Yan &amp; Zhou, Hong-Zhang" docDate="2020" docId="03FA3308FFCA323D1ABE22D6FA312383" docLanguage="en" docName="zootaxa.4748.2.7.pdf" docOrigin="Zootaxa 4748 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Exomis Weise 1889" docType="treatment" docVersion="1" lastPageNumber="352" masterDocId="FFC34B70FFCB323C1A29202BFFAD245E" masterDocTitle="Taxonomy of the leaf beetle genus Exomis Weise (Coleoptera: Chrysomelidae: Clytrini) with description of six new species from China" masterLastPageNumber="364" masterPageNumber="351" pageNumber="352" updateTime="1728995781755" updateUser="juliana" zenodo-license-document="CLOSED">
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<mods:title id="8DDD5DFD16AC7525B2684250E0DBD686">Taxonomy of the leaf beetle genus Exomis Weise (Coleoptera: Chrysomelidae: Clytrini) with description of six new species from China</mods:title>
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<mods:namePart id="8006B4D1432C923472C6EF980159A7B4">Wang, Feng-Yan</mods:namePart>
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<emphasis id="B9275E0CFFCA323D1ABE22D6FE492749" bold="true" box="[151,484,765,791]" pageId="1" pageNumber="352">
Genus
<taxonomicName id="4C53F99DFFCA323D1AC722D6FE492749" authority="Weise, 1889" authorityName="Weise" authorityYear="1889" box="[238,484,765,791]" class="Insecta" family="Chrysomelidae" genus="Exomis" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="352" phylum="Arthropoda" rank="genus">
<emphasis id="B9275E0CFFCA323D1AC722D6FEE42749" bold="true" box="[238,329,765,791]" italics="true" pageId="1" pageNumber="352">Exomis</emphasis>
<bibRefCitation id="EFC2FFEFFFCA323D1B7822D6FE492749" author="Weise, J." box="[337,484,765,791]" pageId="1" pageNumber="352" pagination="560 - 653" refId="ref9504" refString="Weise, J. (1889) Insecta, a CL. G. N. Potanin in China et in Mongolia novissime lecta. IX. Chrysomelidae et Coccinellidae. Horae Societatis Entomologicae Rossicae, 23, 560 - 653." type="journal article" year="1889">Weise, 1889</bibRefCitation>
</taxonomicName>
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<paragraph id="8BEC821EFFCA323D1ABE236AFAB227C9" blockId="1.[151,1436,833,950]" pageId="1" pageNumber="352">
<bibRefCitation id="EFC2FFEFFFCA323D1ABE236AFEE72704" author="Weise, J." box="[151,330,833,858]" pageId="1" pageNumber="352" pagination="560 - 653" refId="ref9504" refString="Weise, J. (1889) Insecta, a CL. G. N. Potanin in China et in Mongolia novissime lecta. IX. Chrysomelidae et Coccinellidae. Horae Societatis Entomologicae Rossicae, 23, 560 - 653." type="journal article" year="1889">Weise, 1889: 563</bibRefCitation>
, 577;
<bibRefCitation id="EFC2FFEFFFCA323D1BA1236AFD102704" author="Jacoby, M. &amp; Clavareau, H." box="[392,701,833,858]" pageId="1" pageNumber="352" pagination="1 - 88" refId="ref9130" refString="Jacoby, M. &amp; Clavareau, H. (1906) Coleoptera Phytophaga, Fam. Chrysomelidae, Subfam. Clytrinae. In: Wytsman, P. (Ed.), Genera Insectorum, Londres &amp; Bruxelles, 49, pp. 1 - 88 + 1 [corrigenda], 5 pl. [1906 - 1907]" type="journal article" year="1906">Jacoby &amp; Clavareau, 1906: 48</bibRefCitation>
(subg. of
<taxonomicName id="4C53F99DFFCA323D190C2369FC472704" authorityName="Lacordaire" authorityYear="1848" box="[805,1002,834,858]" class="Insecta" family="Chrysomelidae" genus="Gynandrophthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="352" phylum="Arthropoda" rank="genus">
<emphasis id="B9275E0CFFCA323D190C2369FC472704" box="[805,1002,834,858]" italics="true" pageId="1" pageNumber="352">Gynandrophthalma</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFC2FFEFFFCA323D1E29236AFB602704" author="Clavareau, H." box="[1024,1229,833,858]" pageId="1" pageNumber="352" pagination="1 - 278" refId="ref8984" refString="Clavareau, H. (1913) Chrysomelidae: 5. Megascelinae, 6. Megalopodinae, 7. Clytrinae, 8. Cryptocephalinae, 9. Chlamydinae, 10. Lamprosomatinae. In: Junk, W. &amp; Schenkling, S. (Eds.), Coleopterorun Catalogus. Pars 53. W. Junk, Berlin, pp. 1 - 278." type="book chapter" year="1913">Clavareau, 1913: 67</bibRefCitation>
