treatments-xml/data/03/F2/DE/03F2DE0CA93CC025FCDE3378FED3E716.xml
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<document ID-DOI="http://doi.org/10.5281/zenodo.5469580" ID-ISSN="1809-127X" ID-Zenodo-Dep="5469580" approvalRequired="71" approvalRequired_for_taxonomicNames="9" approvalRequired_for_textStreams="60" approvalRequired_for_treatments="2" checkinTime="1630327227302" checkinUser="marcus" docAuthor="Cárdenas-Bautista, Johann Stephens, Parada-Alfonso, Jenny Andrea &amp; CarvajalCogollo, Juan E." docDate="2020" docId="03F2DE0CA93CC025FCDE3378FED3E716" docLanguage="en" docName="CheckList.2020.16.4.821-839.pdf" docOrigin="Check List 16 (4)" docSource="http://dx.doi.org/10.15560/16.4.821" docStyle="DocumentStyle:BB30DA0EB5D11E1D6FE264CD32192DE2.1:CheckList.2017-.journal_article.annotatedlist" docStyleId="BB30DA0EB5D11E1D6FE264CD32192DE2" docStyleName="CheckList.2017-.journal_article.annotatedlist" docStyleVersion="1" docTitle="Dichotomius mamillatus" docType="treatment" docVersion="2" lastPageId="10" lastPageNumber="830" masterDocId="FFCBA674A935C02FFFF33644FFE2E676" masterDocTitle="Dung beetles (Scarabaeidae, Scarabaeinae) of the Foothills-Andean Forest strip of Villavicencio, Colombia" masterLastPageNumber="839" masterPageNumber="821" pageId="9" pageNumber="830" updateTime="1630932967158" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Dung beetles (Scarabaeidae, Scarabaeinae) of the Foothills-Andean Forest strip of Villavicencio, Colombia</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Cárdenas-Bautista, Johann Stephens</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Parada-Alfonso, Jenny Andrea</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>CarvajalCogollo, Juan E.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Check List</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="pubDate">
<mods:number>2020-07-03</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>16</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>4</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>821</mods:start>
<mods:end>839</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.15560/16.4.821</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.15560/16.4.821</mods:identifier>
<mods:identifier type="ISSN">1809-127X</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.5469580" ID-Zenodo-Dep="5469580" LSID="urn:lsid:plazi:treatment:03F2DE0CA93CC025FCDE3378FED3E716" httpUri="http://treatment.plazi.org/id/03F2DE0CA93CC025FCDE3378FED3E716" lastPageId="10" lastPageNumber="830" pageId="9" pageNumber="830">
<subSubSection box="[813,1262,1340,1365]" pageId="9" pageNumber="830" type="nomenclature">
<paragraph blockId="9.[813,1262,1340,1365]" box="[813,1262,1340,1365]" pageId="9" pageNumber="830">
<heading box="[813,1262,1340,1365]" fontSize="10" level="2" pageId="9" pageNumber="830" reason="2">
<taxonomicName authority="(Felsche, 1910)" baseAuthorityName="Felsche" baseAuthorityYear="1910" box="[813,1262,1340,1365]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="830" phylum="Arthropoda" rank="species" species="mamillatus">
<emphasis bold="true" box="[813,1262,1340,1365]" inLineHeading="true" pageId="9" pageNumber="830" reason="1">
<emphasis bold="true" box="[813,1088,1340,1364]" italics="true" pageId="9" pageNumber="830">Dichotomius mamillatus</emphasis>
(Felsche, 1910)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="830" type="materials_examined">
<paragraph blockId="9.[813,1420,1382,1707]" pageId="9" pageNumber="830">
<emphasis bold="true" box="[813,1057,1382,1406]" pageId="9" pageNumber="830">Materials examined.</emphasis>
<materialsCitation collectingDate="2018-10-18" collectorName="Vereda El Carmen &amp; Cardenas Johann &amp; Parada Jenny &amp; Crpat" country="Colombia" county="Villavicencio" elevation="730" latitude="4.141161" location="Farm" longLatPrecision="1" longitude="-73.66719" municipality="Vereda El Carmen" pageId="9" pageNumber="830" specimenCount="1" specimenCount-female="1" stateProvince="Meta">
<collectingCountry box="[1072,1215,1382,1406]" name="Colombia" pageId="9" pageNumber="830">COLOMBIA</collectingCountry>
<specimenCount box="[1254,1284,1382,1406]" pageId="9" pageNumber="830" type="female">1♀</specimenCount>
,
<quantity box="[1306,1419,1382,1406]" metricMagnitude="-2" metricUnit="m" metricValue="2.3739999999999997" pageId="9" pageNumber="830" unit="mm" value="23.74">23.74 mm</quantity>
