treatments-xml/data/03/F2/D8/03F2D824FFC890768D2CF8CAFB4903A3.xml

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<mods:title id="7A6A7F7E54A8C37297E9D5F0D1159697">Redescripción de Cochranella megista (Rivero, 1985) y ampliación de la distribución de nueve ranas de cristal (Anura: Centrolenidae) en Colombia</mods:title>
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<paragraph id="8BE46932FFC8907D8D2CF8CAFB790200" blockId="3.[883,1252,1829,1885]" box="[883,1252,1829,1853]" pageId="3" pageNumber="92">
<heading id="D0ACDE5EFFC8907D8D2CF8CAFB790200" bold="true" box="[883,1252,1829,1853]" centered="true" fontSize="9" level="2" pageId="3" pageNumber="92" reason="2">
<taxonomicName id="4C5B12B1FFC8907D8D2CF8CAFB790200" ID-CoL="5ZHFZ" authority="(Rivero, 1985)" authorityName="Y AMPLIA C I ON DE LA DIST R IBU C I ON DE NUE V E R ANAS DE C R ISTAL" baseAuthorityName="RI V E R O" baseAuthorityYear="1985" box="[883,1252,1829,1853]" class="Amphibia" family="Centrolenidae" genus="Cochranella" kingdom="Animalia" order="Anura" pageId="3" pageNumber="92" phylum="Chordata" rank="species" species="megista">
<emphasis id="B92FB520FFC8907D8D2CF8CAFB790200" bold="true" box="[883,1252,1829,1853]" pageId="3" pageNumber="92">
<emphasis id="B92FB520FFC8907D8D2CF8CAFBDE0200" bold="true" box="[883,1091,1829,1853]" italics="true" pageId="3" pageNumber="92">Cochranella megista</emphasis>
(
<bibRefCitation id="EFCA14C3FFC8907D8A0FF8CAFB410200" author="RIVERO, J. A." box="[1104,1244,1829,1853]" pageId="3" pageNumber="92" pagination="334 - 373" refId="ref7316" refString="RIVERO, J. A. 1985. Nuevos Centrolenidos de Colombia y Venezuela. Brenesia, 23: 334 - 373." type="journal article" year="1985">Rivero, 1985</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<paragraph id="8BE46932FFC8907D8DB8F8AAFBED0260" blockId="3.[883,1252,1829,1885]" box="[999,1136,1861,1885]" pageId="3" pageNumber="92">
<emphasis id="B92FB520FFC8907D8DB8F8AAFBED0260" bold="true" box="[999,1136,1861,1885]" pageId="3" pageNumber="92">
<figureCitation id="136075B7FFC8907D8DB8F8AAFBA20260" box="[999,1087,1861,1885]" captionStart="FIGURA 3" captionStartId="3.[208,287,1274,1294]" captionTargetBox="[208,1342,311,1237]" captionTargetId="figure-270@3.[208,1342,311,1237]" captionTargetPageId="3" captionText="FIGURA 3: Fotografía e ilustración dorsal comparativas de (A): Centrolene grandisonae, hembra adulta, MHNCSJ “294” – falso holotipo; (B): Cochranella megista, hembra adulta, MHNCSJ 294 (Holotipo perdido, ilustración tomada de Rivero, 1985); (C): Cochranella megista, macho adulto (Neotipo) MHNCS 291 (Actualmente MHUA 5851); (D): Cochranella megista, macho adulto, ICN 27768." figureDoi="http://doi.org/10.5281/zenodo.13314522" httpUri="https://zenodo.org/record/13314522/files/figure.png" pageId="3" pageNumber="92">Figs. 3C</figureCitation>
, 
<figureCitation id="136075B7FFC8907D8A13F8AAFBED0260" box="[1100,1136,1861,1885]" captionStart-0="FIGURA 4" captionStart-1="FIGURA 5" captionStartId-0="4.[245,324,953,973]" captionStartId-1="4.[245,324,1942,1962]" captionTargetBox-0="[245,1379,311,915]" captionTargetBox-1="[245,793,1187,1905]" captionTargetId-0="figure-185@4.[245,1379,311,915]" captionTargetId-1="figure-161@4.[245,793,1187,1905]" captionTargetPageId-0="4" captionTargetPageId-1="4" captionText-0="FIGURA 4: Fotografía de Cochranella megista, macho adulto (Neotipo), MHNCSJ 291(Actualmente MHUA 5851). A, B = Aspectos dorsal y lateral de la cabeza." captionText-1="FIGURA 5: Fotografía de Cochranella megista, macho adulto (Neotipo), MHNCSJ 291 (Actualmente MHUA 5851). Aspecto ventral de A = Mano; B = Pie." figureDoi-0="http://doi.org/10.5281/zenodo.13314524" figureDoi-1="http://doi.org/10.5281/zenodo.13314526" httpUri-0="https://zenodo.org/record/13314524/files/figure.png" httpUri-1="https://zenodo.org/record/13314526/files/figure.png" pageId="3" pageNumber="92">4-5</figureCitation>
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<subSubSection id="C3413AB9FFC8907A8D46F86AFAA60163" lastPageId="4" lastPageNumber="93" pageId="3" pageNumber="92" type="reference_group">
<paragraph id="8BE46932FFC8907A8D46F86AFAA60163" blockId="3.[793,1342,1925,2014]" lastBlockId="4.[831,1380,1061,1342]" lastPageId="4" lastPageNumber="93" pageId="3" pageNumber="92">
<taxonomicName id="4C5B12B1FFC8907D8D46F86AFB1C02A3" ID-CoL="5XHT7" authority="Rivero, 1985" authorityName="Rivero" authorityYear="1985" box="[793,1153,1925,1950]" class="Amphibia" family="Centrolenidae" genus="Centrolenella" kingdom="Animalia" order="Anura" pageId="3" pageNumber="92" phylum="Chordata" rank="species" species="megistra">
<emphasis id="B92FB520FFC8907D8D46F86AFC6E02A0" box="[793,1011,1925,1949]" italics="true" pageId="3" pageNumber="92">Centrolenella megistra</emphasis>
<bibRefCitation id="EFCA14C3FFC8907D8DA6F86AFB1C02A3" author="RIVERO, J. A." box="[1017,1153,1925,1950]" pageId="3" pageNumber="92" pagination="334 - 373" refId="ref7316" refString="RIVERO, J. A. 1985. Nuevos Centrolenidos de Colombia y Venezuela. Brenesia, 23: 334 - 373." type="journal article" year="1985">Rivero, 1985</bibRefCitation>
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, Brenesia, 23:353. 
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, MHNCSJ 0294, hembra adulta, 40.00 mm LRC, colectado en “Urrao, Parque Las Orquídeas (Polo), 
<quantity id="4CA3C4D7FFCF907A8C6EFBCAFD5F0100" box="[561,706,1061,1086]" metricMagnitude="3" metricUnit="m" metricValue="1.75" metricValueMax="1.8" metricValueMin="1.7" pageId="4" pageNumber="93" unit="m" value="1750.0" valueMax="1800.0" valueMin="1700.0">1700-1800 m</quantity>
, 
<collectingRegion id="499FA7D0FFCF907A8C89FBCAFEC00163" country="Colombia" name="Antioquia" pageId="4" pageNumber="93">Antioquia</collectingRegion>
, 
<collectingCountry id="F34C29A2FFCF907A8F2FFBAAFE4A0163" box="[368,471,1093,1118]" name="Colombia" pageId="4" pageNumber="93">Colombia</collectingCountry>
” = Parque Nacional Natural “Las Orquídeas” (Sitio El Polo), Municipio de Urrao, 
<collectingRegion id="499FA7D0FFCF907A8D9DFBC9FB770103" box="[962,1258,1061,1086]" country="Colombia" name="Antioquia" pageId="4" pageNumber="93">Departamento de Antioquia</collectingRegion>
, 
<collectingCountry id="F34C29A2FFCF907A8AA9FBCAFAFD0103" box="[1270,1376,1061,1086]" name="Colombia" pageId="4" pageNumber="93">Colombia</collectingCountry>
, por Marco A. Serna el 20 de mayo de 1980.
</paragraph>
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<emphasis id="B92FB520FFCF907A8EAAFC56FEC006F0" bold="true" box="[245,349,953,973]" pageId="4" pageNumber="93">FIGURA 4:</emphasis>
Fotografía de 
<emphasis id="B92FB520FFCF907A8F83FC56FD1F06F0" box="[476,642,953,973]" italics="true" pageId="4" pageNumber="93">
<taxonomicName id="4C5B12B1FFCF907A8F83FC56FDE206F0" authorityName="Y AMPLIA C I ON DE LA DIST R IBU C I ON DE NUE V E R ANAS DE C R ISTAL" baseAuthorityName="RI V E R O" baseAuthorityYear="1985" box="[476,639,953,973]" class="Amphibia" family="Centrolenidae" genus="Cochranella" kingdom="Animalia" order="Anura" pageId="4" pageNumber="93" phylum="Chordata" rank="species" species="megista">Cochranella megista</taxonomicName>
,
</emphasis>
macho adulto (Neotipo), MHNCSJ 291(Actualmente MHUA 5851). A, B = Aspectos dorsal y lateral de la cabeza.
</paragraph>
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<subSubSection id="C3413AB9FFCF907A8D60FB8AFC4D02E3" pageId="4" pageNumber="93" type="description">
<paragraph id="8BE46932FFCF907A8D60FB8AFB8501A3" blockId="4.[831,1380,1061,1342]" pageId="4" pageNumber="93">
<taxonomicName id="4C5B12B1FFCF907A8D60FB8AFAC20143" authority="Ruiz-Carranza &amp; Lynch, 1991 a" authorityName="Ruiz-Carranza &amp; Lynch" authorityYear="1991" box="[831,1375,1125,1150]" class="Amphibia" family="Centrolenidae" genus="Cochranella" kingdom="Animalia" order="Anura" pageId="4" pageNumber="93" phylum="Chordata" rank="species" species="megistra">
<emphasis id="B92FB520FFCF907A8D60FB8AFB900140" box="[831,1037,1125,1149]" italics="true" pageId="4" pageNumber="93">Cochranella megistra</emphasis>
Ruiz-Carranza &amp; Lynch, 1991a
</taxonomicName>
, 
<taxonomicName id="4C5B12B1FFCF907A8D27FB6AFC5201A3" box="[888,975,1157,1182]" class="Magnoliopsida" family="Lacistemataceae" genus="Lozania" kingdom="Plantae" order="Malpighiales" pageId="4" pageNumber="101" phylum="Tracheophyta" rank="genus">Lozania</taxonomicName>
., 57:22
</paragraph>
<paragraph id="8BE46932FFCF907A8D60FB4AFBCF01C3" blockId="4.[831,1380,1061,1342]" pageId="4" pageNumber="93">
<taxonomicName id="4C5B12B1FFCF907A8D60FB4AFC6001E3" authority="Ruiz-Carranza, Ardila-Robayo &amp; Lynch, 1996" authorityName="Ruiz-Carranza, Ardila-Robayo &amp; Lynch" authorityYear="1996" class="Amphibia" family="Centrolenidae" genus="Cochranella" kingdom="Animalia" order="Anura" pageId="4" pageNumber="93" phylum="Chordata" rank="species" species="megistra">
<emphasis id="B92FB520FFCF907A8D60FB4AFB960180" box="[831,1035,1189,1213]" italics="true" pageId="4" pageNumber="93">Cochranella megistra</emphasis>
Ruiz-Carranza, Ardila-Robayo &amp; Lynch, 1996
</taxonomicName>
, Rev. Acad. Coloma. Cienc. Exact. Fis. Nat., 20(77):373
</paragraph>
<paragraph id="8BE46932FFCF907A8D60FAEAFBB10003" blockId="4.[831,1380,1061,1342]" pageId="4" pageNumber="93">
<taxonomicName id="4C5B12B1FFCF907A8D60FAEAFB6E0023" authority="Acosta-Galvis, 2000" authorityName="Acosta-Galvis" authorityYear="2000" box="[831,1267,1285,1310]" class="Amphibia" family="Centrolenidae" genus="Cochranella" kingdom="Animalia" order="Anura" pageId="4" pageNumber="93" phylum="Chordata" rank="species" species="megistra">
<emphasis id="B92FB520FFCF907A8D60FAEAFB8C0020" box="[831,1041,1285,1309]" italics="true" pageId="4" pageNumber="93">Cochranella megistra</emphasis>
Acosta-Galvis, 2000
</taxonomicName>
. Biota colombiana., 1(3):8
</paragraph>
<paragraph id="8BE46932FFCF907A8D60FA6AFB9700A0" blockId="4.[831,1034,1413,1437]" box="[831,1034,1413,1437]" pageId="4" pageNumber="93">
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<typeStatus id="54E0D790FFCF907A8D60FA6AFC3D00A0" box="[831,928,1413,1437]" pageId="4" pageNumber="93" type="holotype">Holotipo</typeStatus>
(Perdido)
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE46932FFCF907A8D60FA2AFB440343" blockId="4.[831,1379,1477,1662]" pageId="4" pageNumber="93">
MHNCSJ (Medellín) 0294, una hembra adulta de “Urrao, Parque Las Orquídeas (Polo), 
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, 
<collectingRegion id="499FA7D0FFCF907A8D60F9EAFC340323" box="[831,937,1541,1566]" country="Colombia" name="Antioquia" pageId="4" pageNumber="93">Antioquia</collectingRegion>
, 
<collectingCountry id="F34C29A2FFCF907A8DE4F9EAFBBF0323" box="[955,1058,1541,1566]" name="Colombia" pageId="4" pageNumber="93">Colombia</collectingCountry>
” = Parque Nacional Natural “Las Orquídeas” (Sitio El Polo), municipio de Urrao, 
<collectingRegion id="499FA7D0FFCF907A8D60F9AAFBC10363" box="[831,1116,1605,1630]" country="Colombia" name="Antioquia" pageId="4" pageNumber="93">departamento de Antioquia</collectingRegion>
, 
<collectingCountry id="F34C29A2FFCF907A8A39F9AAFB4D0363" box="[1126,1232,1605,1630]" name="Colombia" pageId="4" pageNumber="93">Colombia</collectingCountry>
, colectado por Marco A. Serna el 20 de mayo de 1980.
</paragraph>
<paragraph id="8BE46932FFCF907A8D60F92AFC0B03E0" blockId="4.[831,918,1733,1757]" box="[831,918,1733,1757]" pageId="4" pageNumber="93">
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<typeStatus id="54E0D790FFCF907A8D60F92AFC0B03E0" box="[831,918,1733,1757]" pageId="4" pageNumber="93" type="neotype">Neotipo</typeStatus>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE46932FFCF907A8D60F8EAFC4D02E3" blockId="4.[831,1379,1797,2014]" pageId="4" pageNumber="93">
MHNCSJ 0291 = (MHUA 5851), un macho adulto de “Urrao, Parque Las Orquídeas (Polo), 
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, 
<collectingRegion id="499FA7D0FFCF907A8D60F8AAFC340263" box="[831,937,1861,1886]" country="Colombia" name="Antioquia" pageId="4" pageNumber="93">Antioquia</collectingRegion>
, 
<collectingCountry id="F34C29A2FFCF907A8DE4F8AAFBBF0263" box="[955,1058,1861,1886]" name="Colombia" pageId="4" pageNumber="93">Colombia</collectingCountry>
” = Parque Nacional Natural “Las Orquídeas” (Sitio El Polo), municipio de Urrao, 
<collectingRegion id="499FA7D0FFCF907A8D60F86AFBC102A3" box="[831,1116,1925,1950]" country="Colombia" name="Antioquia" pageId="4" pageNumber="93">departamento de Antioquia</collectingRegion>
, 
<collectingCountry id="F34C29A2FFCF907A8A39F86AFB4D02A3" box="[1126,1232,1925,1950]" name="Colombia" pageId="4" pageNumber="93">Colombia</collectingCountry>
, colectado por Marco A. Serna el 20 de mayo de 1980. Depositado en el MHUA.
