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<document id="924875E5F98C03DD31A66BD6E5D52BBF" ID-DOI="10.11646/zootaxa.3737.1.4" ID-GBIF-Dataset="3652f971-c438-4c0d-abd0-5c9fe5cae9ad" ID-ISSN="1175-5326" ID-Zenodo-Dep="285228" ID-ZooBank="24EA5BC5-2A87-4FF8-B74C-FF94590A816C" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460699664340" checkinUser="plazi" docAuthor="Kvifte, Gunnar Mikalsen, Ivković, Marija &amp; Klarić, Aleksandra" docDate="2013" docId="03B987A6EA28FFC5FF33FC54B9E83B42" docLanguage="en" docName="zt03737p067.pdf" docOrigin="Zootaxa 3737 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Sycorax silacea Haliday" docType="treatment" docVersion="6" lastPageNumber="60" masterDocId="FF80FFDEEA29FFC6FFA4FFD8BB3A3D1B" masterDocTitle="New records of moth flies (Diptera: Psychodidae) from Croatia, with the description of Berdeniella keroveci sp. nov." masterLastPageNumber="67" masterPageNumber="57" pageNumber="58" updateTime="1698354232509" updateUser="plazi">
<mods:mods id="28516E8E95C641E261A6111BE18EB749" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B2DBB352407CE8B84173E0D372FC47DC">
<mods:title id="597C6F09AF4483A139C2B852E4C939BD">New records of moth flies (Diptera: Psychodidae) from Croatia, with the description of Berdeniella keroveci sp. nov.</mods:title>
</mods:titleInfo>
<mods:name id="82DD71B64C42391BD23CFFDB5D148BEC" type="personal">
<mods:role id="E96BDB7D4D705721ECE205A892B13642">
<mods:roleTerm id="725704E3FB6594288565AE44372EBD46">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="907EA47928E50D555FD00B8DA175D25A">Kvifte, Gunnar Mikalsen</mods:namePart>
</mods:name>
<mods:name id="BA5A4673FBCAD8B4631A83E918494115" type="personal">
<mods:role id="9F0C86D7EF075865957A138B16F7E4FA">
<mods:roleTerm id="5422233164DC5A11D19CD9C609F3E4EA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AAAEE50C376F741C550A94DD4C42FBEA">Ivković, Marija</mods:namePart>
</mods:name>
<mods:name id="37C89A8D757300633967E6C7A1DD2931" type="personal">
<mods:role id="619ACEF9DF838A7BC3553DC8067ABB8A">
<mods:roleTerm id="EF89B90ACA6385B4332203ED6B644E84">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3119E141C145A5746C9348BBEA43F9FA">Klarić, Aleksandra</mods:namePart>
</mods:name>
<mods:typeOfResource id="CCA556BAA2C740915C6870A90612A6E9">text</mods:typeOfResource>
<mods:relatedItem id="AC09E51FFA52BEC47B769630CE9436A9" type="host">
<mods:titleInfo id="4ACBA2B806FC2D63306830851E0BEBF1">
<mods:title id="DE7859C93820BEF41A904454B4034F46">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="6F22C4A9E75227578152DB07A3749AB8">
<mods:date id="FB53504FBE607C7D3D0EBD7165879312">2013</mods:date>
<mods:detail id="2FD81D3D877281FF6F9E76E4E5CC96D2" type="volume">
<mods:number id="112742FF1CA00C0F4BC5761B07A8959F">3737</mods:number>
</mods:detail>
<mods:detail id="315AEEB6CB707482FC26B96B783C950F" type="issue">
<mods:number id="B2F7D151C81EF2FD739AD4271B2961E7">1</mods:number>
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<mods:extent id="660776793BA1077F920E433D03D40F30" unit="page">
<mods:start id="C22489530ECE0CCF9230F75DB821EECF">57</mods:start>
<mods:end id="5D8EC096D4137690258CB3436CC2C174">67</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="C66E279C189434F6FDFC5CF3068CF60E">journal article</mods:classification>
<mods:identifier id="50D1BE64F4C4366DC476A8EFEF7CF318" type="DOI">10.11646/zootaxa.3737.1.4</mods:identifier>
<mods:identifier id="8104F6C0EBAF05A2A69E96CF394F8E74" type="GBIF-Dataset">3652f971-c438-4c0d-abd0-5c9fe5cae9ad</mods:identifier>
<mods:identifier id="375F6910078C10B9B5AF0EB61ADF0AA7" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="A7A8374BA9613CBD4A968F8690AC2A98" type="Zenodo-Dep">285228</mods:identifier>