(subg. of
<taxonomicName id="4C53F99DFFCA323D1F1C2369FA212707" authorityName="Chevrolat in Dejean" authorityYear="1836" box="[1333,1420,834,857]" class="Insecta" family="Lycaenidae" genus="Cyaniris" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="352" phylum="Arthropoda" rank="genus">
<emphasis id="B9275E0CFFCA323D1F1C2369FA212707" box="[1333,1420,834,857]" italics="true" pageId="1" pageNumber="352">Cyaniris</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFC2FFEFFFCA323D1AEE234BFE272726" author="Gressitt, J. L." box="[199,394,864,889]" pageId="1" pageNumber="352" pagination="325 - 376" refId="ref9051" refString="Gressitt, J. L. (1942) Plant-beetles from south and west China. III. Clytrinae, Cryptocephalinae and Chlamydinae. Lingnan Science Journal, 20, 325 - 376, pls. 19 - 22." type="journal article" year="1942">Gressitt, 1942: 328</bibRefCitation>
;
<bibRefCitation id="EFC2FFEFFFCA323D1BBD234BFD1B2726" author="Gressitt, J. L. &amp; Kimoto, S." box="[404,694,864,889]" pageId="1" pageNumber="352" pagination="1 - 299" refId="ref9091" refString="Gressitt, J. L. &amp; Kimoto, S. (1961) The Chrysomelidae (Coleopt.) of China and Korea. Part 1. Pacific insects Monograph, 1 A, 1 - 299." type="journal article" year="1961">Gressitt &amp; Kimoto, 1961: 93</bibRefCitation>
;
<bibRefCitation id="EFC2FFEFFFCA323D18E9234BFC6E2726" author="Seeno, T. N. &amp; Wilcox, J. A." box="[704,963,864,889]" pageId="1" pageNumber="352" pagination="1 - 222" refId="ref9471" refString="Seeno, T. N. &amp; Wilcox, J. A. (1982) Leaf beetle genera (Coleoptera: Chrysomelidae). Entomography, 1, 1 - 222." type="journal article" year="1982">Seeno &amp; Wilcox 1982: 33</bibRefCitation>
(synonym of
<taxonomicName id="4C53F99DFFCA323D1E66234BFB662726" authorityName="Chevrolat in Dejean" authorityYear="1836" box="[1103,1227,864,888]" class="Insecta" family="Chrysomelidae" genus="Smaragdina" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="352" phylum="Arthropoda" rank="genus">
<emphasis id="B9275E0CFFCA323D1E66234BFB662726" box="[1103,1227,864,888]" italics="true" pageId="1" pageNumber="352">Smaragdina</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFC2FFEFFFCA323D1EC9234BFEFB27C9" author="Regalin, R. &amp; Medvedev, L. N." pageId="1" pageNumber="352" pagination="76 - 79" refId="ref9351" refString="Regalin, R. &amp; Medvedev, L. N. (2010 a) New acts and comments. Chrysomelidae: Cryptocephalinae: Clytrini. In: Lobl, I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 6. Chrysomeloidea. Apollo Books, Stenstrup, pp. 76 - 79." type="book chapter" year="2010">Regalin &amp; Medvedev 2010a: 78</bibRefCitation>
(resurrected from synonymy with
<taxonomicName id="4C53F99DFFCA323D189E2354FC9E27C9" authorityName="Chevrolat in Dejean" authorityYear="1836" box="[695,819,895,919]" class="Insecta" family="Chrysomelidae" genus="Smaragdina" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="352" phylum="Arthropoda" rank="genus">
<emphasis id="B9275E0CFFCA323D189E2354FC9E27C9" box="[695,819,895,919]" italics="true" pageId="1" pageNumber="352">Smaragdina</emphasis>
</taxonomicName>
),
<bibRefCitation id="EFC2FFEFFFCA323D19612355FB0D27C9" author="Regalin, R. &amp; Medvedev, L. N." box="[840,1184,894,919]" pageId="1" pageNumber="352" pagination="564 - 580" refId="ref9415" refString="Regalin, R. &amp; Medvedev, L. N. (2010 b) Chrysomelidae: Cryptocephalinae: Clytrini. In: Lobl, I. &amp; Smetana, A. (Ed.), Catalogue of Palaearctic Coleoptera. Vol. 6. Apollo Books, Stenstrup, pp. 564 - 580" type="book chapter" year="2010">Regalin &amp; Medvedev, 2010b: 569</bibRefCitation>
(catalogue).