</paragraph>
</subSubSection>
<caption id="DF2439BAFFCF907A8EAAF879FE7002E0" ID-DOI="http://doi.org/10.5281/zenodo.13314526" ID-Zenodo-Dep="13314526" httpUri="https://zenodo.org/record/13314526/files/figure.png" pageId="4" pageNumber="93" startId="4.[245,324,1942,1962]" targetBox="[245,793,1187,1905]" targetPageId="4" targetType="figure">
<paragraph id="8BE46932FFCF907A8EAAF879FE7002E0" blockId="4.[245,794,1942,2013]" pageId="4" pageNumber="93">
<emphasis id="B92FB520FFCF907A8EAAF879FEFC0297" bold="true" box="[245,353,1942,1962]" pageId="4" pageNumber="93">FIGURA 5:</emphasis>
Fotografía de 
<emphasis id="B92FB520FFCF907A8FB3F879FD0A0297" box="[492,663,1942,1962]" italics="true" pageId="4" pageNumber="93">
<taxonomicName id="4C5B12B1FFCF907A8FB3F879FD0E0297" authorityName="Y AMPLIA C I ON DE LA DIST R IBU C I ON DE NUE V E R ANAS DE C R ISTAL" baseAuthorityName="RI V E R O" baseAuthorityYear="1985" box="[492,659,1942,1962]" class="Amphibia" family="Centrolenidae" genus="Cochranella" kingdom="Animalia" order="Anura" pageId="4" pageNumber="93" phylum="Chordata" rank="species" species="megista">Cochranella megista</taxonomicName>
,
</emphasis>
macho adulto (Neotipo), MHNCSJ 291 (Actualmente MHUA 5851). Aspecto ventral de A = Mano; B = Pie.
</paragraph>
</caption>
<subSubSection id="C3413AB9FFCE90768E8FFEAAFB4903A3" lastPageId="8" lastPageNumber="97" pageId="5" pageNumber="94" type="diagnosis">
<paragraph id="8BE46932FFCE907B8E8FFEAAFEA50460" blockId="5.[208,312,325,349]" box="[208,312,325,349]" pageId="5" pageNumber="94">
<heading id="D0ACDE5EFFCE907B8E8FFEAAFEA50460" box="[208,312,325,349]" fontSize="10" level="3" pageId="5" pageNumber="94" reason="8">
<emphasis id="B92FB520FFCE907B8E8FFEAAFEA50460" bold="true" box="[208,312,325,349]" pageId="5" pageNumber="94">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BE46932FFCE907B8E8FFE69FDD00003" blockId="5.[208,757,389,1342]" pageId="5" pageNumber="94">
Una especie reconocible por los siguientes caracteres: 1) procesos dentígeros y dientes vomerinos presentes; 2) rostro subovoide en vista dorsal, redondeado en vista lateral; 3) tímpano y anillo timpánico visibles, pequeños, orientados posterodorsalmente; 4) en los machos, piel dorsal con abundantes espículas cónicas de distribución uniforme; en las hembras, dorso con verrugas casi planas y escasas espículas; 5) piel ventral granular; región subcloacal débilmente granular, ornamentación cloacal presente como finos puntos blanquecinos justo en el área debajo de la cloaca; 6) peritoneo parietal (1/2 anterior) y pericardio blancos, peritoneos hepático y visceral translúcidos; 7) hígado tetralobulado; 8) espina humeral ausente; 9) fórmula de la palmeadura manual III 2 – 
<superScript id="7C2EC47AFFCE907B8F43FCAAFEA3066E" attach="left" box="[284,318,836,862]" fontSize="6" pageId="5" pageNumber="94">11/2</superScript>
IV; 10) fórmula de la palmeadura pedial I 
<superScript id="7C2EC47AFFCE907B8E8FFC8AFF6C064E" attach="left" box="[208,241,868,894]" fontSize="6" pageId="5" pageNumber="94">11/3</superScript>
– (2-2 
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) II (1 
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1) – 2 
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III (1-1 
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) – (2 
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- 
<superScript id="7C2EC47AFFCE907B8C23FC8AFD02064E" attach="none" box="[636,671,868,894]" fontSize="6" pageId="5" pageNumber="94">21/4</superScript>
) IV 
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– (1-1 
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) V; 11) hilera de pequeños tubérculos sobre los pliegues ulnares y tarsales; 12) excrecencias nupciales tipo I en los machos adultos; prepollex no visible externamente; 13) dedos manuales I y II del mismo tamaño cuando se encuentran adpresos; 14) diámetro del ojo alrededor de 1.5 veces más grande que el tamaño del disco del dedo III; 15) en vida, superficies dorsales del cuerpo y extremidades de color verde con puntos negros o azules; en etanol al 70%, dorso lavanda con puntos azules; huesos verdes en vida; 16) en vida, iris gris pálido con retículo grisáceo; 17) melanóforos sobre los discos de los dedos manuales y pediales; 18) adultos de tamaño grande, machos adultos 
<quantity id="4CA3C4D7FFCE907B8C40FAEAFD290020" box="[543,692,1285,1310]" metricMagnitude="-2" metricUnit="m" metricValue="3.505" metricValueMax="3.7399999999999998" metricValueMin="3.27" pageId="5" pageNumber="94" unit="mm" value="35.05" valueMax="37.4" valueMin="32.7">32.7-37.4 mm</quantity>
LRC, hembras adultas 36.9-40.0 mm LRC.
</paragraph>
<paragraph id="8BE46932FFCE907B8F57FA8AFABD0663" blockId="5.[208,756,1381,2014]" lastBlockId="5.[793,1342,325,862]" pageId="5" pageNumber="94">
<taxonomicName id="4C5B12B1FFCE907B8F57FA8AFE560040" authorityName="Y AMPLIA C I ON DE LA DIST R IBU C I ON DE NUE V E R ANAS DE C R ISTAL" baseAuthorityName="RI V E R O" baseAuthorityYear="1985" box="[264,459,1381,1405]" class="Amphibia" family="Centrolenidae" genus="Cochranella" kingdom="Animalia" order="Anura" pageId="5" pageNumber="94" phylum="Chordata" rank="species" species="megista">
<emphasis id="B92FB520FFCE907B8F57FA8AFE560040" box="[264,459,1381,1405]" italics="true" pageId="5" pageNumber="94">Cochranella megista</emphasis>
</taxonomicName>
se diferencia de los otros dos géneros dentro de 
<taxonomicName id="4C5B12B1FFCE907B8FD0FA6AFDBD00A3" authorityName="Taylor" authorityYear="1951" box="[399,544,1413,1438]" class="Amphibia" family="Centrolenidae" kingdom="Animalia" order="Anura" pageId="5" pageNumber="94" phylum="Chordata" rank="family">Centrolenidae</taxonomicName>
<emphasis id="B92FB520FFCE907B8C77FA6AFD0400A0" box="[552,665,1413,1437]" italics="true" pageId="5" pageNumber="94">
(
<taxonomicName id="4C5B12B1FFCE907B8C70FA6AFD0400A0" authorityName="Jimenez de la Espada" authorityYear="1872" box="[559,665,1413,1437]" class="Amphibia" family="Centrolenidae" genus="Centrolene" kingdom="Animalia" order="Anura" pageId="5" pageNumber="94" phylum="Chordata" rank="genus">Centrolene</taxonomicName>
</emphasis>
y 
<emphasis id="B92FB520FFCE907B8CEBFA6AFEFC0080" italics="true" pageId="5" pageNumber="94">
<taxonomicName id="4C5B12B1FFCE907B8CEBFA6AFEC70080" authorityName="Ruiz-Carranza &amp; Lynch" authorityYear="1991" class="Amphibia" family="Centrolenidae" genus="Hyalinobatrachium" kingdom="Animalia" order="Anura" pageId="5" pageNumber="94" phylum="Chordata" rank="genus">Hyalinobatrachium</taxonomicName>
)
</emphasis>
porque los machos adultos carecen de espinas humeral (presentes en 
<taxonomicName id="4C5B12B1FFCE907B8C56FA2AFDEF00E0" authorityName="Jimenez de la Espada" authorityYear="1872" box="[521,626,1477,1501]" class="Amphibia" family="Centrolenidae" genus="Centrolene" kingdom="Animalia" order="Anura" pageId="5" pageNumber="94" phylum="Chordata" rank="genus">
<emphasis id="B92FB520FFCE907B8C56FA2AFDEF00E0" box="[521,626,1477,1501]" italics="true" pageId="5" pageNumber="94">Centrolene</emphasis>
</taxonomicName>
) y por tener el hígado cubierto por un peritoneo transparente (peritoneo hepático blanco en 
<taxonomicName id="4C5B12B1FFCE907B8FB7F9EAFD350320" authorityName="Ruiz-Carranza &amp; Lynch" authorityYear="1991" box="[488,680,1541,1565]" class="Amphibia" family="Centrolenidae" genus="Hyalinobatrachium" kingdom="Animalia" order="Anura" pageId="5" pageNumber="94" phylum="Chordata" rank="genus">
<emphasis id="B92FB520FFCE907B8FB7F9EAFD350320" box="[488,680,1541,1565]" italics="true" pageId="5" pageNumber="94">Hyalinobatrachium</emphasis>
</taxonomicName>
). En el género 
<emphasis id="B92FB520FFCE907B8F44F9CAFD990300" box="[283,516,1573,1597]" italics="true" pageId="5" pageNumber="94">
<taxonomicName id="4C5B12B1FFCE907B8F44F9CAFE0E0300" authorityName="Taylor" authorityYear="1951" box="[283,403,1573,1597]" class="Amphibia" family="Centrolenidae" genus="Cochranella" kingdom="Animalia" order="Anura" pageId="5" pageNumber="94" phylum="Chordata" rank="genus">Cochranella</taxonomicName>
, 
<taxonomicName id="4C5B12B1FFCE907B8FC1F9C9FD990300" authorityName="Y AMPLIA C I ON DE LA DIST R IBU C I ON DE NUE V E R ANAS DE C R ISTAL" baseAuthorityName="RI V E R O" baseAuthorityYear="1985" box="[414,516,1574,1597]" class="Amphibia" family="Centrolenidae" genus="Cochranella" kingdom="Animalia" order="Anura" pageId="5" pageNumber="94" phylum="Chordata" rank="species" species="megista">C. megista</taxonomicName>
</emphasis>
pertenece al grupo 
<emphasis id="B92FB520FFCE907B8C8EF9C9FF660360" italics="true" pageId="5" pageNumber="94">spinosa</emphasis>
(sensu Ruiz-Carranza &amp; Lynch, 1995a), el cual se distingue de los otros grupos infragenéricos por tener una membrana extensa entre los dedos III y IV de la mano y por carecer de iridóforos en el peritoneo gastrointestinal. Las especies del grupo 
<taxonomicName id="4C5B12B1FFCE907B8C66F92AFF6903C0" baseAuthorityName="Taylor" baseAuthorityYear="1949" class="Amphibia" family="Centrolenidae" genus="Cochranella" kingdom="Animalia" order="Anura" pageId="5" pageNumber="94" phylum="Chordata" rank="species" species="granulosa">
<emphasis id="B92FB520FFCE907B8C66F92AFF6903C0" italics="true" pageId="5" pageNumber="94">Cochranella granulosa</emphasis>
</taxonomicName>
presentan un peritoneo gastrointestinal blanco, mientras que las especies del grupo 
<taxonomicName id="4C5B12B1FFCE907B8C6EF8EAFD690220" baseAuthorityName="Boulenger" baseAuthorityYear="1918" box="[561,756,1797,1821]" class="Amphibia" family="Centrolenidae" genus="Cochranella" kingdom="Animalia" order="Anura" pageId="5" pageNumber="94" phylum="Chordata" rank="species" species="ocellata">
<emphasis id="B92FB520FFCE907B8C6EF8EAFD690220" box="[561,756,1797,1821]" italics="true" pageId="5" pageNumber="94">Cochranella ocellata</emphasis>
</taxonomicName>
(ahora pertenecientes al género 
<taxonomicName id="4C5B12B1FFCE907B8C4EF8CAFD110200" authorityName="Cisneros-Heredia &amp; McDiarmid" authorityYear="2007" box="[529,652,1829,1853]" class="Amphibia" family="Centrolenidae" genus="Nymphargus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="94" phylum="Chordata" rank="genus">
<emphasis id="B92FB520FFCE907B8C4EF8CAFD110200" box="[529,652,1829,1853]" italics="true" pageId="5" pageNumber="94">Nymphargus</emphasis>
</taxonomicName>
Cisneros-Heredia y McDiarmid 2007) tienen una membrana reducida entre los dedos manuales III y IV. De las 22 especies del grupo 
<emphasis id="B92FB520FFCE907B8FCCF869FE4502A0" box="[403,472,1926,1949]" italics="true" pageId="5" pageNumber="94">spinosa</emphasis>
(
<bibRefCitation id="EFCA14C3FFCE907B8FB8F86AFD7502A3" author="GUAYASAMIN, J. M. &amp; BUSTAMANTE, M. R. &amp; ALMEDIA-REINOSO, D. &amp; FUNK, W. C." box="[487,744,1925,1950]" pageId="5" pageNumber="94" pagination="489 - 513" refId="ref7080" refString="GUAYASAMIN, J. M.; BUSTAMANTE, M. R.; ALMEDIA-REINOSO, D. &amp; FUNK, W. C. 2006. Glass frogs (Centrolenidae) of Yanayacu Biological Station, Ecuador, with the description of a new species and comments on centrolenid systematics. Zoological Journal of the Linnean Society, 147: 489 - 513." type="journal article" year="2006">
Guayasamin 
<emphasis id="B92FB520FFCE907B8C2FF869FD3A02A0" box="[624,679,1925,1949]" italics="true" pageId="5" pageNumber="94">et al.,</emphasis>
2006
</bibRefCitation>
), únicamente dos 
<emphasis id="B92FB520FFCE907B8FDFF849FD1B0280" box="[384,646,1958,1981]" italics="true" pageId="5" pageNumber="94">
(
<taxonomicName id="4C5B12B1FFCE907B8FD6F849FE6F0280" authorityName="Y AMPLIA C I ON DE LA DIST R IBU C I ON DE NUE V E R ANAS DE C R ISTAL" baseAuthorityName="RI V E R O" baseAuthorityYear="1985" box="[393,498,1958,1981]" class="Amphibia" family="Centrolenidae" genus="Cochranella" kingdom="Animalia" order="Anura" pageId="5" pageNumber="94" phylum="Chordata" rank="species" species="megista">C. megista</taxonomicName>
, 
<taxonomicName id="4C5B12B1FFCE907B8C5EF849FDE20280" baseAuthorityName="Flores" baseAuthorityYear="1987" box="[513,639,1958,1981]" class="Amphibia" family="Centrolenidae" genus="Cochranella" kingdom="Animalia" order="Anura" pageId="5" pageNumber="94" phylum="Chordata" rank="species" species="ametarsia">C. ametarsia</taxonomicName>
)
</emphasis>
presentan puntos oscuros en el dorso. 