<mods:identifier id="738BF93FB0B9066D846145DACC3A2F42" type="ZooBank">24EA5BC5-2A87-4FF8-B74C-FF94590A816C</mods:identifier>
</mods:mods>
<treatment id="03B987A6EA28FFC5FF33FC54B9E83B42" ID-DOI="http://doi.org/10.5281/zenodo.6164567" ID-GBIF-Taxon="127666428" ID-Zenodo-Dep="6164567" LSID="urn:lsid:plazi:treatment:03B987A6EA28FFC5FF33FC54B9E83B42" httpUri="http://treatment.plazi.org/id/03B987A6EA28FFC5FF33FC54B9E83B42" lastPageId="3" lastPageNumber="60" pageId="1" pageNumber="58">
<subSubSection id="C30A653BEA28FFC7FF33FC54BF2A39D8" pageId="1" pageNumber="58" type="nomenclature">
<paragraph id="8BAF36B0EA28FFC7FF33FC54B9443EBD" blockId="1.[151,638,908,934]" box="[151,638,908,934]" pageId="1" pageNumber="58">
<heading id="D0E781DCEA28FFC7FF33FC54B9443EBD" bold="true" box="[151,638,908,934]" fontSize="11" level="1" pageId="1" pageNumber="58" reason="1">
<emphasis id="B964EAA2EA28FFC7FF33FC54B9443EBD" bold="true" box="[151,638,908,934]" pageId="1" pageNumber="58">
<taxonomicName id="4C104D33EA28FFC7FF33FC54BA873EBD" ID-CoL="53LZ8" authority="Haliday" authorityName="Haliday" box="[151,445,908,934]" class="Insecta" family="Psychodidae" genus="Sycorax" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="58" phylum="Arthropoda" rank="species" species="silacea">
<emphasis id="B964EAA2EA28FFC7FF33FC54BA693EBD" bold="true" box="[151,339,908,934]" italics="true" pageId="1" pageNumber="58">Sycorax silacea</emphasis>
Haliday
</taxonomicName>
<emphasis id="B964EAA2EA28FFC7FE61FC54BAE73EBD" bold="true" box="[453,477,908,934]" italics="true" pageId="1" pageNumber="58">in</emphasis>
Curtis, 1839
</emphasis>
</heading>
</paragraph>
<paragraph id="8BAF36B0EA28FFC7FF33FC0ABACF394C" blockId="1.[151,492,978,1003]" lastBlockId="1.[151,501,1086,1111]" pageId="1" pageNumber="58">
Literature record: Krek (1999)
<emphasis id="B964EAA2EA28FFC7FF33FBE6BACF394C" bold="true" box="[151,501,1086,1111]" pageId="1" pageNumber="58">
*
<taxonomicName id="4C104D33EA28FFC7FF08FBE6BACF394C" authority="Jung, 1953" authorityName="Jung" authorityYear="1953" box="[172,501,1086,1111]" class="Insecta" family="Psychodidae" genus="Sycorax" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="58" phylum="Arthropoda" rank="species" species="tonnoiri">
<emphasis id="B964EAA2EA28FFC7FF08FBE6BA57394C" bold="true" box="[172,365,1086,1111]" italics="true" pageId="1" pageNumber="58">Sycorax tonnoiri</emphasis>
Jung, 1953
</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="8BAF36B0EA28FFC7FF33FB5EBF2A39D8" blockId="1.[151,1436,1158,1291]" pageId="1" pageNumber="58">
First records for
<collectingCountry id="F3077620EA28FFC7FEC4FB5EBA8D3984" box="[352,439,1158,1183]" name="Croatia" pageId="1" pageNumber="58">Croatia</collectingCountry>
: Plitvice Lakes National park, Spring of Crna rijeka (
<geoCoordinate id="EE245077EA28FFC7FB90FB5FBFE33984" box="[1076,1241,1158,1183]" direction="east" orientation="longitude" pageId="1" pageNumber="58" precision="15" value="15.607779">E 15° 36' 28&quot;</geoCoordinate>
<geoCoordinate id="EE245077EA28FFC7FB41FB5FBEB73984" box="[1253,1421,1158,1183]" direction="north" orientation="latitude" pageId="1" pageNumber="58" precision="15" value="44.83722">N 44° 50' 14&quot;</geoCoordinate>
),
<date id="FFAE1070EA28FFC7FF33FB73BA2B39D8" box="[151,273,1195,1219]" pageId="1" pageNumber="58" value="2007-06-28">28.VI.2007</date>
,emergence trap P1,
<specimenCount id="9D16FD39EA28FFC7FE5FFB73B93339D8" box="[507,521,1195,1219]" pageId="1" pageNumber="58" type="generic">1</specimenCount>
♂;
<date id="FFAE1070EA28FFC7FD90FB73B9A739D8" box="[564,669,1195,1219]" pageId="1" pageNumber="58" value="2007-07">VII.2007</date>
, emergence trap P1,
<specimenCount id="9D16FD39EA28FFC7FC22FB73B8AE39D8" box="[902,916,1195,1219]" pageId="1" pageNumber="58" type="generic">1</specimenCount>
♂. MI leg.
