359 lines
45 KiB
XML
359 lines
45 KiB
XML
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<mods:title id="DFCBA0F5852098C1AC9E80203781C07B">Revision of the Aspistomella group of genera (Diptera: Ulidiidae: Pterocallinae: Lipsanini)</mods:title>
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<mods:namePart id="74F5628B2BC643B4B0B448A83D4D0722">Kovac, Damir</mods:namePart>
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<mods:namePart id="341818142B7937452C12204B97FE6C95">Kameneva, Elena P.</mods:namePart>
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<mods:affiliation id="6B918511422941632C95764FFA06B76D">I. I. Schmalhausen Institute of Zoology National Academy of Sciences of Ukraine, Bogdan Chmielnicki St. 15, 01054 Kyiv, Ukraine https: // orcid. org / 0000 - 0002 - 7986 - 9942 & Museum für Naturkunde, Invalidenstr. 43, Berlin, 10115 Germany Corresponding author: valery. korneyev @ gmail. com; https: // orcid. org / 0000 - 0001 - 9631 - 1038</mods:affiliation>
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<mods:affiliation id="E7FF6ADFAAD14B96084F9483FC13242A">Museum für Naturkunde, Invalidenstr. 43, Berlin, 10115 Germany Corresponding author: valery. korneyev @ gmail. com; https: // orcid. org / 0000 - 0001 - 9631 - 1038</mods:affiliation>
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<mods:title id="71041E19841486CEFE4E2FB9BB76062F">Zootaxa</mods:title>
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<emphasis id="B92D442C1E78FFECFF2157B3FE72F8D8" bold="true" box="[151,424,1790,1817]" italics="true" pageId="24" pageNumber="25">Aspistomella crucifera</emphasis>
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(Hendel, 1909)
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<figureCitation id="136284BB1E78FFECFF21566FFF2CF8FD" box="[151,246,1826,1853]" captionStart="FIGURE 6" captionStartId="14.[151,250,1521,1545]" captionTargetBox="[177,1387,245,1460]" captionTargetId="figure-30@14.[151,1436,211,1497]" captionTargetPageId="14" captionText="FIGURE 6. Aspistomella, wing. A, A. sachavaca sp. nov.; B, A. schnusei sp. nov.; C, A. angustifrons (Hendel); D, A. garleppi sp. nov.; E, A. crucifera (Hendel); F, A. quinquincisa sp. nov.; G, A. teresensis sp. nov.; H, A. obliqua sp. nov. Abbreviations: C—costal vein; c—costal cell; M1—first branch of media; m1—first medial cell; ptstg—pterostigma; r1—first radial cell; rm—radial-medial crossvein. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14023163" httpUri="https://zenodo.org/record/14023163/files/figure.png" pageId="24" pageNumber="25">Figs 6 E</figureCitation>
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,
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<figureCitation id="136284BB1E78FFECFEB5566FFEF8F8FC" box="[259,290,1826,1852]" captionStart="FIGURE 13" captionStartId="26.[151,250,1367,1391]" captionTargetBox="[155,1436,185,1343]" captionTargetId="figure-30@26.[151,1436,181,1343]" captionTargetPageId="26" captionText="FIGURE 13. Aspistomella crucifera (Hendel) holotype ♀. A, habitus, left; B, head left; C, head, dorsal; D, mesonotum, dorsal; E, labels." figureDoi="http://doi.org/10.5281/zenodo.14023188" httpUri="https://zenodo.org/record/14023188/files/figure.png" pageId="24" pageNumber="25">13</figureCitation>
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.
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</paragraph>
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<treatmentCitation id="0AF8BE2F1E78FFECFF21562AFDE7F8BF" author="Hendel, F." box="[151,573,1895,1919]" page="262" pageId="24" pageNumber="25" year="1909">
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<taxonomicName id="4C59E3BD1E78FFECFF21562AFDE7F8BF" ID-CoL="4D8MF" authority="Hendel 1909 b: 262" authorityName="Hendel" authorityPageNumber="262" authorityYear="1909" box="[151,573,1895,1919]" class="Insecta" family="Ulidiidae" genus="Paraphyola" kingdom="Animalia" order="Diptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="crucifera">
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<emphasis id="B92D442C1E78FFECFF21562AFEA8F8BF" box="[151,370,1895,1919]" italics="true" pageId="24" pageNumber="25">Paraphyola crucifera</emphasis>
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<bibRefCitation id="EFC8E5CF1E78FFECFECE562AFDE7F8BF" author="Hendel, F." box="[376,573,1895,1919]" pageId="24" pageNumber="25" pagination="247 - 270" refId="ref75134" refString="Hendel, F. (1909 b) Beitrag zur Kenntnis der Ulidiinen (Dipt.). Wiener Entomologische Zeitung, 28, 247 - 270." type="journal article" year="1909">Hendel 1909b: 262</bibRefCitation>
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,
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<treatmentCitation id="0AF8BE2F1E78FFECFDF1562AFD79F8BF" author="Hendel, F." box="[583,675,1895,1919]" page="47" pageId="24" pageNumber="25" year="1910">
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<bibRefCitation id="EFC8E5CF1E78FFECFDF1562AFD79F8BF" author="Hendel, F." box="[583,675,1895,1919]" pageId="24" pageNumber="25" pagination="1 - 76" refId="ref75162" refString="Hendel, F. (1910) Diptera, Fam. Muscaridae. Subfam. Ulidiinae. In: Wytsman, P. (Ed.), Genera Insectorum. Fasc. 106. Wytsman Verl., Bruxelles, pp. 1 - 76, pls. 1 - 4." type="book chapter" year="1910">1910: 47</bibRefCitation>
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;
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<bibRefCitation id="EFC8E5CF1E78FFECFD18562AFC9AF8BF" author="Steyskal, G. C." box="[686,832,1895,1919]" pageId="24" pageNumber="25" refId="ref76283" refString="Steyskal, G. C. (1968) 54. Family Otitidae (Ortalidae; including Pterocallidae, Ulidiidae). In: Vanzolini, P. E. & Papavero, N. (Eds.), A catalogue of the Diptera of Americas south of the United States. Departamento de Zoologia, Secretaria da Agricultura, Sao Paulo, pp. 54.1 - 54.31." type="book" year="1968">Steyskal 1968</bibRefCitation>
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: 54.20.