</paragraph>
</subSubSection>
<subSubSection id="C349D195FFCA323D1ABE23B5FCC527E8" box="[151,872,925,950]" pageId="1" pageNumber="352" type="type_taxon">
<paragraph id="8BEC821EFFCA323D1ABE23B5FCC527E8" blockId="1.[151,1436,833,950]" box="[151,872,925,950]" pageId="1" pageNumber="352">
<emphasis id="B9275E0CFFCA323D1ABE23B5FE8B27E8" bold="true" box="[151,294,925,950]" pageId="1" pageNumber="352">
<typeStatus id="54E83CBCFFCA323D1ABE23B5FF6127E8" box="[151,204,926,950]" pageId="1" pageNumber="352">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="4C53F99DFFCA323D1B0523B5FD3A27E8" authority="Weise, 1889" authorityName="Weise, H" authorityYear="1889" box="[300,663,925,950]" class="Insecta" family="Chrysomelidae" genus="Exomis" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="352" phylum="Arthropoda" rank="species" species="peplopteroides">
<emphasis id="B9275E0CFFCA323D1B0523B5FDBE27E8" box="[300,531,926,950]" italics="true" pageId="1" pageNumber="352">Exomis peplopteroides</emphasis>
<bibRefCitation id="EFC2FFEFFFCA323D183123B6FD3A27E8" author="Weise, J." box="[536,663,925,950]" pageId="1" pageNumber="352" pagination="560 - 653" refId="ref9504" refString="Weise, J. (1889) Insecta, a CL. G. N. Potanin in China et in Mongolia novissime lecta. IX. Chrysomelidae et Coccinellidae. Horae Societatis Entomologicae Rossicae, 23, 560 - 653." type="journal article" year="1889">Weise, 1889</bibRefCitation>
</taxonomicName>
, fixed by monotypy.
</paragraph>
</subSubSection>
<subSubSection id="C349D195FFCA323D1ABE23C8FDBA2139" pageId="1" pageNumber="352" type="diagnosis">
<paragraph id="8BEC821EFFCA323D1ABE23C8FE6420D0" blockId="1.[151,1437,995,1383]" pageId="1" pageNumber="352">
<emphasis id="B9275E0CFFCA323D1ABE23C8FEB927A0" bold="true" box="[151,276,995,1022]" pageId="1" pageNumber="352">Diagnosis:</emphasis>
The genus
<emphasis id="B9275E0CFFCA323D1BBC23CEFE4A27A0" box="[405,487,997,1022]" italics="true" pageId="1" pageNumber="352">Exomis</emphasis>
<bibRefCitation id="EFC2FFEFFFCA323D1BC723CFFDD727A0" author="Weise, J." box="[494,634,996,1022]" pageId="1" pageNumber="352" pagination="560 - 653" refId="ref9504" refString="Weise, J. (1889) Insecta, a CL. G. N. Potanin in China et in Mongolia novissime lecta. IX. Chrysomelidae et Coccinellidae. Horae Societatis Entomologicae Rossicae, 23, 560 - 653." type="journal article" year="1889">Weise, 1889</bibRefCitation>
, can be recognized amongst other
<taxonomicName id="4C53F99DFFCA323D19D523CFFBFE27A0" box="[1020,1107,996,1022]" class="Insecta" family="Chrysomelidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="352" phylum="Arthropoda" rank="tribe" tribe="Clytrini">Clytrini</taxonomicName>
by the following characteristics: body cylindrical, elongate; antennae short, 1
<superScript id="7C262F56FFCA323D18ED242CFD62204B" attach="left" box="[708,719,1031,1045]" fontSize="6" pageId="1" pageNumber="352">st</superScript>
antennomere strongly swollen, 2
<superScript id="7C262F56FFCA323D1E61242CFBF5204B" attach="left" box="[1096,1112,1031,1045]" fontSize="6" pageId="1" pageNumber="352">nd</superScript>
and 3
<superScript id="7C262F56FFCA323D1EB6242CFB00204B" attach="left" box="[1183,1197,1031,1045]" fontSize="6" pageId="1" pageNumber="352">rd</superScript>
antennomeres small; Pronotum convex, as wide as or wider than elytra in males. Scutellum elevated toward posterior angle and above elytral plane. Epipleura pubescent. Legs robust, tarsi short, first tarsomere longer than second or third. Pygidium not exposed in dorsal view.
</paragraph>