<taxonomicName id="4C5B12B1FFCE907B8FA5F82AFD4A02E0" baseAuthorityName="Flores" baseAuthorityYear="1987" box="[506,727,1989,2013]" class="Amphibia" family="Centrolenidae" genus="Cochranella" kingdom="Animalia" order="Anura" pageId="5" pageNumber="94" phylum="Chordata" rank="species" species="ametarsia">
<emphasis id="B92FB520FFCE907B8FA5F82AFD4A02E0" box="[506,727,1989,2013]" italics="true" pageId="5" pageNumber="94">Cochranella ametarsia</emphasis>
</taxonomicName>
se distingue de 
<taxonomicName id="4C5B12B1FFCE907B8DFDFEA9FB950460" authorityName="Y AMPLIA C I ON DE LA DIST R IBU C I ON DE NUE V E R ANAS DE C R ISTAL" baseAuthorityName="RI V E R O" baseAuthorityYear="1985" box="[930,1032,326,349]" class="Amphibia" family="Centrolenidae" genus="Cochranella" kingdom="Animalia" order="Anura" pageId="5" pageNumber="94" phylum="Chordata" rank="species" species="megista">
<emphasis id="B92FB520FFCE907B8DFDFEA9FB950460" box="[930,1032,326,349]" italics="true" pageId="5" pageNumber="94">C. megista</emphasis>
</taxonomicName>
por ser mucho más pequeña (LRC &lt;
<quantity id="4CA3C4D7FFCE907B8D31FE8AFC270443" box="[878,954,357,382]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="94" unit="mm" value="20.0">20 mm</quantity>
) y por tener una distribución restringida a las tierras del piso térmico cálido en la Amazonia. Finalmente, por su morfología externa 
<taxonomicName id="4C5B12B1FFCE907B8A88FE49FAA00480" authorityName="Y AMPLIA C I ON DE LA DIST R IBU C I ON DE NUE V E R ANAS DE C R ISTAL" baseAuthorityName="RI V E R O" baseAuthorityYear="1985" box="[1239,1341,422,445]" class="Amphibia" family="Centrolenidae" genus="Cochranella" kingdom="Animalia" order="Anura" pageId="5" pageNumber="94" phylum="Chordata" rank="species" species="megista">
<emphasis id="B92FB520FFCE907B8A88FE49FAA00480" box="[1239,1341,422,445]" italics="true" pageId="5" pageNumber="94">C. megista</emphasis>
</taxonomicName>
es una especie comparable con 
<emphasis id="B92FB520FFCE907B8A3CFE29FC3904C0" italics="true" pageId="5" pageNumber="94">
<taxonomicName id="4C5B12B1FFCE907B8A3CFE29FB5C04E0" authorityName="Ruiz-Carranza &amp; Lynch" authorityYear="1995" box="[1123,1217,453,477]" class="Amphibia" family="Centrolenidae" genus="Cochranella" kingdom="Animalia" order="Anura" pageId="5" pageNumber="94" phylum="Chordata" rank="species" species="chami">C. chami</taxonomicName>
, 
<taxonomicName id="4C5B12B1FFCE907B8A91FE29FAA704E0" authorityName="Ruiz-Carranza &amp; Lynch" authorityYear="1995" box="[1230,1338,453,477]" class="Amphibia" family="Centrolenidae" genus="Cochranella" kingdom="Animalia" order="Anura" pageId="5" pageNumber="94" phylum="Chordata" rank="species" species="posadae">C. posadae</taxonomicName>
, 
<taxonomicName id="4C5B12B1FFCE907B8D46FE09FC3904C0" baseAuthorityName="Lynch &amp; Duellman" baseAuthorityYear="1973" box="[793,932,485,509]" class="Amphibia" family="Centrolenidae" genus="Cochranella" kingdom="Animalia" order="Anura" pageId="5" pageNumber="94" phylum="Chordata" rank="species" species="megacheira">C. megacheira</taxonomicName>
</emphasis>
y 
<emphasis id="B92FB520FFCE907B8D9BFE09FBA704C0" box="[964,1082,486,509]" italics="true" pageId="5" pageNumber="94">
<taxonomicName id="4C5B12B1FFCE907B8D9BFE09FBAB04C0" authorityName="Ruiz-Carranza &amp; Lynch" authorityYear="1995" box="[964,1078,486,509]" class="Amphibia" family="Centrolenidae" genus="Cochranella" kingdom="Animalia" order="Anura" pageId="5" pageNumber="94" phylum="Chordata" rank="species" species="cristinae">C. cristinae</taxonomicName>
,
</emphasis>
sin embargo, 
<taxonomicName id="4C5B12B1FFCE907B8A8BFE09FAA304C0" authorityName="Ruiz-Carranza &amp; Lynch" authorityYear="1995" box="[1236,1342,485,509]" class="Amphibia" family="Centrolenidae" genus="Cochranella" kingdom="Animalia" order="Anura" pageId="5" pageNumber="94" phylum="Chordata" rank="species" species="posadae">
<emphasis id="B92FB520FFCE907B8A8BFE09FAA304C0" box="[1236,1342,485,509]" italics="true" pageId="5" pageNumber="94">C. posadae</emphasis>
</taxonomicName>
tiene un color dorsal verde claro y la piel dorsal es lisa con escasos gránulos crema; 
<taxonomicName id="4C5B12B1FFCE907B8A60FDC9FB070700" authorityName="Ruiz-Carranza &amp; Lynch" authorityYear="1995" box="[1087,1178,549,573]" class="Amphibia" family="Centrolenidae" genus="Cochranella" kingdom="Animalia" order="Anura" pageId="5" pageNumber="94" phylum="Chordata" rank="species" species="chami">
<emphasis id="B92FB520FFCE907B8A60FDC9FB070700" box="[1087,1178,549,573]" italics="true" pageId="5" pageNumber="94">C. chami</emphasis>
</taxonomicName>
tiene abundantes tubérculos cónicos crema y si bien el tamaño es comparable, la palmeadura manual y pedial es mucho menor y no presenta puntos oscuros en el dorso. La coloración de 
<taxonomicName id="4C5B12B1FFCE907B8DEDFD49FBA10780" baseAuthorityName="Lynch &amp; Duellman" baseAuthorityYear="1973" box="[946,1084,677,701]" class="Amphibia" family="Centrolenidae" genus="Cochranella" kingdom="Animalia" order="Anura" pageId="5" pageNumber="94" phylum="Chordata" rank="species" species="megacheira">
<emphasis id="B92FB520FFCE907B8DEDFD49FBA10780" box="[946,1084,677,701]" italics="true" pageId="5" pageNumber="94">C. megacheira</emphasis>
</taxonomicName>
y 
<taxonomicName id="4C5B12B1FFCE907B8A01FD49FB530780" authorityName="Ruiz-Carranza &amp; Lynch" authorityYear="1995" box="[1118,1230,678,701]" class="Amphibia" family="Centrolenidae" genus="Cochranella" kingdom="Animalia" order="Anura" pageId="5" pageNumber="94" phylum="Chordata" rank="species" species="cristinae">
<emphasis id="B92FB520FFCE907B8A01FD49FB530780" box="[1118,1230,678,701]" italics="true" pageId="5" pageNumber="94">C. cristinae</emphasis>
</taxonomicName>
puede ser similar a la de 
<taxonomicName id="4C5B12B1FFCE907B8DEDFD29FB8507E0" authorityName="Y AMPLIA C I ON DE LA DIST R IBU C I ON DE NUE V E R ANAS DE C R ISTAL" baseAuthorityName="RI V E R O" baseAuthorityYear="1985" box="[946,1048,710,733]" class="Amphibia" family="Centrolenidae" genus="Cochranella" kingdom="Animalia" order="Anura" pageId="5" pageNumber="94" phylum="Chordata" rank="species" species="megista">
<emphasis id="B92FB520FFCE907B8DEDFD29FB8507E0" box="[946,1048,710,733]" italics="true" pageId="5" pageNumber="94">C. megista</emphasis>
</taxonomicName>
(verde con puntos oscuros), pero ambas especies son de menor tamaño (
<emphasis id="B92FB520FFCE907B8AB8FD09FCDF0620" italics="true" pageId="5" pageNumber="94">
<taxonomicName id="4C5B12B1FFCE907B8AB8FD09FCA30620" authorityName="Ruiz-Carranza &amp; Lynch" authorityYear="1995" class="Amphibia" family="Centrolenidae" genus="Cochranella" kingdom="Animalia" order="Anura" pageId="5" pageNumber="94" phylum="Chordata" rank="species" species="cristinae">C. cristinae</taxonomicName>
,
</emphasis>
26.0- 
<quantity id="4CA3C4D7FFCE907B8DDEFCEAFC7D0620" box="[897,992,773,798]" metricMagnitude="-2" metricUnit="m" metricValue="3.11" pageId="5" pageNumber="94" unit="mm" value="31.1">31.1 mm</quantity>
LRC en machos adultos y 
<taxonomicName id="4C5B12B1FFCE907B8AA5FCE9FCF40600" baseAuthorityName="Lynch &amp; Duellman" baseAuthorityYear="1973" class="Amphibia" family="Centrolenidae" genus="Cochranella" kingdom="Animalia" order="Anura" pageId="5" pageNumber="94" phylum="Chordata" rank="species" species="megacheira">
<emphasis id="B92FB520FFCE907B8AA5FCE9FCF40600" italics="true" pageId="5" pageNumber="94">C. megacheira</emphasis>
</taxonomicName>
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LRC en machos adultos) y los rostros son más cortos y truncados en perfil lateral.
</paragraph>
<paragraph id="8BE46932FFCE907B8D46FC4BFB8B0680" blockId="5.[793,1156,932,989]" box="[793,1046,932,957]" pageId="5" pageNumber="94">
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<emphasis id="B92FB520FFCE907B8D46FC4BFB8B0680" bold="true" box="[793,1046,932,957]" pageId="5" pageNumber="94">
Descripción del 
<typeStatus id="54E0D790FFCE907B8D9AFC4AFB8B0680" box="[965,1046,933,957]" pageId="5" pageNumber="94" type="neotype">neotipo</typeStatus>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE46932FFCE907B8D46FC2AFB1906E0" blockId="5.[793,1156,932,989]" box="[793,1156,965,989]" pageId="5" pageNumber="94">
<heading id="D0ACDE5EFFCE907B8D46FC2AFB1906E0" box="[793,1156,965,989]" fontSize="10" level="3" pageId="5" pageNumber="94" reason="8">
<emphasis id="B92FB520FFCE907B8D46FC2AFB1906E0" bold="true" box="[793,1156,965,989]" pageId="5" pageNumber="94">(MHNCSJ 0291, = MHUA 5851)</emphasis>
</heading>
</paragraph>
<paragraph id="8BE46932FFCE907B8D0DFBEAFB1D02E0" blockId="5.[793,1342,1029,2014]" pageId="5" pageNumber="94">
Macho adulto de tamaño grande (LRC = 
<quantity id="4CA3C4D7FFCE907B8D32FBCAFC520103" box="[877,975,1061,1086]" metricMagnitude="-2" metricUnit="m" metricValue="3.3899999999999997" pageId="5" pageNumber="94" unit="mm" value="33.9">33.9 mm</quantity>
). Cabeza más ancha que el cuerpo, anchura cefálica 26.8% de la longitud rostro-cloacal; rostro subovoide en vista dorsal, truncado, débilmente inclinado en vista lateral (
<figureCitation id="136075B7FFCE907B8A48FB6AFBCF01A3" box="[1047,1106,1157,1182]" captionStart="FIGURA 4" captionStartId="4.[245,324,953,973]" captionTargetBox="[245,1379,311,915]" captionTargetId="figure-185@4.[245,1379,311,915]" captionTargetPageId="4" captionText="FIGURA 4: Fotografía de Cochranella megista, macho adulto (Neotipo), MHNCSJ 291(Actualmente MHUA 5851). A, B = Aspectos dorsal y lateral de la cabeza." figureDoi="http://doi.org/10.5281/zenodo.13314524" httpUri="https://zenodo.org/record/13314524/files/figure.png" pageId="5" pageNumber="94">Fig. 4</figureCitation>
A-B). Distancia ojo-narina 61.7% del diámetro anteroposterior del ojo; región loreal ligeramente cóncava, descendiendo abruptamente hacia el labio superior que es redondeado, no expandido. Narinas algo prominentes dirigidas anterolateralmente, con débil depresión entre ellas y con aberturas externas elipsoideas; 
<emphasis id="B92FB520FFCE907B8DA9FAAAFB0A0060" box="[1014,1175,1349,1373]" italics="true" pageId="5" pageNumber="94">canthus rostralis</emphasis>
redondos, algo pronunciados y ligeramente cóncavos. Ojos grandes, dirigidos anterolateralmente en ángulo de 
<emphasis id="B92FB520FFCE907B8A9EFA69FB4B00A0" box="[1217,1238,1414,1437]" italics="true" pageId="5" pageNumber="94">ca</emphasis>
45°, no se extienden más allá del borde externo del labio superior cuando la cabeza se observa en vista ventral, diámetro anteroposterior del ojo equivalente al 51.6% de la longitud cefálica; anchura del párpado superior equivalente al 70.6% de la distancia interorbital. Tímpano y anillo timpánico visibles en sus 3/4 inferiores, de posición posterodorsolateral, cuyo diámetro equivale al 19.1% del diámetro ocular, pliegue supratimpánico poco visible, débilmente engrosado, se extiende desde el ángulo posterior de la órbita hasta cerca del nivel de la inserción del brazo. Aberturas nasales internas grandes, ovoideas; odontóforos y dientes vomerinos presentes; lengua redondeada, su anchura algo mayor que su longitud, con el borde posterior ligeramente escotado, márgenes posteriores y laterales no adheridos al piso de la boca; hendiduras vocales dirigidas de la base posterior lateral de la lengua a los ángulos mandibulares; saco vocal sencillo, medial, externo.
</paragraph>
<paragraph id="8BE46932FFCD90788C2CFF02FC7B043C" blockId="6.[627,998,237,257]" box="[627,998,237,257]" pageId="6" pageNumber="95">PA P ÉIS AVULSOS DE ZOOLOGIA, 48(12), 2008</paragraph>
<paragraph id="8BE46932FFCD90788B11FF02FAFE043C" blockId="6.[1358,1379,237,257]" box="[1358,1379,237,257]" pageId="6" pageNumber="95">95</paragraph>
<paragraph id="8BE46932FFCD90788F71FEAAFDCE0660" blockId="6.[245,794,325,1310]" pageId="6" pageNumber="95">
Espina humeral ausente; antebrazo moderadamente robusto, con un bajo pliegue ulnar en el que se apenas se distinguen una serie de tubérculos y puntos muy bajos; dedo manual I igual que el II, extremo del dedo IV no alcanza la base del disco del dedo III, dedos manuales anchos con discos de borde distal truncado y contorneado por un fino reborde lateral dérmico, de diámetro transverso más ancho que los artejos dístales correspondientes; sin membranas entre los dedos internos de la mano, entre los dedos externos la fórmula modal corresponde a III 2 – 
<superScript id="7C2EC47AFFCD90788C32FD6AFD1207AE" attach="left" box="[621,655,644,670]" fontSize="6" pageId="6" pageNumber="95">11/2</superScript>
IV (
<figureCitation id="136075B7FFCD90788CE2FD6AFC9007A3" box="[701,781,645,670]" captionStart="FIGURA 5" captionStartId="4.[245,324,1942,1962]" captionTargetBox="[245,793,1187,1905]" captionTargetId="figure-161@4.[245,793,1187,1905]" captionTargetPageId="4" captionText="FIGURA 5: Fotografía de Cochranella megista, macho adulto (Neotipo), MHNCSJ 291 (Actualmente MHUA 5851). Aspecto ventral de A = Mano; B = Pie." figureDoi="http://doi.org/10.5281/zenodo.13314526" httpUri="https://zenodo.org/record/13314526/files/figure.png" pageId="6" pageNumber="95">Fig. 5A</figureCitation>
); tubérculos subarticulares grandes, redondos, sencillos, los dístales de mayor tamaño que los básales; tubérculos supernumerarios manuales numerosos, redondos, localizados en la base de los dedos; tubérculo palmar más largo que ancho, tenar elíptico igual que el palmar; excrecencias nupciales tipo I.