</paragraph>
</subSubSection>
<subSubSection id="C30A653BEA28FFC7FF63FB16BA94386C" pageId="1" pageNumber="58" type="discussion">
<paragraph id="8BAF36B0EA28FFC7FF63FB16B9CD3810" blockId="1.[151,1436,1158,1291]" pageId="1" pageNumber="58">
<emphasis id="B964EAA2EA28FFC7FF63FB16BA7539FC" bold="true" box="[199,335,1230,1255]" pageId="1" pageNumber="58">Comments.</emphasis>
<taxonomicName id="4C104D33EA28FFC7FEFDFB17B92C39FD" box="[345,534,1231,1254]" class="Insecta" family="Psychodidae" genus="Sycorax" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="58" phylum="Arthropoda" rank="species" species="tonnoiri">
<emphasis id="B964EAA2EA28FFC7FEFDFB17B92C39FD" box="[345,534,1231,1254]" italics="true" pageId="1" pageNumber="58">Sycorax tonnoiri</emphasis>
</taxonomicName>
is widespread in Central and Southern Europe. Previous Balkan records of
<taxonomicName id="4C104D33EA28FFC7FA23FB17BBCB3810" class="Insecta" family="Psychodidae" genus="Sycorax" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="58" phylum="Arthropoda" rank="species" species="tonnoiri">
<emphasis id="B964EAA2EA28FFC7FA23FB17BBCB3810" italics="true" pageId="1" pageNumber="58">S. tonnoiri</emphasis>
</taxonomicName>
are from
<collectingCountry id="F3077620EA28FFC7FEFBFB2AB9653810" box="[351,607,1266,1291]" name="Bosnia and Herzegovina" pageId="1" pageNumber="58">Bosnia &amp; Herzegovina</collectingCountry>
(Krek 1999).
</paragraph>
<paragraph id="8BAF36B0EA28FFC7FF33FA86BA94386C" blockId="1.[151,430,1374,1399]" box="[151,430,1374,1399]" pageId="1" pageNumber="58">
<emphasis id="B964EAA2EA28FFC7FF33FA86BA94386C" bold="true" box="[151,430,1374,1399]" pageId="1" pageNumber="58">
Subfamily
<taxonomicName id="4C104D33EA28FFC7FEBDFA86BA94386C" box="[281,430,1374,1399]" class="Insecta" family="Psychodidae" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="58" phylum="Arthropoda" rank="subFamily" subFamily="Psychodinae">Psychodinae</taxonomicName>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C30A653BEA28FFC7FF33FA7EB8343B30" pageId="1" pageNumber="58" type="description">
<paragraph id="8BAF36B0EA28FFC7FF33FA7EB8343B30" blockId="1.[151,367,1446,1471]" lastBlockId="1.[151,782,1554,1579]" pageId="1" pageNumber="58">
<emphasis id="B964EAA2EA28FFC7FF33FA7EB8343B30" bold="true" pageId="1" pageNumber="58">
Tribe Pericomaini
<taxonomicName id="4C104D33EA28FFC7FF33F9CAB9943B30" authority="Kvifte, Ivkovic &amp; Klaric" authorityName="Kvifte, Ivkovic &amp; Klaric" box="[151,686,1554,1579]" class="Insecta" family="Psychodidae" genus="Berdeniella" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="58" phylum="Arthropoda" rank="species" species="keroveci" status="sp. nov.">
<emphasis id="B964EAA2EA28FFC7FF33F9CABABE3B30" bold="true" box="[151,388,1554,1579]" italics="true" pageId="1" pageNumber="58">Berdeniella keroveci</emphasis>
Kvifte, Ivković &amp; Klarić
</taxonomicName>
<taxonomicNameLabel id="A25757D9EA28FFC7FD11F9CAB8343B30" box="[693,782,1554,1579]" pageId="1" pageNumber="58" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C30A653BEA28FFC7FF33F982B8793AC0" pageId="1" pageNumber="58" type="materials_examined">
<paragraph id="8BAF36B0EA28FFC7FF33F982B93E3A18" blockId="1.[151,1436,1626,2011]" pageId="1" pageNumber="58">
<emphasis id="B964EAA2EA28FFC7FF33F982BA7F3B68" bold="true" box="[151,325,1626,1651]" pageId="1" pageNumber="58">
<typeStatus id="54AB8812EA28FFC7FF33F982BBE83B68" box="[151,210,1626,1651]" pageId="1" pageNumber="58">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54AB8812EA28FFC7FEEFF982BA893B68" box="[331,435,1626,1651]" pageId="1" pageNumber="58" type="holotype">Holotype</typeStatus>
♂:
<collectingCountry id="F3077620EA28FFC7FE7AF983B9663B68" box="[478,604,1627,1651]" name="Croatia" pageId="1" pageNumber="58">CROATIA</collectingCountry>
: Plitvice Lakes National park, Spring of Bijela rijeka (
<geoCoordinate id="EE245077EA28FFC7FB18F983BE683B68" box="[1212,1362,1626,1651]" direction="east" orientation="longitude" pageId="1" pageNumber="58" precision="15" value="15.561945">E 15° 33' 43&quot;</geoCoordinate>
<geoCoordinate id="EE245077EA28FFC7FAFEF983BBCA3B8C" direction="north" orientation="latitude" pageId="1" pageNumber="58" precision="15" value="44.83472">N 44° 50' 05&quot;</geoCoordinate>
),
<date id="FFAE1070EA28FFC7FEA9F9A7BA4F3B8C" box="[269,373,1663,1687]" pageId="1" pageNumber="58" value="2010-07">VII.2010</date>
, (emergence trap P4), M. Ivković leg.