<paragraph id="8BEC821EFFCA323D1AEE24B3FDBA2139" blockId="1.[151,1437,995,1383]" pageId="1" pageNumber="352">
The aedeagi in this genus are different from other genera: elongate, tegmen situated at apical part of the median lobe, basal half of median lobe behind tegmen feebly sclerotized or membranous. The characteristics of the aedeagi are not as reliable as in other genera of the tribe
<taxonomicName id="4C53F99DFFCA323D189024CBFCBD20A4" box="[697,784,1248,1274]" class="Insecta" family="Chrysomelidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="352" phylum="Arthropoda" rank="tribe" tribe="Clytrini">Clytrini</taxonomicName>
for species recognition. The female spermatheca and kotpresse sclerites are useful species diagnosing structures. The following determination key to species is a balanced one, and easy to use. For exact species recognition it is helpful to consider both genital characters of both sexes and external morphological structures.
</paragraph>
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<emphasis id="B9275E0CFFCA323D1ABE2585FCE12199" bold="true" box="[151,844,1453,1480]" pageId="1" pageNumber="352">
Key to Chinese species of the genus
<taxonomicName id="4C53F99DFFCA323D187F2586FCE12199" authority="Weise, 1889" authorityName="Weise" authorityYear="1889" box="[598,844,1453,1479]" class="Insecta" family="Chrysomelidae" genus="Exomis" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="352" phylum="Arthropoda" rank="genus">
<emphasis id="B9275E0CFFCA323D187F2586FD1C2199" bold="true" box="[598,689,1453,1479]" italics="true" pageId="1" pageNumber="352">Exomis</emphasis>
<bibRefCitation id="EFC2FFEFFFCA323D18902586FCE12199" author="Weise, J." box="[697,844,1453,1479]" pageId="1" pageNumber="352" pagination="560 - 653" refId="ref9504" refString="Weise, J. (1889) Insecta, a CL. G. N. Potanin in China et in Mongolia novissime lecta. IX. Chrysomelidae et Coccinellidae. Horae Societatis Entomologicae Rossicae, 23, 560 - 653." type="journal article" year="1889">Weise, 1889</bibRefCitation>
</taxonomicName>
</emphasis>
</paragraph>
<key id="FFC2D52FFFCA323D1ABE25D9FA312383" pageId="1" pageNumber="352">
<keyStep id="30A79ABBFFCA323D1ABE25D9FA31227A" pageId="1" pageNumber="352">
<paragraph id="8BEC821EFFCA323D1ABE25D9FA312259" blockId="1.[151,1437,1522,2013]" box="[151,1436,1522,1544]" pageId="1" pageNumber="352">
<keyLead id="30A2212BFFCA323D1ABE25D9FA312259" box="[151,1436,1522,1544]" pageId="1" pageNumber="352">1. Elytra pubescent...................................................................................... 2</keyLead>
</paragraph>
<paragraph id="8BEC821EFFCA323D1ABE263BFA31227A" blockId="1.[151,1437,1522,2013]" box="[151,1436,1551,1573]" pageId="1" pageNumber="352">
<keyLead id="30A2212BFFCA323D1ABE263BFA31227A" box="[151,1436,1551,1573]" pageId="1" pageNumber="352">- Elytra glabrous, at most lateral sides with a short row of setae.................................................. 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A79ABBFFCA323D1ABE2606FA312201" pageId="1" pageNumber="352">
<paragraph id="8BEC821EFFCA323D1ABE2606FA31221C" blockId="1.[151,1437,1522,2013]" box="[151,1436,1580,1603]" pageId="1" pageNumber="352">
<keyLead id="30A2212BFFCA323D1ABE2606FA31221C" box="[151,1436,1580,1603]" pageId="1" pageNumber="352">
2. Pronotum pubescent (male body length
<quantity id="4CAB2FFBFFCA323D18482607FD00221C" box="[609,685,1580,1602]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="1" pageNumber="352" unit="mm" value="5.1">5.1 mm</quantity>
; female unknown)..............................