</paragraph>
<paragraph id="8BE46932FFCD90788F71FC8AFB920763" blockId="6.[245,794,325,1310]" lastBlockId="6.[831,1380,325,862]" pageId="6" pageNumber="95">
Extremidades posteriores delgadas, longitud tibial equivalente al 57.8% de la longitud rostro-cloacal, cuando el miembro posterior se halla adpreso hacia delante, el talón sobrepasa ligeramente el extremo rostral; cuando los miembros posteriores se hallan flexados hacia delante y los posteriores hacia atrás, las rodillas se sobreponen a los codos; cuando los muslos se hallan perpendiculares al plano sagital del cuerpo y las piernas flexadas hacia dentro, los talones se sobreponen entre si; pliegue tarsal con pequeños tubérculos; tubérculo metatarsal externo no evidenciable; tubérculo metatarsal interno ovalado, dos veces más largo que ancho, externo no evidenciable, tubérculos subarticulares grandes, redondos, prominentes, tubérculos supernumerarios plantares abundantes y pequeños, redondos, situados en la base de los dedos; discos de los dedos pediales más pequeños que los manuales, redondeados, con tendencia a ser truncados, con fino reborde, ligeramente mas anchos que los artejos dístales correspondientes; extensas membranas entre los dedos del pie, fórmula de la palmeadura pedial I 
<superScript id="7C2EC47AFFCD90788D60FDCAFCFF070E" attach="left" box="[831,866,548,574]" fontSize="6" pageId="6" pageNumber="95">11/3</superScript>
– (2-2 
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) II (1 
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1) – 2 
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III (1-1 
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) – (2 
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- 
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) IV 
<superScript id="7C2EC47AFFCD90788B1EFDCAFAFE070E" attach="none" box="[1345,1379,548,574]" fontSize="6" pageId="6" pageNumber="95">21/4</superScript>
– (1-1 
<superScript id="7C2EC47AFFCD90788D20FDABFC1A076E" attach="none" box="[895,903,580,595]" fontSize="6" pageId="6" pageNumber="95">+</superScript>
) V (
<figureCitation id="136075B7FFCD90788DECFDAAFB9F0763" box="[947,1026,581,606]" captionStart="FIGURA 5" captionStartId="4.[245,324,1942,1962]" captionTargetBox="[245,793,1187,1905]" captionTargetId="figure-161@4.[245,793,1187,1905]" captionTargetPageId="4" captionText="FIGURA 5: Fotografía de Cochranella megista, macho adulto (Neotipo), MHNCSJ 291 (Actualmente MHUA 5851). Aspecto ventral de A = Mano; B = Pie." figureDoi="http://doi.org/10.5281/zenodo.13314526" httpUri="https://zenodo.org/record/13314526/files/figure.png" pageId="6" pageNumber="95">Fig. 5B</figureCitation>
).
</paragraph>
<paragraph id="8BE46932FFCD90788D27FD8AFAF90663" blockId="6.[831,1380,325,862]" pageId="6" pageNumber="95">Piel de las superficies expuestas con pequeñas espículas dispuestas de manera uniforme sobre las superficies dorsales del ejemplar, algunas cónicas; garganta lisa, vientre y superficie posteroventral del muslo densamente granular; vaina cloacal no levantada, horizontal, dirigida posteriomedialmente y situada a nivel superior de los muslos, con un par de tubérculos en la región inferior de los muslos justo debajo de la cloaca.</paragraph>
<paragraph id="8BE46932FFCD90788D60FC4BFB4F0680" blockId="6.[831,1234,932,957]" box="[831,1234,932,957]" pageId="6" pageNumber="95">
<heading id="D0ACDE5EFFCD90788D60FC4BFB4F0680" box="[831,1234,932,957]" fontSize="10" level="3" pageId="6" pageNumber="95" reason="8">
<emphasis id="B92FB520FFCD90788D60FC4BFB4F0680" bold="true" box="[831,1234,932,957]" pageId="6" pageNumber="95">
Coloración en etanol 70% del 
<typeStatus id="54E0D790FFCD90788ADEFC4AFB4F0680" box="[1153,1234,933,957]" pageId="6" pageNumber="95" type="neotype">neotipo</typeStatus>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE46932FFCD90788D27FC0AFAC40163" blockId="6.[831,1380,997,1118]" pageId="6" pageNumber="95">
Dorsalmente todas la superficies castaño con motas café oscuro y puntos oscuros casi negros, algunos en correspondencia con las espículas levantadas (
<figureCitation id="136075B7FFCD90788D19FBAAFC350163" box="[838,936,1093,1118]" captionStart="FIGURA 3" captionStartId="3.[208,287,1274,1294]" captionTargetBox="[208,1342,311,1237]" captionTargetId="figure-270@3.[208,1342,311,1237]" captionTargetPageId="3" captionText="FIGURA 3: Fotografía e ilustración dorsal comparativas de (A): Centrolene grandisonae, hembra adulta, MHNCSJ “294” – falso holotipo; (B): Cochranella megista, hembra adulta, MHNCSJ 294 (Holotipo perdido, ilustración tomada de Rivero, 1985); (C): Cochranella megista, macho adulto (Neotipo) MHNCS 291 (Actualmente MHUA 5851); (D): Cochranella megista, macho adulto, ICN 27768." figureDoi="http://doi.org/10.5281/zenodo.13314522" httpUri="https://zenodo.org/record/13314522/files/figure.png" pageId="6" pageNumber="95">Fig. 3C y</figureCitation>
<figureCitation id="136075B7FFCD90788DF1FBAAFC260163" box="[942,955,1093,1118]" captionStart="FIGURA 4" captionStartId="4.[245,324,953,973]" captionTargetBox="[245,1379,311,915]" captionTargetId="figure-185@4.[245,1379,311,915]" captionTargetPageId="4" captionText="FIGURA 4: Fotografía de Cochranella megista, macho adulto (Neotipo), MHNCSJ 291(Actualmente MHUA 5851). A, B = Aspectos dorsal y lateral de la cabeza." figureDoi="http://doi.org/10.5281/zenodo.13314524" httpUri="https://zenodo.org/record/13314524/files/figure.png" pageId="6" pageNumber="95">4</figureCitation>
A-B). Ventralmente café claro uniforme.
</paragraph>
<paragraph id="8BE46932FFCD90788D60FB4BFC3F0181" blockId="6.[831,930,1188,1212]" box="[831,930,1188,1212]" pageId="6" pageNumber="95">
<heading id="D0ACDE5EFFCD90788D60FB4BFC3F0181" box="[831,930,1188,1212]" fontSize="10" level="3" pageId="6" pageNumber="95" reason="8">
<emphasis id="B92FB520FFCD90788D60FB4BFC3F0181" bold="true" box="[831,930,1188,1212]" pageId="6" pageNumber="95">Variación</emphasis>
</heading>
</paragraph>
<paragraph id="8BE46932FFCD90798D27FB09FF78043C" blockId="6.[831,1379,1253,1310]" lastBlockId="7.[208,229,237,257]" lastPageId="7" lastPageNumber="96" pageId="6" pageNumber="95">
Las variaciones en las dimensiones corporales pueden ser observadas en la 
<tableCitation id="C6D95C89FFCD90788A2EFAEAFB230023" box="[1137,1214,1285,1310]" captionStart="TABLA 1" captionStartId="6.[245,311,1384,1404]" captionTargetBox="[257,1375,1536,2005]" captionTargetId="graphics-547@6.[245,1379,1527,2008]" captionText="TABLA 1: Variación en las dimensiones corporales (mm) de los adultos de Cochranella megista. Abreviaturas en Español: Longitud rostro cloacal (LRC), Longitud de la cabeza (LC), Ancho de la cabeza (AC), Diámetro horizontal del ojo (DO), Distancia ojo-narina (DON), Ancho del párpado superior (APS), Distancia interorbital (DIO), Diámetro transversal del tímpano (DTT), Diámetro del disco manual III (Ddd III), Longitud de la tibia (LT), Longitud de la mano (LM), Longitud del pie (LP). Las dimensiones del espécimen MHNCSJ 294 fueron tomadas de la publicación original (Rivero, 1985)." httpUri="http://table.plazi.org/id/DF2439BAFFCD90788EAAFA87FD4D00DE" pageId="6" pageNumber="95" tableUuid="DF2439BAFFCD90788EAAFA87FD4D00DE">Tabla 1</tableCitation>
. En general las 96
</paragraph>
<caption id="DF2439BAFFCD90788EAAFA87FD4D00DE" ID-Table-UUID="DF2439BAFFCD90788EAAFA87FD4D00DE" httpUri="http://table.plazi.org/id/DF2439BAFFCD90788EAAFA87FD4D00DE" pageId="6" pageNumber="95" startId="6.[245,311,1384,1404]" targetBox="[257,1375,1536,2005]" targetIsTable="true" targetPageId="6" targetType="table">
<paragraph id="8BE46932FFCD90788EAAFA87FD4D00DE" blockId="6.[245,1379,1384,1507]" pageId="6" pageNumber="95">
<emphasis id="B92FB520FFCD90788EAAFA87FED30041" bold="true" box="[245,334,1384,1404]" pageId="6" pageNumber="95">TABLA 1:</emphasis>
Variación en las dimensiones corporales (mm) de los adultos de 
<emphasis id="B92FB520FFCD90788D3AFA87FB970041" box="[869,1034,1384,1404]" italics="true" pageId="6" pageNumber="95">
<taxonomicName id="4C5B12B1FFCD90788D3AFA87FB9B0041" authorityName="Y AMPLIA C I ON DE LA DIST R IBU C I ON DE NUE V E R ANAS DE C R ISTAL" baseAuthorityName="RI V E R O" baseAuthorityYear="1985" box="[869,1030,1384,1404]" class="Amphibia" family="Centrolenidae" genus="Cochranella" kingdom="Animalia" order="Anura" pageId="6" pageNumber="95" phylum="Chordata" rank="species" species="megista">Cochranella megista</taxonomicName>
.
</emphasis>
Abreviaturas en Español: Longitud rostro cloacal (LRC), Longitud de la cabeza (LC), Ancho de la cabeza (AC), Diámetro horizontal del ojo (DO), Distancia ojo-narina (DON), Ancho del párpado superior (APS), Distancia interorbital (DIO), Diámetro transversal del tímpano (DTT), Diámetro del disco manual III (Ddd III), Longitud de la tibia (LT), Longitud de la mano (LM), Longitud del pie (LP). Las dimensiones del espécimen MHNCSJ 294 fueron tomadas de la publicación original (
<bibRefCitation id="EFCA14C3FFCD90788C09FA20FD5B00DE" author="RIVERO, J. A." box="[598,710,1487,1507]" pageId="6" pageNumber="95" pagination="334 - 373" refId="ref7316" refString="RIVERO, J. A. 1985. Nuevos Centrolenidos de Colombia y Venezuela. Brenesia, 23: 334 - 373." type="journal article" year="1985">Rivero, 1985</bibRefCitation>
).