<typeStatus id="54AB8812EA28FFC7FCFEF9A7B8F13B8C" box="[858,971,1663,1687]" pageId="1" pageNumber="58" type="paratype">Paratypes</typeStatus>
: same locality as
<typeStatus id="54AB8812EA28FFC7FB0AF9A6BE293B8C" box="[1198,1299,1662,1687]" pageId="1" pageNumber="58" type="holotype">holotype</typeStatus>
,
<date id="FFAE1070EA28FFC7FA82F9A7BEA23B8C" box="[1318,1432,1663,1687]" pageId="1" pageNumber="58" value="2010-08">VIII.2010</date>
, emergence trap P1,1 ♂;
<date id="FFAE1070EA28FFC7FE01F97BB93E3BA0" box="[421,516,1699,1723]" pageId="1" pageNumber="58" value="2009-06">VI.2009</date>
, emergence trap P3,
<specimenCount id="9D16FD39EA28FFC7FD48F97BB9C23BA0" box="[748,760,1699,1723]" pageId="1" pageNumber="58" type="generic">1</specimenCount>
♂;
<date id="FFAE1070EA28FFC7FCBAF97BB8473BA0" box="[798,893,1699,1723]" pageId="1" pageNumber="58" value="2010-06">VI.2010</date>
, emergence trap P3,
<specimenCount id="9D16FD39EA28FFC7FBC1F97BBF4B3BA0" box="[1125,1137,1699,1723]" pageId="1" pageNumber="58" type="generic">1</specimenCount>
♂. Plitvice Lakes National park, Spring of Crna rijeka (
<geoCoordinate id="EE245077EA28FFC7FE71F91FB9573BC4" box="[469,621,1734,1759]" direction="east" orientation="longitude" pageId="1" pageNumber="58" precision="15" value="15.607779">E 15° 36' 28&quot;</geoCoordinate>
<geoCoordinate id="EE245077EA28FFC7FDD1F91FB82B3BC4" box="[629,785,1734,1759]" direction="north" orientation="latitude" pageId="1" pageNumber="58" precision="15" value="44.83722">N 44° 50' 14&quot;</geoCoordinate>
),
<date id="FFAE1070EA28FFC7FC83F91EB8BF3BC4" box="[807,901,1734,1759]" pageId="1" pageNumber="58" value="2006-06">VI.2006</date>
, emergence trap P4,
<specimenCount id="9D16FD39EA28FFC7FBD5F91FBF473BC4" box="[1137,1149,1735,1759]" pageId="1" pageNumber="58" type="generic">1</specimenCount>
♂;
<date id="FFAE1070EA28FFC7FB07F91FBE223BC4" box="[1187,1304,1735,1759]" pageId="1" pageNumber="58" value="2011-05-29">29.V.2011</date>
, emergence trap P4,
<specimenCount id="9D16FD39EA28FFC7FF51F933BA3B3A18" box="[245,257,1771,1795]" pageId="1" pageNumber="58" type="generic">1</specimenCount>
♂. All leg. M. Ivković.
</paragraph>
<paragraph id="8BAF36B0EA28FFC7FF63F8D6B8793AC0" blockId="1.[151,1436,1626,2011]" pageId="1" pageNumber="58">
<emphasis id="B964EAA2EA28FFC7FF63F8D6BAEE3A3C" bold="true" box="[199,468,1806,1831]" pageId="1" pageNumber="58">Diagnostic characters.</emphasis>
<taxonomicName id="4C104D33EA28FFC7FE79F8D6B9FC3A3C" box="[477,710,1806,1831]" class="Insecta" family="Psychodidae" genus="Berdeniella" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="58" phylum="Arthropoda" rank="species" species="keroveci">
<emphasis id="B964EAA2EA28FFC7FE79F8D6B9FC3A3C" box="[477,710,1806,1831]" italics="true" pageId="1" pageNumber="58">Berdeniella keroveci</emphasis>
</taxonomicName>
can be distinguished from all other
<taxonomicName id="4C104D33EA28FFC7FBCAF8D6BFCA3A3C" box="[1134,1264,1806,1831]" class="Insecta" family="Psychodidae" genus="Berdeniella" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="B964EAA2EA28FFC7FBCAF8D6BFCA3A3C" box="[1134,1264,1806,1831]" italics="true" pageId="1" pageNumber="58">Berdeniella</emphasis>
</taxonomicName>
species on the following combination of characters: Phallic sheath with deep U-shaped median incision, gonostylus s-shaped, surstylus with one distally thickened tenaculum and one to two spines. The very similar
<taxonomicName id="4C104D33EA28FFC7FB15F88EBA163A88" authority="Krek, 1967" authorityName="Krek" authorityYear="1967" class="Insecta" family="Psychodidae" genus="Berdeniella" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="58" phylum="Arthropoda" rank="species" species="vaillanti">
<emphasis id="B964EAA2EA28FFC7FB15F88EBEA63A74" box="[1201,1436,1878,1903]" italics="true" pageId="1" pageNumber="58">Berdeniella vaillanti</emphasis>
(Krek, 1967)
</taxonomicName>
differs by having the gonocoxite with less pronounced curvature, the aedeagus with the median moveable appendage more slender and its base evenly arched, the lobes of the phallic sheath shorter and the surstyli shorter and stouter (Krek 1967, 1999; Wagner 1978).