<taxonomicName id="4C53F99DFFCA323D1EE52606FAE7221C" authority="Wang &amp; Zhou, 2020" authorityName="Wang &amp; Zhou" authorityYear="2020" box="[1228,1354,1581,1602]" class="Insecta" family="Chrysomelidae" genus="Exomis" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="352" phylum="Arthropoda" rank="species" species="pubescens" status="sp. nov.">
<emphasis id="B9275E0CFFCA323D1EE52606FAE7221C" box="[1228,1354,1581,1602]" italics="true" pageId="1" pageNumber="352">E. pubescens</emphasis>
</taxonomicName>
<emphasis id="B9275E0CFFCA323D1F792606FA31221C" bold="true" box="[1360,1436,1581,1602]" pageId="1" pageNumber="352">
<taxonomicNameLabel id="A214E377FFCA323D1F792606FA31221C" box="[1360,1436,1581,1602]" pageId="1" pageNumber="352" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BEC821EFFCA323D1ABE2661FA312201" blockId="1.[151,1437,1522,2013]" box="[151,1436,1610,1632]" pageId="1" pageNumber="352">
<keyLead id="30A2212BFFCA323D1ABE2661FA312201" box="[151,1436,1610,1632]" pageId="1" pageNumber="352">
- Pronotum glabrous (male body length
<quantity id="4CAB2FFBFFCA323D187D2661FD672201" box="[596,714,1610,1632]" metricMagnitude="-3" metricUnit="m" metricValue="6.85" metricValueMax="6.9" metricValueMin="6.8" pageId="1" pageNumber="352" unit="mm" value="6.85" valueMax="6.9" valueMin="6.8">6.8–6.9 mm</quantity>
; female unknown)...........................
<taxonomicName id="4C53F99DFFCA323D1EEF2660FAE42201" authority="Wang &amp; Zhou, 2020" authorityName="Wang &amp; Zhou" authorityYear="2020" box="[1222,1353,1610,1632]" class="Insecta" family="Chrysomelidae" genus="Exomis" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="352" phylum="Arthropoda" rank="species" species="pubipennis" status="sp. nov.">
<emphasis id="B9275E0CFFCA323D1EEF2660FAE42201" box="[1222,1353,1610,1632]" italics="true" pageId="1" pageNumber="352">E. pubipennis</emphasis>
</taxonomicName>
<emphasis id="B9275E0CFFCA323D1F792661FA312201" bold="true" box="[1360,1436,1610,1631]" pageId="1" pageNumber="352">
<taxonomicNameLabel id="A214E377FFCA323D1F792661FA312201" box="[1360,1436,1610,1631]" pageId="1" pageNumber="352" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A79ABBFFCA323D1ABE264CFA3122C4" pageId="1" pageNumber="352">
<paragraph id="8BEC821EFFCA323D1ABE264CFA312223" blockId="1.[151,1437,1522,2013]" box="[151,1436,1639,1661]" pageId="1" pageNumber="352">
<keyLead id="30A2212BFFCA323D1ABE264CFA312223" box="[151,1436,1639,1661]" pageId="1" pageNumber="352">