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</paragraph>
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<emphasis id="B92D442C1E78FFECFF2156E0FE0DF807" bold="true" box="[151,471,1965,1991]" pageId="24" pageNumber="25">Material examined. Type.</emphasis>
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<materialsCitation id="3B3192631E78FFECFE5056E0FD31F82B" collectionCode="SMTD" collectorName="Schnuse" country="Peru" location="Peru" pageId="24" pageNumber="25" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
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<typeStatus id="54E2269C1E78FFECFE5056E0FD94F807" box="[486,590,1965,1991]" pageId="24" pageNumber="25" type="holotype">Holotype</typeStatus>
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<specimenCount id="9D5F53B71E78FFECFDEA56E0FDAEF807" box="[604,628,1965,1991]" count="1" pageId="24" pageNumber="25" type="female">♀</specimenCount>
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:
|
||
<emphasis id="B92D442C1E78FFECFD3F56E0FD19F807" bold="true" box="[649,707,1965,1991]" pageId="24" pageNumber="25">
|
||
<collectingCountry id="F34ED8AE1E78FFECFD3F56E0FD19F807" box="[649,707,1965,1991]" name="Peru" pageId="24" pageNumber="25">Peru</collectingCountry>
|
||
</emphasis>
|
||
: “Peru—Meshagua / 29.9.03 / Urubambafl.” [
|
||
<collectorName id="26ACFDE81E78FFECFB4256E0FA8EF807" box="[1268,1364,1965,1991]" pageId="24" pageNumber="25">Schnuse</collectorName>
|
||
leg.], “
|
||
<taxonomicName id="4C59E3BD1E78FFECFF17569CFEF8F82B" authorityName="Hendel" authorityYear="1909" box="[161,290,2001,2027]" class="Insecta" family="Ulidiidae" genus="Paraphyola" kingdom="Animalia" order="Diptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">Paraphyola</taxonomicName>
|
||
\ crucifera H. / det.
|
||
<taxonomicName id="4C59E3BD1E78FFECFDB5569CFDABF82B" authority="(SMTD)" authorityName="Hendel" authorityYear="1909" baseAuthorityName="SMTD" box="[515,625,2001,2027]" class="Insecta" family="Ulidiidae" genus="Axiologina" kingdom="Animalia" order="Diptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="hendel">F. Hendel</taxonomicName>
|
||
” (
|
||
<collectionCode id="ED4800FB1E78FFECFD3D569CFD05F82B" box="[651,735,2001,2027]" pageId="24" pageNumber="25">SMTD</collectionCode>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C343CBB51E79FFEDFF7151DBFBE3FDFD" pageId="25" pageNumber="26" type="diagnosis">
|
||
<paragraph id="8BE6983E1E79FFEDFF7151DBFBD6FE35" blockId="25.[151,1437,150,1977]" pageId="25" pageNumber="26">
|
||
<emphasis id="B92D442C1E79FFEDFF7151DBFEE0FF70" bold="true" box="[199,314,150,176]" pageId="25" pageNumber="26">Diagnosis</emphasis>
|
||
.