</paragraph>
</subSubSection>
<caption id="DF6F6638EA2BFFC4FF33F95FB91B3A03" httpUri="https://zenodo.org/record/285229/files/figure.png" pageId="2" pageNumber="59" targetBox="[219,1370,209,1639]" targetPageId="2">
<paragraph id="8BAF36B0EA2BFFC4FF33F95FB91B3A03" blockId="2.[151,1436,1671,1816]" pageId="2" pageNumber="59">
<emphasis id="B964EAA2EA2BFFC4FF33F95FBA2E3B87" bold="true" box="[151,276,1671,1693]" pageId="2" pageNumber="59">FIGURE 1.</emphasis>
<emphasis id="B964EAA2EA2BFFC4FEB8F95FBACE3B86" box="[284,500,1671,1693]" italics="true" pageId="2" pageNumber="59">
<taxonomicName id="4C104D33EA2BFFC4FEB8F95FBACA3B86" box="[284,496,1671,1693]" class="Insecta" family="Psychodidae" genus="Berdeniella" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="keroveci" status="sp. nov.">Berdeniella keroveci</taxonomicName>
,
</emphasis>
<emphasis id="B964EAA2EA2BFFC4FE58F95FB9773B87" bold="true" box="[508,589,1671,1692]" pageId="2" pageNumber="59">
<taxonomicNameLabel id="A25757D9EA2BFFC4FE58F95FB9773B87" box="[508,589,1671,1692]" pageId="2" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male.
<emphasis id="B964EAA2EA2BFFC4FD38F95FB98E3B87" bold="true" box="[668,692,1671,1692]" pageId="2" pageNumber="59">A.</emphasis>
Wing.
<emphasis id="B964EAA2EA2BFFC4FCA5F95FB8233B87" bold="true" box="[769,793,1671,1692]" pageId="2" pageNumber="59">B.</emphasis>
Head.
<emphasis id="B964EAA2EA2BFFC4FCC0F95FB8473B86" bold="true" box="[868,893,1671,1693]" pageId="2" pageNumber="59">C.</emphasis>
Epandrium, gonopods and surstyli.
<emphasis id="B964EAA2EA2BFFC4FB56F95FBE313B87" bold="true" box="[1266,1291,1671,1692]" pageId="2" pageNumber="59">D.</emphasis>
aedeagus and phallic sheath. Scale bars for A–D: 0.2 mm, 0.04, 0.04 mm and 0.02 mm, respectively. Abbreviations: b, basiphallus; bmma, base of moveable median appendage; e, epiproct; ep, epandrium; gcx, gonocoxite; gst, gonostylus; hyp, hypandrium; lps, lobes of phallic sheath; mma, moveable median appendage; pso, pseudospiracular opening; rp, rigid phallus; sst, surstylus; t, tenaculum; vep, ventral epandrial plate
</paragraph>
</caption>
<subSubSection id="C30A653BEA2BFFC5FF63F8E7BEA73E7E" lastPageId="3" lastPageNumber="60" pageId="2" pageNumber="59" type="description">
<paragraph id="8BAF36B0EA2BFFC5FF63F8E7BEA73E7E" blockId="2.[151,1436,1855,2024]" lastBlockId="3.[151,1437,151,1625]" lastPageId="3" lastPageNumber="60" pageId="2" pageNumber="59">
<emphasis id="B964EAA2EA2BFFC4FF63F8E7BA633A43" bold="true" box="[199,345,1855,1880]" pageId="2" pageNumber="59">Description.</emphasis>
Male (n = 6, except measurements which are n = 4). Head (
<figureCitation id="132B2A35EA2BFFC4FC52F8E7BF0F3A43" box="[1014,1077,1855,1880]" captionStart="FIGURE 1" captionStartId="2.[151,250,1671,1693]" captionTargetBox="[219,1370,209,1639]" captionTargetId="figure@2.[211,1376,195,1647]" captionTargetPageId="2" captionText="FIGURE 1. Berdeniella keroveci, sp. nov., male. A. Wing. B. Head. C. Epandrium, gonopods and surstyli. D. aedeagus and phallic sheath. Scale bars for A – D: 0.2 mm, 0.04, 0.04 mm and 0.02 mm, respectively. Abbreviations: b, basiphallus; bmma, base of moveable median appendage; e, epiproct; ep, epandrium; gcx, gonocoxite; gst, gonostylus; hyp, hypandrium; lps, lobes of phallic sheath; mma, moveable median appendage; pso, pseudospiracular opening; rp, rigid phallus; sst, surstylus; t, tenaculum; vep, ventral epandrial plate" httpUri="https://zenodo.org/record/285229/files/figure.png" pageId="2" pageNumber="59">Fig. 1</figureCitation>