(mean);
<collectingRegion box="[907,965,1416,1439]" country="Colombia" name="Meta" pageId="9" pageNumber="830">Meta</collectingRegion>
,
<collectingCounty box="[982,1129,1416,1440]" pageId="9" pageNumber="830">Villavicencio</collectingCounty>
,
<collectorName box="[1146,1362,1416,1440]" pageId="9" pageNumber="830">
<collectingMunicipality box="[1146,1362,1416,1440]" pageId="9" pageNumber="830">Vereda El Carmen</collectingMunicipality>
</collectorName>
, “La Loma”
<location LSID="urn:lsid:plazi:treatment:03F2DE0CA93CC025FCDE3378FED3E716:8E8439C1A93CC026FC7F33EEFC2AE3B7" box="[908,968,1450,1473]" country="Colombia" county="Villavicencio" latitude="4.141161" longLatPrecision="1" longitude="-73.66719" municipality="Vereda El Carmen" name="Farm" pageId="9" pageNumber="830" stateProvince="Meta">Farm</location>
;
<geoCoordinate box="[993,1153,1448,1473]" degrees="04" direction="north" minutes="08" orientation="latitude" pageId="9" pageNumber="830" precision="1" seconds="28.18" value="4.141161">04°0828.18″N</geoCoordinate>
,
<geoCoordinate box="[1178,1355,1448,1473]" degrees="073" direction="west" minutes="40" orientation="longitude" pageId="9" pageNumber="830" precision="1" seconds="01.90" value="-73.66719">073°4001.90″W</geoCoordinate>
;
<elevation metricMagnitude="2" metricUnit="m" metricValue="7.3" pageId="9" pageNumber="830" unit="m" value="730.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="7.3" pageId="9" pageNumber="830" unit="m" value="730.0">730 m</quantity>
a.s.l.
</elevation>
;
<date box="[918,1069,1483,1507]" pageId="9" pageNumber="830" value="2018-10-18">
<collectingDate box="[918,1069,1483,1507]" pageId="9" pageNumber="830" value="2018-10-18">18 Oct. 2018</collectingDate>
</date>
;
<collectorName box="[1087,1279,1483,1507]" pageId="9" pageNumber="830">Cárdenas Johann</collectorName>
and
<collectorName pageId="9" pageNumber="830">Parada Jenny</collectorName>
leg.; wooded pasture;
<collectorName box="[1141,1206,1516,1540]" pageId="9" pageNumber="830">Crpat</collectorName>
3; UPTC-In-00054
</materialsCitation>
. •
<materialsCitation collectingDate="2018-10-17" collectorName="Vereda El Carmen &amp; Cardenas Johann &amp; Parada Jenny &amp; Crbrt" country="Colombia" county="Villavicencio" elevation="774" latitude="4.139403" location="Cano Blanco - Cano Buque" longLatPrecision="1" longitude="-73.67005" municipality="Vereda El Carmen" pageId="9" pageNumber="830" specimenCount="1" specimenCount-male="1" stateProvince="Meta">
<specimenCount box="[831,862,1550,1574]" pageId="9" pageNumber="830" type="male">1♂</specimenCount>
,
<quantity box="[876,982,1550,1574]" metricMagnitude="-2" metricUnit="m" metricValue="2.3739999999999997" pageId="9" pageNumber="830" unit="mm" value="23.74">23.74 mm</quantity>
(mean);
<collectingRegion box="[1081,1139,1550,1573]" country="Colombia" name="Meta" pageId="9" pageNumber="830">Meta</collectingRegion>
,
<collectingCounty box="[1152,1299,1549,1573]" pageId="9" pageNumber="830">Villavicencio</collectingCounty>
,
<collectorName pageId="9" pageNumber="830">
<collectingMunicipality pageId="9" pageNumber="830">Vereda El Carmen</collectingMunicipality>
</collectorName>
,
<location LSID="urn:lsid:plazi:treatment:03F2DE0CA93CC025FCDE3378FED3E716:8E8439C1A93CC026FC68306BFB31E030" box="[923,1235,1583,1607]" country="Colombia" county="Villavicencio" latitude="4.139403" longLatPrecision="1" longitude="-73.67005" municipality="Vereda El Carmen" name="Cano Blanco - Cano Buque" pageId="9" pageNumber="830" stateProvince="Meta">Caño BlancoCaño Buque</location>
;
<geoCoordinate box="[1256,1416,1581,1606]" degrees="04" direction="north" minutes="08" orientation="latitude" pageId="9" pageNumber="830" precision="1" seconds="21.85" value="4.139403">04°0821.85″N</geoCoordinate>
,
<geoCoordinate box="[813,989,1615,1640]" degrees="073" direction="west" minutes="40" orientation="longitude" pageId="9" pageNumber="830" precision="1" seconds="12.19" value="-73.67005">073°4012.19″W</geoCoordinate>
;
<elevation box="[1004,1137,1616,1640]" metricMagnitude="2" metricUnit="m" metricValue="7.74" pageId="9" pageNumber="830" unit="m" value="774.0">
<quantity box="[1004,1075,1616,1640]" metricMagnitude="2" metricUnit="m" metricValue="7.74" pageId="9" pageNumber="830" unit="m" value="774.0">774 m</quantity>
a.s.l.