3. Elytra almost entirely metallic green or blue, with a fulvous apical spot (5.4–7.0 mm).................
<taxonomicName id="4C53F99DFFCA323D1EDB2643FAE42222" authority="Wang &amp; Zhou, 2020" authorityName="Wang &amp; Zhou" authorityYear="2020" box="[1266,1353,1639,1661]" class="Insecta" family="Chrysomelidae" genus="Exomis" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="352" phylum="Arthropoda" rank="species" species="viridis" status="sp. nov.">
<emphasis id="B9275E0CFFCA323D1EDB2643FAE42222" box="[1266,1353,1639,1661]" italics="true" pageId="1" pageNumber="352">E. viridis</emphasis>
</taxonomicName>
<emphasis id="B9275E0CFFCA323D1F792643FA312223" bold="true" box="[1360,1436,1640,1661]" pageId="1" pageNumber="352">
<taxonomicNameLabel id="A214E377FFCA323D1F792643FA312223" box="[1360,1436,1640,1661]" pageId="1" pageNumber="352" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BEC821EFFCA323D1ABE26AEFA3122C4" blockId="1.[151,1437,1522,2013]" box="[151,1436,1669,1691]" pageId="1" pageNumber="352">
<keyLead id="30A2212BFFCA323D1ABE26AEFA3122C4" box="[151,1436,1669,1691]" pageId="1" pageNumber="352">- Elytra without metallic tinge............................................................................ 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A79ABBFFCA323D1ABE2689FA31228B" pageId="1" pageNumber="352">
<paragraph id="8BEC821EFFCA323D1ABE2689FA3122E6" blockId="1.[151,1437,1522,2013]" box="[151,1436,1698,1720]" pageId="1" pageNumber="352">
<keyLead id="30A2212BFFCA323D1ABE2689FA3122E6" box="[151,1436,1698,1720]" pageId="1" pageNumber="352">4. Elytra completely fulvous, without dark markings, at most an indistinct spot at humerus............................. 5</keyLead>
</paragraph>
<paragraph id="8BEC821EFFCA323D1ABE26EBFA31228B" blockId="1.[151,1437,1522,2013]" box="[151,1436,1727,1749]" pageId="1" pageNumber="352">
<keyLead id="30A2212BFFCA323D1ABE26EBFA31228B" box="[151,1436,1727,1749]" pageId="1" pageNumber="352">- Elytra black with fulvous markings, or fulvous with black markings............................................. 7</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A79ABBFFCA323D1ABE26F7FA312351" pageId="1" pageNumber="352">
<paragraph id="8BEC821EFFCA323D1ABE26F7FA3122AD" blockId="1.[151,1437,1522,2013]" box="[151,1436,1756,1779]" pageId="1" pageNumber="352">
<keyLead id="30A2212BFFCA323D1ABE26F7FA3122AD" box="[151,1436,1756,1779]" pageId="1" pageNumber="352">5. Abdomen and thorax entirely dark........................................................................ 6</keyLead>
</paragraph>
<paragraph id="8BEC821EFFCA323D1ABE26D1FA312351" blockId="1.[151,1437,1522,2013]" box="[151,1436,1786,1808]" pageId="1" pageNumber="352">
<keyLead id="30A2212BFFCA323D1ABE26D1FA312351" box="[151,1436,1786,1808]" pageId="1" pageNumber="352">
- Venter with episternum and epimeron fulvous, body length
<quantity id="4CAB2FFBFFCA323D193426D1FC3D2351" box="[797,912,1786,1808]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.3" metricValueMin="3.7" pageId="1" pageNumber="352" unit="mm" value="4.5" valueMax="5.3" valueMin="3.7">3.7–5.3 mm</quantity>
.........................