</paragraph>
</caption>
<paragraph id="8BE46932FFCD90788F5EF9F5FACD02E8" pageId="6" pageNumber="95">
<table id="F95B9B92FFCD6F818F5EF9EFFAC202E8" box="[257,1375,1536,2005]" gridcols="13" gridrows="14" pageId="6" pageNumber="95">
<tr id="356B6B70FFCD6F818F5EF9EFFAC2037A" box="[257,1375,1536,1607]" gridrow="0" pageId="6" pageNumber="95">
<th id="76BA020CFFCD6F818F5EF9EFFEC7037A" box="[257,346,1536,1607]" gridcol="0" gridrow="0" pageId="6" pageNumber="95">Espécimen</th>
<th id="76BA020CFFCD6F818F32F9EFFE65037A" box="[365,504,1536,1607]" gridcol="1" gridrow="0" pageId="6" pageNumber="95">MHNCSJ 291 = MHNUA 5851 (Neotipo)</th>
<th id="76BA020CFFCD6F818C55F9EFFDF9037A" box="[522,612,1536,1607]" gridcol="2" gridrow="0" pageId="6" pageNumber="95">MHNCSJ 294 (Holotipo)</th>
<th id="76BA020CFFCD6F818C27F9EFFD2F037A" box="[632,690,1536,1607]" gridcol="3" gridrow="0" pageId="6" pageNumber="95">ICN 27763</th>
<th id="76BA020CFFCD6F818C9BF9EFFD63037A" box="[708,766,1536,1607]" gridcol="4" gridrow="0" pageId="6" pageNumber="95">ICN 27764</th>
<th id="76BA020CFFCD6F818D50F9EFFCD4037A" box="[783,841,1536,1607]" gridcol="5" gridrow="0" pageId="6" pageNumber="95">ICN 27765</th>
<th id="76BA020CFFCD6F818D04F9EFFC08037A" box="[859,917,1536,1607]" gridcol="6" gridrow="0" pageId="6" pageNumber="95">ICN 27766</th>
<th id="76BA020CFFCD6F818DF8F9EFFC7C037A" box="[935,993,1536,1607]" gridcol="7" gridrow="0" pageId="6" pageNumber="95">ICN 27767</th>
<th id="76BA020CFFCD6F818DADF9EFFBB1037A" box="[1010,1068,1536,1607]" gridcol="8" gridrow="0" pageId="6" pageNumber="95">ICN 27768</th>
<th id="76BA020CFFCD6F818A67F9EFFBE1037A" box="[1080,1148,1536,1607]" gridcol="9" gridrow="0" pageId="6" pageNumber="95">ICN 17242</th>
<th id="76BA020CFFCD6F818ADBF9EFFB55037A" box="[1156,1224,1536,1607]" gridcol="10" gridrow="0" pageId="6" pageNumber="95">ICN 17243</th>
<th id="76BA020CFFCD6F818A8AF9EFFA92037A" box="[1237,1295,1536,1607]" gridcol="11" gridrow="0" pageId="6" pageNumber="95">ICN 17244</th>
<th id="76BA020CFFCD6F818B44F9EFFAC2037A" box="[1307,1375,1536,1607]" gridcol="12" gridrow="0" pageId="6" pageNumber="95">ICN 28796</th>
</tr>
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<td id="76BA020CFFCD6F818C55F9BDFDF9035B" box="[522,612,1618,1638]" gridcol="2" gridrow="1" pageId="6" pageNumber="95">Hembra</td>
<td id="76BA020CFFCD6F818C27F9BDFD2F035B" box="[632,690,1618,1638]" gridcol="3" gridrow="1" pageId="6" pageNumber="95">Macho</td>
<td id="76BA020CFFCD6F818C9BF9BDFD63035B" box="[708,766,1618,1638]" gridcol="4" gridrow="1" pageId="6" pageNumber="95">Macho</td>
<td id="76BA020CFFCD6F818D50F9BDFCD4035B" box="[783,841,1618,1638]" gridcol="5" gridrow="1" pageId="6" pageNumber="95">Macho</td>
<td id="76BA020CFFCD6F818D04F9BDFC08035B" box="[859,917,1618,1638]" gridcol="6" gridrow="1" pageId="6" pageNumber="95">Macho</td>
<td id="76BA020CFFCD6F818DF8F9BDFC7C035B" box="[935,993,1618,1638]" gridcol="7" gridrow="1" pageId="6" pageNumber="95">Macho</td>
<td id="76BA020CFFCD6F818DADF9BDFBB1035B" box="[1010,1068,1618,1638]" gridcol="8" gridrow="1" pageId="6" pageNumber="95">Macho</td>
<td id="76BA020CFFCD6F818A67F9BDFBE1035B" box="[1080,1148,1618,1638]" gridcol="9" gridrow="1" pageId="6" pageNumber="95">Hembra</td>
<td id="76BA020CFFCD6F818ADBF9BDFB55035B" box="[1156,1224,1618,1638]" gridcol="10" gridrow="1" pageId="6" pageNumber="95">Hembra</td>
<td id="76BA020CFFCD6F818A8AF9BDFA92035B" box="[1237,1295,1618,1638]" gridcol="11" gridrow="1" pageId="6" pageNumber="95">Macho</td>
<td id="76BA020CFFCD6F818B44F9BDFAC2035B" box="[1307,1375,1618,1638]" gridcol="12" gridrow="1" pageId="6" pageNumber="95">Hembra</td>
</tr>
<tr id="356B6B70FFCD6F818F5EF99FFAC203B9" box="[257,1375,1648,1668]" gridrow="2" pageId="6" pageNumber="95">
<th id="76BA020CFFCD6F818F5EF99FFEC703B9" box="[257,346,1648,1668]" gridcol="0" gridrow="2" pageId="6" pageNumber="95">LRC</th>
<td id="76BA020CFFCD6F818F32F99FFE6503B9" box="[365,504,1648,1668]" gridcol="1" gridrow="2" pageId="6" pageNumber="95">33.9</td>
<td id="76BA020CFFCD6F818C55F99FFDF903B9" box="[522,612,1648,1668]" gridcol="2" gridrow="2" pageId="6" pageNumber="95">40.0</td>
<td id="76BA020CFFCD6F818C27F99FFD2F03B9" box="[632,690,1648,1668]" gridcol="3" gridrow="2" pageId="6" pageNumber="95">35.4</td>
<td id="76BA020CFFCD6F818C9BF99FFD6303B9" box="[708,766,1648,1668]" gridcol="4" gridrow="2" pageId="6" pageNumber="95">34.9</td>
<td id="76BA020CFFCD6F818D50F99FFCD403B9" box="[783,841,1648,1668]" gridcol="5" gridrow="2" pageId="6" pageNumber="95">33.5</td>
<td id="76BA020CFFCD6F818D04F99FFC0803B9" box="[859,917,1648,1668]" gridcol="6" gridrow="2" pageId="6" pageNumber="95">32.7</td>
<td id="76BA020CFFCD6F818DF8F99FFC7C03B9" box="[935,993,1648,1668]" gridcol="7" gridrow="2" pageId="6" pageNumber="95">35.9</td>
<td id="76BA020CFFCD6F818DADF99FFBB103B9" box="[1010,1068,1648,1668]" gridcol="8" gridrow="2" pageId="6" pageNumber="95">35.7</td>
<td id="76BA020CFFCD6F818A67F99FFBE103B9" box="[1080,1148,1648,1668]" gridcol="9" gridrow="2" pageId="6" pageNumber="95">37.3</td>
<td id="76BA020CFFCD6F818ADBF99FFB5503B9" box="[1156,1224,1648,1668]" gridcol="10" gridrow="2" pageId="6" pageNumber="95">38.0</td>
<td id="76BA020CFFCD6F818A8AF99FFA9203B9" box="[1237,1295,1648,1668]" gridcol="11" gridrow="2" pageId="6" pageNumber="95">36.9</td>
<td id="76BA020CFFCD6F818B44F99FFAC203B9" box="[1307,1375,1648,1668]" gridcol="12" gridrow="2" pageId="6" pageNumber="95">37.4</td>
</tr>
<tr id="356B6B70FFCD6F818F5EF960FAC2039E" box="[257,1375,1679,1699]" gridrow="3" pageId="6" pageNumber="95">
<th id="76BA020CFFCD6F818F5EF960FEC7039E" box="[257,346,1679,1699]" gridcol="0" gridrow="3" pageId="6" pageNumber="95">LC</th>
<td id="76BA020CFFCD6F818F32F960FE65039E" box="[365,504,1679,1699]" gridcol="1" gridrow="3" pageId="6" pageNumber="95">9.1</td>
<td id="76BA020CFFCD6F818C55F960FDF9039E" box="[522,612,1679,1699]" gridcol="2" gridrow="3" pageId="6" pageNumber="95">11.0</td>
<td id="76BA020CFFCD6F818C27F960FD2F039E" box="[632,690,1679,1699]" gridcol="3" gridrow="3" pageId="6" pageNumber="95">8.9</td>
<td id="76BA020CFFCD6F818C9BF960FD63039E" box="[708,766,1679,1699]" gridcol="4" gridrow="3" pageId="6" pageNumber="95">8.2</td>
<td id="76BA020CFFCD6F818D50F960FCD4039E" box="[783,841,1679,1699]" gridcol="5" gridrow="3" pageId="6" pageNumber="95">9.4</td>
<td id="76BA020CFFCD6F818D04F960FC08039E" box="[859,917,1679,1699]" gridcol="6" gridrow="3" pageId="6" pageNumber="95">8.9</td>
<td id="76BA020CFFCD6F818DF8F960FC7C039E" box="[935,993,1679,1699]" gridcol="7" gridrow="3" pageId="6" pageNumber="95">9.3</td>
<td id="76BA020CFFCD6F818DADF960FBB1039E" box="[1010,1068,1679,1699]" gridcol="8" gridrow="3" pageId="6" pageNumber="95">8.9</td>
<td id="76BA020CFFCD6F818A67F960FBE1039E" box="[1080,1148,1679,1699]" gridcol="9" gridrow="3" pageId="6" pageNumber="95">9.2</td>
<td id="76BA020CFFCD6F818ADBF960FB55039E" box="[1156,1224,1679,1699]" gridcol="10" gridrow="3" pageId="6" pageNumber="95">10.5</td>
<td id="76BA020CFFCD6F818A8AF960FA92039E" box="[1237,1295,1679,1699]" gridcol="11" gridrow="3" pageId="6" pageNumber="95">8.8</td>
<td id="76BA020CFFCD6F818B44F960FAC2039E" box="[1307,1375,1679,1699]" gridcol="12" gridrow="3" pageId="6" pageNumber="95">9.3</td>
</tr>
<tr id="356B6B70FFCD6F818F5EF942FAC203FC" box="[257,1375,1709,1729]" gridrow="4" pageId="6" pageNumber="95">
<th id="76BA020CFFCD6F818F5EF942FEC703FC" box="[257,346,1709,1729]" gridcol="0" gridrow="4" pageId="6" pageNumber="95">AC</th>
<td id="76BA020CFFCD6F818F32F942FE6503FC" box="[365,504,1709,1729]" gridcol="1" gridrow="4" pageId="6" pageNumber="95">10.8</td>
<td id="76BA020CFFCD6F818C55F942FDF903FC" box="[522,612,1709,1729]" gridcol="2" gridrow="4" pageId="6" pageNumber="95">12.1</td>
<td id="76BA020CFFCD6F818C27F942FD2F03FC" box="[632,690,1709,1729]" gridcol="3" gridrow="4" pageId="6" pageNumber="95">11.5</td>
<td id="76BA020CFFCD6F818C9BF942FD6303FC" box="[708,766,1709,1729]" gridcol="4" gridrow="4" pageId="6" pageNumber="95">11.6</td>
<td id="76BA020CFFCD6F818D50F942FCD403FC" box="[783,841,1709,1729]" gridcol="5" gridrow="4" pageId="6" pageNumber="95">11.2</td>
<td id="76BA020CFFCD6F818D04F942FC0803FC" box="[859,917,1709,1729]" gridcol="6" gridrow="4" pageId="6" pageNumber="95">11.0</td>
<td id="76BA020CFFCD6F818DF8F942FC7C03FC" box="[935,993,1709,1729]" gridcol="7" gridrow="4" pageId="6" pageNumber="95">11.5</td>
<td id="76BA020CFFCD6F818DADF942FBB103FC" box="[1010,1068,1709,1729]" gridcol="8" gridrow="4" pageId="6" pageNumber="95">11.6</td>
<td id="76BA020CFFCD6F818A67F942FBE103FC" box="[1080,1148,1709,1729]" gridcol="9" gridrow="4" pageId="6" pageNumber="95">12.0</td>
<td id="76BA020CFFCD6F818ADBF942FB5503FC" box="[1156,1224,1709,1729]" gridcol="10" gridrow="4" pageId="6" pageNumber="95">12.4</td>
<td id="76BA020CFFCD6F818A8AF942FA9203FC" box="[1237,1295,1709,1729]" gridcol="11" gridrow="4" pageId="6" pageNumber="95">12.0</td>
<td id="76BA020CFFCD6F818B44F942FAC203FC" box="[1307,1375,1709,1729]" gridcol="12" gridrow="4" pageId="6" pageNumber="95">11.8</td>
</tr>
<tr id="356B6B70FFCD6F818F5EF923FAC203DD" box="[257,1375,1740,1760]" gridrow="5" pageId="6" pageNumber="95">
<th id="76BA020CFFCD6F818F5EF923FEC703DD" box="[257,346,1740,1760]" gridcol="0" gridrow="5" pageId="6" pageNumber="95">DO</th>
<td id="76BA020CFFCD6F818F32F923FE6503DD" box="[365,504,1740,1760]" gridcol="1" gridrow="5" pageId="6" pageNumber="95">4.7</td>
<td id="76BA020CFFCD6F818C55F923FDF903DD" box="[522,612,1740,1760]" gridcol="2" gridrow="5" pageId="6" pageNumber="95">—</td>
<td id="76BA020CFFCD6F818C27F923FD2F03DD" box="[632,690,1740,1760]" gridcol="3" gridrow="5" pageId="6" pageNumber="95">4.3</td>
<td id="76BA020CFFCD6F818C9BF923FD6303DD" box="[708,766,1740,1760]" gridcol="4" gridrow="5" pageId="6" pageNumber="95">4.4</td>
<td id="76BA020CFFCD6F818D50F923FCD403DD" box="[783,841,1740,1760]" gridcol="5" gridrow="5" pageId="6" pageNumber="95">4.6</td>
<td id="76BA020CFFCD6F818D04F923FC0803DD" box="[859,917,1740,1760]" gridcol="6" gridrow="5" pageId="6" pageNumber="95">4.4</td>
<td id="76BA020CFFCD6F818DF8F923FC7C03DD" box="[935,993,1740,1760]" gridcol="7" gridrow="5" pageId="6" pageNumber="95">4.8</td>
<td id="76BA020CFFCD6F818DADF923FBB103DD" box="[1010,1068,1740,1760]" gridcol="8" gridrow="5" pageId="6" pageNumber="95">4.3</td>
<td id="76BA020CFFCD6F818A67F923FBE103DD" box="[1080,1148,1740,1760]" gridcol="9" gridrow="5" pageId="6" pageNumber="95">4.5</td>
<td id="76BA020CFFCD6F818ADBF923FB5503DD" box="[1156,1224,1740,1760]" gridcol="10" gridrow="5" pageId="6" pageNumber="95">4.5</td>
<td id="76BA020CFFCD6F818A8AF923FA9203DD" box="[1237,1295,1740,1760]" gridcol="11" gridrow="5" pageId="6" pageNumber="95">4.8</td>
<td id="76BA020CFFCD6F818B44F923FAC203DD" box="[1307,1375,1740,1760]" gridcol="12" gridrow="5" pageId="6" pageNumber="95">4.4</td>
</tr>
<tr id="356B6B70FFCD6F818F5EF904FAC203C2" box="[257,1375,1771,1791]" gridrow="6" pageId="6" pageNumber="95">
<th id="76BA020CFFCD6F818F5EF904FEC703C2" box="[257,346,1771,1791]" gridcol="0" gridrow="6" pageId="6" pageNumber="95">DON</th>
<td id="76BA020CFFCD6F818F32F904FE6503C2" box="[365,504,1771,1791]" gridcol="1" gridrow="6" pageId="6" pageNumber="95">2.9</td>
<td id="76BA020CFFCD6F818C55F904FDF903C2" box="[522,612,1771,1791]" gridcol="2" gridrow="6" pageId="6" pageNumber="95">3.5</td>
<td id="76BA020CFFCD6F818C27F904FD2F03C2" box="[632,690,1771,1791]" gridcol="3" gridrow="6" pageId="6" pageNumber="95">3.3</td>
<td id="76BA020CFFCD6F818C9BF904FD6303C2" box="[708,766,1771,1791]" gridcol="4" gridrow="6" pageId="6" pageNumber="95">3.0</td>
<td id="76BA020CFFCD6F818D50F904FCD403C2" box="[783,841,1771,1791]" gridcol="5" gridrow="6" pageId="6" pageNumber="95">3.0</td>
<td id="76BA020CFFCD6F818D04F904FC0803C2" box="[859,917,1771,1791]" gridcol="6" gridrow="6" pageId="6" pageNumber="95">3.2</td>
<td id="76BA020CFFCD6F818DF8F904FC7C03C2" box="[935,993,1771,1791]" gridcol="7" gridrow="6" pageId="6" pageNumber="95">3.1</td>