</elevation>
;
<date box="[1155,1303,1616,1640]" pageId="9" pageNumber="830" value="2018-10-17">
<collectingDate box="[1155,1303,1616,1640]" pageId="9" pageNumber="830" value="2018-10-17">17 Oct. 2018</collectingDate>
</date>
;
<collectorName pageId="9" pageNumber="830">Cárdenas Johann</collectorName>
and
<collectorName box="[950,1098,1650,1674]" pageId="9" pageNumber="830">Parada Jenny</collectorName>
leg.; riparian forest;
<collectorName box="[1342,1402,1650,1674]" pageId="9" pageNumber="830">Crbrt</collectorName>
4; UPTC-In-00055
</materialsCitation>
.
</paragraph>
<paragraph blockId="9.[813,1420,1724,1983]" pageId="9" pageNumber="830">
<emphasis bold="true" box="[813,974,1724,1748]" pageId="9" pageNumber="830">Identification.</emphasis>
This species can be distinguished from other
<taxonomicName box="[878,1103,1758,1782]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="830" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis box="[878,1016,1758,1782]" italics="true" pageId="9" pageNumber="830">Dichotomius</emphasis>
species
</taxonomicName>
by the head with transverse roughness in the clypeus, gena and frons, the presence of a short, thick and blunt horn at the height of the gena with two small horns adjacent to the base of the central horn; pronotum with two small protuberances in the anterior medial region (
<bibRefCitation author="Sarmiento-Garces R &amp; Amat-Garcia G" pageId="9" pageNumber="830" refId="ref18606" refString="Sarmiento-Garces R, Amat-Garcia G (2014) Escarabajos del genero Dichotomius Hope, 1838 (Scarabaeidae, Scarabaeinae) en Colombia. Universidad Nacional de Colombia, Bogota, 134 pp." type="book" year="2014">Sarmiento-Gárces and Amat-García 2014</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection lastPageId="10" lastPageNumber="831" pageId="9" pageNumber="830" type="distribution">
<paragraph blockId="9.[813,1419,2000,2091]" lastBlockId="10.[167,776,160,352]" lastPageId="10" lastPageNumber="831" pageId="9" pageNumber="830">
<emphasis bold="true" box="[813,961,2000,2024]" pageId="9" pageNumber="830">Distribution.</emphasis>
The species is distributed in the Colombian and Ecuadorian Amazon, where it is a very wellrepresented species, extending to the foothills up to
<quantity box="[168,248,160,184]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="10" pageNumber="831" unit="m" value="1300.0">1300 m</quantity>
a.s.l. (
<bibRefCitation author="Sarmiento-Garces R &amp; Amat-Garcia G" box="[317,771,160,184]" pageId="10" pageNumber="831" pagination="285 - 296" refId="ref18562" refString="Sarmiento-Garces R, Amat-Garcia G (2009) Escarabajos del genero Dichotomius Hope 1838 (Scarabaeidae: Scarabaeinae) en la Amazonia Colombiana. Revista de la Academia Colombiana de Ciencias Exactas Fisicas y Naturales 33 (127): 285 - 296." type="journal article" year="2009">Sarmiento-Gárces and Amat-García 2009</bibRefCitation>
;
<bibRefCitation author="Sarmiento-Garces R &amp; Amat-Garcia G" box="[167,624,194,218]" pageId="10" pageNumber="831" refId="ref18606" refString="Sarmiento-Garces R, Amat-Garcia G (2014) Escarabajos del genero Dichotomius Hope, 1838 (Scarabaeidae, Scarabaeinae) en Colombia. Universidad Nacional de Colombia, Bogota, 134 pp." type="book" year="2014">Sarmiento-Gárces and Amat-García 2014</bibRefCitation>
). In the study area, it was observed between
<quantity box="[499,620,227,251]" metricMagnitude="2" metricUnit="m" metricValue="8.0" metricValueMax="8.5" metricValueMin="7.5" pageId="10" pageNumber="831" unit="m" value="800.0" valueMax="850.0" valueMin="750.0">750850 m</quantity>
a.s.l. and seen in the habitats of riparian forest, high secondary forest, and wooded grassland, although it was rare species (59 individuals).
</paragraph>
</subSubSection>
</treatment>
</document>