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<document id="FC11BF06866CB337EE13785F523D6DA4" ID-DOI="10.11646/zootaxa.5538.5.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="14239741" ID-ZooBank="6AF05486-CCAC-4FB3-9D72-7A5D4FDB66D8" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1732829771073" checkinUser="plazi" docAuthor="Golovatch, Sergei I., Korotayeva, Alyona M. &amp; Martínez-Torres, Daniela" docDate="2024" docId="03CB3B556A036C05FF466F01A4E348D7" docLanguage="en" docName="zootaxa.5538.5.2.pdf" docOrigin="Zootaxa 5538 (5)" docSource="http://dx.doi.org/10.11646/zootaxa.5360.3.8" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nyssodesmus python" docType="treatment" docVersion="4" lastPageNumber="436" masterDocId="FFF2432D6A086C09FFD16A01A7704D18" masterDocTitle="Three species of the millipede family Platyrhacidae from Costa Rica: a new Barydesmus Cook, 1896 and two oldest Nyssodesmus Cook, 1896 (Diplopoda Polydesmida)" masterLastPageNumber="438" masterPageNumber="424" pageNumber="435" updateTime="1733244892161" updateUser="felipe" zenodo-license-document="CLOSED">
<mods:mods id="0AADBC153B1052FFBD97F515F3F9CDC3" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="098F6E2516EEFBBF184D36C4956A9450">
<mods:title id="DA0A38E5A83A50DB81D1643ECFB349E1">Three species of the millipede family Platyrhacidae from Costa Rica: a new Barydesmus Cook, 1896 and two oldest Nyssodesmus Cook, 1896 (Diplopoda Polydesmida)</mods:title>
</mods:titleInfo>
<mods:name id="7295DDD193CAFADD842E61959CC4CB2A" type="personal">
<mods:role id="3BBF642970AFEEA0E6172D21F5366A1C">
<mods:roleTerm id="2BB29E307200975AA01852043260FC5A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="593D3E019FAF0584D9B4BD7A24C1CADF">Golovatch, Sergei I.</mods:namePart>
<mods:affiliation id="92077B09D83BA0632CE8C6229C29ABD9">Institute of Ecology and Evolution, Russian Academy of Sciences, Moscow, Russia</mods:affiliation>
</mods:name>
<mods:name id="6F21A75B7667005311AA36A7946FFBD3" type="personal">
<mods:role id="FA18FA6EFF45881E1E5E375E1697DFB7">
<mods:roleTerm id="343144F6AE300BEBBD6504C123B8354D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B88737EE0D9654A9132D7AF0AC11DC2B">Korotayeva, Alyona M.</mods:namePart>
<mods:nameIdentifier id="411467040CA396F4136CA08DAA028D48" type="ORCID">0009-0009-2716-5140</mods:nameIdentifier>
<mods:affiliation id="0E0234F02B3004D42F7C85A0F5EF4BAC">Institute of Ecology and Evolution, Russian Academy of Sciences, Moscow, Russia</mods:affiliation>
<mods:nameIdentifier id="0DB46678668579E29AF3FD5C4D4394DC" type="email">alenacorotaeva2017@mail.ru</mods:nameIdentifier>
</mods:name>
<mods:name id="2B652F298AD3E1A5DEE2F3F15404DC0E" type="personal">
<mods:role id="596028F36E7D603C577663C50DC1873C">
<mods:roleTerm id="C313CE44658275A142CC37B667C5999F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="129ABC53ECAE4C223B39BA4EF3DA7335">Martínez-Torres, Daniela</mods:namePart>
<mods:nameIdentifier id="DC7A39A0256122D3525F8C02E000914B" type="ORCID">0000-0001-8865-927X</mods:nameIdentifier>
<mods:affiliation id="139D3AEF3C144D1985C47CEC8CEA0839">Grupo de Investigación en Aracnología y Miriapodología (GAM-UN), Instituto de Ciencias Naturales, Universidad Nacional de Colombia, Edificio 425, Oficina 105, Bogotá, Colombia; Departamento de Biología, Facultad de Ciencias, Universidad Nacional de Colombia, Bogotá, Colombia</mods:affiliation>
<mods:nameIdentifier id="DED9673C3CE2909BF0CD1087E22882C6" type="email">sdmartinezt@unal.edu.co</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="4891696D2AC2C31FF34EE475A43FEFE5">text</mods:typeOfResource>
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<mods:title id="C5561BAE4F742754AE4E9FD8DA12F4E0">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="09E2CAC87AC590DCD2EAB9B052FAB954">
<mods:date id="A07B91B7DA2DD6B19740D2AA81B50A60">2024</mods:date>
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<mods:number id="BA8D6634907B25755C9A602D88A827DF">2024-11-19</mods:number>
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<mods:number id="050DADBEE7911A96DC82E203AA43A1CD">5538</mods:number>
</mods:detail>
<mods:detail id="E52F78595D629915A9A9B76DCE37565C" type="issue">
<mods:number id="B1060CF162EB404A369F6DB0D56E74A0">5</mods:number>
</mods:detail>
<mods:extent id="609F85E5166FFA45EE414F9EF835B2F8" unit="page">
<mods:start id="88347352CDCA34342B9C5FB550F8631D">424</mods:start>
<mods:end id="A18646A7A218ECA45CA8C8FBA52885B3">438</mods:end>
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<mods:url id="21D8B34DA4723C1966558B383DE118BE">http://dx.doi.org/10.11646/zootaxa.5360.3.8</mods:url>
</mods:location>
<mods:classification id="40FBC4802A2B96B2A512FAF49A07A44B">journal article</mods:classification>
<mods:identifier id="231A3252E6ED258CB82D6DBCFA4A7680" type="DOI">10.11646/zootaxa.5538.5.2</mods:identifier>
<mods:identifier id="86D7CCD1E8F785DE33BC814DCFDFB291" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="F89DBD434FD0339DC67474F8DE294F46" type="Zenodo-Dep">14239741</mods:identifier>
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<treatment id="03CB3B556A036C05FF466F01A4E348D7" ID-DOI="http://doi.org/10.5281/zenodo.14248171" ID-Zenodo-Dep="14248171" LSID="urn:lsid:plazi:treatment:03CB3B556A036C05FF466F01A4E348D7" httpUri="http://treatment.plazi.org/id/03CB3B556A036C05FF466F01A4E348D7" lastPageId="12" lastPageNumber="436" pageId="11" pageNumber="435">
<subSubSection id="C378D9C86A036C02FF466F01A51B4803" box="[151,619,1280,1307]" pageId="11" pageNumber="435" type="nomenclature">
<paragraph id="8BDD8A436A036C02FF466F01A51B4803" blockId="11.[151,619,1280,1307]" box="[151,619,1280,1307]" pageId="11" pageNumber="435">
<heading id="D0953D2F6A036C02FF466F01A51B4803" bold="true" box="[151,619,1280,1307]" fontSize="11" level="1" pageId="11" pageNumber="435" reason="1">
<taxonomicName id="4C62F1C06A036C02FF466F01A51B4803" ID-CoL="487SN" authority="(Peters, 1865)" baseAuthorityName="Peters" baseAuthorityYear="1865" box="[151,619,1280,1307]" class="Diplopoda" family="Platyrhacidae" genus="Nyssodesmus" isUncertain="true" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="435" phylum="Arthropoda" rank="species" species="python">
<emphasis id="B91656516A036C02FF466F01A51B4803" bold="true" box="[151,619,1280,1307]" pageId="11" pageNumber="435">
<emphasis id="B91656516A036C02FF466F01A6484803" bold="true" box="[151,312,1280,1307]" italics="true" pageId="11" pageNumber="435">Nyssodesmus</emphasis>
cf.