|
||
<taxonomicName id="4C59E3BD1E79FFEDFEF151DAFDE3FF71" authorityName="Hendel" authorityYear="1909" box="[327,569,151,177]" class="Insecta" family="Ulidiidae" genus="Paraphyola" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="crucifera">
|
||
<emphasis id="B92D442C1E79FFEDFEF151DAFDE3FF71" box="[327,569,151,177]" italics="true" pageId="25" pageNumber="26">Paraphyola crucifera</emphasis>
|
||
</taxonomicName>
|
||
shares with
|
||
<taxonomicName id="4C59E3BD1E79FFEDFD7451DAFCB2FF71" authorityName="Kovac & Kameneva & Korneyev & Araújo & Savaris & Smit & Schneider & Schreiber & Korneyev" authorityYear="2024" baseAuthorityName="Hendel" baseAuthorityYear="1909" box="[706,872,151,177]" class="Insecta" family="Ulidiidae" genus="Aspistomella" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="angustifrons">
|
||
<emphasis id="B92D442C1E79FFEDFD7451DAFCB2FF71" box="[706,872,151,177]" italics="true" pageId="25" pageNumber="26">P. angustifrons</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C59E3BD1E79FFEDFCC351DAFBFAFF70" authorityName="Kovac & Kameneva & Korneyev & Araújo & Savaris & Smit & Schneider & Schreiber & Korneyev" authorityYear="2024" box="[885,1056,151,176]" class="Insecta" family="Ulidiidae" genus="Aspistomella" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="quinquincisa">
|
||
<emphasis id="B92D442C1E79FFEDFCC351DAFBFAFF70" box="[885,1056,151,176]" italics="true" pageId="25" pageNumber="26">P. quinquincisa</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C59E3BD1E79FFEDFBEA51DAFB39FF70" authorityName="Kovac & Kameneva & Korneyev & Araújo & Savaris & Smit & Schneider & Schreiber & Korneyev" authorityYear="2024" box="[1116,1251,151,176]" class="Insecta" family="Ulidiidae" genus="Aspistomella" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="teresensis">
|
||
<emphasis id="B92D442C1E79FFEDFBEA51DAFB39FF70" box="[1116,1251,151,176]" italics="true" pageId="25" pageNumber="26">P. teresensis</emphasis>
|
||
</taxonomicName>
|
||
a comparatively short head, narrow frons and wings, a wing pattern with a widely dark apical half, as well as wing venation, and chaetotaxy, It differs from
|
||
<taxonomicName id="4C59E3BD1E79FFEDFE795192FDA3FF39" authorityName="Kovac & Kameneva & Korneyev & Araújo & Savaris & Smit & Schneider & Schreiber & Korneyev" authorityYear="2024" baseAuthorityName="Hendel" baseAuthorityYear="1909" box="[463,633,223,249]" class="Insecta" family="Ulidiidae" genus="Aspistomella" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="angustifrons">
|
||
<emphasis id="B92D442C1E79FFEDFE795192FDA3FF39" box="[463,633,223,249]" italics="true" pageId="25" pageNumber="26">P. angustifrons</emphasis>
|
||
</taxonomicName>
|
||
in having yellow coxae, femora and tarsi (in
|
||
<taxonomicName id="4C59E3BD1E79FFEDFB225192FAE4FF39" authorityName="Kovac & Kameneva & Korneyev & Araújo & Savaris & Smit & Schneider & Schreiber & Korneyev" authorityYear="2024" baseAuthorityName="Hendel" baseAuthorityYear="1909" box="[1172,1342,223,249]" class="Insecta" family="Ulidiidae" genus="Aspistomella" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="angustifrons">
|
||
<emphasis id="B92D442C1E79FFEDFB225192FAE4FF39" box="[1172,1342,223,249]" italics="true" pageId="25" pageNumber="26">P. angustifrons</emphasis>
|
||
</taxonomicName>
|
||
, largely black) and hyaline spot in posterior margin of cell m
|
||
<subScript id="17DD9A7B1E79FFEDFD5D5058FD29FEE3" attach="left" box="[747,755,277,291]" fontSize="6" pageId="25" pageNumber="26">4</subScript>
|
||
. It is most similar to
|
||
<taxonomicName id="4C59E3BD1E79FFEDFC5E504EFB4FFEDC" authorityName="Kovac & Kameneva & Korneyev & Araújo & Savaris & Smit & Schneider & Schreiber & Korneyev" authorityYear="2024" box="[1000,1173,259,284]" class="Insecta" family="Ulidiidae" genus="Aspistomella" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="quinquincisa">
|
||
<emphasis id="B92D442C1E79FFEDFC5E504EFB4FFEDC" box="[1000,1173,259,284]" italics="true" pageId="25" pageNumber="26">P. quinquincisa</emphasis>
|
||
</taxonomicName>
|
||
in having wing pattern with apical dark, differing from it by having 4 hyaline incisions: cell r
|
||
<subScript id="17DD9A7B1E79FFEDFC0A5074FC1EFE87" attach="left" box="[956,964,313,327]" fontSize="6" pageId="25" pageNumber="26">1</subScript>
|
||
with 2 marginal hyaline marks narrowed posteriorly, and the hyaline mark in cell m
|
||
<subScript id="17DD9A7B1E79FFEDFD325010FD56FEAB" attach="left" box="[644,652,349,363]" fontSize="6" pageId="25" pageNumber="26">1</subScript>
|
||
only reaching vein M
|
||
<subScript id="17DD9A7B1E79FFEDFC385010FC4CFEAB" attach="left" box="[910,918,349,363]" fontSize="6" pageId="25" pageNumber="26">1</subScript>
|
||
(in,
|
||
<taxonomicName id="4C59E3BD1E79FFEDFC665006FBA5FEA4" authorityName="Kovac & Kameneva & Korneyev & Araújo & Savaris & Smit & Schneider & Schreiber & Korneyev" authorityYear="2024" box="[976,1151,331,356]" class="Insecta" family="Ulidiidae" genus="Aspistomella" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="quinquincisa">