b) about as wide as long; vertex wide, equal to nearly half of total head length; eyebridge of four facet rows, separated by 3–4 facet diameters; ocular bristles present at sides only; interocular suture broadly U-shaped, with angular arches, occasionally reduced at middle; frontal scar patch oval with broad posterior band reaching to interocular suture; palp comprised of four segments, length of palp segments 64–80 (74): 98–108 (104): 98–118 (110): 130–172 (156); terminal palp segment striated; labellum bulbous, elongate-ovoid; antennae comprised of 16 segments; scape cylindrical, pedicel elongate-ovoid, flagellomeres fusiform; ascoids apparently absent; terminal flagellomere with stout apiculus; apiculus one third the length and half the width of the node from which it arises; length of antennal segments 72–82 (76): 62–66 (64): 54–58 (56): 40–44 (42): 38–42 (40): 36–42 (40): 38–42 (40): 36–38 (38): 36–42 (38): 36– 40 (36): 30–36 (34): 30–36 (32): 28–34 (32): 24–26 (24): 24–28 (26): 32–36 (34). Thorax with scutum, humeri, anepisternum, scutellum and lower two thirds of katepimeron densely setose, other sclerites mostly bare; legs without special features. Wing (
<figureCitation id="132B2A35EA2AFFC5FE58FEB7B9013C93" box="[508,571,367,392]" captionStart="FIGURE 1" captionStartId="2.[151,250,1671,1693]" captionTargetBox="[219,1370,209,1639]" captionTargetId="figure@2.[211,1376,195,1647]" captionTargetPageId="2" captionText="FIGURE 1. Berdeniella keroveci, sp. nov., male. A. Wing. B. Head. C. Epandrium, gonopods and surstyli. D. aedeagus and phallic sheath. Scale bars for A – D: 0.2 mm, 0.04, 0.04 mm and 0.02 mm, respectively. Abbreviations: b, basiphallus; bmma, base of moveable median appendage; e, epiproct; ep, epandrium; gcx, gonocoxite; gst, gonostylus; hyp, hypandrium; lps, lobes of phallic sheath; mma, moveable median appendage; pso, pseudospiracular opening; rp, rigid phallus; sst, surstylus; t, tenaculum; vep, ventral epandrial plate" httpUri="https://zenodo.org/record/285229/files/figure.png" pageId="3" pageNumber="60">Fig. 1</figureCitation>
a) 2.26–2.4 (2.33) mm long, ovate; wing tip between R4 and R5; pterostigmal area slightly darkened, membrane otherwise clear; Sc extending to origin of R2+3; radial and medial forks about in line with apex of CuA2, but with radial fork placed slightly distal; medial fork weakened at base of M
<quantity id="4CE89B55EA2AFFC5FA91FE14BE663CCD" box="[1333,1372,445,474]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="3" pageNumber="60" unit="in" value="2.0">2 in</quantity>
most specimens; jugum V-shaped. Terminalia with hypandrium narrow, laterally wider than at middle, often with lateral unpigmented spots or apertures; gonocoxites (
<figureCitation id="132B2A35EA2AFFC5FD1DFDD0B9C43F3A" box="[697,766,520,545]" captionStart="FIGURE 1" captionStartId="2.[151,250,1671,1693]" captionTargetBox="[219,1370,209,1639]" captionTargetId="figure@2.[211,1376,195,1647]" captionTargetPageId="2" captionText="FIGURE 1. Berdeniella keroveci, sp. nov., male. A. Wing. B. Head. C. Epandrium, gonopods and surstyli. D. aedeagus and phallic sheath. Scale bars for A – D: 0.2 mm, 0.04, 0.04 mm and 0.02 mm, respectively. Abbreviations: b, basiphallus; bmma, base of moveable median appendage; e, epiproct; ep, epandrium; gcx, gonocoxite; gst, gonostylus; hyp, hypandrium; lps, lobes of phallic sheath; mma, moveable median appendage; pso, pseudospiracular opening; rp, rigid phallus; sst, surstylus; t, tenaculum; vep, ventral epandrial plate" httpUri="https://zenodo.org/record/285229/files/figure.png" pageId="3" pageNumber="60">Fig. 1</figureCitation>
c) about twice as long as broad; shorter than gonostyli; gonostyli (
<figureCitation id="132B2A35EA2AFFC5FEB4FDF4BA6B3F5E" box="[272,337,556,581]" captionStart="FIGURE 1" captionStartId="2.[151,250,1671,1693]" captionTargetBox="[219,1370,209,1639]" captionTargetId="figure@2.[211,1376,195,1647]" captionTargetPageId="2" captionText="FIGURE 1. Berdeniella keroveci, sp. nov., male. A. Wing. B. Head. C. Epandrium, gonopods and surstyli. D. aedeagus and phallic sheath. Scale bars for A – D: 0.2 mm, 0.04, 0.04 mm and 0.02 mm, respectively. Abbreviations: b, basiphallus; bmma, base of moveable median appendage; e, epiproct; ep, epandrium; gcx, gonocoxite; gst, gonostylus; hyp, hypandrium; lps, lobes of phallic sheath; mma, moveable median appendage; pso, pseudospiracular opening; rp, rigid phallus; sst, surstylus; t, tenaculum; vep, ventral epandrial plate" httpUri="https://zenodo.org/record/285229/files/figure.png" pageId="3" pageNumber="60">Fig. 1</figureCitation>