<emphasis id="B91656516A036C02FEB46F01A6C84803" bold="true" box="[357,440,1280,1307]" italics="true" pageId="11" pageNumber="435">python</emphasis>
(
<bibRefCitation id="EFF3F7B26A036C02FE196F00A5124803" author="Peters, W." box="[456,610,1281,1307]" pageId="11" pageNumber="435" pagination="529 - 551" refId="ref7857" refString="Peters, W. (1865) Ubersicht der im Konigl. zoologischen Museum befindlichen Myriopoden aus der Familie der Polydesmi, so wie Beschreibungen einer neuen Gattung, Trachyjulus, der Juli und neuer Arten der Gattung Siphonophora. Monatsberichte der Koniglichen Preussischen Akademie der Wissenschaften zu Berlin, 1864, 529 - 551." type="journal article" year="1865">Peters, 1865</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C378D9C86A036C02FF466F48A4EE4A44" pageId="11" pageNumber="435" type="materials_examined">
<paragraph id="8BDD8A436A036C02FF466F48A4EE4A44" blockId="11.[151,1437,1353,2028]" pageId="11" pageNumber="435">
<emphasis id="B91656516A036C02FF466F48A6F3487B" bold="true" box="[151,387,1353,1379]" pageId="11" pageNumber="435">Material examined.</emphasis>
<specimenCount id="9D6441CA6A036C02FE5A6F48A6AB487B" box="[395,475,1353,1379]" count="1" pageId="11" pageNumber="435" type="male">1 male</specimenCount>
,
<specimenCount id="9D6441CA6A036C02FE386F48A53C487B" box="[489,588,1353,1379]" count="1" pageId="11" pageNumber="435" type="female">1 female</specimenCount>
(
<collectionCode id="ED7312866A036C02FD8C6F48A5D7487B" box="[605,679,1353,1379]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="11" pageNumber="435" type="Museum">FSCA</collectionCode>
,
<specimenCode id="DBC422386A036C02FD656F4BA458487B" box="[692,808,1353,1380]" collectionCode="DMT" pageId="11" pageNumber="435">DMT 134</specimenCode>
),
<collectingCountry id="F375CAD36A036C02FCEF6F48A4CD487B" box="[830,957,1353,1379]" name="Costa Rica" pageId="11" pageNumber="435">Costa Rica</collectingCountry>
,
<collectingRegion id="49A644A16A036C02FC1B6F48A3E2487B" box="[970,1170,1353,1379]" country="Costa Rica" name="Heredia" pageId="11" pageNumber="435">Heredia Province</collectingRegion>
,
<location id="8EBDDC986A036C02FB4E6F48A7A8489F" LSID="urn:lsid:plazi:treatment:03CB3B556A036C05FF466F01A4E348D7:8EBDDC986A036C02FB4E6F48A7A8489F" country="Costa Rica" latitude="10.416667" longLatPrecision="1" longitude="-84.0" name="Estacion Biologica La Selva" pageId="11" pageNumber="435" stateProvince="Heredia">Estación Biologica La Selva</location>
,
<geoCoordinate id="EE56EC846A036C02FF376F6CA6F0489F" box="[230,384,1389,1415]" degrees="10.416667" direction="north" orientation="latitude" pageId="11" pageNumber="435" precision="1" value="10.416667">10.416667°N</geoCoordinate>
, -
<geoCoordinate id="EE56EC846A036C02FE446F6CA544489F" box="[405,564,1389,1415]" degrees="84.000000" direction="west" orientation="longitude" pageId="11" pageNumber="435" precision="1" value="-84.0">84.000000°W</geoCoordinate>
,
<elevation id="004F6D706A036C02FD936F6CA585489F" box="[578,757,1389,1416]" metricMagnitude="2" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="11" pageNumber="435" unit="m" value="100.0" valueMax="150.0" valueMin="50.0">
<quantity id="4C9A27A66A036C02FD936F6CA5C64890" box="[578,694,1389,1416]" metricMagnitude="2" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="11" pageNumber="435" unit="m" value="100.0" valueMax="150.0" valueMin="50.0">50–150 m</quantity>
a.s.l.
</elevation>
, in copula, side of trail,
<date id="FFDCAC836A036C02FBC76F6CA3FD489F" box="[1046,1165,1389,1415]" pageId="11" pageNumber="435" value="2001-02-14">
<collectingDate id="EF98556B6A036C02FBC76F6CA3FD489F" box="[1046,1165,1389,1415]" pageId="11" pageNumber="435" value="2001-02-14">14.II.2001</collectingDate>
</date>
,
<collectorName id="2697EF956A036C02FB4D6F6CA24D489F" box="[1180,1341,1389,1415]" pageId="11" pageNumber="435">G.B. Edwards</collectorName>
leg.,
<determinerName id="7E0166876A036C02FA506F6FA63C48B3" pageId="11" pageNumber="435">D. Martínez-Torres</determinerName>
det., 2016;
<specimenCount id="9D6441CA6A036C02FE096F90A55848B3" box="[472,552,1425,1451]" count="1" pageId="11" pageNumber="435" type="male">1 male</specimenCount>
,
<specimenCount id="9D6441CA6A036C02FDE76F90A5E948B3" box="[566,665,1425,1451]" count="1" pageId="11" pageNumber="435" type="female">1 female</specimenCount>
(
<specimenCode id="DBC422386A036C02FD7B6F90A44A48B4" box="[682,826,1425,1452]" collectionCode="MCZ" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="11" pageNumber="435" type="Museum">MCZ 44105</specimenCode>
,
<specimenCode id="DBC422386A036C02FC966F93A4CC48B3" box="[839,956,1425,1452]" collectionCode="DMT" pageId="11" pageNumber="435">DMT 366</specimenCode>
),
<collectingRegion id="49A644A16A036C02FC036F90A3E948B3" box="[978,1177,1425,1451]" country="Costa Rica" name="Heredia" pageId="11" pageNumber="435">Heredia Province</collectingRegion>
, near
<collectingMunicipality id="6BB910396A036C02FB316F93A20248B3" box="[1248,1394,1425,1452]" pageId="11" pageNumber="435">Puerto Viejo</collectingMunicipality>
,
<location id="8EBDDC986A036C02FAAE6F93A7A848D7" LSID="urn:lsid:plazi:treatment:03CB3B556A036C05FF466F01A4E348D7:8EBDDC986A036C02FAAE6F93A7A848D7" country="Costa Rica" latitude="10.434769" longLatPrecision="1" longitude="-84.01891" municipality="Puerto Viejo" name="La Selva" pageId="11" pageNumber="435" stateProvince="Heredia">La Selva</location>
,
<geoCoordinate id="EE56EC846A036C02FF366FB4A6F048D7" box="[231,384,1461,1487]" degrees="10.434769" direction="north" orientation="latitude" pageId="11" pageNumber="435" precision="1" value="10.434769">10.434769°N</geoCoordinate>
, -
<geoCoordinate id="EE56EC846A036C02FE476FB4A54448D7" box="[406,564,1461,1487]" degrees="84.018914" direction="west" orientation="longitude" pageId="11" pageNumber="435" precision="1" value="-84.01891">84.018914°W</geoCoordinate>
,
<elevation id="004F6D706A036C02FD926FB4A5CF48D7" box="[579,703,1461,1488]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="11" pageNumber="435" unit="m" value="60.0">
<quantity id="4C9A27A66A036C02FD926FB4A50F48C8" box="[579,639,1461,1488]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="11" pageNumber="435" unit="m" value="60.0">60 m</quantity>
a.s.l.
