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<document id="8D55DA5DEC1A509EA66A69D7CD8685B4" ID-DOI="10.5281/zenodo.6391953" ID-ISSN="1942-1354" ID-Zenodo-Dep="6391953" ID-ZooBank="B6862B03-E882-437A-B9E0-3B5FB1443602" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1648504685522" checkinUser="felipe" docAuthor="Skelley, Paul E. &amp; Keller, Oliver" docDate="2022" docId="03D5014CFFF3FFF6FF1ECD933D9A61A4" docLanguage="en" docName="InsectaMundi.2022.0916.1-11.pdf" docOrigin="Insecta Mundi 2022 (916)" docStyle="DocumentStyle:F9976E9B7F8A094D890004515A7A0592.3:InsectaMundi.2020-.journal_article" docStyleId="F9976E9B7F8A094D890004515A7A0592" docStyleName="InsectaMundi.2020-.journal_article" docStyleVersion="3" docTitle="Haroldiellus lansbergei" docType="treatment" docVersion="3" lastPageNumber="3" masterDocId="FFEC7934FFF7FFF2FFDECD463F236519" masterDocTitle="A third species of Haroldiellus Gordon and Skelley, 2007 from Mexico and Central America (Coleoptera: Scarabaeidae: Aphodiinae: Aphodiini)" masterLastPageNumber="11" masterPageNumber="1" pageNumber="3" updateTime="1699308862729" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="8DFA3B348FC69613EB9F0B0324C2D9F8">A third species of Haroldiellus Gordon and Skelley, 2007 from Mexico and Central America (Coleoptera: Scarabaeidae: Aphodiinae: Aphodiini)</mods:title>
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<mods:namePart id="CD41B15C5F975DA47A8FE9AB9B84C00E">Skelley, Paul E.</mods:namePart>
<mods:affiliation id="D87697A6C8853308231CE5B272DBFA89">Florida State Collection of Arthropods Florida Department of Agriculture and Consumer Services P. O. Box 147100 Gainesville, FL 32614 - 7100</mods:affiliation>
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<mods:namePart id="563F01EBC00DF18D11081ABFA4920351">Keller, Oliver</mods:namePart>
<mods:affiliation id="9F6B47934D05AF33B2A2E9F256FDBCE1">Florida State Collection of Arthropods Florida Department of Agriculture and Consumer Services P. O. Box 147100 Gainesville, FL 32614 - 7100</mods:affiliation>
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<treatment id="03D5014CFFF3FFF6FF1ECD933D9A61A4" ID-DOI="http://doi.org/10.5281/zenodo.6391961" ID-Zenodo-Dep="6391961" LSID="urn:lsid:plazi:treatment:03D5014CFFF3FFF6FF1ECD933D9A61A4" httpUri="http://treatment.plazi.org/id/03D5014CFFF3FFF6FF1ECD933D9A61A4" lastPageNumber="3" pageId="4" pageNumber="3">
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<paragraph id="8BC3B05AFFF3FFF6FF1ECD933D7965EB" blockId="4.[192,602,213,276]" box="[192,602,213,242]" pageId="4" pageNumber="3">
<heading id="D08B0736FFF3FFF6FF1ECD933D7965EB" box="[192,602,213,242]" fontSize="12" level="2" pageId="4" pageNumber="3" reason="2">
<taxonomicName id="4C7CCBD9FFF3FFF6FF1ECD933D7965EB" authority="(Harold)" baseAuthorityName="Harold" baseAuthorityYear="1874" box="[192,602,213,242]" class="Insecta" family="Aphodiidae" genus="Haroldiellus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="lansbergei">
<emphasis id="B9086C48FFF3FFF6FF1ECD933D7965EB" bold="true" box="[192,602,213,242]" pageId="4" pageNumber="3">
<emphasis id="B9086C48FFF3FFF6FF1ECD933EC765EB" bold="true" box="[192,484,213,242]" italics="true" pageId="4" pageNumber="3">Haroldiellus lansbergei</emphasis>
(Harold)
</emphasis>
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<subSubSection id="C366E3D1FFF3FFF6FF1ECDBF3E4B640D" box="[192,360,249,276]" pageId="4" pageNumber="3" type="description">
<paragraph id="8BC3B05AFFF3FFF6FF1ECDBF3E4B640D" blockId="4.[192,602,213,276]" box="[192,360,249,276]" pageId="4" pageNumber="3">
<figureCitation id="1347ACDFFFF3FFF6FF1ECDBF3E0B640D" box="[192,296,249,276]" captionStart="Figures 15" captionStartId="8.[192,279,1279,1305]" captionTargetBox="[192,1440,213,1249]" captionTargetId="figure-66@8.[192,1440,213,1249]" captionTargetPageId="8" captionText="Figures 15. Haroldiellus species. 1) H. sallei dorsal habitus. 2) H. lansbergei dorsal habitus. 3) H. woodruffi, holotype, dorsal habitus. 4) H. sallei protibia dorsal surface with punctures. 5) H. woodruffi protibia dorsal surface without punctures. Scale line = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6391957" httpUri="https://zenodo.org/record/6391957/files/figure.png" pageId="4" pageNumber="3">Figures 2</figureCitation>
,