|
||
<emphasis id="B92D442C1E79FFEDFC665006FBA5FEA4" box="[976,1151,331,356]" italics="true" pageId="25" pageNumber="26">P. quinquincisa</emphasis>
|
||
</taxonomicName>
|
||
, crossing vein M
|
||
<subScript id="17DD9A7B1E79FFEDFAFF5010FA8BFEAB" attach="left" box="[1353,1361,349,363]" fontSize="6" pageId="25" pageNumber="26">1</subScript>
|
||
to the middle of cell r
|
||
<subScript id="17DD9A7B1E79FFEDFEFD50CCFEBFFE4F" attach="left" box="[331,357,385,399]" fontSize="6" pageId="25" pageNumber="26">4+5</subScript>
|
||
). It differs from
|
||
<taxonomicName id="4C59E3BD1E79FFEDFD9A5022FD62FE48" authorityName="Kovac & Kameneva & Korneyev & Araújo & Savaris & Smit & Schneider & Schreiber & Korneyev" authorityYear="2024" box="[556,696,367,392]" class="Insecta" family="Ulidiidae" genus="Aspistomella" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="teresensis">
|
||
<emphasis id="B92D442C1E79FFEDFD9A5022FD62FE48" box="[556,696,367,392]" italics="true" pageId="25" pageNumber="26">P. teresensis</emphasis>
|
||
</taxonomicName>
|
||
by cell r
|
||
<subScript id="17DD9A7B1E79FFEDFC9550CCFCF1FE4F" attach="left" box="[803,811,385,399]" fontSize="6" pageId="25" pageNumber="26">1</subScript>
|
||
with 2 marginal hyaline marks, the first one reaching vein M
|
||
<subScript id="17DD9A7B1E79FFEDFF5E50E8FF2FFE73" attach="left" box="[232,245,421,435]" fontSize="6" pageId="25" pageNumber="26">1,</subScript>
|
||
the second one reaching the anterior half of cell r
|
||
<subScript id="17DD9A7B1E79FFEDFC9A50E8FC9CFE73" attach="left" box="[812,838,421,435]" fontSize="6" pageId="25" pageNumber="26">4+5</subScript>
|
||
and cell m
|
||
<subScript id="17DD9A7B1E79FFEDFC7250E8FC16FE73" attach="left" box="[964,972,421,435]" fontSize="6" pageId="25" pageNumber="26">1</subScript>
|
||
with submarginal hyaline mark reaching only vein M
|
||
<subScript id="17DD9A7B1E79FFEDFE945084FEF0FE17" attach="left" box="[290,298,457,471]" fontSize="6" pageId="25" pageNumber="26">1</subScript>
|
||
(in
|
||
<taxonomicName id="4C59E3BD1E79FFEDFEEC50FAFE39FE10" authorityName="Kovac & Kameneva & Korneyev & Araújo & Savaris & Smit & Schneider & Schreiber & Korneyev" authorityYear="2024" box="[346,483,439,464]" class="Insecta" family="Ulidiidae" genus="Aspistomella" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="teresensis">
|
||
<emphasis id="B92D442C1E79FFEDFEEC50FAFE39FE10" box="[346,483,439,464]" italics="true" pageId="25" pageNumber="26">P. teresensis</emphasis>
|
||
</taxonomicName>
|
||
, cell r
|
||
<subScript id="17DD9A7B1E79FFEDFD9A5084FDEEFE17" attach="left" box="[556,564,457,471]" fontSize="6" pageId="25" pageNumber="26">1</subScript>
|
||
with one cuneiform hyaline incision extending into cell dm and cell m
|
||
<subScript id="17DD9A7B1E79FFEDFAEC5084FAB8FE17" attach="left" box="[1370,1378,457,471]" fontSize="6" pageId="25" pageNumber="26">1</subScript>
|
||
with cuneiform hyaline incision extending at least to vein R
|
||
<subScript id="17DD9A7B1E79FFEDFD4A50A0FCCCFE3B" attach="left" box="[764,790,493,507]" fontSize="6" pageId="25" pageNumber="26">4+5</subScript>
|
||
or even into cell r
|
||
<subScript id="17DD9A7B1E79FFEDFC5450A0FC21FE3B" attach="left" box="[994,1019,493,507]" fontSize="6" pageId="25" pageNumber="26">2+3</subScript>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE6983E1E79FFEDFF7150B2FBE3FDFD" blockId="25.[151,1437,150,1977]" pageId="25" pageNumber="26">
|
||
It also differs from
|
||
<taxonomicName id="4C59E3BD1E79FFEDFE1150B2FDFEFDD9" authorityName="Kovac & Kameneva & Korneyev & Araújo & Savaris & Smit & Schneider & Schreiber & Korneyev" authorityYear="2024" box="[423,548,511,537]" class="Insecta" family="Ulidiidae" genus="Aspistomella" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="schnusei">
|
||
<emphasis id="B92D442C1E79FFEDFE1150B2FDFEFDD9" box="[423,548,511,537]" italics="true" pageId="25" pageNumber="26">P. schnusei</emphasis>
|
||
</taxonomicName>
|
||
by short epistome and wing apical half widely dark (in
|
||
<taxonomicName id="4C59E3BD1E79FFEDFB1C50B2FAFDFDD9" authorityName="Kovac & Kameneva & Korneyev & Araújo & Savaris & Smit & Schneider & Schreiber & Korneyev" authorityYear="2024" box="[1194,1319,511,537]" class="Insecta" family="Ulidiidae" genus="Aspistomella" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="schnusei">
|
||
<emphasis id="B92D442C1E79FFEDFB1C50B2FAFDFDD9" box="[1194,1319,511,537]" italics="true" pageId="25" pageNumber="26">P. schnusei</emphasis>
|
||
</taxonomicName>
|
||
, epistome strongly produced anteriorly and wing widely hyaline with narrow bands apically).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C343CBB51E79FFEDFF71530BFACEF879" pageId="25" pageNumber="26" type="description">
|
||
<paragraph id="8BE6983E1E79FFEDFF71530BFD88FB99" blockId="25.[151,1437,150,1977]" pageId="25" pageNumber="26">
|
||
<emphasis id="B92D442C1E79FFEDFF71530BFE83FDA0" bold="true" box="[199,345,582,608]" pageId="25" pageNumber="26">Description.</emphasis>
|
||
Male.