<td id="76BA020CFFCD6F818DADF904FBB103C2" box="[1010,1068,1771,1791]" gridcol="8" gridrow="6" pageId="6" pageNumber="95">3.1</td>
<td id="76BA020CFFCD6F818A67F904FBE103C2" box="[1080,1148,1771,1791]" gridcol="9" gridrow="6" pageId="6" pageNumber="95">3.6</td>
<td id="76BA020CFFCD6F818ADBF904FB5503C2" box="[1156,1224,1771,1791]" gridcol="10" gridrow="6" pageId="6" pageNumber="95">3.3</td>
<td id="76BA020CFFCD6F818A8AF904FA9203C2" box="[1237,1295,1771,1791]" gridcol="11" gridrow="6" pageId="6" pageNumber="95">3.0</td>
<td id="76BA020CFFCD6F818B44F904FAC203C2" box="[1307,1375,1771,1791]" gridcol="12" gridrow="6" pageId="6" pageNumber="95">3.6</td>
</tr>
<tr id="356B6B70FFCD6F818F5EF8E6FAC20220" box="[257,1375,1801,1821]" gridrow="7" pageId="6" pageNumber="95">
<th id="76BA020CFFCD6F818F5EF8E6FEC70220" box="[257,346,1801,1821]" gridcol="0" gridrow="7" pageId="6" pageNumber="95">APS</th>
<td id="76BA020CFFCD6F818F32F8E6FE650220" box="[365,504,1801,1821]" gridcol="1" gridrow="7" pageId="6" pageNumber="95">2.4</td>
<td id="76BA020CFFCD6F818C55F8E6FDF90220" box="[522,612,1801,1821]" gridcol="2" gridrow="7" pageId="6" pageNumber="95">2.4</td>
<td id="76BA020CFFCD6F818C27F8E6FD2F0220" box="[632,690,1801,1821]" gridcol="3" gridrow="7" pageId="6" pageNumber="95">2.2</td>
<td id="76BA020CFFCD6F818C9BF8E6FD630220" box="[708,766,1801,1821]" gridcol="4" gridrow="7" pageId="6" pageNumber="95">2.4</td>
<td id="76BA020CFFCD6F818D50F8E6FCD40220" box="[783,841,1801,1821]" gridcol="5" gridrow="7" pageId="6" pageNumber="95">2.6</td>
<td id="76BA020CFFCD6F818D04F8E6FC080220" box="[859,917,1801,1821]" gridcol="6" gridrow="7" pageId="6" pageNumber="95">2.7</td>
<td id="76BA020CFFCD6F818DF8F8E6FC7C0220" box="[935,993,1801,1821]" gridcol="7" gridrow="7" pageId="6" pageNumber="95">2.6</td>
<td id="76BA020CFFCD6F818DADF8E6FBB10220" box="[1010,1068,1801,1821]" gridcol="8" gridrow="7" pageId="6" pageNumber="95">2.1</td>
<td id="76BA020CFFCD6F818A67F8E6FBE10220" box="[1080,1148,1801,1821]" gridcol="9" gridrow="7" pageId="6" pageNumber="95">2.7</td>
<td id="76BA020CFFCD6F818ADBF8E6FB550220" box="[1156,1224,1801,1821]" gridcol="10" gridrow="7" pageId="6" pageNumber="95">2.4</td>
<td id="76BA020CFFCD6F818A8AF8E6FA920220" box="[1237,1295,1801,1821]" gridcol="11" gridrow="7" pageId="6" pageNumber="95">2.5</td>
<td id="76BA020CFFCD6F818B44F8E6FAC20220" box="[1307,1375,1801,1821]" gridcol="12" gridrow="7" pageId="6" pageNumber="95">2.5</td>
</tr>
<tr id="356B6B70FFCD6F818F5EF8C7FAC20201" box="[257,1375,1832,1852]" gridrow="8" pageId="6" pageNumber="95">
<th id="76BA020CFFCD6F818F5EF8C7FEC70201" box="[257,346,1832,1852]" gridcol="0" gridrow="8" pageId="6" pageNumber="95">DIO</th>
<td id="76BA020CFFCD6F818F32F8C7FE650201" box="[365,504,1832,1852]" gridcol="1" gridrow="8" pageId="6" pageNumber="95">3.4</td>
<td id="76BA020CFFCD6F818C55F8C7FDF90201" box="[522,612,1832,1852]" gridcol="2" gridrow="8" pageId="6" pageNumber="95">3.3</td>
<td id="76BA020CFFCD6F818C27F8C7FD2F0201" box="[632,690,1832,1852]" gridcol="3" gridrow="8" pageId="6" pageNumber="95">3.3</td>
<td id="76BA020CFFCD6F818C9BF8C7FD630201" box="[708,766,1832,1852]" gridcol="4" gridrow="8" pageId="6" pageNumber="95">3.3</td>
<td id="76BA020CFFCD6F818D50F8C7FCD40201" box="[783,841,1832,1852]" gridcol="5" gridrow="8" pageId="6" pageNumber="95">3.1</td>
<td id="76BA020CFFCD6F818D04F8C7FC080201" box="[859,917,1832,1852]" gridcol="6" gridrow="8" pageId="6" pageNumber="95">3.3</td>
<td id="76BA020CFFCD6F818DF8F8C7FC7C0201" box="[935,993,1832,1852]" gridcol="7" gridrow="8" pageId="6" pageNumber="95">3.3</td>
<td id="76BA020CFFCD6F818DADF8C7FBB10201" box="[1010,1068,1832,1852]" gridcol="8" gridrow="8" pageId="6" pageNumber="95">3.1</td>
<td id="76BA020CFFCD6F818A67F8C7FBE10201" box="[1080,1148,1832,1852]" gridcol="9" gridrow="8" pageId="6" pageNumber="95">3.5</td>
<td id="76BA020CFFCD6F818ADBF8C7FB550201" box="[1156,1224,1832,1852]" gridcol="10" gridrow="8" pageId="6" pageNumber="95">3.6</td>
<td id="76BA020CFFCD6F818A8AF8C7FA920201" box="[1237,1295,1832,1852]" gridcol="11" gridrow="8" pageId="6" pageNumber="95">3.3</td>
<td id="76BA020CFFCD6F818B44F8C7FAC20201" box="[1307,1375,1832,1852]" gridcol="12" gridrow="8" pageId="6" pageNumber="95">3.6</td>
</tr>
<tr id="356B6B70FFCD6F818F5EF8A9FAC20267" box="[257,1375,1862,1882]" gridrow="9" pageId="6" pageNumber="95">
<th id="76BA020CFFCD6F818F5EF8A9FEC70267" box="[257,346,1862,1882]" gridcol="0" gridrow="9" pageId="6" pageNumber="95">DTT</th>
<td id="76BA020CFFCD6F818F32F8A9FE650267" box="[365,504,1862,1882]" gridcol="1" gridrow="9" pageId="6" pageNumber="95">0.9</td>
<td id="76BA020CFFCD6F818C55F8A9FDF90267" box="[522,612,1862,1882]" gridcol="2" gridrow="9" pageId="6" pageNumber="95">—</td>
<td id="76BA020CFFCD6F818C27F8A9FD2F0267" box="[632,690,1862,1882]" gridcol="3" gridrow="9" pageId="6" pageNumber="95">0.9</td>
<td id="76BA020CFFCD6F818C9BF8A9FD630267" box="[708,766,1862,1882]" gridcol="4" gridrow="9" pageId="6" pageNumber="95">0.7</td>
<td id="76BA020CFFCD6F818D50F8A9FCD40267" box="[783,841,1862,1882]" gridcol="5" gridrow="9" pageId="6" pageNumber="95">0.8</td>
<td id="76BA020CFFCD6F818D04F8A9FC080267" box="[859,917,1862,1882]" gridcol="6" gridrow="9" pageId="6" pageNumber="95">0.8</td>
<td id="76BA020CFFCD6F818DF8F8A9FC7C0267" box="[935,993,1862,1882]" gridcol="7" gridrow="9" pageId="6" pageNumber="95">1.0</td>
<td id="76BA020CFFCD6F818DADF8A9FBB10267" box="[1010,1068,1862,1882]" gridcol="8" gridrow="9" pageId="6" pageNumber="95">0.9</td>
<td id="76BA020CFFCD6F818A67F8A9FBE10267" box="[1080,1148,1862,1882]" gridcol="9" gridrow="9" pageId="6" pageNumber="95">0.9</td>
<td id="76BA020CFFCD6F818ADBF8A9FB550267" box="[1156,1224,1862,1882]" gridcol="10" gridrow="9" pageId="6" pageNumber="95">0.9</td>
<td id="76BA020CFFCD6F818A8AF8A9FA920267" box="[1237,1295,1862,1882]" gridcol="11" gridrow="9" pageId="6" pageNumber="95">0.9</td>
<td id="76BA020CFFCD6F818B44F8A9FAC20267" box="[1307,1375,1862,1882]" gridcol="12" gridrow="9" pageId="6" pageNumber="95">1.0</td>
</tr>
<tr id="356B6B70FFCD6F818F5EF88AFAC20244" box="[257,1375,1893,1913]" gridrow="10" pageId="6" pageNumber="95">
<th id="76BA020CFFCD6F818F5EF88AFEC70244" box="[257,346,1893,1913]" gridcol="0" gridrow="10" pageId="6" pageNumber="95">Ddd III</th>
<td id="76BA020CFFCD6F818F32F88AFE650244" box="[365,504,1893,1913]" gridcol="1" gridrow="10" pageId="6" pageNumber="95">—</td>
<td id="76BA020CFFCD6F818C55F88AFDF90244" box="[522,612,1893,1913]" gridcol="2" gridrow="10" pageId="6" pageNumber="95">—</td>
<td id="76BA020CFFCD6F818C27F88AFD2F0244" box="[632,690,1893,1913]" gridcol="3" gridrow="10" pageId="6" pageNumber="95">2.9</td>
<td id="76BA020CFFCD6F818C9BF88AFD630244" box="[708,766,1893,1913]" gridcol="4" gridrow="10" pageId="6" pageNumber="95">2.9</td>
<td id="76BA020CFFCD6F818D50F88AFCD40244" box="[783,841,1893,1913]" gridcol="5" gridrow="10" pageId="6" pageNumber="95">2.8</td>
<td id="76BA020CFFCD6F818D04F88AFC080244" box="[859,917,1893,1913]" gridcol="6" gridrow="10" pageId="6" pageNumber="95">2.8</td>
<td id="76BA020CFFCD6F818DF8F88AFC7C0244" box="[935,993,1893,1913]" gridcol="7" gridrow="10" pageId="6" pageNumber="95">2.8</td>
<td id="76BA020CFFCD6F818DADF88AFBB10244" box="[1010,1068,1893,1913]" gridcol="8" gridrow="10" pageId="6" pageNumber="95">2.7</td>
<td id="76BA020CFFCD6F818A67F88AFBE10244" box="[1080,1148,1893,1913]" gridcol="9" gridrow="10" pageId="6" pageNumber="95">2.3</td>
<td id="76BA020CFFCD6F818ADBF88AFB550244" box="[1156,1224,1893,1913]" gridcol="10" gridrow="10" pageId="6" pageNumber="95">2.3</td>
<td id="76BA020CFFCD6F818A8AF88AFA920244" box="[1237,1295,1893,1913]" gridcol="11" gridrow="10" pageId="6" pageNumber="95">2.7</td>
<td id="76BA020CFFCD6F818B44F88AFAC20244" box="[1307,1375,1893,1913]" gridcol="12" gridrow="10" pageId="6" pageNumber="95">2.8</td>
</tr>
<tr id="356B6B70FFCD6F818F5EF86BFAC202A5" box="[257,1375,1924,1944]" gridrow="11" pageId="6" pageNumber="95">
<th id="76BA020CFFCD6F818F5EF86BFEC702A5" box="[257,346,1924,1944]" gridcol="0" gridrow="11" pageId="6" pageNumber="95">LT</th>
<td id="76BA020CFFCD6F818F32F86BFE6502A5" box="[365,504,1924,1944]" gridcol="1" gridrow="11" pageId="6" pageNumber="95">19.6</td>
<td id="76BA020CFFCD6F818C55F86BFDF902A5" box="[522,612,1924,1944]" gridcol="2" gridrow="11" pageId="6" pageNumber="95">23.0</td>
<td id="76BA020CFFCD6F818C27F86BFD2F02A5" box="[632,690,1924,1944]" gridcol="3" gridrow="11" pageId="6" pageNumber="95">21.1</td>
<td id="76BA020CFFCD6F818C9BF86BFD6302A5" box="[708,766,1924,1944]" gridcol="4" gridrow="11" pageId="6" pageNumber="95">20.7</td>
<td id="76BA020CFFCD6F818D50F86BFCD402A5" box="[783,841,1924,1944]" gridcol="5" gridrow="11" pageId="6" pageNumber="95">20.5</td>
<td id="76BA020CFFCD6F818D04F86BFC0802A5" box="[859,917,1924,1944]" gridcol="6" gridrow="11" pageId="6" pageNumber="95">19.5</td>
<td id="76BA020CFFCD6F818DF8F86BFC7C02A5" box="[935,993,1924,1944]" gridcol="7" gridrow="11" pageId="6" pageNumber="95">20.6</td>
<td id="76BA020CFFCD6F818DADF86BFBB102A5" box="[1010,1068,1924,1944]" gridcol="8" gridrow="11" pageId="6" pageNumber="95">21.0</td>
<td id="76BA020CFFCD6F818A67F86BFBE102A5" box="[1080,1148,1924,1944]" gridcol="9" gridrow="11" pageId="6" pageNumber="95">23.5</td>
<td id="76BA020CFFCD6F818ADBF86BFB5502A5" box="[1156,1224,1924,1944]" gridcol="10" gridrow="11" pageId="6" pageNumber="95">23.0</td>
<td id="76BA020CFFCD6F818A8AF86BFA9202A5" box="[1237,1295,1924,1944]" gridcol="11" gridrow="11" pageId="6" pageNumber="95">22.0</td>
<td id="76BA020CFFCD6F818B44F86BFAC202A5" box="[1307,1375,1924,1944]" gridcol="12" gridrow="11" pageId="6" pageNumber="95">22.0</td>
</tr>
<tr id="356B6B70FFCD6F818F5EF84DFAC2028B" box="[257,1375,1954,1974]" gridrow="12" pageId="6" pageNumber="95">
<th id="76BA020CFFCD6F818F5EF84DFEC7028B" box="[257,346,1954,1974]" gridcol="0" gridrow="12" pageId="6" pageNumber="95">LM</th>
<td id="76BA020CFFCD6F818F32F84DFE65028B" box="[365,504,1954,1974]" gridcol="1" gridrow="12" pageId="6" pageNumber="95">12.8</td>
<td id="76BA020CFFCD6F818C55F84DFDF9028B" box="[522,612,1954,1974]" gridcol="2" gridrow="12" pageId="6" pageNumber="95">—</td>
<td id="76BA020CFFCD6F818C27F84DFD2F028B" box="[632,690,1954,1974]" gridcol="3" gridrow="12" pageId="6" pageNumber="95">12.8</td>
<td id="76BA020CFFCD6F818C9BF84DFD63028B" box="[708,766,1954,1974]" gridcol="4" gridrow="12" pageId="6" pageNumber="95">—</td>
<td id="76BA020CFFCD6F818D50F84DFCD4028B" box="[783,841,1954,1974]" gridcol="5" gridrow="12" pageId="6" pageNumber="95">12.0</td>
<td id="76BA020CFFCD6F818D04F84DFC08028B" box="[859,917,1954,1974]" gridcol="6" gridrow="12" pageId="6" pageNumber="95">12.1</td>
<td id="76BA020CFFCD6F818DF8F84DFC7C028B" box="[935,993,1954,1974]" gridcol="7" gridrow="12" pageId="6" pageNumber="95">13.0</td>
<td id="76BA020CFFCD6F818DADF84DFBB1028B" box="[1010,1068,1954,1974]" gridcol="8" gridrow="12" pageId="6" pageNumber="95">12.3</td>
<td id="76BA020CFFCD6F818A67F84DFBE1028B" box="[1080,1148,1954,1974]" gridcol="9" gridrow="12" pageId="6" pageNumber="95">12.8</td>
<td id="76BA020CFFCD6F818ADBF84DFB55028B" box="[1156,1224,1954,1974]" gridcol="10" gridrow="12" pageId="6" pageNumber="95">12.8</td>
<td id="76BA020CFFCD6F818A8AF84DFA92028B" box="[1237,1295,1954,1974]" gridcol="11" gridrow="12" pageId="6" pageNumber="95">13.0</td>
<td id="76BA020CFFCD6F818B44F84DFAC2028B" box="[1307,1375,1954,1974]" gridcol="12" gridrow="12" pageId="6" pageNumber="95">10.7</td>
</tr>
<tr id="356B6B70FFCD6F818F5EF82EFAC202E8" box="[257,1375,1985,2005]" gridrow="13" pageId="6" pageNumber="95">
<th id="76BA020CFFCD6F818F5EF82EFEC702E8" box="[257,346,1985,2005]" gridcol="0" gridrow="13" pageId="6" pageNumber="95">LP</th>
<td id="76BA020CFFCD6F818F32F82EFE6502E8" box="[365,504,1985,2005]" gridcol="1" gridrow="13" pageId="6" pageNumber="95">15.9</td>
<td id="76BA020CFFCD6F818C55F82EFDF902E8" box="[522,612,1985,2005]" gridcol="2" gridrow="13" pageId="6" pageNumber="95">17.5</td>
<td id="76BA020CFFCD6F818C27F82EFD2F02E8" box="[632,690,1985,2005]" gridcol="3" gridrow="13" pageId="6" pageNumber="95">17.1</td>