<figureCitation id="1347ACDFFFF3FFF6FEEDCDBC3E61640D" box="[307,322,250,276]" captionStart="Figures 69" captionStartId="9.[195,282,992,1018]" captionTargetBox="[192,1440,213,962]" captionTargetId="figure-11@9.[192,1440,213,962]" captionTargetPageId="9" captionText="Figures 69. Heads of Haroldiellus species. 6) H. sallei. 7) H. lansbergei. 8) H. woodruffi female allotype. 9) H. woodruffi male holotype. Scale line = 0.5 mm (all figures)." figureDoi="http://doi.org/10.5281/zenodo.6391959" httpUri="https://zenodo.org/record/6391959/files/figure.png" pageId="4" pageNumber="3">7</figureCitation>
,
<figureCitation id="1347ACDFFFF3FFF6FE93CDBC3E4B640D" box="[333,360,250,276]" captionStart="Figure 10" captionStartId="10.[192,268,866,892]" captionTargetBox="[192,1440,213,836]" captionTargetId="figure-86@10.[192,1440,213,836]" captionTargetPageId="10" captionText="Figure 10. Distribution map of Haroldiellus lansbergei (blue) as currently known from specimen labels and literature records. Chiapas, Mexico is striped because it is a literature record that may represent the new species H. woodruffi described herein. Map made with SimpleMappr (Shorthouse 2010)." figureDoi="http://doi.org/10.5281/zenodo.6391963" httpUri="https://zenodo.org/record/6391963/files/figure.png" pageId="4" pageNumber="3">10</figureCitation>
</paragraph>
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<subSubSection id="C366E3D1FFF3FFF6FF3ECC643C4C64AC" pageId="4" pageNumber="3" type="reference_group">
<paragraph id="8BC3B05AFFF3FFF6FF3ECC643DC96441" blockId="4.[224,1440,290,437]" pageId="4" pageNumber="3">
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<treatmentCitation id="0ADD964BFFF3FFF6FF3ECC643D756423" author="Harold E." box="[224,598,290,314]" page="179" pageId="4" pageNumber="3" year="1874">
<taxonomicName id="4C7CCBD9FFF3FFF6FF3ECC643D756423" authority="Harold 1874: 179" authorityName="Harold" authorityPageNumber="179" authorityYear="1874" box="[224,598,290,314]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="lansbergei">
<emphasis id="B9086C48FFF3FFF6FF3ECC643E856423" box="[224,422,290,314]" italics="true" pageId="4" pageNumber="3">Aphodius lansbergei</emphasis>
<bibRefCitation id="EFEDCDABFFF3FFF6FE75CC643D756423" author="Harold E." box="[427,598,290,314]" pageId="4" pageNumber="3" pagination="177 - 208" refId="ref5369" refString="Harold E. 1874. Beitrage zur Kenntnis einiger coprophagen Lamellicornien. (Achtes Stuck). Berliner Entomologische Zeitschrift 18: 177 - 208." type="journal article" year="1874">Harold 1874: 179</bibRefCitation>
</taxonomicName>
</treatmentCitation>
~
<treatmentCitation id="0ADD964BFFF3FFF6FDB1CC643DDD6423" author="Bates HW" box="[623,766,290,314]" page="84" pageId="4" pageNumber="3" year="1887">
<bibRefCitation id="EFEDCDABFFF3FFF6FDB1CC643DDD6423" author="Bates HW" box="[623,766,290,314]" pageId="4" pageNumber="3" pagination="25 - 160" refId="ref4902" refString="Bates HW. 1887. Biologia Centrali-Americana. Insecta. Coleoptera: Lamellicornia. (Copridae, Aphodiidae, Orphnidae, Hybosoridae, Geotrupidae, Trogidae, Aclopidae, Chasmatopteridae, Melolonthidae) 2: 25 - 160." type="journal article" year="1887">Bates 1887: 84</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0ADD964BFFF3FFF6FCD9CC643CCA6423" author="Blackwelder RE" box="[775,1001,290,314]" page="212" pageId="4" pageNumber="3" year="1944">
<bibRefCitation id="EFEDCDABFFF3FFF6FCD9CC643CCA6423" author="Blackwelder RE" box="[775,1001,290,314]" pageId="4" pageNumber="3" pagination="189 - 341" refId="ref4964" refString="Blackwelder RE. 1944. Checklist of the Coleopterous insects of Mexico, Central America, the West Indies and South America. Part II. Bulletin of the United States National Museum 185: i - iii + 189 - 341." type="journal article" year="1944">Blackwelder 1944: 212</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0ADD964BFFF3FFF6FC2FCC643BCB6423" author="Dellacasa M &amp; Gordon RD &amp; Dellacasa G." box="[1009,1256,290,314]" page="158" pageId="4" pageNumber="3" year="1998">
<bibRefCitation id="EFEDCDABFFF3FFF6FC2FCC643BCB6423" author="Dellacasa M &amp; Gordon RD &amp; Dellacasa G." box="[1009,1256,290,314]" pageId="4" pageNumber="3" pagination="139 - 162" refId="ref5126" refString="Dellacasa M, Gordon RD, Dellacasa G. 1998. The types of Aphodius species described by Harold from Mexico with description of a new genus. Acta Zoologica Mexicana 74: 139 - 162." type="journal article" year="1998">Dellacasa et al. 1998: 158</bibRefCitation>
</treatmentCitation>
(habitus, epipharynx and aedeagus figured;
<typeStatus id="54C70EF8FFF3FFF6FDD2CC063D4B6441" box="[524,616,320,344]" pageId="4" pageNumber="3" type="lectotype">lectotype</typeStatus>
designated).