|
||
<emphasis id="B92D442C1E79FFEDFE10530AFE3CFDA1" bold="true" box="[422,486,583,609]" pageId="25" pageNumber="26">Head</emphasis>
|
||
(
|
||
<figureCitation id="136284BB1E79FFEDFE42530AFD59FDA1" box="[500,643,583,609]" captionStart="FIGURE 13" captionStartId="26.[151,250,1367,1391]" captionTargetBox="[155,1436,185,1343]" captionTargetId="figure-30@26.[151,1436,181,1343]" captionTargetPageId="26" captionText="FIGURE 13. Aspistomella crucifera (Hendel) holotype ♀. A, habitus, left; B, head left; C, head, dorsal; D, mesonotum, dorsal; E, labels." figureDoi="http://doi.org/10.5281/zenodo.14023188" httpUri="https://zenodo.org/record/14023188/files/figure.png" pageId="25" pageNumber="26">Figs 13 B–C</figureCitation>
|
||
) ratio (length: height: width) = 1: 1.5: 2, frons, face above transverse fold, parafacial, and gena reddish or yellowish brown, epistome and occiput mostly black. Frons very narrow, slightly narrowed posteriorly, 0.95× as long (from lunule to anterior ocellus) or 1.8× (from lunule to inner vertical seta) as wide (at lunule), with subshining, brown or black ocellar triangle and brown vertical plates; orbits, vertex, gena and occiput silver-white microtrichose; parafacial narrow, 0.25–0.3× broader than postpedicel, reddish-yellow, sparsely white microtrichose. Frontal plates with 8–9 tiny parafrontal setulae and 5–7 frontal setae on each side, frontal vitta sparsely whitish microtrichose, with 2–4 pro- or inclinate interfrontal setae (
|
||
<figureCitation id="136284BB1E79FFEDFB105252FAC1FCF9" box="[1190,1307,799,825]" captionStart="FIGURE 13" captionStartId="26.[151,250,1367,1391]" captionTargetBox="[155,1436,185,1343]" captionTargetId="figure-30@26.[151,1436,181,1343]" captionTargetPageId="26" captionText="FIGURE 13. Aspistomella crucifera (Hendel) holotype ♀. A, habitus, left; B, head left; C, head, dorsal; D, mesonotum, dorsal; E, labels." figureDoi="http://doi.org/10.5281/zenodo.14023188" httpUri="https://zenodo.org/record/14023188/files/figure.png" pageId="25" pageNumber="26">Fig. 13 C</figureCitation>
|
||
). Eye 1.5× higher than long. Face; yellow, white microtrichose above epistome; epistome moderately produced anteriorly and separated from upper part of face by shallow fold, brown, sparsely white microtrichose, with metallic bluish sheen. Clypeus brown, sparsely white microtrichose, subshining, 0.2× higher than epistome height. Gena brownish yellow, with long genal seta and 6–7 additional long peristomal setae anterior of it, 0.5–0.7× longer than genal seta (
|
||
<figureCitation id="136284BB1E79FFEDFAFA52E2FF76FC2D" captionStart="FIGURE 13" captionStartId="26.[151,250,1367,1391]" captionTargetBox="[155,1436,185,1343]" captionTargetId="figure-30@26.[151,1436,181,1343]" captionTargetPageId="26" captionText="FIGURE 13. Aspistomella crucifera (Hendel) holotype ♀. A, habitus, left; B, head left; C, head, dorsal; D, mesonotum, dorsal; E, labels." figureDoi="http://doi.org/10.5281/zenodo.14023188" httpUri="https://zenodo.org/record/14023188/files/figure.png" pageId="25" pageNumber="26">Fig. 13 B</figureCitation>
|
||
). Occiput black, with cyan metallic sheen, greyish microtrichose. Antenna yellow; scape and pedicel with black setulae; postpedicel yellow, whitish microtrichose, 1.9× longer than wide, apically rounded; arista black except yellow at base, almost bare. Mouthparts brown to black, prementum black, sparsely microtrichose. Palp yellow to brown, as described for
|
||
<taxonomicName id="4C59E3BD1E79FFEDFE135572FD91FB99" authorityName="Kovac & Kameneva & Korneyev & Araújo & Savaris & Smit & Schneider & Schreiber & Korneyev" authorityYear="2024" baseAuthorityName="Hendel" baseAuthorityYear="1909" box="[421,587,1087,1113]" class="Insecta" family="Ulidiidae" genus="Aspistomella" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="angustifrons">
|
||