<td id="76BA020CFFCD6F818C9BF82EFD6302E8" box="[708,766,1985,2005]" gridcol="4" gridrow="13" pageId="6" pageNumber="95">—</td>
<td id="76BA020CFFCD6F818D50F82EFCD402E8" box="[783,841,1985,2005]" gridcol="5" gridrow="13" pageId="6" pageNumber="95">16.5</td>
<td id="76BA020CFFCD6F818D04F82EFC0802E8" box="[859,917,1985,2005]" gridcol="6" gridrow="13" pageId="6" pageNumber="95">15.6</td>
<td id="76BA020CFFCD6F818DF8F82EFC7C02E8" box="[935,993,1985,2005]" gridcol="7" gridrow="13" pageId="6" pageNumber="95">17.3</td>
<td id="76BA020CFFCD6F818DADF82EFBB102E8" box="[1010,1068,1985,2005]" gridcol="8" gridrow="13" pageId="6" pageNumber="95">17.4</td>
<td id="76BA020CFFCD6F818A67F82EFBE102E8" box="[1080,1148,1985,2005]" gridcol="9" gridrow="13" pageId="6" pageNumber="95">17.5</td>
<td id="76BA020CFFCD6F818ADBF82EFB5502E8" box="[1156,1224,1985,2005]" gridcol="10" gridrow="13" pageId="6" pageNumber="95">17.2</td>
<td id="76BA020CFFCD6F818A8AF82EFA9202E8" box="[1237,1295,1985,2005]" gridcol="11" gridrow="13" pageId="6" pageNumber="95">17.4</td>
<td id="76BA020CFFCD6F818B44F82EFAC202E8" box="[1307,1375,1985,2005]" gridcol="12" gridrow="13" pageId="6" pageNumber="95">18.1</td>
</tr>
</table>
</paragraph>
<paragraph id="8BE46932FFCC90798FBAFF02FBB5043C" blockId="7.[485,1064,237,257]" box="[485,1064,237,257]" pageId="7" pageNumber="96">
RADA, M. &amp; GUAYASAMIN, J.M.: 
<taxonomicName id="4C5B12B1FFCC90798CAEFF02FBB5043C" authority="REDES C RI P TION" box="[753,1064,237,257]" class="Amphibia" family="Centrolenidae" genus="Cochranella" kingdom="Animalia" order="Anura" pageId="7" pageNumber="96" phylum="Chordata" rank="species" species="megista">
<emphasis id="B92FB520FFCC90798CAEFF02FC3D05C2" box="[753,928,237,257]" italics="true" pageId="7" pageNumber="96">
<smallCapsWord id="8D02FFEEFFCC90798CAEFF02FCC105C2" baselines="252,251" box="[753,860,237,257]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Cochranella" pageId="7" pageNumber="96">COCHRANELLA</smallCapsWord>
<smallCapsWord id="8D02FFEEFFCC90798D3DFF1EFC3D05C2" baselines="251" box="[866,928,241,255]" lowerCaseFontSize="6" mainFontSize="8" normCase="lower" normString="megista" pageId="7" pageNumber="96">MEGISTA</smallCapsWord>
</emphasis>
REDES C RI P TION
</taxonomicName>
</paragraph>
<caption id="DF2439BAFFCC90798E8FFCB5FB280653" ID-DOI="http://doi.org/10.5281/zenodo.13314528" ID-Zenodo-Dep="13314528" box="[208,1205,858,878]" httpUri="https://zenodo.org/record/13314528/files/figure.png" pageId="7" pageNumber="96" startId="7.[208,287,858,878]" targetBox="[208,1342,311,821]" targetPageId="7" targetType="figure">
<paragraph id="8BE46932FFCC90798E8FFCB5FB280653" blockId="7.[208,1205,858,878]" box="[208,1205,858,878]" pageId="7" pageNumber="96">
<emphasis id="B92FB520FFCC90798E8FFCB5FEA90653" bold="true" box="[208,308,858,878]" pageId="7" pageNumber="96">FIGURA 6:</emphasis>
Fotografía de 
<emphasis id="B92FB520FFCC90798FF3FCB5FDD20653" box="[428,591,858,878]" italics="true" pageId="7" pageNumber="96">
<taxonomicName id="4C5B12B1FFCC90798FF3FCB5FDD10653" box="[428,588,858,878]" class="Amphibia" family="Centrolenidae" genus="Cochranella" kingdom="Animalia" order="Anura" pageId="7" pageNumber="96" phylum="Chordata" rank="species" species="megista">Cochranella megista</taxonomicName>
,
</emphasis>
macho adulto, ICN 27766. (A, B = Aspectos dorsal y lateral de la cabeza).
</paragraph>
</caption>
<paragraph id="8BE46932FFCC90798E8FFC2AFE1F0043" blockId="7.[208,756,965,1406]" pageId="7" pageNumber="96">
características morfológicas son escasamente invariables, sin embargo, y tal vez como producto del estado de preservación en el que se encuentra el 
<typeStatus id="54E0D790FFCC90798C27FBEAFD5A0123" box="[632,711,1029,1054]" pageId="7" pageNumber="96" type="neotype">neotipo</typeStatus>
con respecto a material adicional referido de la especie depositado en el ICN, se registra una débil diferencia en dos características que a continuación se mencionan: 1) en vista lateral el rostro de los ejemplares referidos de 
<taxonomicName id="4C5B12B1FFCC90798EAEFB49FEC50180" box="[241,344,1190,1213]" class="Amphibia" family="Centrolenidae" genus="Cochranella" kingdom="Animalia" order="Anura" pageId="7" pageNumber="96" phylum="Chordata" rank="species" species="megista">
<emphasis id="B92FB520FFCC90798EAEFB49FEC50180" box="[241,344,1190,1213]" italics="true" pageId="7" pageNumber="96">C. megista</emphasis>
</taxonomicName>
presentan un contorno redondeado en comparación con el 
<typeStatus id="54E0D790FFCC90798FFBFB2AFE6E01E3" box="[420,499,1221,1246]" pageId="7" pageNumber="96" type="neotype">neotipo</typeStatus>
(
<figureCitation id="136075B7FFCC90798C5CFB2AFDCE01E3" box="[515,595,1221,1246]" captionStart="FIGURA 4" captionStartId="4.[245,324,953,973]" captionTargetBox="[245,1379,311,915]" captionTargetId="figure-185@4.[245,1379,311,915]" captionTargetPageId="4" captionText="FIGURA 4: Fotografía de Cochranella megista, macho adulto (Neotipo), MHNCSJ 291(Actualmente MHUA 5851). A, B = Aspectos dorsal y lateral de la cabeza." figureDoi="http://doi.org/10.5281/zenodo.13314524" httpUri="https://zenodo.org/record/13314524/files/figure.png" pageId="7" pageNumber="96">Fig. 4A</figureCitation>
, 
<figureCitation id="136075B7FFCC90798C00FB2AFD1C01E3" box="[607,641,1221,1246]" captionStart="FIGURA 6" captionStartId="7.[208,287,858,878]" captionTargetBox="[208,1342,311,821]" captionTargetId="figure-275@7.[208,1342,311,821]" captionTargetPageId="7" captionText="FIGURA 6: Fotografía de Cochranella megista, macho adulto, ICN 27766. (A, B = Aspectos dorsal y lateral de la cabeza)." figureDoi="http://doi.org/10.5281/zenodo.13314528" httpUri="https://zenodo.org/record/13314528/files/figure.png" pageId="7" pageNumber="96">6A</figureCitation>
); 2) la presencia de abundantes espículas cónicas de distribución uniforme en los machos en estado reproductivo (
<figureCitation id="136075B7FFCC90798E88FACAFE8E0003" box="[215,275,1317,1342]" captionStart="FIGURA 4" captionStartId="4.[245,324,953,973]" captionTargetBox="[245,1379,311,915]" captionTargetId="figure-185@4.[245,1379,311,915]" captionTargetPageId="4" captionText="FIGURA 4: Fotografía de Cochranella megista, macho adulto (Neotipo), MHNCSJ 291(Actualmente MHUA 5851). A, B = Aspectos dorsal y lateral de la cabeza." figureDoi="http://doi.org/10.5281/zenodo.13314524" httpUri="https://zenodo.org/record/13314524/files/figure.png" pageId="7" pageNumber="96">Fig. 4</figureCitation>
A-B; 6A-B; 7-8), en contraposición en el 
<typeStatus id="54E0D790FFCC90798C9AFAC9FF670063" pageId="7" pageNumber="96" type="neotype">neotipo</typeStatus>
en donde el dorso exhibe una menor cantidad de espículas cónicas.
</paragraph>
<paragraph id="8BE46932FFCC90798E8FFA2BFD1100E0" blockId="7.[208,652,1476,1501]" box="[208,652,1476,1501]" pageId="7" pageNumber="96">
<heading id="D0ACDE5EFFCC90798E8FFA2BFD1100E0" bold="true" box="[208,652,1476,1501]" centered="true" fontSize="9" level="3" pageId="7" pageNumber="96" reason="2">
<emphasis id="B92FB520FFCC90798E8FFA2BFD1100E0" bold="true" box="[208,652,1476,1501]" pageId="7" pageNumber="96">
Coloración en vida de 
<taxonomicName id="4C5B12B1FFCC90798FE3FA2AFD1100E0" box="[444,652,1477,1501]" class="Amphibia" family="Centrolenidae" genus="Cochranella" kingdom="Animalia" order="Anura" pageId="7" pageNumber="96" phylum="Chordata" rank="species" species="megista">
<emphasis id="B92FB520FFCC90798FE3FA2AFD1100E0" bold="true" box="[444,652,1477,1501]" italics="true" pageId="7" pageNumber="96">Cochranella megista</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE46932FFCC90798F57F9EAFE060383" blockId="7.[208,756,1541,1726]" pageId="7" pageNumber="96">(Basadas en el ejemplar ICN 28796, hembra adulta, notas de John D. Lynch, junio 28 de 1991). Dorso verde con puntos negros, iris gris pálido con retículo grisáceo; pericardio y mitad anterior del peritoneo parietal color blanco, peritoneo visceral translúcido; huesos verdes.</paragraph>
<paragraph id="8BE46932FFCC90798E8FF8EBFECB0221" blockId="7.[208,342,1796,1820]" box="[208,342,1796,1820]" pageId="7" pageNumber="96">
<heading id="D0ACDE5EFFCC90798E8FF8EBFECB0221" box="[208,342,1796,1820]" fontSize="10" level="3" pageId="7" pageNumber="96" reason="8">
<emphasis id="B92FB520FFCC90798E8FF8EBFECB0221" bold="true" box="[208,342,1796,1820]" pageId="7" pageNumber="96">Distribución</emphasis>
</heading>
</paragraph>
<paragraph id="8BE46932FFCC90798F57F8AAFD6902E3" blockId="7.[208,756,1861,2014]" pageId="7" pageNumber="96">
<taxonomicName id="4C5B12B1FFCC90798F57F8AAFE520260" box="[264,463,1861,1885]" class="Amphibia" family="Centrolenidae" genus="Cochranella" kingdom="Animalia" order="Anura" pageId="7" pageNumber="96" phylum="Chordata" rank="species" species="megista">
<emphasis id="B92FB520FFCC90798F57F8AAFE520260" box="[264,463,1861,1885]" italics="true" pageId="7" pageNumber="96">Cochranella megista</emphasis>
</taxonomicName>
es conocida para cuatro localidades en la Cordillera Occidental de 
<collectingCountry id="F34C29A2FFCC90798CD3F88AFD6E0243" box="[652,755,1893,1918]" name="Colombia" pageId="7" pageNumber="96">Colombia</collectingCountry>
con límite norte en las inmediaciones del municipio de Urrao, 
<collectingRegion id="499FA7D0FFCC90798F66F84AFDC70283" box="[313,602,1957,1982]" country="Colombia" name="Antioquia" pageId="7" pageNumber="96">departamento de Antioquia</collectingRegion>
, 
<quantity id="4CA3C4D7FFCC90798C3AF84AFD690280" box="[613,756,1957,1982]" metricMagnitude="3" metricUnit="m" metricValue="1.75" metricValueMax="1.8" metricValueMin="1.7" pageId="7" pageNumber="96" unit="m" value="1750.0" valueMax="1800.0" valueMin="1700.0">1700-1800 m</quantity>
y límite sur en el departamento de 
<collectingRegion id="499FA7D0FFCC90798C16F82AFD7202E3" box="[585,751,1989,2014]" country="Colombia" name="Valle del Cauca" pageId="7" pageNumber="96">Valle del Cauca</collectingRegion>
,
</paragraph>
<paragraph id="8BE46932FFCC90798D46FC2AFB9D0183" blockId="7.[793,1342,965,1214]" pageId="7" pageNumber="96">
cercanía con el 
<collectingRegion id="499FA7D0FFCC90798D9AFC2AFB9006E3" box="[965,1037,965,990]" country="Colombia" name="Choco" pageId="7" pageNumber="96">Chocó</collectingRegion>
, km 20.2 al NW por la carretera Los Galápagos-La Carbonera, 
<quantity id="4CA3C4D7FFCC90798AF9FC0AFB6706C0" box="[1190,1274,997,1022]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="7" pageNumber="96" unit="m" value="2100.0">2100 m</quantity>
. También ha sido coleccionada en el municipio de Pueblo Rico, km 7 por la carretera La Selva-La Repetidora, 
<quantity id="4CA3C4D7FFCC90798D46FBAAFC350160" box="[793,936,1093,1118]" metricMagnitude="3" metricUnit="m" metricValue="1.85" metricValueMax="2.0" metricValueMin="1.7" pageId="7" pageNumber="96" unit="m" value="1850.0" valueMax="2000.0" valueMin="1700.0">1700-2000 m</quantity>
y en el 
<collectingRegion id="499FA7D0FFCC90798DA0FBAAFA9B0163" box="[1023,1286,1093,1118]" country="Colombia" name="Choco" pageId="7" pageNumber="96">departamento del Chocó</collectingRegion>
, municipio de Carmen de Atrato, km 18 por la carretera que conduce de Carmen de Atrato a la vereda Guaduas, 
<quantity id="4CA3C4D7FFCC90798D0AFB4AFC3B0180" box="[853,934,1189,1214]" metricMagnitude="3" metricUnit="m" metricValue="1.88" pageId="7" pageNumber="96" unit="m" value="1880.0">1880 m</quantity>
(
<figureCitation id="136075B7FFCC90798DEBFB4AFC6F0183" box="[948,1010,1189,1214]" captionStart="FIGURA 9" captionStartId="9.[208,287,1967,1987]" captionTargetBox="[208,1342,519,1927]" captionTargetId="figure-46@9.[208,1342,514,1931]" captionTargetPageId="9" captionText="FIGURA 9: Mapa de Colombia mostrando las localidades conocidas (círculo) de Cochranella megista (Rivero, 1985). La localidad tipo es indicada como un triángulo." figureDoi="http://doi.org/10.5281/zenodo.13314536" httpUri="https://zenodo.org/record/13314536/files/figure.png" pageId="7" pageNumber="96">Fig. 9</figureCitation>
).
</paragraph>
<caption id="DF2439BAFFCC90798D46F840FB0A02E0" ID-DOI="http://doi.org/10.5281/zenodo.13314530" ID-Zenodo-Dep="13314530" httpUri="https://zenodo.org/record/13314530/files/figure.png" pageId="7" pageNumber="96" startId="7.[793,872,1967,1987]" targetBox="[793,1341,1251,1931]" targetPageId="7" targetType="figure">
<paragraph id="8BE46932FFCC90798D46F840FB0A02E0" blockId="7.[793,1342,1967,2013]" pageId="7" pageNumber="96">
<emphasis id="B92FB520FFCC90798D46F840FCE302FE" bold="true" box="[793,894,1967,1987]" pageId="7" pageNumber="96">FIGURA 7:</emphasis>
Fotografía de 
<emphasis id="B92FB520FFCC90798DABF840FB0502FE" box="[1012,1176,1967,1987]" italics="true" pageId="7" pageNumber="96">
<taxonomicName id="4C5B12B1FFCC90798DABF840FB0902FE" box="[1012,1172,1967,1987]" class="Amphibia" family="Centrolenidae" genus="Cochranella" kingdom="Animalia" order="Anura" pageId="7" pageNumber="96" phylum="Chordata" rank="species" species="megista">Cochranella megista</taxonomicName>
,
</emphasis>
macho adulto, ICN 27766. Aspecto ventral de A = Mano; B = Pie.
</paragraph>
</caption>
<paragraph id="8BE46932FFC390768C2CFF02FC7B043C" blockId="8.[627,998,237,257]" box="[627,998,237,257]" pageId="8" pageNumber="97">PA P ÉIS AVULSOS DE ZOOLOGIA, 48(12), 2008</paragraph>
<paragraph id="8BE46932FFC390768B11FF02FAFE043C" blockId="8.[1358,1379,237,257]" box="[1358,1379,237,257]" pageId="8" pageNumber="97">97</paragraph>
<caption id="DF2439BAFFC390768EAAFC49FA920687" ID-DOI="http://doi.org/10.5281/zenodo.13314534" ID-Zenodo-Dep="13314534" box="[245,1295,934,954]" httpUri="https://zenodo.org/record/13314534/files/figure.png" pageId="8" pageNumber="97" startId="8.[245,324,934,954]" targetBox="[245,1379,311,897]" targetPageId="8" targetType="figure">
<paragraph id="8BE46932FFC390768EAAFC49FA920687" blockId="8.[245,1295,934,954]" box="[245,1295,934,954]" pageId="8" pageNumber="97">
<emphasis id="B92FB520FFC390768EAAFC49FEC60687" bold="true" box="[245,347,934,954]" pageId="8" pageNumber="97">FIGURA 8:</emphasis>
Fotografía en perfil dorsal (A) y ventral (B) de 
<emphasis id="B92FB520FFC390768C81FC49FC140687" box="[734,905,934,954]" italics="true" pageId="8" pageNumber="97">
<taxonomicName id="4C5B12B1FFC390768C81FC49FC190687" authorityName="Acosta-Galvis" authorityYear="2000" box="[734,900,934,954]" class="Amphibia" family="Centrolenidae" genus="Cochranella" kingdom="Animalia" order="Anura" pageId="8" pageNumber="97" phylum="Chordata" rank="species" species="megistra">Cochranella megistra</taxonomicName>
,
</emphasis>
macho adulto, LRC = 35.68 mm, ICN 27768.
</paragraph>
</caption>
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<emphasis id="B92FB520FFC390768EAAFC0AFDF606C0" bold="true" box="[245,619,996,1021]" pageId="8" pageNumber="97">Aspectos ecológicos y reproductivos</emphasis>
</heading>
</paragraph>
<paragraph id="8BE46932FFC390768F71FBCAFC880043" blockId="8.[245,794,1061,1406]" pageId="8" pageNumber="97">
Gran parte de los adultos de 
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<emphasis id="B92FB520FFC390768C01FBCAFE950160" italics="true" pageId="8" pageNumber="97">Cochranella megista</emphasis>
</taxonomicName>
fueron encontrados en la vegetación asociada a los arroyos en donde los machos vocalizaban cerca de las posturas (ramas cercanas), principalmente en hojas de “Araceas” (
<quantity id="4CA3C4D7FFC390768F3CFB4AFE750183" box="[355,488,1189,1214]" metricMagnitude="0" metricUnit="m" metricValue="1.65" metricValueMax="1.8" metricValueMin="1.5" pageId="8" pageNumber="97" unit="m" value="1.65" valueMax="1.8" valueMin="1.5">1.50-1.80 m</quantity>
). Sin embargo, algunos ejemplares de la especie (ICN 17242-4, notas de campo de Maria 
<taxonomicName id="4C5B12B1FFC390768F07FB0AFDD701C3" authority="Ardila-Robayo" authorityName="Ardila-Robayo" box="[344,586,1253,1278]" class="Arachnida" family="Phalangiidae" genus="Cristina" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="97" phylum="Arthropoda" rank="genus">Cristina Ardila-Robayo</taxonomicName>
, agosto 29 de 1987) se hallaron lejos de los cuerpos de agua, internados en el bosque maduro (
<quantity id="4CA3C4D7FFC390768F82FACAFDAC0000" box="[477,561,1317,1342]" metricMagnitude="0" metricUnit="m" metricValue="1.25" metricValueMax="2.0" metricValueMin="0.5" pageId="8" pageNumber="97" unit="m" value="1.25" valueMax="2.0" valueMin="0.5">0.5-2 m</quantity>
sobre el suelo), lo que sugiere una posible migración de los adultos durante la época no reproductiva hacia el interior del bosque.
</paragraph>
<paragraph id="8BE46932FFC390768EAAFA2AFD7800E0" blockId="8.[245,741,1477,1533]" box="[245,741,1477,1501]" pageId="8" pageNumber="97">
<heading id="D0ACDE5EFFC390768EAAFA2AFD7800E0" bold="true" box="[245,741,1477,1501]" centered="true" fontSize="9" level="3" pageId="8" pageNumber="97" reason="2">
<emphasis id="B92FB520FFC390768EAAFA2AFD7800E0" bold="true" box="[245,741,1477,1501]" pageId="8" pageNumber="97">
Ejemplares adicionales referidos de 
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<emphasis id="B92FB520FFC390768C36FA2AFD7800E0" bold="true" box="[617,741,1477,1501]" italics="true" pageId="8" pageNumber="97">Cochranella</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE46932FFC390768EAAFA0AFEDF00C0" blockId="8.[245,741,1477,1533]" box="[245,322,1509,1533]" pageId="8" pageNumber="97">
<emphasis id="B92FB520FFC390768EAAFA0AFEDF00C0" bold="true" box="[245,322,1509,1533]" italics="true" pageId="8" pageNumber="97">megista</emphasis>
</paragraph>
<paragraph id="8BE46932FFC390768F71F9C9FB990183" blockId="8.[245,794,1573,2014]" lastBlockId="8.[831,1379,997,1214]" pageId="8" pageNumber="97">
<collectingRegion id="499FA7D0FFC390768F71F9C9FD170303" box="[302,650,1573,1598]" country="Colombia" name="Antioquia" pageId="8" pageNumber="97">Departamento de Antioquia</collectingRegion>
, municipio de Urrao, Parque Nacional Natural “Las Orquídeas” (Polo). 
<quantity id="4CA3C4D7FFC390768FD4F98AFD870340" box="[395,538,1637,1662]" metricMagnitude="3" metricUnit="m" metricValue="1.75" metricValueMax="1.8" metricValueMin="1.7" pageId="8" pageNumber="97" unit="m" value="1750.0" valueMax="1800.0" valueMin="1700.0">1700-1800 m</quantity>
(
<emphasis id="B92FB520FFC390768C6EF989FDDB0340" box="[561,582,1638,1661]" italics="true" pageId="8" pageNumber="97">ca</emphasis>
<geoCoordinate id="EE6F0FF5FFC390768C0CF98AFC9E0340" box="[595,771,1637,1662]" degrees="6" direction="north" minutes="18" orientation="latitude" pageId="8" pageNumber="97" precision="1" seconds="36.0" value="6.3100004">6°18’36.0 Norte</geoCoordinate>
/ 
<geoCoordinate id="EE6F0FF5FFC390768EAAF96AFE3203A3" box="[245,431,1669,1694]" degrees="76" direction="west" minutes="7" orientation="longitude" pageId="8" pageNumber="97" precision="1" seconds="48.8" value="-76.130226">76°7’48.8” Oeste</geoCoordinate>
), macho MHNCSJ 0291 (
<typeStatus id="54E0D790FFC390768C8AF96AFE8F0383" pageId="8" pageNumber="97" type="neotype">Neotipo</typeStatus>
), coleccionado con el 
<typeStatus id="54E0D790FFC390768FA3F94AFDCA0383" box="[508,599,1701,1726]" pageId="8" pageNumber="97" type="holotype">holotipo</typeStatus>
; Departamento de 
<collectingRegion id="499FA7D0FFC390768EAAF92AFE0103E3" box="[245,412,1733,1758]" country="Colombia" name="Valle del Cauca" pageId="8" pageNumber="97">Valle del Cauca</collectingRegion>
, Límite con el 
<collectingRegion id="499FA7D0FFC390768C1EF92AFD1403E3" box="[577,649,1733,1758]" country="Colombia" name="Choco" pageId="8" pageNumber="97">Chocó</collectingRegion>
, km 20.2 por la carretera Los Galápagos-La Carbonera, 
<quantity id="4CA3C4D7FFC390768CFBF90AFD6803C0" box="[676,757,1765,1790]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="8" pageNumber="97" unit="m" value="2100.0">2100 m</quantity>
(
<emphasis id="B92FB520FFC390768D5BF909FC8403C0" box="[772,793,1766,1789]" italics="true" pageId="8" pageNumber="97">ca</emphasis>
<geoCoordinate id="EE6F0FF5FFC390768EAAF8EAFE030220" box="[245,414,1797,1822]" degrees="4" direction="north" minutes="5" orientation="latitude" pageId="8" pageNumber="97" precision="1" seconds="22.5" value="4.0895834">4°5’22.5” Norte</geoCoordinate>
/ 
<geoCoordinate id="EE6F0FF5FFC390768FE7F8EAFDE00223" box="[440,637,1797,1822]" degrees="76" direction="west" minutes="32" orientation="longitude" pageId="8" pageNumber="97" precision="1" seconds="13.2" value="-76.537">76°32’13.2” Oeste</geoCoordinate>
), hembra ICN 28796 cols. Pedro M. Ruiz-Carranza &amp; John D. Lynch, junio 28 de 1991; 
<collectingRegion id="499FA7D0FFC390768FAFF8A9FC8B0263" box="[496,790,1861,1886]" country="Colombia" name="Risaralda" pageId="8" pageNumber="97">Departamento de Risaralda</collectingRegion>
, municipio de Pueblo Rico, km 7 por la carretera La Selva-La Repetidora, 
<quantity id="4CA3C4D7FFC390768FBFF86AFDF202A0" box="[480,623,1925,1950]" metricMagnitude="3" metricUnit="m" metricValue="1.85" metricValueMax="2.0" metricValueMin="1.7" pageId="8" pageNumber="97" unit="m" value="1850.0" valueMax="2000.0" valueMin="1700.0">1700-2000 m</quantity>
(
<emphasis id="B92FB520FFC390768CD9F869FD0602A0" box="[646,667,1926,1949]" italics="true" pageId="8" pageNumber="97">ca</emphasis>
<geoCoordinate id="EE6F0FF5FFC390768CF5F86AFEAF0280" degrees="5" direction="north" minutes="14" orientation="latitude" pageId="8" pageNumber="97" precision="1" seconds="16.8" value="5.238">5°14’16.8” Norte</geoCoordinate>
/ 
<geoCoordinate id="EE6F0FF5FFC390768F17F84AFE6D0283" box="[328,496,1957,1982]" degrees="76" direction="west" minutes="2" orientation="longitude" pageId="8" pageNumber="97" precision="1" seconds="9.6" value="-76.036">76°2’9.6” Oeste</geoCoordinate>
), machos ICN 27763-8 cols. Maria Cristina Ardila-R., abril 15 de 1991. Departamento de 
<collectingRegion id="499FA7D0FFC390768DF1FC0AFC6B06C3" box="[942,1014,997,1022]" country="Colombia" name="Choco" pageId="8" pageNumber="97">Chocó</collectingRegion>
, municipio de Carmen de Atrato, km 18 por la carretera que conduce de Carmen de Atrato a la vereda Guaduas, 
<quantity id="4CA3C4D7FFC390768A36FBCAFB270100" box="[1129,1210,1061,1086]" metricMagnitude="3" metricUnit="m" metricValue="1.88" pageId="8" pageNumber="97" unit="m" value="1880.0">1880 m</quantity>
(
<emphasis id="B92FB520FFC390768A95FBC9FB420100" box="[1226,1247,1062,1085]" italics="true" pageId="8" pageNumber="97">ca</emphasis>
<geoCoordinate id="EE6F0FF5FFC390768AB7FBCAFAFE0100" box="[1256,1379,1061,1086]" degrees="5" direction="north" minutes="47" orientation="latitude" pageId="8" pageNumber="97" precision="925" value="5.7833333">5 47’ Norte</geoCoordinate>
/ 
<geoCoordinate id="EE6F0FF5FFC390768D10FBAAFC450163" box="[847,984,1093,1118]" degrees="76" direction="west" minutes="20" orientation="longitude" pageId="8" pageNumber="97" precision="925" value="-76.333336">76 20’ Oeste</geoCoordinate>
), hembras ICN 17242-3, macho ICN 17244 cols. Maria Cristina Ardila-R., Pedro M. Ruiz-Carranza, Ricardo Sánchez &amp; Gustavo Susatama., agosto 29 de 1987.
</paragraph>
<paragraph id="8BE46932FFC390768A52FAEAFB080020" blockId="8.[1037,1173,1285,1309]" box="[1037,1173,1285,1309]" pageId="8" pageNumber="97">
<heading id="D0ACDE5EFFC390768A52FAEAFB080020" box="[1037,1173,1285,1309]" centered="true" fontSize="10" level="3" pageId="8" pageNumber="97" reason="8">
<emphasis id="B92FB520FFC390768A52FAEAFB080020" bold="true" box="[1037,1173,1285,1309]" pageId="8" pageNumber="97">Comentarios</emphasis>
</heading>
</paragraph>
<paragraph id="8BE46932FFC390768D27FAAAFB4903A3" blockId="8.[831,1380,1349,1694]" pageId="8" pageNumber="97">
A continuación reportamos algunas localidades adicionales de nueve ranas centrolénidas previamente descritas para 
<collectingCountry id="F34C29A2FFC390768DADFA6AFBC100A3" box="[1010,1116,1413,1438]" name="Colombia" pageId="8" pageNumber="97">Colombia</collectingCountry>
, producto del trabajo de campo llevado a cabo durante los últimos cinco años por parte del personal de Conservación Internacional-Colombia y de la revisión de algunos ejemplares depositados en el Museo de Historia Natural de la Universidad Nacional de 
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, Universidad del 
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, 
<collectingRegion id="499FA7D0FFC390768DD2F9AAFC690363" box="[909,1012,1605,1630]" country="Colombia" name="Antioquia" pageId="8" pageNumber="97">Antioquia</collectingRegion>
y de la Pontificia Universidad Javeriana. Estos registros amplían los rangos altitudinales y geopolíticos de las siguientes especies:
</paragraph>
</subSubSection>
</treatment>
</document>