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</paragraph>
<paragraph id="8BC3B05AFFF3FFF6FF3ECC193C12646E" blockId="4.[224,1440,290,437]" box="[224,817,351,375]" pageId="4" pageNumber="3">
<treatmentCitationGroup id="AB6C9774FFF3FFF6FF3ECC193C12646E" box="[224,817,351,375]" pageId="4" pageNumber="3">
<taxonomicName id="4C7CCBD9FFF3FFF6FF3ECC193D76646E" authority="Harold" authorityName="Harold" authorityYear="1874" box="[224,597,351,375]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="lansbergei" subGenus="Bodilus">
<emphasis id="B9086C48FFF3FFF6FF3ECC193E1D646E" box="[224,318,351,375]" italics="true" pageId="4" pageNumber="3">Aphodius</emphasis>
(
<emphasis id="B9086C48FFF3FFF6FE93CC193EB4646E" box="[333,407,351,375]" italics="true" pageId="4" pageNumber="3">Bodilus</emphasis>
)
<emphasis id="B9086C48FFF3FFF6FE7BCC193D2B646E" box="[421,520,351,375]" italics="true" pageId="4" pageNumber="3">lansbergei</emphasis>
Harold
</taxonomicName>
~
<treatmentCitation id="0ADD964BFFF3FFF6FDAECC193C0E646E" author="Schmidt A." box="[624,813,351,375]" page="123" pageId="4" pageNumber="3" year="1913">
<bibRefCitation id="EFEDCDABFFF3FFF6FDAECC193C0E646E" author="Schmidt A." box="[624,813,351,375]" pageId="4" pageNumber="3" pagination="117 - 178" refId="ref5655" refString="Schmidt A. 1913. Erster Versuch einer Einteilung der exotischen Aphodien in Subgenera und als Anhang einige Neubeschreibungen. Archiv fur Naturgeschichte, Abteilung A 79: 117 - 178." type="journal article" year="1913">Schmidt 1913: 123</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph id="8BC3B05AFFF3FFF6FF3ECC383C3C648F" blockId="4.[224,1440,290,437]" box="[224,799,382,406]" pageId="4" pageNumber="3">
<treatmentCitationGroup id="AB6C9774FFF3FFF6FF3ECC383C3C648F" box="[224,799,382,406]" pageId="4" pageNumber="3">
<taxonomicName id="4C7CCBD9FFF3FFF6FF3ECC383D21648F" authority="(Harold)" baseAuthorityName="Harold" box="[224,514,382,406]" class="Insecta" family="Aphodiidae" genus="Agrilinus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="lansbergei">
<emphasis id="B9086C48FFF3FFF6FF3ECC383E87648F" box="[224,420,382,406]" italics="true" pageId="4" pageNumber="3">Agrilinus lansbergei</emphasis>
(Harold)
</taxonomicName>
~
<treatmentCitation id="0ADD964BFFF3FFF6FDC3CC383C38648F" author="Dellacasa M &amp; Gordon RD &amp; Dellacasa G." box="[541,795,382,406]" page="161" pageId="4" pageNumber="3" year="2002">
<bibRefCitation id="EFEDCDABFFF3FFF6FDC3CC383C38648F" author="Dellacasa M &amp; Gordon RD &amp; Dellacasa G." box="[541,795,382,406]" pageId="4" pageNumber="3" pagination="155 - 223" refId="ref5221" refString="Dellacasa M, Gordon RD, Dellacasa G. 2002. Aphodiinae described or recorded by Bates in Biologia Centrali-Americana (Coleoptera Scarabaeoidea: Aphodiidae). Acta Zoologica Mexicana 86: 155 - 223." type="journal article" year="2002">Dellacasa et al. 2002: 161</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph id="8BC3B05AFFF3FFF6FF3ECCDA3C4C64AC" blockId="4.[224,1440,290,437]" box="[224,879,412,437]" pageId="4" pageNumber="3">
<treatmentCitationGroup id="AB6C9774FFF3FFF6FF3ECCDA3C4C64AC" box="[224,879,412,437]" pageId="4" pageNumber="3">
<taxonomicName id="4C7CCBD9FFF3FFF6FF3ECCDA3D0364AD" authority="(Harold)" baseAuthorityName="Harold" baseAuthorityYear="1874" box="[224,544,412,436]" class="Insecta" family="Aphodiidae" genus="Haroldiellus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="lansbergei">
<emphasis id="B9086C48FFF3FFF6FF3ECCDA3EE264AD" box="[224,449,412,436]" italics="true" pageId="4" pageNumber="3">Haroldiellus lansbergei</emphasis>
(Harold)
</taxonomicName>
~
<treatmentCitation id="0ADD964BFFF3FFF6FDE4CCDA3C4864AC" author="Gordon RD &amp; Skelley PE" box="[570,875,412,437]" page="270" pageId="4" pageNumber="3" year="2007">