<emphasis id="B92D442C1E79FFEDFE135572FD91FB99" box="[421,587,1087,1113]" italics="true" pageId="25" pageNumber="26">P. angustifrons</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BE6983E1E79FFEDFF71552EFDCEFAF1" blockId="25.[151,1437,150,1977]" pageId="25" pageNumber="26">
|
||
<emphasis id="B92D442C1E79FFEDFF71552EFEFAFBBD" bold="true" box="[199,288,1123,1149]" pageId="25" pageNumber="26">Thorax</emphasis>
|
||
brown to black, silvery subshining, with sparse white microtrichia not hiding underlying cuticle. Mesonotal scutum (
|
||
<figureCitation id="136284BB1E79FFEDFEC155CAFE3CFB61" box="[375,486,1159,1185]" captionStart="FIGURE 13" captionStartId="26.[151,250,1367,1391]" captionTargetBox="[155,1436,185,1343]" captionTargetId="figure-30@26.[151,1436,181,1343]" captionTargetPageId="26" captionText="FIGURE 13. Aspistomella crucifera (Hendel) holotype ♀. A, habitus, left; B, head left; C, head, dorsal; D, mesonotum, dorsal; E, labels." figureDoi="http://doi.org/10.5281/zenodo.14023188" httpUri="https://zenodo.org/record/14023188/files/figure.png" pageId="25" pageNumber="26">Fig. 13 D</figureCitation>
|
||
) 1.4–1.5× longer than wide; black setulose, with 6–7 rows of setulae between rows of dorsocentral setulae; acrostichal prescutellar seta present, 0.4–0.5× longer than posterior dorsocentral seta (prescutellar area partly destroyed with pin). Other setae as described for the genus, black. Scutellum dorsally very flattened, brown, subshining, sparsely microtrichose, devoid of setulae, with silver sheen. Subscutellum and mediotergite as in
|
||
<taxonomicName id="4C59E3BD1E79FFEDFED0545AFDD7FAF1" authorityName="Kovac & Kameneva & Korneyev & Araújo & Savaris & Smit & Schneider & Schreiber & Korneyev" authorityYear="2024" baseAuthorityName="Hendel" baseAuthorityYear="1909" box="[358,525,1303,1329]" class="Insecta" family="Ulidiidae" genus="Aspistomella" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="angustifrons">
|
||
<emphasis id="B92D442C1E79FFEDFED0545AFDD7FAF1" box="[358,525,1303,1329]" italics="true" pageId="25" pageNumber="26">P. angustifrons</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BE6983E1E79FFEDFF715477FDE5F921" blockId="25.[151,1437,150,1977]" pageId="25" pageNumber="26">
|
||
<emphasis id="B92D442C1E79FFEDFF715477FED2FA94" bold="true" box="[199,264,1338,1364]" pageId="25" pageNumber="26">Wing</emphasis>
|
||
(
|
||
<figureCitation id="136284BB1E79FFEDFEA15476FEA8FA95" box="[279,370,1339,1365]" captionStart="FIGURE 6" captionStartId="14.[151,250,1521,1545]" captionTargetBox="[177,1387,245,1460]" captionTargetId="figure-30@14.[151,1436,211,1497]" captionTargetPageId="14" captionText="FIGURE 6. Aspistomella, wing. A, A. sachavaca sp. nov.; B, A. schnusei sp. nov.; C, A. angustifrons (Hendel); D, A. garleppi sp. nov.; E, A. crucifera (Hendel); F, A. quinquincisa sp. nov.; G, A. teresensis sp. nov.; H, A. obliqua sp. nov. Abbreviations: C—costal vein; c—costal cell; M1—first branch of media; m1—first medial cell; ptstg—pterostigma; r1—first radial cell; rm—radial-medial crossvein. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14023163" httpUri="https://zenodo.org/record/14023163/files/figure.png" pageId="25" pageNumber="26">Fig. 6 E</figureCitation>
|
||
) about
|
||
<quantity id="4CA135DB1E79FFEDFE735476FDDCFA95" box="[453,518,1339,1365]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="25" pageNumber="26" unit="mm" value="4.0">4 mm</quantity>
|
||
(
|
||
<specimenCount id="9D5F53B71E79FFEDFDA15476FDF3FA95" box="[535,553,1339,1365]" count="1" pageId="25" pageNumber="26" type="female">♀</specimenCount>
|
||
) long (
|
||
<typeStatus id="54E2269C1E79FFEDFDCC5476FD04FA95" box="[634,734,1339,1365]" pageId="25" pageNumber="26" type="holotype">holotype</typeStatus>
|
||
with both wings broken off), 3.3× longer than wide; basicostal cell hyaline; costal cell straight, 5.2× longer than wide, brown in basal 1/8 of length; pterostigma entirely brown, narrow triangular, 2.7–3× longer than wide, vein R
|
||