<taxonomicName id="4C53F99DFFCA323D1EE326D0FAE42351" authority="Wang &amp; Zhou, 2020" authorityName="Wang &amp; Zhou" authorityYear="2020" box="[1226,1353,1786,1808]" class="Insecta" family="Chrysomelidae" genus="Exomis" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="352" phylum="Arthropoda" rank="species" species="deqinensis" status="sp. nov.">
<emphasis id="B9275E0CFFCA323D1EE326D0FAE42351" box="[1226,1353,1786,1808]" italics="true" pageId="1" pageNumber="352">E. deqinensis</emphasis>
</taxonomicName>
<emphasis id="B9275E0CFFCA323D1F7926D1FA312351" bold="true" box="[1360,1436,1786,1807]" pageId="1" pageNumber="352">
<taxonomicNameLabel id="A214E377FFCA323D1F7926D1FA312351" box="[1360,1436,1786,1807]" pageId="1" pageNumber="352" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A79ABBFFCA323D1ABE273CFA312314" pageId="1" pageNumber="352">
<paragraph id="8BEC821EFFCA323D1ABE273CFA302373" blockId="1.[151,1437,1522,2013]" box="[151,1437,1815,1837]" pageId="1" pageNumber="352">
<keyLead id="30A2212BFFCA323D1ABE273CFA302373" box="[151,1437,1815,1837]" pageId="1" pageNumber="352">
6. Legs entirely black........................................................................
<taxonomicName id="4C53F99DFFCA323D1ED42733FA302373" authority="Pic" authorityName="Pic" box="[1277,1437,1815,1837]" class="Insecta" family="Chrysomelidae" genus="Exomis" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="352" phylum="Arthropoda" rank="species" species="degenevei">
<emphasis id="B9275E0CFFCA323D1ED42733FAD52372" box="[1277,1400,1815,1837]" italics="true" pageId="1" pageNumber="352">E. degenevei</emphasis>
Pic
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BEC821EFFCA323D1ABE271EFA312314" blockId="1.[151,1437,1522,2013]" box="[151,1436,1844,1867]" pageId="1" pageNumber="352">
<keyLead id="30A2212BFFCA323D1ABE271EFA312314" box="[151,1436,1844,1867]" pageId="1" pageNumber="352">
- Femora except for black apex orange, body length
<quantity id="4CAB2FFBFFCA323D18E3271FFC902314" box="[714,829,1844,1866]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="5.4" metricValueMin="4.6" pageId="1" pageNumber="352" unit="mm" value="5.0" valueMax="5.4" valueMin="4.6">4.6–5.4 mm</quantity>
..........
<taxonomicName id="4C53F99DFFCA323D1990271EFA992315" authority="(Lopatin &amp; Konstantinov)" authorityName="Wang &amp; Zhou" authorityYear="2020" baseAuthorityName="Lopatin &amp; Konstantinov" box="[953,1332,1845,1867]" class="Insecta" family="Chrysomelidae" genus="Smaragdina" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="352" phylum="Arthropoda" rank="species" species="oblongum" status="comb. nov.">
<emphasis id="B9275E0CFFCA323D1990271EFB992314" box="[953,1076,1845,1866]" italics="true" pageId="1" pageNumber="352">E. oblongum</emphasis>
(Lopatin &amp; Konstantinov)
</taxonomicName>
<taxonomicNameLabel id="A214E377FFCA323D1F13271EFA312314" box="[1338,1436,1845,1866]" pageId="1" pageNumber="352" rank="species">
<emphasis id="B9275E0CFFCA323D1F13271EFA3B2314" bold="true" box="[1338,1430,1845,1866]" pageId="1" pageNumber="352">com. nov</emphasis>
.
</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A79ABBFFCA323D1ABE2779FA3123FC" pageId="1" pageNumber="352">
<paragraph id="8BEC821EFFCA323D1ABE2779FA3123DB" blockId="1.[151,1437,1522,2013]" pageId="1" pageNumber="352">
<keyLead id="30A2212BFFCA323D1ABE2779FA3123DB" pageId="1" pageNumber="352">
7. Elytra black, each elytron with a round fulvous spot at apex, pronotum fulvous with a central black spot extending both to anterior and posterior margins, body length
<quantity id="4CAB2FFBFFCA323D18AE2744FD5123DB" box="[647,764,1903,1925]" metricMagnitude="-3" metricUnit="m" metricValue="5.15" metricValueMax="5.7" metricValueMin="4.6" pageId="1" pageNumber="352" unit="mm" value="5.15" valueMax="5.7" valueMin="4.6">4.6–5.7 mm</quantity>
).......................................