<bibRefCitation id="EFEDCDABFFF3FFF6FDE4CCDA3C4864AC" author="Gordon RD &amp; Skelley PE" box="[570,875,412,437]" pageId="4" pageNumber="3" pagination="1 - 580" refId="ref5257" refString="Gordon RD, Skelley PE. 2007. A monograph of the Aphodiini inhabiting the United States and Canada (Coleoptera: Scarabaeidae: Aphodiinae). Memoirs of the American Entomological Institute (Gainesville) 79: i - iv + 1 - 580." type="journal article" year="2007">Gordon and Skelley 2007: 270</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
</subSubSection>
<subSubSection id="C366E3D1FFF3FFF6FF1ECC823EC7675D" pageId="4" pageNumber="3" type="diagnosis">
<paragraph id="8BC3B05AFFF3FFF6FF1ECC823EC7675D" blockId="4.[192,1441,452,580]" pageId="4" pageNumber="3">
<emphasis id="B9086C48FFF3FFF6FF1ECC823E1F64C7" bold="true" box="[192,316,452,478]" pageId="4" pageNumber="3">Diagnosis.</emphasis>
Length 4.35.0 mm, width
<quantity id="4C841DBFFFF3FFF6FDABCC833DDD64C6" box="[629,766,453,479]" metricMagnitude="-3" metricUnit="m" metricValue="2.15" metricValueMax="2.4" metricValueMin="1.9" pageId="4" pageNumber="3" unit="mm" value="2.15" valueMax="2.4" valueMin="1.9">1.92.4 mm</quantity>
. A member of
<taxonomicName id="4C7CCBD9FFF3FFF6FC79CC823B0F64C7" authorityName="Gordon and Skelley" authorityYear="2007" box="[935,1068,452,478]" class="Insecta" family="Aphodiidae" genus="Haroldiellus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B9086C48FFF3FFF6FC79CC823B0F64C7" box="[935,1068,452,478]" italics="true" pageId="4" pageNumber="3">Haroldiellus</emphasis>
</taxonomicName>
distinguished from other species by: Elytral striae weakly impressed, intervals weakly convex, interval color pale, lacking distinct markings (
<figureCitation id="1347ACDFFFF3FFF6FAABCCA03FF2673B" captionStart="Figures 15" captionStartId="8.[192,279,1279,1305]" captionTargetBox="[192,1440,213,1249]" captionTargetId="figure-66@8.[192,1440,213,1249]" captionTargetPageId="8" captionText="Figures 15. Haroldiellus species. 1) H. sallei dorsal habitus. 2) H. lansbergei dorsal habitus. 3) H. woodruffi, holotype, dorsal habitus. 4) H. sallei protibia dorsal surface with punctures. 5) H. woodruffi protibia dorsal surface without punctures. Scale line = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6391957" httpUri="https://zenodo.org/record/6391957/files/figure.png" pageId="4" pageNumber="3">Fig. 2</figureCitation>
). Female lacking median frontal tubercle; male median frontal tubercle transverse (
<figureCitation id="1347ACDFFFF3FFF6FBB1CF4E3B92673A" box="[1135,1201,520,547]" captionStart="Figures 69" captionStartId="9.[195,282,992,1018]" captionTargetBox="[192,1440,213,962]" captionTargetId="figure-11@9.[192,1440,213,962]" captionTargetPageId="9" captionText="Figures 69. Heads of Haroldiellus species. 6) H. sallei. 7) H. lansbergei. 8) H. woodruffi female allotype. 9) H. woodruffi male holotype. Scale line = 0.5 mm (all figures)." figureDoi="http://doi.org/10.5281/zenodo.6391959" httpUri="https://zenodo.org/record/6391959/files/figure.png" pageId="4" pageNumber="3">Fig. 7</figureCitation>
). Protibia with dorsal surface lacking punctures.
</paragraph>
</subSubSection>
<subSubSection id="C366E3D1FFF3FFF6FF1ECF153A696789" pageId="4" pageNumber="3" type="distribution">
<paragraph id="8BC3B05AFFF3FFF6FF1ECF153A696789" blockId="4.[192,1441,595,656]" pageId="4" pageNumber="3">
<emphasis id="B9086C48FFF3FFF6FF1ECF153E796774" bold="true" box="[192,346,595,621]" pageId="4" pageNumber="3">Distribution.</emphasis>
(
<figureCitation id="1347ACDFFFF3FFF6FEB4CF123E986777" box="[362,443,596,622]" captionStart="Figure 10" captionStartId="10.[192,268,866,892]" captionTargetBox="[192,1440,213,836]" captionTargetId="figure-86@10.[192,1440,213,836]" captionTargetPageId="10" captionText="Figure 10. Distribution map of Haroldiellus lansbergei (blue) as currently known from specimen labels and literature records. Chiapas, Mexico is striped because it is a literature record that may represent the new species H. woodruffi described herein. Map made with SimpleMappr (Shorthouse 2010)." figureDoi="http://doi.org/10.5281/zenodo.6391963" httpUri="https://zenodo.org/record/6391963/files/figure.png" pageId="4" pageNumber="3">Fig. 10</figureCitation>
).
<emphasis id="B9086C48FFF3FFF6FE0FCF153D056774" bold="true" box="[465,550,595,621]" pageId="4" pageNumber="3">
<collectingCountry id="F36BF0CAFFF3FFF6FE0FCF153D056774" box="[465,550,595,621]" name="Mexico" pageId="4" pageNumber="3">Mexico</collectingCountry>
</emphasis>
(
<collectingRegion id="49B87EB8FFF3FFF6FDEACF123DB16777" box="[564,658,596,622]" country="Mexico" name="Chiapas" pageId="4" pageNumber="3">Chiapas</collectingRegion>
,
<collectingRegion id="49B87EB8FFF3FFF6FD40CF123DD96777" box="[670,762,596,622]" country="Mexico" name="Hidalgo" pageId="4" pageNumber="3">Hidalgo</collectingRegion>
,
<collectingRegion id="49B87EB8FFF3FFF6FCDBCF123CA76777" box="[773,900,596,622]" country="Mexico" name="Michoacan de Ocampo" pageId="4" pageNumber="3">Michoacán</collectingRegion>
,
<collectingRegion id="49B87EB8FFF3FFF6FC4ECF123CC66777" box="[912,997,596,622]" country="Mexico" name="Oaxaca" pageId="4" pageNumber="3">Oaxaca</collectingRegion>
,
<collectingRegion id="49B87EB8FFF3FFF6FC2FCF123B466777" box="[1009,1125,596,622]" country="Mexico" name="Queretaro de Arteaga" pageId="4" pageNumber="3">Querétaro</collectingRegion>
,
<collectingRegion id="49B87EB8FFF3FFF6FBAFCF123A3C6777" box="[1137,1311,596,622]" country="Mexico" name="San Luis Potosi" pageId="4" pageNumber="3">San Luis Potosí</collectingRegion>
,
<collectingRegion id="49B87EB8FFF3FFF6FAF5CF123AB66777" box="[1323,1429,596,622]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="4" pageNumber="3">Veracruz</collectingRegion>
). Records here are repeated from
<bibRefCitation id="EFEDCDABFFF3FFF6FDFBCF333C2A6789" author="Dellacasa M &amp; Gordon RD &amp; Dellacasa G." box="[549,777,629,656]" pageId="4" pageNumber="3" pagination="155 - 223" refId="ref5221" refString="Dellacasa M, Gordon RD, Dellacasa G. 2002. Aphodiinae described or recorded by Bates in Biologia Centrali-Americana (Coleoptera Scarabaeoidea: Aphodiidae). Acta Zoologica Mexicana 86: 155 - 223." type="journal article" year="2002">Dellacasa et al. 2002</bibRefCitation>
;
<bibRefCitation id="EFEDCDABFFF3FFF6FCCDCF333CF96789" author="Skelley PE &amp; Dellacasa M &amp; Dellacasa G &amp; Gordon RD" box="[787,986,629,656]" pageId="4" pageNumber="3" pagination="1 - 14" refId="ref5812" refString="Skelley PE, Dellacasa M, Dellacasa G, Gordon RD. 2007. Checklist of the Aphodiini of Mexico, Central and South America (Coleoptera: Scarabaeidae: Aphodiinae). Insecta Mundi 0014: 1 - 14." type="book chapter" year="2007">Skelley et al. 2007</bibRefCitation>
;
<bibRefCitation id="EFEDCDABFFF3FFF6FC3BCF333B976789" author="Halffter G &amp; Verdu JR &amp; Moreno CE &amp; Halffter V." box="[997,1204,629,656]" pageId="4" pageNumber="3" pagination="54 - 60" refId="ref5303" refString="Halffter G, Verdu JR, Moreno CE, Halffter V. 2012. Historical and ecological determinants of dung beetle assemblages in two arid zones of central Mexico. Journal of Arid Environments 76: 54 - 60." type="journal article" year="2012">Halffter et al. 2012</bibRefCitation>
;
<bibRefCitation id="EFEDCDABFFF3FFF6FB61CF333A656789" author="Minor P." box="[1215,1350,629,656]" pageId="4" pageNumber="3" pagination="493 - 514" refId="ref5476" refString="Minor P. 2017. New state records and updated checklist of Aphodiini and Eupariini (Coleoptera: Scarabaeidae: Aphodiinae) from Mexico. Zootaxa 4244 (4): 493 - 514." type="journal article" year="2017">Minor 2017</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C366E3D1FFF3FFF6FF1ECFD93C6D6115" pageId="4" pageNumber="3" type="materials_examined">
<paragraph id="8BC3B05AFFF3FFF6FF1ECFD93C6D6115" blockId="4.[191,1441,671,1036]" pageId="4" pageNumber="3">
<emphasis id="B9086C48FFF3FFF6FF1ECFD93E8767A0" bold="true" box="[192,420,671,697]" pageId="4" pageNumber="3">Material examined.</emphasis>
<collectingCountry id="F36BF0CAFFF3FFF6FE72CFD93D3F67A0" box="[428,540,671,697]" name="Mexico" pageId="4" pageNumber="3">MEXICO</collectingCountry>
:
<collectingRegion id="49B87EB8FFF3FFF6FDFECFD93D5967A0" box="[544,634,671,697]" country="Mexico" name="Chiapas" pageId="4" pageNumber="3">Chiapas</collectingRegion>
/
<quantity id="4C841DBFFFF3FFF6FD54CFE63DF667A0" box="[650,725,671,698]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="3" unit="km" value="10.0">10 km</quantity>
. e. LaTrinitaria/
<date id="FFC2969AFFF3FFF6FC4DCFE63B6967A0" box="[915,1098,671,698]" pageId="4" pageNumber="3" value="1967-08-14">August 14, 1967</date>
/ under dung/ W.W. Gibson // TAMU-ENTO/ X0081443/ Barcode //
<taxonomicName id="4C7CCBD9FFF3FFF6FDAACF873DDB67C2" authorityName="Gordon and Skelley" authorityYear="2007" box="[628,760,705,731]" class="Insecta" family="Aphodiidae" genus="Haroldiellus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B9086C48FFF3FFF6FDAACF873DDB67C2" box="[628,760,705,731]" italics="true" pageId="4" pageNumber="3">Haroldiellus</emphasis>
</taxonomicName>
/
<taxonomicName id="4C7CCBD9FFF3FFF6FCD6CF873C5567C2" baseAuthorityName="Harold" baseAuthorityYear="1874" box="[776,886,705,731]" class="Insecta" family="Aphodiidae" genus="Haroldiellus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="lansbergei">
<emphasis id="B9086C48FFF3FFF6FCD6CF873C5567C2" box="[776,886,705,731]" italics="true" pageId="4" pageNumber="3">lansbergei</emphasis>
</taxonomicName>
/ det.P.E.Skelley 2021 (1
<collectionCode id="ED6D289FFFF3FFF6FB57CF873BFD67C2" box="[1161,1246,705,731]" pageId="4" pageNumber="3">TAMU</collectionCode>
);
<collectingCountry id="F36BF0CAFFF3FFF6FB2FCF873A4367C2" box="[1265,1376,705,731]" name="Mexico" pageId="4" pageNumber="3">MEXICO</collectingCountry>
: State of/
<collectingRegion id="49B87EB8FFF3FFF6FF38CFA53E6167E4" box="[230,322,739,765]" country="Mexico" name="Hidalgo" pageId="4" pageNumber="3">Hidalgo</collectingRegion>
, 1Km.N / Jacala, 5000Ft. // R.E.Woodruff/ 24-X-70/ Cow Dung (4
<collectionCode id="ED6D289FFFF3FFF6FBC3CFA53B4767E4" box="[1053,1124,739,765]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="4" pageNumber="3" type="Museum">FSCA</collectionCode>
);
<collectingCountry id="F36BF0CAFFF3FFF6FBABCFA53BC767E4" box="[1141,1252,739,765]" name="Mexico" pageId="4" pageNumber="3">MEXICO</collectingCountry>
:
<collectingRegion id="49B87EB8FFF3FFF6FB33CFA53A6B67E4" box="[1261,1352,739,765]" country="Mexico" name="Hidalgo" pageId="4" pageNumber="3">Hidalgo</collectingRegion>
/ Puerto de la Zora/
<date id="FFC2969AFFF3FFF6FE9DCE433E9F6606" box="[323,444,773,799]" pageId="4" pageNumber="3" value="1970-10-24">24-X-1970</date>
, light/ R. E.Woodruff (1
<collectionCode id="ED6D289FFFF3FFF6FD0BCE433C3F6606" box="[725,796,773,799]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="4" pageNumber="3" type="Museum">FSCA</collectionCode>
); E. DO
<collectingRegion id="49B87EB8FFF3FFF6FCA0CE433B0A6606" box="[894,1065,773,799]" country="Mexico" name="Michoacan de Ocampo" pageId="4" pageNumber="3">MICHOACAN</collectingRegion>
/ rd. Méx-37 Uruapan / Nueva
<collectingCountry id="F36BF0CAFFF3FFF6FF1FCE613FD96658" box="[193,250,807,833]" name="Italy" pageId="4" pageNumber="3">Italia</collectingCountry>
/ La Gallina dint. // m 1250-
<date id="FFC2969AFFF3FFF6FDE5CE603D946658" box="[571,695,806,833]" pageId="4" pageNumber="3" value="2001-07-23">23.07.2001</date>
/ N19°19 -W102°04 / M. Dellacasa &amp; / I. Martinez leg. (1
<collectionCode id="ED6D289FFFF3FFF6FA8ECE613AB76658" box="[1360,1428,807,833]" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34935" lsid="urn:lsid:biocol.org:col:34935" name="Canadian Museum of Nature" pageId="4" pageNumber="3" type="Museum">CMN</collectionCode>
);
<collectingCountry id="F36BF0CAFFF3FFF6FF1FCE0F3E13667A" box="[193,304,841,867]" name="Mexico" pageId="4" pageNumber="3">MEXICO</collectingCountry>
:
<collectingRegion id="49B87EB8FFF3FFF6FEE2CE0F3E92667A" box="[316,433,841,867]" country="Mexico" name="Queretaro de Arteaga" pageId="4" pageNumber="3">Querétaro</collectingRegion>
; Mpio:/ Jalpan; nr.Valle Verde,/ Bio.St.El Pilon/
<geoCoordinate id="EE48D69DFFF3FFF6FC0BCE0F3B7B667A" box="[981,1112,841,869]" degrees="21" direction="north" minutes="27" orientation="latitude" pageId="4" pageNumber="3" precision="15" seconds="41" value="21.46139">21°2741″N</geoCoordinate>
,
<geoCoordinate id="EE48D69DFFF3FFF6FBBBCE0F3BCE667A" box="[1125,1261,841,869]" degrees="99" direction="west" minutes="10" orientation="longitude" pageId="4" pageNumber="3" precision="15" seconds="41" value="-99.178055">99°1041″W</geoCoordinate>
/
<date id="FFC2969AFFF3FFF6FB20CE0F3ABF667A" box="[1278,1436,841,867]" pageId="4" pageNumber="3" value="2013-12-11">11-DEC-2013</date>
, 1284m/ P.Skelley, P.Kovarik, R. Jones (1
<collectionCode id="ED6D289FFFF3FFF6FDA7CE2C3DE3669D" box="[633,704,874,900]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="4" pageNumber="3" type="Museum">FSCA</collectionCode>
); Rancho Pextlán/ (Xico)/
<geoCoordinate id="EE48D69DFFF3FFF6FC3ACE2C3B5A669D" box="[996,1145,874,903]" degrees="19" direction="north" minutes="25" orientation="latitude" pageId="4" pageNumber="3" precision="1" seconds="28.9" value="19.424694">19°2528.9″N</geoCoordinate>
/
<geoCoordinate id="EE48D69DFFF3FFF6FB56CE2C3A3B669D" box="[1160,1304,874,903]" degrees="97" direction="west" minutes="01" orientation="longitude" pageId="4" pageNumber="3" precision="1" seconds="43.0" value="-97.02862">97°0143.0W</geoCoordinate>
/
<date id="FFC2969AFFF3FFF6FAF8CE2C3A83669C" box="[1318,1440,874,901]" pageId="4" pageNumber="3" value="2005-03-19">19.III.2005</date>
- m 1465 //
<collectingCountry id="F36BF0CAFFF3FFF6FE94CECA3E9466BF" box="[330,439,908,934]" name="Mexico" pageId="4" pageNumber="3">MEXICO</collectingCountry>
/ Edo.
<collectingRegion id="49B87EB8FFF3FFF6FDDDCECA3D4A66BF" box="[515,617,908,934]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="4" pageNumber="3">Veracruz</collectingRegion>
/ Leg. González O. &amp;/ Suarez T./
<emphasis id="B9086C48FFF3FFF6FC3ACECB3B4A66BF" box="[996,1129,908,934]" italics="true" pageId="4" pageNumber="3">excr. bovino</emphasis>
//
<taxonomicName id="4C7CCBD9FFF3FFF6FB55CECA3A3666BF" authorityName="Gordon and Skelley" authorityYear="2007" box="[1163,1301,908,934]" class="Insecta" family="Aphodiidae" genus="Haroldiellus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus">Haroldiellus</taxonomicName>
/ lansbergei/ (
<bibRefCitation id="EFEDCDABFFF3FFF6FF14CEE83E4666D1" author="Harold E." box="[202,357,942,968]" pageId="4" pageNumber="3" pagination="177 - 208" refId="ref5369" refString="Harold E. 1874. Beitrage zur Kenntnis einiger coprophagen Lamellicornien. (Achtes Stuck). Berliner Entomologische Zeitschrift 18: 177 - 208." type="journal article" year="1874">Harold, 1874</bibRefCitation>
)/ Det. Dellacasa, 2011 (1
<collectionCode id="ED6D289FFFF3FFF6FD41CEE83DC566D1" box="[671,742,942,968]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="4" pageNumber="3" type="Museum">FSCA</collectionCode>
); Rancho Pextlan/ Mpio. Xico/
<geoCoordinate id="EE48D69DFFF3FFF6FB81CEE83BD166D1" box="[1119,1266,942,970]" degrees="19" direction="north" minutes="25" orientation="latitude" pageId="4" pageNumber="3" precision="1" seconds="28.9" value="19.424694">19°2528.9″N</geoCoordinate>
/
<geoCoordinate id="EE48D69DFFF3FFF6FAD9CEE83AB466D1" box="[1287,1431,942,970]" degrees="97" direction="west" minutes="01" orientation="longitude" pageId="4" pageNumber="3" precision="1" seconds="43.0" value="-97.02862">97°0143.0W</geoCoordinate>
/
<date id="FFC2969AFFF3FFF6FF1ECE963E1966F2" box="[192,314,976,1003]" pageId="4" pageNumber="3" value="2005-09-12">12.IX.2005</date>
- m 1465 //
<collectingCountry id="F36BF0CAFFF3FFF6FE1ECE963D0E66F3" box="[448,557,976,1002]" name="Mexico" pageId="4" pageNumber="3">MEXICO</collectingCountry>
/ Edo.
<collectingRegion id="49B87EB8FFF3FFF6FDADCE963DFA66F3" box="[627,729,976,1002]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="4" pageNumber="3">Veracruz</collectingRegion>
/ Leg. Cruz M. Cabrero F. &amp; Suarez T./
<emphasis id="B9086C48FFF3FFF6FB5BCE973BCC66F3" box="[1157,1263,977,1002]" italics="true" pageId="4" pageNumber="3">excr. vaca</emphasis>
//
<taxonomicName id="4C7CCBD9FFF3FFF6FAD0CE963ABB66F3" authorityName="Gordon and Skelley" authorityYear="2007" box="[1294,1432,976,1002]" class="Insecta" family="Aphodiidae" genus="Haroldiellus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus">Haroldiellus</taxonomicName>
/ lansbergei/ (
<bibRefCitation id="EFEDCDABFFF3FFF6FE96CEB43EFD6115" author="Harold E." box="[328,478,1010,1036]" pageId="4" pageNumber="3" pagination="177 - 208" refId="ref5369" refString="Harold E. 1874. Beitrage zur Kenntnis einiger coprophagen Lamellicornien. (Achtes Stuck). Berliner Entomologische Zeitschrift 18: 177 - 208." type="journal article" year="1874">Harold, 1874</bibRefCitation>
)/ Det. Dellacasa, 2011 (1
<collectionCode id="ED6D289FFFF3FFF6FD25CEB43C616115" box="[763,834,1010,1036]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="4" pageNumber="3" type="Museum">FSCA</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C366E3D1FFF3FFF6FF1EC95A3D9A61A4" pageId="4" pageNumber="3" type="discussion">
<paragraph id="8BC3B05AFFF3FFF6FF1EC95A3D9A61A4" blockId="4.[191,1442,1051,1213]" pageId="4" pageNumber="3">
<emphasis id="B9086C48FFF3FFF6FF1EC95A3E0C612F" bold="true" box="[192,303,1052,1078]" pageId="4" pageNumber="3">Remarks.</emphasis>
The specimen from
<collectingRegion id="49B87EB8FFF3FFF6FDC5C95A3DAD612F" box="[539,654,1052,1078]" country="Mexico" name="Queretaro de Arteaga" pageId="4" pageNumber="3">Querétaro</collectingRegion>
is slightly larger than other specimens presently available. It also has the discal intervals of the elytra darkened, appearing as a single dark mark. Other morphological features like the more flattened intervals and lack of protibial punctures indicate it is
<taxonomicName id="4C7CCBD9FFF3FFF6FC1BC9263B766160" baseAuthorityName="Harold" baseAuthorityYear="1874" box="[965,1109,1119,1146]" class="Insecta" family="Aphodiidae" genus="Haroldiellus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="lansbergei">
<emphasis id="B9086C48FFF3FFF6FC1BC9263CFA6160" box="[965,985,1120,1145]" italics="true" pageId="4" pageNumber="3">H</emphasis>
.
<emphasis id="B9086C48FFF3FFF6FC39C9193B766160" box="[999,1109,1119,1145]" italics="true" pageId="4" pageNumber="3">lansbergei</emphasis>
</taxonomicName>
as presently understood. The
<collectingRegion id="49B87EB8FFF3FFF6FF1FC9C73E386182" box="[193,283,1153,1179]" country="Mexico" name="Chiapas" pageId="4" pageNumber="3">Chiapas</collectingRegion>
(
<collectingCountry id="F36BF0CAFFF3FFF6FEF7C9C73EA36185" box="[297,384,1153,1180]" name="Mexico" pageId="4" pageNumber="3">Mexico</collectingCountry>
) record published in
<bibRefCitation id="EFEDCDABFFF3FFF6FDB3C9C73C266182" author="Minor P." box="[621,773,1153,1179]" pageId="4" pageNumber="3" pagination="493 - 514" refId="ref5476" refString="Minor P. 2017. New state records and updated checklist of Aphodiini and Eupariini (Coleoptera: Scarabaeidae: Aphodiinae) from Mexico. Zootaxa 4244 (4): 493 - 514." type="journal article" year="2017">Minor (2017)</bibRefCitation>
represents most likely the new species described herein, but needs to be re-examined based on its locality.
</paragraph>
</subSubSection>
</treatment>
</document>