c) oblong, subconical, broadly bulbous at base, with elongate sinously curved and pointed apex, with pair of ventral setae inserted subapically; aedeagus (
<figureCitation id="132B2A35EA2AFFC5FCB1FD88B8693F72" box="[789,851,592,617]" captionStart="FIGURE 1" captionStartId="2.[151,250,1671,1693]" captionTargetBox="[219,1370,209,1639]" captionTargetId="figure@2.[211,1376,195,1647]" captionTargetPageId="2" captionText="FIGURE 1. Berdeniella keroveci, sp. nov., male. A. Wing. B. Head. C. Epandrium, gonopods and surstyli. D. aedeagus and phallic sheath. Scale bars for A – D: 0.2 mm, 0.04, 0.04 mm and 0.02 mm, respectively. Abbreviations: b, basiphallus; bmma, base of moveable median appendage; e, epiproct; ep, epandrium; gcx, gonocoxite; gst, gonostylus; hyp, hypandrium; lps, lobes of phallic sheath; mma, moveable median appendage; pso, pseudospiracular opening; rp, rigid phallus; sst, surstylus; t, tenaculum; vep, ventral epandrial plate" httpUri="https://zenodo.org/record/285229/files/figure.png" pageId="3" pageNumber="60">Fig. 1</figureCitation>
d) with basiphallus narrow both in dorsal and lateral views; distal phallic appendages consisting of a moveable median morphoventral appendage and a rigid morphodorsal phallus consisting of two fused phallomeres; phallic sheath distally with two elongate lobes; base of moveable median appendage shaped like a broad arrow; epandrium (
<figureCitation id="132B2A35EA2AFFC5FC38FD64B8E63FCE" box="[924,988,700,725]" captionStart="FIGURE 1" captionStartId="2.[151,250,1671,1693]" captionTargetBox="[219,1370,209,1639]" captionTargetId="figure@2.[211,1376,195,1647]" captionTargetPageId="2" captionText="FIGURE 1. Berdeniella keroveci, sp. nov., male. A. Wing. B. Head. C. Epandrium, gonopods and surstyli. D. aedeagus and phallic sheath. Scale bars for A – D: 0.2 mm, 0.04, 0.04 mm and 0.02 mm, respectively. Abbreviations: b, basiphallus; bmma, base of moveable median appendage; e, epiproct; ep, epandrium; gcx, gonocoxite; gst, gonostylus; hyp, hypandrium; lps, lobes of phallic sheath; mma, moveable median appendage; pso, pseudospiracular opening; rp, rigid phallus; sst, surstylus; t, tenaculum; vep, ventral epandrial plate" httpUri="https://zenodo.org/record/285229/files/figure.png" pageId="3" pageNumber="60">Fig. 1</figureCitation>
c) about 1.5 times as long as wide, with paired pseudospiracular openings; ventral epandrial plate rod-shaped, indistinct; proctiger small, with hypoproct diamond-shaped, setose and epiproct small, ovoid; medial surfaces of surstyli basally (flanking proctiger) darker and more setose than rest of surstyli; surstyli each with one tenaculum and two spines inserted subapically; some specimens with only a single spine; tenaculum about 5 times as long as wide; apex of tenaculum expanded, smooth.
</paragraph>
</subSubSection>
<subSubSection id="C30A653BEA2AFFC5FF63FCA8BF1A3EB6" pageId="3" pageNumber="60" type="etymology">
<paragraph id="8BAF36B0EA2AFFC5FF63FCA8BF1A3EB6" blockId="3.[151,1437,151,1625]" pageId="3" pageNumber="60">
<emphasis id="B964EAA2EA2AFFC5FF63FCA8BA7C3E92" bold="true" box="[199,326,880,905]" pageId="3" pageNumber="60">Etymology</emphasis>
. Named after Mladen Kerovec, University of Zagreb, in gratitude for his support of MI's research in general and the Plitvice Lakes National Park emergence research in particular.
</paragraph>
</subSubSection>
<subSubSection id="C30A653BEA2AFFC5FF63FC60B9A63846" pageId="3" pageNumber="60" type="distribution">
<paragraph id="8BAF36B0EA2AFFC5FF63FC60B96D3EEE" blockId="3.[151,1437,151,1625]" pageId="3" pageNumber="60">
<emphasis id="B964EAA2EA2AFFC5FF63FC60BA633ECA" bold="true" box="[199,345,952,977]" pageId="3" pageNumber="60">Distribution</emphasis>
.