<taxonomicName id="4C53F99DFFCA323D1EC5275BFAE723DA" authority="Wang &amp; Zhou, 2020" authorityName="Wang &amp; Zhou" authorityYear="2020" box="[1260,1354,1903,1925]" class="Insecta" family="Chrysomelidae" genus="Exomis" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="352" phylum="Arthropoda" rank="species" species="huangi" status="sp. nov.">
<emphasis id="B9275E0CFFCA323D1EC5275BFAE723DA" box="[1260,1354,1903,1925]" italics="true" pageId="1" pageNumber="352">E. huangi</emphasis>
</taxonomicName>
<emphasis id="B9275E0CFFCA323D1F79275BFA3123DB" bold="true" box="[1360,1436,1904,1925]" pageId="1" pageNumber="352">
<taxonomicNameLabel id="A214E377FFCA323D1F79275BFA3123DB" box="[1360,1436,1904,1925]" pageId="1" pageNumber="352" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BEC821EFFCA323D1ABE27A6FA3123FC" blockId="1.[151,1437,1522,2013]" box="[151,1436,1933,1955]" pageId="1" pageNumber="352">
<keyLead id="30A2212BFFCA323D1ABE27A6FA3123FC" box="[151,1436,1933,1955]" pageId="1" pageNumber="352">- Elytra fulvous with black humeral spot or several black markings............................................... 8</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A79ABBFFCA323D1ABE2781FA312383" pageId="1" pageNumber="352">
<paragraph id="8BEC821EFFCA323D1ABE2781FA31239E" blockId="1.[151,1437,1522,2013]" box="[151,1436,1962,1984]" pageId="1" pageNumber="352">
<keyLead id="30A2212BFFCA323D1ABE2781FA31239E" box="[151,1436,1962,1984]" pageId="1" pageNumber="352">
8 Elytra fulvous with several black markings (
<quantity id="4CAB2FFBFFCA323D18A22781FCAC23E1" box="[651,769,1962,1984]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="6.7" metricValueMin="5.3" pageId="1" pageNumber="352" unit="mm" value="6.0" valueMax="6.7" valueMin="5.3">5.3–6.7 mm</quantity>
)...................................
<taxonomicName id="4C53F99DFFCA323D1E9D2780FA31239E" authority="Weise" authorityName="Weise, H" authorityYear="1889" box="[1204,1436,1962,1984]" class="Insecta" family="Chrysomelidae" genus="Exomis" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="352" phylum="Arthropoda" rank="species" species="peplopteroides">
<emphasis id="B9275E0CFFCA323D1E9D2780FAF123E1" box="[1204,1372,1962,1984]" italics="true" pageId="1" pageNumber="352">E. peplopteroides</emphasis>
Weise
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BEC821EFFCA323D1ABE27E3FA312383" blockId="1.[151,1437,1522,2013]" box="[151,1436,1991,2013]" pageId="1" pageNumber="352">
<keyLead id="30A2212BFFCA323D1ABE27E3FA312383" box="[151,1436,1991,2013]" pageId="1" pageNumber="352">
- Each elytron with a black humeral spot, legs mostly fulvous (4.6–6.0 mm)............................
<taxonomicName id="4C53F99DFFCA323D1ED327E3FAE42383" authority="Wang &amp; Zhou, 2020" authorityName="Wang &amp; Zhou" authorityYear="2020" box="[1274,1353,1992,2013]" class="Insecta" family="Chrysomelidae" genus="Exomis" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="352" phylum="Arthropoda" rank="species" species="tanae" status="sp. nov.">
<emphasis id="B9275E0CFFCA323D1ED327E3FAE42383" box="[1274,1353,1992,2013]" italics="true" pageId="1" pageNumber="352">E. tanae</emphasis>
</taxonomicName>
<emphasis id="B9275E0CFFCA323D1F7927E3FA312383" bold="true" box="[1360,1436,1992,2013]" pageId="1" pageNumber="352">
<taxonomicNameLabel id="A214E377FFCA323D1F7927E3FA312383" box="[1360,1436,1992,2013]" pageId="1" pageNumber="352" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>
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