<subScript id="17DD9A7B1E79FFEDFD5454D8FD30FA63" attach="left" box="[738,746,1429,1443]" fontSize="6" pageId="25" pageNumber="26">1</subScript>
|
||
bare, ending proximally of crossvein dm-m level; vein R
|
||
<subScript id="17DD9A7B1E79FFEDFA3554D8FA46FA63" attach="left" box="[1411,1436,1429,1443]" fontSize="6" pageId="25" pageNumber="26">2+3</subScript>
|
||
slightly arcuate in basal half, straight at apex. Crossvein r-m distally of middle of cell dm. Cell r
|
||
<subScript id="17DD9A7B1E79FFEDFB4354F4FAD5FA07" attach="left" box="[1269,1295,1465,1479]" fontSize="6" pageId="25" pageNumber="26">4+5</subScript>
|
||
6.5× longer than wide. Cell m
|
||
<subScript id="17DD9A7B1E79FFEDFEEB5490FEBFFA2B" attach="left" box="[349,357,1501,1515]" fontSize="6" pageId="25" pageNumber="26">1</subScript>
|
||
narrow triangular; ultimate section of M
|
||
<subScript id="17DD9A7B1E79FFEDFC9B5490FCEFFA2B" attach="left" box="[813,821,1501,1515]" fontSize="6" pageId="25" pageNumber="26">1</subScript>
|
||
4.7× longer than crossvein dm-m and 3.2× longer than penultimate section. Vein CuA Z-shaped, forming moderately short posteroapical lobe of cell cua along vein CuP, 1.25× longer than its anterior shoulder. Wing pattern with short subbasal crossband from humeral vein through cell cua into alula; wing distally of cells c, bm, cua and radial fork dark brown with 4 large hyaline areas: 2 hyaline subtriangular incisions at anterior margin from cell r
|
||
<subScript id="17DD9A7B1E79FFEDFD545720FD30F9BB" attach="left" box="[738,746,1645,1659]" fontSize="6" pageId="25" pageNumber="26">1</subScript>
|
||
through cell r
|
||
<subScript id="17DD9A7B1E79FFEDFC3E5720FC7BF9BB" attach="left" box="[904,929,1645,1659]" fontSize="6" pageId="25" pageNumber="26">2+3</subScript>
|
||
into base of cell r
|
||
<subScript id="17DD9A7B1E79FFEDFBDD5720FB5FF9BB" attach="left" box="[1131,1157,1645,1659]" fontSize="6" pageId="25" pageNumber="26">4+5</subScript>
|
||
distally of crossvein r-m and 2 rounded or subrectangular hyaline spots in apical parts of cells dm and cua
|
||
<subScript id="17DD9A7B1E79FFEDFB8257DCFBE6F95F" attach="left" box="[1076,1084,1681,1695]" fontSize="6" pageId="25" pageNumber="26">1</subScript>
|
||
and basal part of cell m
|
||
<subScript id="17DD9A7B1E79FFEDFAE457DCFA80F95F" attach="left" box="[1362,1370,1681,1695]" fontSize="6" pageId="25" pageNumber="26">1</subScript>
|
||
; dark pattern separating these hyaline areas having shape of cross; wing apex dark without hyaline marks. Calypters white, with white cilia. Halter yellow.
|
||
</paragraph>
|
||
<paragraph id="8BE6983E1E79FFEDFF7157A6FD27F88D" blockId="25.[151,1437,150,1977]" pageId="25" pageNumber="26">
|
||
<emphasis id="B92D442C1E79FFEDFF7157A6FF25F8C5" bold="true" box="[199,255,1771,1797]" pageId="25" pageNumber="26">Legs</emphasis>
|
||
(
|
||
<figureCitation id="136284BB1E79FFEDFEB957A6FEA6F8C5" box="[271,380,1771,1797]" captionStart="FIGURE 13" captionStartId="26.[151,250,1367,1391]" captionTargetBox="[155,1436,185,1343]" captionTargetId="figure-30@26.[151,1436,181,1343]" captionTargetPageId="26" captionText="FIGURE 13. Aspistomella crucifera (Hendel) holotype ♀. A, habitus, left; B, head left; C, head, dorsal; D, mesonotum, dorsal; E, labels." figureDoi="http://doi.org/10.5281/zenodo.14023188" httpUri="https://zenodo.org/record/14023188/files/figure.png" pageId="25" pageNumber="26">Fig. 13 A</figureCitation>
|
||
) yellow; black setose and setulose. Fore femur uniformly setulose, with subapical row of 2–4 posteroventral setae as long as femur width. Mid femur anteriorly with short suberect setulae; mid tibia ventrally with single spur-like seta 3–5× longer than tibia width.
|
||
</paragraph>
|
||
<paragraph id="8BE6983E1E79FFEDFF71561BFD67F8B1" blockId="25.[151,1437,150,1977]" box="[199,701,1878,1905]" pageId="25" pageNumber="26">
|
||
<emphasis id="B92D442C1E79FFEDFF71561BFEE6F8B0" bold="true" box="[199,316,1878,1904]" pageId="25" pageNumber="26">Abdomen</emphasis>
|
||
brown to black, sparsely setulose.