<taxonomicName id="4C104D33EA2AFFC5FECEFC60B96E3ECA" box="[362,596,952,977]" class="Insecta" family="Psychodidae" genus="Berdeniella" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="60" phylum="Arthropoda" rank="species" species="keroveci">
<emphasis id="B964EAA2EA2AFFC5FECEFC60B96E3ECA" box="[362,596,952,977]" italics="true" pageId="3" pageNumber="60">Berdeniella keroveci</emphasis>
</taxonomicName>
has thus far only been collected from two springs in the Plitvice Lakes National Park watershed in Lika region.
</paragraph>
<paragraph id="8BAF36B0EA2AFFC5FF63FBD8B9A63846" blockId="3.[151,1437,151,1625]" pageId="3" pageNumber="60">
<emphasis id="B964EAA2EA2AFFC5FF63FBD8BA193902" bold="true" box="[199,291,1024,1049]" pageId="3" pageNumber="60">Habitat</emphasis>
. The Bijela rijeka and Crna rijeka streams are situated in the Plitvice Lakes National park in
<collectingCountry id="F3077620EA2AFFC5FAE7FBD8BEA33902" box="[1347,1433,1024,1049]" name="Croatia" pageId="3" pageNumber="60">Croatia</collectingCountry>
. The Plitvice Lakes system is supplied with water from the Matica stream formed by merging of the streams Crna rijeka and Bijela rijeka. The spring of Bijela rijeka is located at an altitude of
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a.s.l. (
<geoCoordinate id="EE245077EA2AFFC5FB49FB91BE44397A" box="[1261,1406,1096,1121]" direction="north" orientation="latitude" pageId="3" pageNumber="60" precision="15" value="44.83472">N 44°50'05&quot;</geoCoordinate>
<geoCoordinate id="EE245077EA2AFFC5FA2FFB91BA30399E" direction="east" orientation="longitude" pageId="3" pageNumber="60" precision="15" value="15.561945">E 15°33'43&quot;</geoCoordinate>
). It is a rheocrene spring surrounded by open canopy vegetation, and dries out only during extremely dry years (Marušić &amp; Ćuruvija, 1990/1991). Aquatic vegetation is abundant in the spring and especially during autumn and winter allochthonous organic material is abundant due to slower stream flow. On the other hand Crna rijeka spring is a typical rheocrene spring surrounded with dense canopy vegetation. It is located at
<quantity id="4CE89B55EA2AFFC5FB47FB00BE1439EB" box="[1251,1326,1240,1265]" metricMagnitude="2" metricUnit="m" metricValue="6.8" pageId="3" pageNumber="60" unit="m" value="680.0">680 m</quantity>
a.s.l., (
<geoCoordinate id="EE245077EA2AFFC5FA22FB01BA3C380E" direction="north" orientation="latitude" pageId="3" pageNumber="60" precision="15" value="44.83722">N 44°50'14&quot;</geoCoordinate>
<geoCoordinate id="EE245077EA2AFFC5FEAAFB25BAA0380E" box="[270,410,1276,1301]" direction="east" orientation="longitude" pageId="3" pageNumber="60" precision="15" value="15.607779">E 15°36'28&quot;</geoCoordinate>
). It has the highest discharge rate in the National park Plitvice Lakes, even during very dry years (Marušić &amp; Ćuruvija, 1990/1991). The water is fast and turbulent, but very shortly the spring divides in two parts and the velocity of the water diminishes.
</paragraph>
</subSubSection>
<subSubSection id="C30A653BEA2AFFC5FF63FAB0B9E83B42" pageId="3" pageNumber="60" type="description">
<paragraph id="8BAF36B0EA2AFFC5FF63FAB0B9E83B42" blockId="3.[151,1437,151,1625]" pageId="3" pageNumber="60">
<emphasis id="B964EAA2EA2AFFC5FF63FAB0BA0D389A" bold="true" box="[199,311,1384,1409]" pageId="3" pageNumber="60">Variation</emphasis>
. The examined specimens from Crna rijeka spring appear to be smaller than those from Bijela rijeka spring, but the differences are not statistically significant. The measurements reported in the description are from the
<typeStatus id="54AB8812EA2AFFC5FF60FA69BBCE38D2" box="[196,244,1457,1481]" pageId="3" pageNumber="60">type</typeStatus>
locality in Bijele rijeka spring. The two specimens from Crna rijeka spring appear to be similar, but smaller. The measurements for these specimens are, for the palps 64–66: 92–92: 96–98: 146–162; for the antennae 68–80: 60–60: 48–50: 34–36: 32–36: 32–36: 30–36: 30–34: 32–34: 30–30: 28–30: 26–28: 28–28: 22–22: 24–26: 32–32; and for the wings 1,86–2,14. It is likely that this variation is due to the differences in environmental conditions between the two springs.
</paragraph>
</subSubSection>
</treatment>
</document>
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