</elevation>
,
<date id="FFDCAC836A036C02FD1E6FB4A42148D7" box="[719,849,1461,1487]" pageId="11" pageNumber="435" value="1979-06-21">
<collectingDate id="EF98556B6A036C02FD1E6FB4A42148D7" box="[719,849,1461,1487]" pageId="11" pageNumber="435" value="1979-06-21">21.VI.1979</collectingDate>
</date>
,
<collectorName id="2697EF956A036C02FC8E6FB7A37048D7" box="[863,1024,1461,1488]" pageId="11" pageNumber="435">J. Coddington</collectorName>
leg.,
<determinerName id="7E0166876A036C02FB956FB7A26F48D7" box="[1092,1311,1461,1488]" pageId="11" pageNumber="435">D. Martínez-Torres</determinerName>
det., 2015 (https://mczbase.mcz.harvard.edu/guid/
<collectionCode id="ED7312866A036C02FD9E6FD8A5FE48EB" box="[591,654,1497,1523]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="11" pageNumber="435" type="Museum">MCZ</collectionCode>
:IZ:44105);
<specimenCount id="9D6441CA6A036C02FCCB6FD8A41948EB" box="[794,873,1497,1523]" count="1" pageId="11" pageNumber="435" type="male">1 male</specimenCount>
(
<collectionCode id="ED7312866A036C02FCAD6FD8A49648EB" box="[892,998,1497,1523]" pageId="11" pageNumber="435">NCMNH</collectionCode>
<specimenCode id="DBC422386A036C02FC216FD8A34F48EC" box="[1008,1087,1497,1524]" collectionCode="NCMNH" pageId="11" pageNumber="435">A3657</specimenCode>
,
<specimenCode id="DBC422386A036C02FB9F6FDBA3C648EB" box="[1102,1206,1497,1524]" collectionCode="DMT" pageId="11" pageNumber="435">DMT 64</specimenCode>
),
<collectingRegion id="49A644A16A036C02FB1F6FD8A2E848EB" box="[1230,1432,1497,1523]" country="Costa Rica" name="Heredia" pageId="11" pageNumber="435">Heredia Province</collectingRegion>
,
<collectingMunicipality id="6BB910396A036C02FF466FFCA6444B0F" box="[151,308,1533,1559]" pageId="11" pageNumber="435">Rio Sarapiqui</collectingMunicipality>
, above road from
<location id="8EBDDC986A036C02FE2E6FFFA4464B0F" LSID="urn:lsid:plazi:treatment:03CB3B556A036C05FF466F01A4E348D7:8EBDDC986A036C02FE2E6FFFA4464B0F" box="[511,822,1533,1560]" country="Costa Rica" latitude="10.416667" longLatPrecision="1" longitude="-84.0" municipality="Rio Sarapiqui" name="La Selva Biological Station" pageId="11" pageNumber="435" stateProvince="Heredia">La Selva Biological Station</location>
,
<geoCoordinate id="EE56EC846A036C02FC906FFCA4AA4B0F" box="[833,986,1533,1559]" degrees="10.416667" direction="north" orientation="latitude" pageId="11" pageNumber="435" precision="1" value="10.416667">10.416667°N</geoCoordinate>
, -
<geoCoordinate id="EE56EC846A036C02FC3D6FFCA3FB4B0F" box="[1004,1163,1533,1559]" degrees="84.000000" direction="west" orientation="longitude" pageId="11" pageNumber="435" precision="1" value="-84.0">84.000000°W</geoCoordinate>
,
<elevation id="004F6D706A036C02FB476FFCA2694B0F" box="[1174,1305,1533,1560]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="435" unit="m" value="100.0">
<quantity id="4C9A27A66A036C02FB476FFCA3AD4B00" box="[1174,1245,1533,1560]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="435" unit="m" value="100.0">100 m</quantity>
a.s.l.
</elevation>
,
<date id="FFDCAC836A036C02FAF46FFCA2E74B0F" box="[1317,1431,1533,1559]" pageId="11" pageNumber="435" value="1981-03-09">
<collectingDate id="EF98556B6A036C02FAF46FFCA2E74B0F" box="[1317,1431,1533,1559]" pageId="11" pageNumber="435" value="1981-03-09">9.III.1981</collectingDate>
</date>
,
<collectorName id="2697EF956A036C02FF466C23A6534B23" box="[151,291,1569,1596]" pageId="11" pageNumber="435">R.E. Ashton</collectorName>
leg.,
<determinerName id="7E0166876A036C02FEB86C23A5344B23" box="[361,580,1569,1596]" pageId="11" pageNumber="435">D. Martínez-Torres</determinerName>
det., 2015;
<specimenCount id="9D6441CA6A036C02FD046C20A4574B23" box="[725,807,1569,1595]" count="1" pageId="11" pageNumber="435" type="male">1 male</specimenCount>
,
<specimenCount id="9D6441CA6A036C02FCE76C20A4EB4B23" box="[822,923,1569,1595]" count="1" pageId="11" pageNumber="435" type="female">1 female</specimenCount>
(
<specimenCode id="DBC422386A036C02FC7C6C20A3304B23" box="[941,1088,1569,1595]" collectionCode="MCZ" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="11" pageNumber="435" type="Museum">MCZ 41246</specimenCode>
,
<specimenCode id="DBC422386A036C02FB9F6C23A3B44B23" box="[1102,1220,1569,1596]" collectionCode="DMT" pageId="11" pageNumber="435">DMT 363</specimenCode>
),
<collectingRegion id="49A644A16A036C02FB0D6C20A2E84B23" box="[1244,1432,1569,1595]" country="Costa Rica" name="Limon" pageId="11" pageNumber="435">Limón Province</collectingRegion>
,
<location id="8EBDDC986A036C02FF466C44A78D4B47" LSID="urn:lsid:plazi:treatment:03CB3B556A036C05FF466F01A4E348D7:8EBDDC986A036C02FF466C44A78D4B47" box="[151,253,1605,1631]" country="Costa Rica" latitude="10.215878" longLatPrecision="1" longitude="-83.786606" name="Guapiles" pageId="11" pageNumber="435" stateProvince="Limon">Guapiles</location>
,
<geoCoordinate id="EE56EC846A036C02FED86C44A6C14B78" box="[265,433,1605,1632]" degrees="10.2158780" direction="north" orientation="latitude" pageId="11" pageNumber="435" precision="1" value="10.215878">10.2158780°N</geoCoordinate>
, -
<geoCoordinate id="EE56EC846A036C02FE126C44A5124B47" box="[451,610,1605,1631]" degrees="83.786609" direction="west" orientation="longitude" pageId="11" pageNumber="435" precision="1" value="-83.786606">83.786609°W</geoCoordinate>
,
<elevation id="004F6D706A036C02FDBF6C44A5824B47" box="[622,754,1605,1632]" metricMagnitude="2" metricUnit="m" metricValue="2.56" pageId="11" pageNumber="435" unit="m" value="256.0">
<quantity id="4C9A27A66A036C02FDBF6C44A5C64B78" box="[622,694,1605,1632]" metricMagnitude="2" metricUnit="m" metricValue="2.56" pageId="11" pageNumber="435" unit="m" value="256.0">256 m</quantity>
a.s.l.
</elevation>
,
<date id="FFDCAC836A036C02FD2E6C44A42D4B47" box="[767,861,1605,1631]" pageId="11" pageNumber="435" value="1924-03">
<collectingDate id="EF98556B6A036C02FD2E6C44A42D4B47" box="[767,861,1605,1631]" pageId="11" pageNumber="435" value="1924-03">III.1924</collectingDate>
</date>
,
<collectorName id="2697EF956A036C02FCB86C44A4B54B47" box="[873,965,1605,1631]" pageId="11" pageNumber="435">Barbour</collectorName>
and
<collectorName id="2697EF956A036C02FC2D6C44A33D4B47" box="[1020,1101,1605,1631]" pageId="11" pageNumber="435">Brooks</collectorName>
leg.,
<determinerName id="7E0166876A036C02FB5D6C47A2144B47" box="[1164,1380,1605,1632]" pageId="11" pageNumber="435">D. Martínez-Torres</determinerName>
det., 2016 (https://mczbase.mcz.harvard.edu/guid/
<collectionCode id="ED7312866A036C02FD416C68A5BF4B9B" box="[656,719,1641,1667]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="11" pageNumber="435" type="Museum">MCZ</collectionCode>
:IZ:41246);
<specimenCount id="9D6441CA6A036C02FC886C68A4D64B9B" box="[857,934,1641,1667]" count="1" pageId="11" pageNumber="435" type="male">1 male</specimenCount>
(
<specimenCode id="DBC422386A036C02FC666C68A3374B9C" box="[951,1095,1641,1668]" collectionCode="MCZ" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="11" pageNumber="435" type="Museum">MCZ 45362</specimenCode>
,
<specimenCode id="DBC422386A036C02FB856C6BA3B84B9B" box="[1108,1224,1641,1668]" collectionCode="DMT" pageId="11" pageNumber="435">DMT 371</specimenCode>
),
<collectingRegion id="49A644A16A036C02FB0F6C68A2E84B9B" box="[1246,1432,1641,1667]" country="Costa Rica" name="Limon" pageId="11" pageNumber="435">Limón Province</collectingRegion>
,
<location id="8EBDDC986A036C02FF466C8CA6394BB0" LSID="urn:lsid:plazi:treatment:03CB3B556A036C05FF466F01A4E348D7:8EBDDC986A036C02FF466C8CA6394BB0" box="[151,329,1677,1704]" country="Costa Rica" latitude="10.284663" longLatPrecision="1" longitude="-83.41015" name="Hamburg Farm" pageId="11" pageNumber="435" stateProvince="Limon">Hamburg Farm</location>
,
<geoCoordinate id="EE56EC846A036C02FE896C8CA5704BBF" box="[344,512,1677,1703]" degrees="10.2846630" direction="north" orientation="latitude" pageId="11" pageNumber="435" precision="1" value="10.284663">10.2846630°N</geoCoordinate>
, -
<geoCoordinate id="EE56EC846A036C02FDC76C8CA5C44BBF" box="[534,692,1677,1703]" degrees="83.410149" direction="west" orientation="longitude" pageId="11" pageNumber="435" precision="1" value="-83.41015">83.410149°W</geoCoordinate>
,
<elevation id="004F6D706A036C02FD126C8CA4304BBF" box="[707,832,1677,1704]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="435" unit="m" value="10.0">
<quantity id="4C9A27A66A036C02FD126C8CA4704BB0" box="[707,768,1677,1704]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="435" unit="m" value="10.0">10 m</quantity>
a.s.l.
</elevation>
,
<collectorName id="2697EF956A036C02FC806C8CA4EC4BBF" box="[849,924,1677,1703]" pageId="11" pageNumber="435">Dodge</collectorName>
leg.,
<determinerName id="7E0166876A036C02FC306C8FA3CC4BBF" box="[993,1212,1677,1704]" pageId="11" pageNumber="435">D. Martínez-Torres</determinerName>
det., 2016 (https:// mczbase.mcz.harvard.edu/guid/
<collectionCode id="ED7312866A036C02FE296CB0A5474BD3" box="[504,567,1713,1739]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="11" pageNumber="435" type="Museum">MCZ</collectionCode>
:IZ:45362);
<specimenCount id="9D6441CA6A036C02FD6E6CB0A47A4BD3" box="[703,778,1713,1739]" count="1" pageId="11" pageNumber="435" type="male">1 male</specimenCount>
(
<specimenCode id="DBC422386A036C02FCC96CB0A4D64BD3" box="[792,934,1713,1739]" collectionCode="MCZ" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="11" pageNumber="435" type="Museum">MCZ 46480</specimenCode>
,
<specimenCode id="DBC422386A036C02FC606CB3A3544BD3" box="[945,1060,1713,1740]" collectionCode="DMT" pageId="11" pageNumber="435">DMT 403</specimenCode>
),
<collectingRegion id="49A644A16A036C02FBE66CB0A39E4BD3" box="[1079,1262,1713,1739]" country="Costa Rica" name="Limon" pageId="11" pageNumber="435">Limón Province</collectingRegion>
,
<collectingCounty id="62BCF2CF6A036C02FB286CB3A2E84BD3" box="[1273,1432,1713,1740]" pageId="11" pageNumber="435">Las Mercedes</collectingCounty>
, between
<collectingMunicipality id="6BB910396A036C02FF2A6CD4A6284BF7" box="[251,344,1749,1775]" pageId="11" pageNumber="435">Squirres</collectingMunicipality>
and
<location id="8EBDDC986A036C02FE5F6CD4A6844BF7" LSID="urn:lsid:plazi:treatment:03CB3B556A036C05FF466F01A4E348D7:8EBDDC986A036C02FE5F6CD4A6844BF7" box="[398,500,1749,1775]" country="Costa Rica" county="Las Mercedes" latitude="10.175627" longLatPrecision="1" longitude="-83.611534" municipality="Squirres" name="Guapiles" pageId="11" pageNumber="435" stateProvince="Limon">Guapiles</location>
,
<geoCoordinate id="EE56EC846A036C02FE2E6CD4A5D74BE8" box="[511,679,1749,1776]" degrees="10.1756270" direction="north" orientation="latitude" pageId="11" pageNumber="435" precision="1" value="10.175627">10.1756270°N</geoCoordinate>
, -
<geoCoordinate id="EE56EC846A036C02FD686CD4A4264BE8" box="[697,854,1749,1776]" degrees="83.611534" direction="west" orientation="longitude" pageId="11" pageNumber="435" precision="1" value="-83.611534">83.611534°W</geoCoordinate>
,
<elevation id="004F6D706A036C02FCB06CD4A4A64BF7" box="[865,982,1749,1776]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="11" pageNumber="435" unit="m" value="50.0">
<quantity id="4C9A27A66A036C02FCB06CD4A4EA4BE8" box="[865,922,1749,1776]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="11" pageNumber="435" unit="m" value="50.0">50 m</quantity>
a.s.l.
</elevation>
,
<date id="FFDCAC836A036C02FC336CD4A3484BF7" box="[994,1080,1749,1775]" pageId="11" pageNumber="435" value="1984-10">
<collectingDate id="EF98556B6A036C02FC336CD4A3484BF7" box="[994,1080,1749,1775]" pageId="11" pageNumber="435" value="1984-10">X.1984</collectingDate>
</date>
,
<collectorName id="2697EF956A036C02FB936CD7A3BF4BF7" box="[1090,1231,1749,1776]" pageId="11" pageNumber="435">W. Eberhard</collectorName>
leg.,
<determinerName id="7E0166876A036C02FADD6CD7A7AE4A0C" pageId="11" pageNumber="435">D. Martínez-Torres</determinerName>
det., 2016 (https://mczbase.mcz.harvard.edu/guid/
<collectionCode id="ED7312866A036C02FCF36CF8A4114A0B" box="[802,865,1785,1811]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="11" pageNumber="435" type="Museum">MCZ</collectionCode>
:IZ:46480);
<specimenCount id="9D6441CA6A036C02FC3E6CF8A34F4A0B" box="[1007,1087,1785,1811]" count="1" pageId="11" pageNumber="435" type="male">1 male</specimenCount>
(
<collectionCode id="ED7312866A036C02FB836CF8A3CC4A0B" box="[1106,1212,1785,1811]" pageId="11" pageNumber="435">NCMNH</collectionCode>
<specimenCode id="DBC422386A036C02FB166CF8A2664A0C" box="[1223,1302,1785,1812]" collectionCode="NCMNH" pageId="11" pageNumber="435">A3655</specimenCode>
,
<specimenCode id="DBC422386A036C02FAF76CFBA2FF4A0C" box="[1318,1423,1785,1812]" collectionCode="DMT" pageId="11" pageNumber="435">DMT 65</specimenCode>
),
<collectingCountry id="F375CAD36A036C02FF466D1CA6644A2F" box="[151,276,1821,1847]" name="Costa Rica" pageId="11" pageNumber="435">Costa Rica</collectingCountry>
,
<collectingRegion id="49A644A16A036C02FECE6D1FA5764A2F" box="[287,518,1821,1848]" country="Costa Rica" name="Puntarenas" pageId="11" pageNumber="435">Puntarenas Province</collectingRegion>
,
<locationDeviation id="9A6F93616A036C02FDC36D1CA4F64A2F" box="[530,902,1821,1848]" location="Pan Am Highway" pageId="11" pageNumber="435">
<quantity id="4C9A27A66A036C02FDC36D1CA5294A2F" box="[530,601,1821,1848]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="11" pageNumber="435" unit="km" value="25.0">25 km</quantity>
NNE of Pan Am Highway
</locationDeviation>
,
<collectingMunicipality id="6BB910396A036C02FC416D1CA3674A2F" box="[912,1047,1821,1847]" pageId="11" pageNumber="435">Monteverde</collectingMunicipality>
Сloud
<location id="8EBDDC986A036C02FBB86D1FA2634A20" LSID="urn:lsid:plazi:treatment:03CB3B556A036C05FF466F01A4E348D7:8EBDDC986A036C02FBB86D1FA2634A20" box="[1129,1299,1822,1848]" country="Costa Rica" municipality="Monteverde" name="Forest Reserve" pageId="11" pageNumber="435" stateProvince="Puntarenas">Forest Reserve</location>
,
<elevation id="004F6D706A036C02FACF6D1CA79A4A43" metricMagnitude="3" metricUnit="m" metricValue="1.65" metricValueMax="1.7" metricValueMin="1.6" pageId="11" pageNumber="435" unit="m" value="1650.0" valueMax="1700.0" valueMin="1600.0">
<quantity id="4C9A27A66A036C02FACF6D1CA7DD4A44" metricMagnitude="3" metricUnit="m" metricValue="1.65" metricValueMax="1.7" metricValueMin="1.6" pageId="11" pageNumber="435" unit="m" value="1650.0" valueMax="1700.0" valueMin="1600.0">1600–1700 m</quantity>
a.s.l.
</elevation>
,
<date id="FFDCAC836A036C02FF276D40A6074A43" box="[246,375,1857,1883]" pageId="11" pageNumber="435" value="1981-03-10">
<collectingDate id="EF98556B6A036C02FF276D40A6074A43" box="[246,375,1857,1883]" pageId="11" pageNumber="435" value="1981-03-10">10.III.1981</collectingDate>
</date>
,
<collectorName id="2697EF956A036C02FE536D43A57A4A43" box="[386,522,1857,1884]" pageId="11" pageNumber="435">R.E. Ashton</collectorName>
leg.,
<determinerName id="7E0166876A036C02FD996D43A4504A43" box="[584,800,1857,1884]" pageId="11" pageNumber="435">D. Martínez-Torres</determinerName>
det., 2015.
</paragraph>
</subSubSection>
<subSubSection id="C378D9C86A036C05FF166D64A66B4C91" lastPageId="12" lastPageNumber="436" pageId="11" pageNumber="435" type="description">
<paragraph id="8BDD8A436A036C05FF166D64A66B4C91" blockId="11.[151,1437,1353,2028]" lastBlockId="12.[151,1437,151,393]" lastPageId="12" lastPageNumber="436" pageId="11" pageNumber="435">
<emphasis id="B91656516A036C02FF166D64A6904A67" bold="true" box="[199,480,1893,1919]" pageId="11" pageNumber="435">Brief descriptive notes.</emphasis>
Length ca
<quantity id="4C9A27A66A036C02FDB66D64A5814A98" box="[615,753,1893,1920]" metricMagnitude="-2" metricUnit="m" metricValue="8.5" metricValueMax="10.0" metricValueMin="7.0" pageId="11" pageNumber="435" unit="mm" value="85.0" valueMax="100.0" valueMin="70.0">70–100 mm</quantity>
(male, female) (
<figureCitation id="135996C66A036C02FC656D64A3704A98" box="[948,1024,1893,1920]" captionStart="FIGURE 5" captionStartId="6.[152,255,1896,1920]" captionTargetBox="[237,1351,1011,1870]" captionTargetId="figure-56@6.[237,1351,1011,1870]" captionTargetPageId="6" captionText="FIGURE 5. Nyssodesmus alboalatus Cook, 1896. Male from Farm Hamburg, Waldbodan (VMNH, DMT 98). A. Habitus, dorsal view. Labels reading “Costa Rica, Farm Hamburg, Waldbodan, Feb. 11, 1934, F. Nevermann leg.” and “DMT 98”. Scale bars: 1.0 mm (B, C)." figureDoi="http://doi.org/10.5281/zenodo.14239757" httpUri="https://zenodo.org/record/14239757/files/figure.png" pageId="11" pageNumber="435">Figs 5</figureCitation>
,
<figureCitation id="135996C66A036C02FBDE6D64A3394A67" box="[1039,1097,1893,1919]" captionStart-0="FIGURE 8" captionStart-1="FIGURE 9" captionStart-2="FIGURE 10" captionStart-3="FIGURE 11" captionStart-4="FIGURE 12" captionStart-5="FIGURE 13" captionStartId-0="9.[152,254,931,953]" captionStartId-1="9.[152,254,1924,1946]" captionStartId-2="10.[152,255,947,971]" captionStartId-3="10.[152,255,1940,1964]" captionStartId-4="11.[152,255,1103,1127]" captionStartId-5="12.[152,255,1102,1126]" captionTargetBox-0="[265,1320,179,903]" captionTargetBox-1="[259,1327,1052,1894]" captionTargetBox-2="[365,1223,179,921]" captionTargetBox-3="[371,1218,1065,1914]" captionTargetBox-4="[219,1372,198,1076]" captionTargetBox-5="[215,1372,414,1076]" captionTargetId-0="figure-21@9.[265,1323,179,903]" captionTargetId-1="figure-80@9.[259,1327,1046,1896]" captionTargetId-2="figure-17@10.[365,1223,179,921]" captionTargetId-3="figure-77@10.[369,1218,1065,1914]" captionTargetId-4="figure-21@11.[215,1372,198,1076]" captionTargetId-5="figure-137@12.[215,1372,414,1076]" captionTargetPageId-0="9" captionTargetPageId-1="9" captionTargetPageId-2="10" captionTargetPageId-3="10" captionTargetPageId-4="11" captionTargetPageId-5="12" captionText-0="FIGURE 8. Nyssodesmus python (Peters, 1865). From up to down, two males and one female from La Virgen, Heredia Prov. (NCMNH A3654, DMT 63), together with the original labels. A. Habitus, dorsal view. B, C. Left gonopod, lateral and mesal views, respectively. Abbreviations: lb, lateral branch; sl, solenomere. Photographed not to scale." captionText-1="FIGURE 9. Nyssodesmus python (Peters, 1865). Two males from Monteverde, Puntarenas Prov. (VMNH, DMT 95), together with the original labels. A. Habitus, dorsal view. B, C. Left gonopod, mesal and lateral views, respectively. Abbreviations: lb, lateral branch; sl, solenomere. Scale bar: 1 mm (B, C)." captionText-2="FIGURE 10. Nyssodesmus python (Peters, 1865). Two males and one female from Finca La Selva near Puerto Viejo, Heredia Prov. (VMNH, DMT 93), together with the original labels. A. Habitus, dorsal view. B, C. Left gonopod, lateral and mesal views, respectively. Abbreviations: lb, lateral branch; sl, solenomere. Scale bar: 1 mm (B, C)." captionText-3="FIGURE 11. Nyssodesmus python (Peters, 1865). Male from La Selva Biological Station, Heredia Prov. (ZISP 8-2023). A, B. Habitus, dorsal and ventral views, respectively. C, D. Right gonopod, lateral and mesal views, respectively. Photographed not to scale." captionText-4="FIGURE 12. Nyssodesmus python (Peters, 1865). Male from La Selva Biological Station, Heredia Prov. (ZISP MYR_DIP_ 0000203). A, B. Habitus, dorsal and ventral views, respectively. C, D. Right gonopod, lateral and mesal views, respectively. Photographed not to scale." captionText-5="FIGURE 13. Nyssodesmus python (Peters, 1865). Live colour patterns in nature.A, B. Individual females. C, D. Paired couples (male on top). Photographed not to scale. Courtesy flickr.com›photos/tags/nyssodesmuspython/" figureDoi-0="http://doi.org/10.5281/zenodo.14239767" figureDoi-1="http://doi.org/10.5281/zenodo.14239771" figureDoi-2="http://doi.org/10.5281/zenodo.14239773" figureDoi-3="http://doi.org/10.5281/zenodo.14239777" figureDoi-4="http://doi.org/10.5281/zenodo.14239781" figureDoi-5="http://doi.org/10.5281/zenodo.14239783" httpUri-0="https://zenodo.org/record/14239767/files/figure.png" httpUri-1="https://zenodo.org/record/14239771/files/figure.png" httpUri-2="https://zenodo.org/record/14239773/files/figure.png" httpUri-3="https://zenodo.org/record/14239777/files/figure.png" httpUri-4="https://zenodo.org/record/14239781/files/figure.png" httpUri-5="https://zenodo.org/record/14239783/files/figure.png" pageId="11" pageNumber="435">8–13</figureCitation>
), width of midbody pro- and metazona up to 6.2 and
<quantity id="4C9A27A66A036C02FE716D88A5764ABC" box="[416,518,1929,1956]" metricMagnitude="-2" metricUnit="m" metricValue="1.22" pageId="11" pageNumber="435" unit="mm" value="12.2">12.2 mm</quantity>
, respectively (male). Live coloration (
<figureCitation id="135996C66A036C02FC7E6D88A48F4ABB" box="[943,1023,1929,1955]" captionStart="FIGURE 13" captionStartId="12.[152,255,1102,1126]" captionTargetBox="[215,1372,414,1076]" captionTargetId="figure-137@12.[215,1372,414,1076]" captionTargetPageId="12" captionText="FIGURE 13. Nyssodesmus python (Peters, 1865). Live colour patterns in nature.A, B. Individual females. C, D. Paired couples (male on top). Photographed not to scale. Courtesy flickr.com›photos/tags/nyssodesmuspython/" figureDoi="http://doi.org/10.5281/zenodo.14239783" httpUri="https://zenodo.org/record/14239783/files/figure.png" pageId="11" pageNumber="435">Fig. 13</figureCitation>
) usually variegated, vivid and stripy, light to dark brown, always with a wide, light, red-brown to pallid, mid-dorsal, axial stripe often accompanied by a vague, narrow, darker, greyish axial line; pro- and metaterga contrasting light mid-dorsally, paraterga usually light as well, but dark/infuscate only at or near bases of paraterga, occasionally dark also almost until their lateral margins, these being orange to pallid. Dry or alcohol material (
<figureCitation id="135996C66A046C05FD3D6ABAA4194DCD" box="[748,873,187,213]" captionStart="FIGURE 6" captionStartId="7.[152,255,1711,1735]" captionTargetBox="[227,1360,675,1685]" captionTargetId="figure-178@7.[227,1360,675,1685]" captionTargetPageId="7" captionText="FIGURE 6. Nyssodesmus python (Peters, 1865).Female holotype(ZMB 8). A, B. Habitus, dorsal andventral views, respectively. Scale bar: 10 mm. C. Right paratergum 10, dorsal view, photographed not to scale. D. Original labels. Courtesy O. Werb." figureDoi="http://doi.org/10.5281/zenodo.14239761" httpUri="https://zenodo.org/record/14239761/files/figure.png" pageId="12" pageNumber="436">Figs 6A, B</figureCitation>
,
<figureCitation id="135996C66A046C05FCA56ABAA4E84DCD" box="[884,920,187,213]" captionStart="FIGURE 8" captionStartId="9.[152,254,931,953]" captionTargetBox="[265,1320,179,903]" captionTargetId="figure-21@9.[265,1323,179,903]" captionTargetPageId="9" captionText="FIGURE 8. Nyssodesmus python (Peters, 1865). From up to down, two males and one female from La Virgen, Heredia Prov. (NCMNH A3654, DMT 63), together with the original labels. A. Habitus, dorsal view. B, C. Left gonopod, lateral and mesal views, respectively. Abbreviations: lb, lateral branch; sl, solenomere. Photographed not to scale." figureDoi="http://doi.org/10.5281/zenodo.14239767" httpUri="https://zenodo.org/record/14239767/files/figure.png" pageId="12" pageNumber="436">8A</figureCitation>
,
<figureCitation id="135996C66A046C05FC756ABAA4B94DCD" box="[932,969,187,213]" captionStart="FIGURE 9" captionStartId="9.[152,254,1924,1946]" captionTargetBox="[259,1327,1052,1894]" captionTargetId="figure-80@9.[259,1327,1046,1896]" captionTargetPageId="9" captionText="FIGURE 9. Nyssodesmus python (Peters, 1865). Two males from Monteverde, Puntarenas Prov. (VMNH, DMT 95), together with the original labels. A. Habitus, dorsal view. B, C. Left gonopod, mesal and lateral views, respectively. Abbreviations: lb, lateral branch; sl, solenomere. Scale bar: 1 mm (B, C)." figureDoi="http://doi.org/10.5281/zenodo.14239771" httpUri="https://zenodo.org/record/14239771/files/figure.png" pageId="12" pageNumber="436">9A</figureCitation>
,
<figureCitation id="135996C66A046C05FC026ABAA3764DCD" box="[979,1030,187,213]" captionStart="FIGURE 10" captionStartId="10.[152,255,947,971]" captionTargetBox="[365,1223,179,921]" captionTargetId="figure-17@10.[365,1223,179,921]" captionTargetPageId="10" captionText="FIGURE 10. Nyssodesmus python (Peters, 1865). Two males and one female from Finca La Selva near Puerto Viejo, Heredia Prov. (VMNH, DMT 93), together with the original labels. A. Habitus, dorsal view. B, C. Left gonopod, lateral and mesal views, respectively. Abbreviations: lb, lateral branch; sl, solenomere. Scale bar: 1 mm (B, C)." figureDoi="http://doi.org/10.5281/zenodo.14239773" httpUri="https://zenodo.org/record/14239773/files/figure.png" pageId="12" pageNumber="436">10A</figureCitation>
,
<figureCitation id="135996C66A046C05FBC06ABAA3134DCD" box="[1041,1123,187,213]" captionStart="FIGURE 11" captionStartId="10.[152,255,1940,1964]" captionTargetBox="[371,1218,1065,1914]" captionTargetId="figure-77@10.[369,1218,1065,1914]" captionTargetPageId="10" captionText="FIGURE 11. Nyssodesmus python (Peters, 1865). Male from La Selva Biological Station, Heredia Prov. (ZISP 8-2023). A, B. Habitus, dorsal and ventral views, respectively. C, D. Right gonopod, lateral and mesal views, respectively. Photographed not to scale." figureDoi="http://doi.org/10.5281/zenodo.14239777" httpUri="https://zenodo.org/record/14239777/files/figure.png" pageId="12" pageNumber="436">11A, B</figureCitation>
,
<figureCitation id="135996C66A046C05FBBF6ABAA3CF4DCD" box="[1134,1215,187,213]" captionStart="FIGURE 12" captionStartId="11.[152,255,1103,1127]" captionTargetBox="[219,1372,198,1076]" captionTargetId="figure-21@11.[215,1372,198,1076]" captionTargetPageId="11" captionText="FIGURE 12. Nyssodesmus python (Peters, 1865). Male from La Selva Biological Station, Heredia Prov. (ZISP MYR_DIP_ 0000203). A, B. Habitus, dorsal and ventral views, respectively. C, D. Right gonopod, lateral and mesal views, respectively. Photographed not to scale." figureDoi="http://doi.org/10.5281/zenodo.14239781" httpUri="https://zenodo.org/record/14239781/files/figure.png" pageId="12" pageNumber="436">12A, B</figureCitation>
) usually apparently faded, up to uniformly greyish to light beige (
<figureCitation id="135996C66A046C05FD466ADEA5854DE1" box="[663,757,223,249]" captionStart="FIGURE 6" captionStartId="7.[152,255,1711,1735]" captionTargetBox="[227,1360,675,1685]" captionTargetId="figure-178@7.[227,1360,675,1685]" captionTargetPageId="7" captionText="FIGURE 6. Nyssodesmus python (Peters, 1865).Female holotype(ZMB 8). A, B. Habitus, dorsal andventral views, respectively. Scale bar: 10 mm. C. Right paratergum 10, dorsal view, photographed not to scale. D. Original labels. Courtesy O. Werb." figureDoi="http://doi.org/10.5281/zenodo.14239761" httpUri="https://zenodo.org/record/14239761/files/figure.png" pageId="12" pageNumber="436">Figs 6A</figureCitation>
,
<figureCitation id="135996C66A046C05FCD06ADEA4454DE1" box="[769,821,223,249]" captionStart="FIGURE 10" captionStartId="10.[152,255,947,971]" captionTargetBox="[365,1223,179,921]" captionTargetId="figure-17@10.[365,1223,179,921]" captionTargetPageId="10" captionText="FIGURE 10. Nyssodesmus python (Peters, 1865). Two males and one female from Finca La Selva near Puerto Viejo, Heredia Prov. (VMNH, DMT 93), together with the original labels. A. Habitus, dorsal view. B, C. Left gonopod, lateral and mesal views, respectively. Abbreviations: lb, lateral branch; sl, solenomere. Scale bar: 1 mm (B, C)." figureDoi="http://doi.org/10.5281/zenodo.14239773" httpUri="https://zenodo.org/record/14239773/files/figure.png" pageId="12" pageNumber="436">10A</figureCitation>
). Metatergal granulations typically indistinct (
<figureCitation id="135996C66A046C05FAEB6ADEA2E84DE1" box="[1338,1432,223,249]" captionStart="FIGURE 6" captionStartId="7.[152,255,1711,1735]" captionTargetBox="[227,1360,675,1685]" captionTargetId="figure-178@7.[227,1360,675,1685]" captionTargetPageId="7" captionText="FIGURE 6. Nyssodesmus python (Peters, 1865).Female holotype(ZMB 8). A, B. Habitus, dorsal andventral views, respectively. Scale bar: 10 mm. C. Right paratergum 10, dorsal view, photographed not to scale. D. Original labels. Courtesy O. Werb." figureDoi="http://doi.org/10.5281/zenodo.14239761" httpUri="https://zenodo.org/record/14239761/files/figure.png" pageId="12" pageNumber="436">Figs 6A</figureCitation>
,
<figureCitation id="135996C66A046C05FF466B02A79A4C05" box="[151,234,259,285]" captionStart="FIGURE 11" captionStartId="10.[152,255,1940,1964]" captionTargetBox="[371,1218,1065,1914]" captionTargetId="figure-77@10.[369,1218,1065,1914]" captionTargetPageId="10" captionText="FIGURE 11. Nyssodesmus python (Peters, 1865). Male from La Selva Biological Station, Heredia Prov. (ZISP 8-2023). A, B. Habitus, dorsal and ventral views, respectively. C, D. Right gonopod, lateral and mesal views, respectively. Photographed not to scale." figureDoi="http://doi.org/10.5281/zenodo.14239777" httpUri="https://zenodo.org/record/14239777/files/figure.png" pageId="12" pageNumber="436">11A, B</figureCitation>
,
<figureCitation id="135996C66A046C05FF276B02A6394C05" box="[246,329,259,285]" captionStart="FIGURE 12" captionStartId="11.[152,255,1103,1127]" captionTargetBox="[219,1372,198,1076]" captionTargetId="figure-21@11.[215,1372,198,1076]" captionTargetPageId="11" captionText="FIGURE 12. Nyssodesmus python (Peters, 1865). Male from La Selva Biological Station, Heredia Prov. (ZISP MYR_DIP_ 0000203). A, B. Habitus, dorsal and ventral views, respectively. C, D. Right gonopod, lateral and mesal views, respectively. Photographed not to scale." figureDoi="http://doi.org/10.5281/zenodo.14239781" httpUri="https://zenodo.org/record/14239781/files/figure.png" pageId="12" pageNumber="436">12A, B</figureCitation>
). Ozopore on paratergum 10 lying ca 1–2 its diameter off lateral margin (
<figureCitation id="135996C66A046C05FB536B02A3A44C05" box="[1154,1236,259,285]" captionStart="FIGURE 6" captionStartId="7.[152,255,1711,1735]" captionTargetBox="[227,1360,675,1685]" captionTargetId="figure-178@7.[227,1360,675,1685]" captionTargetPageId="7" captionText="FIGURE 6. Nyssodesmus python (Peters, 1865).Female holotype(ZMB 8). A, B. Habitus, dorsal andventral views, respectively. Scale bar: 10 mm. C. Right paratergum 10, dorsal view, photographed not to scale. D. Original labels. Courtesy O. Werb." figureDoi="http://doi.org/10.5281/zenodo.14239761" httpUri="https://zenodo.org/record/14239761/files/figure.png" pageId="12" pageNumber="436">Fig 6C</figureCitation>
). Gonopods (
<figureCitation id="135996C66A046C05FABB6B02A7D74C59" captionStart="FIGURE 7" captionStartId="8.[152,255,1503,1527]" captionTargetBox="[227,1360,666,1477]" captionTargetId="figure-216@8.[227,1360,666,1477]" captionTargetPageId="8" captionText="FIGURE 7. Nyssodesmus python (Peters, 1865). A, B. Right gonopod of holotype of N. nigricaudus Chamberlin, 1922 (USNM 843, DMT 27), mesal and lateral views, respectively. Abbreviations: lb, lateral branch; sl, solenomere." figureDoi="http://doi.org/10.5281/zenodo.14239765" httpUri="https://zenodo.org/record/14239765/files/figure.png" pageId="12" pageNumber="436">Figs 7</figureCitation>
,
<figureCitation id="135996C66A046C05FF626B26A7874C59" box="[179,247,295,321]" captionStart="FIGURE 8" captionStartId="9.[152,254,931,953]" captionTargetBox="[265,1320,179,903]" captionTargetId="figure-21@9.[265,1323,179,903]" captionTargetPageId="9" captionText="FIGURE 8. Nyssodesmus python (Peters, 1865). From up to down, two males and one female from La Virgen, Heredia Prov. (NCMNH A3654, DMT 63), together with the original labels. A. Habitus, dorsal view. B, C. Left gonopod, lateral and mesal views, respectively. Abbreviations: lb, lateral branch; sl, solenomere. Photographed not to scale." figureDoi="http://doi.org/10.5281/zenodo.14239767" httpUri="https://zenodo.org/record/14239767/files/figure.png" pageId="12" pageNumber="436">8B, C</figureCitation>
,
<figureCitation id="135996C66A046C05FED36B26A6364C59" box="[258,326,295,321]" captionStart="FIGURE 9" captionStartId="9.[152,254,1924,1946]" captionTargetBox="[259,1327,1052,1894]" captionTargetId="figure-80@9.[259,1327,1046,1896]" captionTargetPageId="9" captionText="FIGURE 9. Nyssodesmus python (Peters, 1865). Two males from Monteverde, Puntarenas Prov. (VMNH, DMT 95), together with the original labels. A. Habitus, dorsal view. B, C. Left gonopod, mesal and lateral views, respectively. Abbreviations: lb, lateral branch; sl, solenomere. Scale bar: 1 mm (B, C)." figureDoi="http://doi.org/10.5281/zenodo.14239771" httpUri="https://zenodo.org/record/14239771/files/figure.png" pageId="12" pageNumber="436">9B, C</figureCitation>
,
<figureCitation id="135996C66A046C05FE806B26A6D44C59" box="[337,420,295,321]" captionStart="FIGURE 11" captionStartId="10.[152,255,1940,1964]" captionTargetBox="[371,1218,1065,1914]" captionTargetId="figure-77@10.[369,1218,1065,1914]" captionTargetPageId="10" captionText="FIGURE 11. Nyssodesmus python (Peters, 1865). Male from La Selva Biological Station, Heredia Prov. (ZISP 8-2023). A, B. Habitus, dorsal and ventral views, respectively. C, D. Right gonopod, lateral and mesal views, respectively. Photographed not to scale." figureDoi="http://doi.org/10.5281/zenodo.14239777" httpUri="https://zenodo.org/record/14239777/files/figure.png" pageId="12" pageNumber="436">11C, D</figureCitation>
,
<figureCitation id="135996C66A046C05FE7E6B26A5704C59" box="[431,512,295,321]" captionStart="FIGURE 12" captionStartId="11.[152,255,1103,1127]" captionTargetBox="[219,1372,198,1076]" captionTargetId="figure-21@11.[215,1372,198,1076]" captionTargetPageId="11" captionText="FIGURE 12. Nyssodesmus python (Peters, 1865). Male from La Selva Biological Station, Heredia Prov. (ZISP MYR_DIP_ 0000203). A, B. Habitus, dorsal and ventral views, respectively. C, D. Right gonopod, lateral and mesal views, respectively. Photographed not to scale." figureDoi="http://doi.org/10.5281/zenodo.14239781" httpUri="https://zenodo.org/record/14239781/files/figure.png" pageId="12" pageNumber="436">12C, D</figureCitation>
) virtually same as in
<taxonomicName id="4C62F1C06A046C05FD206B26A4F74C59" authorityName="Cook" authorityYear="1896" box="[753,903,295,321]" class="Diplopoda" family="Platyrhacidae" genus="Nyssodesmus" kingdom="Animalia" order="Polydesmida" pageId="12" pageNumber="436" phylum="Arthropoda" rank="species" species="alboalatus">
<emphasis id="B91656516A046C05FD206B26A4F74C59" box="[753,903,295,321]" italics="true" pageId="12" pageNumber="436">N. alboalatus</emphasis>
</taxonomicName>
, acropodite likewise ribbon-shaped and rapidly bent mesad, divided into a rudimentary solenomere (
<emphasis id="B91656516A046C05FD0C6B4AA5804C7D" bold="true" box="[733,752,331,357]" pageId="12" pageNumber="436">sl</emphasis>
) and a longer, slightly sigmoid, rounded to subtruncate lateral branch (
<emphasis id="B91656516A046C05FF256B6EA67B4C91" bold="true" box="[244,267,367,393]" pageId="12" pageNumber="436">lb</emphasis>
).
</paragraph>
</subSubSection>
<caption id="DF1DDACB6A046C05FF496E4FA3144993" ID-DOI="http://doi.org/10.5281/zenodo.14239783" ID-Zenodo-Dep="14239783" httpUri="https://zenodo.org/record/14239783/files/figure.png" pageId="12" pageNumber="436" startId="12.[152,255,1102,1126]" targetBox="[215,1372,414,1076]" targetPageId="12" targetType="figure">
<paragraph id="8BDD8A436A046C05FF496E4FA3144993" blockId="12.[151,1436,1102,1163]" pageId="12" pageNumber="436">
<emphasis id="B91656516A046C05FF496E4FA654497E" bold="true" box="[152,292,1102,1126]" pageId="12" pageNumber="436">FIGURE 13.</emphasis>
<taxonomicName id="4C62F1C06A046C05FEF86E4EA5E0497E" authority="(Peters, 1865)" baseAuthorityName="Peters" baseAuthorityYear="1865" box="[297,656,1102,1127]" class="Diplopoda" family="Platyrhacidae" genus="Nyssodesmus" kingdom="Animalia" order="Polydesmida" pageId="12" pageNumber="436" phylum="Arthropoda" rank="species" species="python">
<emphasis id="B91656516A046C05FEF86E4EA68B497F" box="[297,507,1103,1127]" italics="true" pageId="12" pageNumber="436">Nyssodesmus python</emphasis>
(
<bibRefCitation id="EFF3F7B26A046C05FDD66E4EA5F8497E" author="Peters, W." box="[519,648,1102,1127]" pageId="12" pageNumber="436" pagination="529 - 551" refId="ref7857" refString="Peters, W. (1865) Ubersicht der im Konigl. zoologischen Museum befindlichen Myriopoden aus der Familie der Polydesmi, so wie Beschreibungen einer neuen Gattung, Trachyjulus, der Juli und neuer Arten der Gattung Siphonophora. Monatsberichte der Koniglichen Preussischen Akademie der Wissenschaften zu Berlin, 1864, 529 - 551." type="journal article" year="1865">Peters, 1865</bibRefCitation>
)
</taxonomicName>
. Live colour patterns in nature.
<emphasis id="B91656516A046C05FC1A6E4FA370497F" bold="true" box="[971,1024,1102,1127]" pageId="12" pageNumber="436">A, B.</emphasis>
Individual females.
<emphasis id="B91656516A046C05FB1C6E4FA274497F" bold="true" box="[1229,1284,1102,1127]" pageId="12" pageNumber="436">C, D.</emphasis>
Paired couples (male on top). Photographed not to scale. Courtesy
<emphasis id="B91656516A046C05FD706E73A47E4992" bold="true" box="[673,782,1138,1162]" pageId="12" pageNumber="436">flickr.com</emphasis>
›photos/tags/nyssodesmuspython/
</paragraph>
</caption>
<subSubSection id="C378D9C86A046C05FF166EDCA4E348D7" pageId="12" pageNumber="436" type="discussion">
<paragraph id="8BDD8A436A046C05FF166EDCA4D2489F" blockId="12.[151,1436,1245,1487]" pageId="12" pageNumber="436">
<emphasis id="B91656516A046C05FF166EDCA64B49EF" bold="true" box="[199,315,1245,1271]" pageId="12" pageNumber="436">Remarks.</emphasis>
Because the above alcohol samples referred to as
<taxonomicName id="4C62F1C06A046C05FCBF6EDCA30A49EF" baseAuthorityName="Peters" baseAuthorityYear="1865" box="[878,1146,1245,1271]" class="Diplopoda" family="Platyrhacidae" genus="Nyssodesmus" isUncertain="true" kingdom="Animalia" order="Polydesmida" pageId="12" pageNumber="436" phylum="Arthropoda" rank="species" species="python">
<emphasis id="B91656516A046C05FCBF6EDCA37349EF" box="[878,1027,1245,1271]" italics="true" pageId="12" pageNumber="436">Nyssodesmus</emphasis>
cf.
<emphasis id="B91656516A046C05FBFF6EDCA30A49EF" box="[1070,1146,1245,1271]" italics="true" pageId="12" pageNumber="436">python</emphasis>
</taxonomicName>
tend to show some minor deviations from the typical patterns, such as a little more distinct metatergal granulations and/or more strongly dentate lateral margins of some paraterga, and/or a little better discernible solenomere vestige, they all are provisionally treated, and mapped (
<figureCitation id="135996C66A046C05FE5C6F48A690487B" box="[397,480,1353,1379]" captionStart="FIGURE 14" captionStartId="13.[152,255,847,871]" captionTargetBox="[151,1436,179,821]" captionTargetId="figure-21@13.[151,1436,179,821]" captionTargetPageId="13" captionText="FIGURE 14. Distributions of Barydesmus truncatus sp. nov. (yellow triangle), Nyssodesmus python (Peters, 1865) (black square), N. cf. python (Peters, 1865) (blank square), and N. alboalatus Cook, 1896, type locality (yellow circle) and additional record (orange circle)." figureDoi="http://doi.org/10.5281/zenodo.14239785" httpUri="https://zenodo.org/record/14239785/files/figure.png" pageId="12" pageNumber="436">Fig. 14</figureCitation>
), as
<taxonomicName id="4C62F1C06A046C05FDC66F4BA5DA487B" baseAuthorityName="Peters" baseAuthorityYear="1865" box="[535,682,1353,1379]" class="Diplopoda" family="Platyrhacidae" genus="Nyssodesmus" isUncertain="true" kingdom="Animalia" order="Polydesmida" pageId="12" pageNumber="436" phylum="Arthropoda" rank="species" species="python">
<emphasis id="B91656516A046C05FDC66F4BA55A487B" box="[535,554,1354,1379]" italics="true" pageId="12" pageNumber="436">N</emphasis>
. cf.
<emphasis id="B91656516A046C05FD8F6F48A5DA487B" box="[606,682,1353,1379]" italics="true" pageId="12" pageNumber="436">python</emphasis>
</taxonomicName>
, separately from
<taxonomicName id="4C62F1C06A046C05FCA36F4BA490487B" baseAuthorityName="Peters" baseAuthorityYear="1865" box="[882,992,1353,1379]" class="Diplopoda" family="Platyrhacidae" genus="Nyssodesmus" kingdom="Animalia" order="Polydesmida" pageId="12" pageNumber="436" phylum="Arthropoda" rank="species" species="python">
<emphasis id="B91656516A046C05FCA36F4BA490487B" box="[882,992,1353,1379]" italics="true" pageId="12" pageNumber="436">N. python</emphasis>
</taxonomicName>
, even though they appear to be largely strictly sympatric or even syntopic with doubtless
<taxonomicName id="4C62F1C06A046C05FD1A6F6FA448489F" baseAuthorityName="Peters" baseAuthorityYear="1865" box="[715,824,1389,1415]" class="Diplopoda" family="Platyrhacidae" genus="Nyssodesmus" kingdom="Animalia" order="Polydesmida" pageId="12" pageNumber="436" phylum="Arthropoda" rank="species" species="python">
<emphasis id="B91656516A046C05FD1A6F6FA448489F" box="[715,824,1389,1415]" italics="true" pageId="12" pageNumber="436">N. python</emphasis>
</taxonomicName>
material.
</paragraph>
<paragraph id="8BDD8A436A046C05FF166F90A4E348D7" blockId="12.[151,1436,1245,1487]" pageId="12" pageNumber="436">
Basically, sympatry or syntopy is known to be very common amongst Neotropical
<taxonomicName id="4C62F1C06A046C05FBAF6F90A26948B3" authorityName="Pocock" authorityYear="1895" box="[1150,1305,1425,1451]" class="Diplopoda" family="Platyrhacidae" kingdom="Animalia" order="Polydesmida" pageId="12" pageNumber="436" phylum="Arthropoda" rank="family">Platyrhacidae</taxonomicName>
, with up to several congeners often co-occurring together (e.g.,
<bibRefCitation id="EFF3F7B26A046C05FD0E6FB4A4F748D7" author="Hoffman, R. L." box="[735,903,1461,1487]" pageId="12" pageNumber="436" pagination="1 - 584" refId="ref7592" refString="Hoffman, R. L. (1999) Checklist of the millipeds of North and Middle America. Virginia Museum of Natural History Special Publication, No. 8, 1 - 584." type="book chapter" year="1999">Hoffman 1999</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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