|
||
</paragraph>
|
||
<paragraph id="8BE6983E1E79FFEDFF715636FE54F855" blockId="25.[151,1437,150,1977]" box="[199,398,1915,1941]" pageId="25" pageNumber="26">Male: not known.</paragraph>
|
||
<paragraph id="8BE6983E1E79FFEDFF7156D2FACEF879" blockId="25.[151,1437,150,1977]" box="[199,1300,1950,1977]" pageId="25" pageNumber="26">
|
||
<emphasis id="B92D442C1E79FFEDFF7156D2FE7BF878" bold="true" box="[199,417,1950,1977]" pageId="25" pageNumber="26">Female terminalia</emphasis>
|
||
(not dissected): aculeus with ovoid, long-setulose cercal unit (partly exposed).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF26C8B61E7AFFEEFF21541AFF2DFA53" ID-DOI="http://doi.org/10.5281/zenodo.14023188" ID-Zenodo-Dep="14023188" httpUri="https://zenodo.org/record/14023188/files/figure.png" pageId="26" pageNumber="27" startId="26.[151,250,1367,1391]" targetBox="[155,1436,185,1343]" targetPageId="26" targetType="figure">
|
||
<paragraph id="8BE6983E1E7AFFEEFF21541AFF2DFA53" blockId="26.[151,1436,1367,1427]" pageId="26" pageNumber="27">
|
||
<emphasis id="B92D442C1E7AFFEEFF21541AFEC5FAAF" bold="true" box="[151,287,1367,1391]" pageId="26" pageNumber="27">FIGURE 13.</emphasis>
|
||
<taxonomicName id="4C59E3BD1E7AFFEEFE93541AFDB0FAAF" authority="(Hendel)" authorityName="Kovac & Kameneva & Korneyev & Araújo & Savaris & Smit & Schneider & Schreiber & Korneyev" authorityYear="2024" baseAuthorityName="Hendel" baseAuthorityYear="1909" box="[293,618,1367,1391]" class="Insecta" family="Ulidiidae" genus="Aspistomella" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="crucifera">
|
||
<emphasis id="B92D442C1E7AFFEEFE93541AFDD1FAAF" box="[293,523,1367,1391]" italics="true" pageId="26" pageNumber="27">Aspistomella crucifera</emphasis>
|
||
(Hendel)
|
||
</taxonomicName>
|
||
holotype ♀.
|
||
<emphasis id="B92D442C1E7AFFEEFD58541AFCDAFAAF" bold="true" box="[750,768,1367,1391]" pageId="26" pageNumber="27">A</emphasis>
|
||
, habitus, left;
|
||
<emphasis id="B92D442C1E7AFFEEFC26541AFC7BFAAF" bold="true" box="[912,929,1367,1391]" pageId="26" pageNumber="27">B</emphasis>
|
||
, head left;
|
||
<emphasis id="B92D442C1E7AFFEEFBA7541AFBF9FAAF" bold="true" box="[1041,1059,1367,1391]" pageId="26" pageNumber="27">C</emphasis>
|
||
, head, dorsal;
|
||
<emphasis id="B92D442C1E7AFFEEFB00541AFB12FAAF" bold="true" box="[1206,1224,1367,1391]" pageId="26" pageNumber="27">D</emphasis>
|
||
, mesonotum, dorsal;
|
||
<emphasis id="B92D442C1E7AFFEEFF215436FF72FA53" bold="true" box="[151,168,1403,1427]" pageId="26" pageNumber="27">E</emphasis>
|
||
, labels.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C343CBB51E7AFFEEFF7154ABFD9EF988" pageId="26" pageNumber="27" type="discussion">
|
||
<paragraph id="8BE6983E1E7AFFEEFF7154ABFD9EF988" blockId="26.[151,1437,1509,1608]" pageId="26" pageNumber="27">
|
||
<emphasis id="B92D442C1E7AFFEEFF7154ABFEEEF9C0" bold="true" box="[199,308,1510,1536]" pageId="26" pageNumber="27">Remarks</emphasis>
|
||
. The specimen is partially damaged: both wings are broken off. A pencil sketch of the wing pattern was drawn by VK in 2001, before the curator glued it back to the thorax with fish glue as in the photo (
|
||
<figureCitation id="136284BB1E7AFFEEFA905744FF74F988" captionStart="FIGURE 13" captionStartId="26.[151,250,1367,1391]" captionTargetBox="[155,1436,185,1343]" captionTargetId="figure-30@26.[151,1436,181,1343]" captionTargetPageId="26" captionText="FIGURE 13. Aspistomella crucifera (Hendel) holotype ♀. A, habitus, left; B, head left; C, head, dorsal; D, mesonotum, dorsal; E, labels." figureDoi="http://doi.org/10.5281/zenodo.14023188" httpUri="https://zenodo.org/record/14023188/files/figure.png" pageId="26" pageNumber="27">Figs 13 A, D</figureCitation>
|
||
); it was later redrawn as in
|
||
<figureCitation id="136284BB1E7AFFEEFE555760FD9AF988" box="[483,576,1581,1608]" captionStart="FIGURE 6" captionStartId="14.[151,250,1521,1545]" captionTargetBox="[177,1387,245,1460]" captionTargetId="figure-30@14.[151,1436,211,1497]" captionTargetPageId="14" captionText="FIGURE 6. Aspistomella, wing. A, A. sachavaca sp. nov.; B, A. schnusei sp. nov.; C, A. angustifrons (Hendel); D, A. garleppi sp. nov.; E, A. crucifera (Hendel); F, A. quinquincisa sp. nov.; G, A. teresensis sp. nov.; H, A. obliqua sp. nov. Abbreviations: C—costal vein; c—costal cell; M1—first branch of media; m1—first medial cell; ptstg—pterostigma; r1—first radial cell; rm—radial-medial crossvein. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14023163" httpUri="https://zenodo.org/record/14023163/files/figure.png" pageId="26" pageNumber="27">Fig. 6 E</figureCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |