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<document id="56F1ECC410A9FF97FE8791518060CC0B" ID-CLB-Dataset="51794" ID-GBIF-Dataset="3098b92e-9f08-4f18-8cde-3c5830d7f16c" checkinTime="1427463568664" checkinUser="donat" docAuthor="John S. Lapolla, Robert J. Kallal &amp; Seán G. Brady" docDate="2012" docId="039C9A6B641094531566D7886570F9AD" docLanguage="en" docName="27998.pdf" docOrigin="Systematic Entomology 37" docTitle="Zatania electra Lapolla, Kallal &amp; Brady, 2012, sp.n." docType="treatment" docVersion="8" lastPageNumber="209" masterDocId="FFA5E2136416945A1658D32F6769FF94" masterDocTitle="A new ant genus from the Greater Antilles and Central America, Zatania (Hymenoptera: Formicidae), exemplifies the utility of male and molecular character systems" masterLastPageNumber="214" masterPageNumber="200" pageNumber="206" updateTime="1732752850568" updateUser="ExternalLinkService">
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<mods:title id="5AD44631752D89F8A58C1902149FEA6C">A new ant genus from the Greater Antilles and Central America, Zatania (Hymenoptera: Formicidae), exemplifies the utility of male and molecular character systems</mods:title>
</mods:titleInfo>
<mods:name id="04091C08F46B3AAEDFD160995CDC32A7" type="personal">
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<mods:roleTerm id="657F87ACEFEBCF704C5FC457CF3FDB00">Author</mods:roleTerm>
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<mods:namePart id="6017FFC4E7D10646F7CA77B9A9C448DC">John S. Lapolla</mods:namePart>
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<mods:namePart id="261061DCAC13A6C8FE558C2C86F3391A">Robert J. Kallal</mods:namePart>
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<mods:name id="2253D125107474915BD649CF5766137C" type="personal">
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<mods:roleTerm id="0FE919BAF9FD48B8F45BD9C56B0AF11C">Author</mods:roleTerm>
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<mods:namePart id="21369BC4E6255BEC33B576EA9F664061">Seán G. Brady</mods:namePart>
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<mods:title id="54FAD339A0520EC8F0C46107CA1E06F9">Systematic Entomology</mods:title>
</mods:titleInfo>
<mods:part id="DBAD35F1170A83C21D3B557A40F9E37A">
<mods:date id="AA661EB6B082DBB0FB40E9DEFC099D18">2012</mods:date>
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<mods:number id="C7F989AF67801072B1FDAC05648C232F">37</mods:number>
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<mods:classification id="F92C6EA150B43E5E6DD55EBAEE30E954">journal article</mods:classification>
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<treatment id="039C9A6B641094531566D7886570F9AD" ID-DOI="http://doi.org/10.5281/zenodo.6179232" ID-GBIF-Taxon="114923629" ID-Zenodo-Dep="6179232" LSID="urn:lsid:zoobank.org:act:8D0D7E92-39FB-493A-B414-358C67789848" httpUri="http://treatment.plazi.org/id/039C9A6B641094531566D7886570F9AD" lastPageId="9" lastPageNumber="209" pageId="6" pageNumber="206" scope_family="Formicidae" scope_order="Hymenoptera">
<subSubSection id="C32F78F66410945C1566D7886375FB28" box="[830,1052,1187,1212]" pageId="6" pageNumber="206" type="nomenclature">
<paragraph id="8B8A2B7D6410945C1566D7886375FB28" box="[830,1052,1187,1212]" pageId="6" pageNumber="206">
<taxonomicName id="4C3550FE6410945C1566D78864BBFB28" box="[830,978,1191,1212]" class="Insecta" family="Formicidae" genus="Zatania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="206" phylum="Arthropoda" rank="species" species="electra" status="sp.n.">
<emphasis id="B941F76F6410945C1566D78864BBFB28" box="[830,978,1191,1212]" italics="true" pageId="6" pageNumber="206">Zatania electra</emphasis>
</taxonomicName>
†,
<emphasis id="B941F76F6410945C15B4D7886375FB28" bold="true" box="[1004,1052,1191,1212]" pageId="6" pageNumber="206">
<taxonomicNameLabel id="A2724A146410945C15B4D7886375FB28" box="[1004,1052,1191,1212]" pageId="6" pageNumber="206">sp.n.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C32F78F66410945C1566D7EB638CFB4D" box="[830,1253,1220,1241]" pageId="6" pageNumber="206" type="description">
<paragraph id="8B8A2B7D6410945C1566D7EB638CFB4D" box="[830,1253,1220,1241]" pageId="6" pageNumber="206">
[
<figureCitation id="130E37F86410945C151CD7EB64F8FB4D" box="[836,913,1220,1241]" captionStart="Fig. 30" captionStartId="9.[151,185,927,947]" captionTargetBox="[159,749,174,892]" captionTargetId="figure@9.[159,751,174,894]" captionTargetPageId="9" captionText="Fig. 30. Zatania electra †: A, photograph of holotype worker; B, illustration of holotype worker." pageId="6" pageNumber="206">Figs 30</figureCitation>
a, b and 31b (worker), 31a (male)]
</paragraph>
</subSubSection>
<subSubSection id="C32F78F66410945C1566D7D26382FADA" pageId="6" pageNumber="206" type="materials_examined">
<paragraph id="8B8A2B7D6410945C1566D7D26382FADA" pageId="6" pageNumber="206">Holotype worker, Dominican Amber fossil, AMNH DR10- 159; one paratype worker, Dominican Amber fossil (MCZC); and one paratype male, AMNH PB-118-25.</paragraph>
</subSubSection>
<subSubSection id="C32F78F66410945C150CD6496315FA67" pageId="6" pageNumber="206" type="diagnosis">
<paragraph id="8B8A2B7D6410945C150CD6496315FA67" pageId="6" pageNumber="206">
<emphasis id="B941F76F6410945C150CD649636AFAEF" box="[852,1027,1382,1403]" italics="true" pageId="6" pageNumber="206">Worker diagnosis.</emphasis>
Dominican amber fossil species. Measurements (
<emphasis id="B941F76F6410945C15BAD6AB6486FA0D" bold="true" box="[994,1007,1412,1433]" italics="true" pageId="6" pageNumber="206">n</emphasis>
= 1): TL 2.80 mm; HW 0.57 mm; HL 0.73 mm; EL 0.17 mm; SL 1.18 mm; PW 0.41 mm; WL 1.14 mm; PrFL 0.84 mm; PrFW 0.16 mm; GL 0.94 mm. Indices: CI 78; REL 23; SI 207.
</paragraph>
</subSubSection>
<subSubSection id="C32F78F6641094521566D50A63E1F8DD" lastPageId="8" lastPageNumber="208" pageId="6" pageNumber="206" type="description">
<paragraph id="8B8A2B7D6410945C1566D50A636AF9AF" box="[830,1027,1573,1595]" pageId="6" pageNumber="206">
<heading id="D0C29C116410945C1566D50A636AF9AF" bold="true" box="[830,1027,1573,1595]" fontSize="9" level="1" pageId="6" pageNumber="206" reason="6">
<emphasis id="B941F76F6410945C1566D50A636AF9AF" bold="true" box="[830,1027,1573,1595]" pageId="6" pageNumber="206">Species description</emphasis>
</heading>
</paragraph>
<paragraph id="8B8A2B7D6410945C150CD56B62F4F8DD" pageId="6" pageNumber="206">
<emphasis id="B941F76F6410945C150CD56B64F4F9CD" box="[852,925,1604,1625]" italics="true" pageId="6" pageNumber="206">Worker.</emphasis>
Coloration difficult to ascertain in amber matrix, but appears to be either uniformly dark or reddish brown. Head distinctly longer than wide (CI 78) and strongly oval in shape; mandibles seemingly with five teeth; posterolateral corners of head indistinct and rounded; posterior margin short; scattered macrosetae on head; difficult to ascertain if pubescence is present. Scapes surpass posterior margin by about length of first six or seven funicular segments; scapes with thin, short erect setae, difficult to ascertain if pubescence present.
</paragraph>
<caption id="DF4A7BF56411945D16CFD43564EFF8DC" pageId="7" pageNumber="207" targetBox="[231,1347,174,1785]" targetPageId="7">
<paragraph id="8B8A2B7D6411945D16CFD43564EFF8DC" pageId="7" pageNumber="207">
<emphasis id="B941F76F6411945D16CFD4356669F8B9" bold="true" box="[151,256,1818,1837]" pageId="7" pageNumber="207">Figs 421.</emphasis>
<taxonomicName id="4C3550FE6411945D1752D4346625F8B9" box="[266,332,1819,1837]" class="Insecta" family="Formicidae" genus="Zatania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="207" phylum="Arthropoda" rank="genus">
<emphasis id="B941F76F6411945D1752D4346625F8B9" box="[266,332,1819,1837]" italics="true" pageId="7" pageNumber="207">Zatania</emphasis>
</taxonomicName>
workers in various views (left to right: dorsal, full frontal, lateral and scape): 47,
<taxonomicName id="4C3550FE6411945D1211D434626AF8B9" box="[1097,1283,1819,1837]" class="Insecta" family="Formicidae" genus="Zatania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="207" phylum="Arthropoda" rank="species" species="albimaculata">
<emphasis id="B941F76F6411945D1211D434626AF8B9" box="[1097,1283,1819,1837]" italics="true" pageId="7" pageNumber="207">Zatania albimaculata</emphasis>
</taxonomicName>
; 811,
<taxonomicName id="4C3550FE6411945D1309D43467A3F8DC" class="Insecta" family="Formicidae" genus="Zatania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="207" phylum="Arthropoda" rank="species" species="cisipa">
<emphasis id="B941F76F6411945D1309D43467A3F8DC" italics="true" pageId="7" pageNumber="207">Zatania cisipa</emphasis>
</taxonomicName>
; 1215,
<taxonomicName id="4C3550FE6411945D1746D41A66D2F8DC" box="[286,443,1845,1864]" class="Insecta" family="Formicidae" genus="Zatania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="207" phylum="Arthropoda" rank="species" species="gibberosa">
<emphasis id="B941F76F6411945D1746D41A66D2F8DC" box="[286,443,1845,1864]" italics="true" pageId="7" pageNumber="207">Zatania gibberosa</emphasis>
</taxonomicName>
; 1619,
<taxonomicName id="4C3550FE6411945D1457D41A65F7F8DC" box="[527,670,1845,1864]" class="Insecta" family="Formicidae" genus="Zatania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="207" phylum="Arthropoda" rank="species" species="gloriosa">
<emphasis id="B941F76F6411945D1457D41A65F7F8DC" box="[527,670,1845,1864]" italics="true" pageId="7" pageNumber="207">Zatania gloriosa</emphasis>
</taxonomicName>
; 2021,
<taxonomicName id="4C3550FE6411945D14AAD41A64EBF8DC" box="[754,898,1845,1864]" class="Insecta" family="Formicidae" genus="Zatania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="207" phylum="Arthropoda" rank="species" species="karstica">Zatania karstica</taxonomicName>
.
</paragraph>
</caption>
<caption id="DF4A7BF5641E945216F9D66663EBFAE3" pageId="8" pageNumber="208" targetBox="[312,1284,174,1320]" targetPageId="8">
<paragraph id="8B8A2B7D641E945216F9D66663EBFAE3" pageId="8" pageNumber="208">
<emphasis id="B941F76F641E945216F9D666667CFAC8" bold="true" box="[161,277,1353,1372]" pageId="8" pageNumber="208">Figs 2229.</emphasis>
<taxonomicName id="4C3550FE641E94521747D66566F5FAC9" box="[287,412,1354,1373]" class="Insecta" family="Formicidae" genus="Zatania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="208" phylum="Arthropoda" rank="species" species="cisipa">
<emphasis id="B941F76F641E94521747D66566F5FAC9" box="[287,412,1354,1373]" italics="true" pageId="8" pageNumber="208">Zatania cisipa</emphasis>
</taxonomicName>
reproductives: 22, queen in full-frontal view; 23, queen in lateral view; 24, male in full-frontal view; 25, male in lateral view; 2629, male genitalia (dorsal, ventral, ventral close-up of digitus and cuspis, and lateral parameres).
</paragraph>
</caption>
<paragraph id="8B8A2B7D641E945216F9D6826572F9CC" pageId="8" pageNumber="208">Eyes relatively small, but distinctly convex. Pronotum with scattered erect setae; erect setae on notum; propodeal erect setae shorter than other notal setae; distinct constriction present immediately behind pronotum; metanotal area long; dorsal face of propodeum dome-like, higher than remainder of notum; gaster with scattered erect macrosetae.</paragraph>
<paragraph id="8B8A2B7D641E945216EED55E6605F911" box="[182,364,1648,1669]" pageId="8" pageNumber="208">
<emphasis id="B941F76F641E945216EED55E6795F911" box="[182,252,1649,1669]" italics="true" pageId="8" pageNumber="208">Queen.</emphasis>
Unknown.
</paragraph>
<paragraph id="8B8A2B7D641E945216EED5B164C7F945" pageId="8" pageNumber="208">
<emphasis id="B941F76F641E945216EED5B16786F927" box="[182,239,1694,1715]" italics="true" pageId="8" pageNumber="208">Male.</emphasis>
Coloration difficult to ascertain in amber matrix but appears to be either uniformly dark or reddish brown. Head slightly wider than long, dominated by large, convex compound eyes that take up most of lateral margin of head; mandibles, with prominent, pointed apical tooth; masticatory margin partly obscured, but seemingly with indistinct denticles, similar to what is observed in
<taxonomicName id="4C3550FE641E94521228D6826380FA56" box="[1136,1257,1453,1474]" class="Insecta" family="Formicidae" genus="Prenolepis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="208" phylum="Arthropoda" rank="species" species="gibberosa">
<emphasis id="B941F76F641E94521228D6826380FA56" box="[1136,1257,1453,1474]" italics="true" pageId="8" pageNumber="208">Z. gibberosa</emphasis>
</taxonomicName>
males; head with scattered erect macrosetae; difficult to ascertain if pubescence present. Scapes surpass posterior margin by about length of first five or six funicular segments; scapes with short, thin erect setae. Mesosoma modified for presence of wing; erect setae scattered across mesonotum and metanotum; propodeum indistinct with very short dorsal face and long declivitous face. Gaster with scattered erect setae; genitalia with long, narrow parameres, similar to what is observed in
<taxonomicName id="4C3550FE641E94521342D5B16204F926" box="[1306,1389,1694,1714]" class="Insecta" family="Formicidae" genus="Paratrechina" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="208" phylum="Arthropoda" rank="species" species="cisipa">
<emphasis id="B941F76F641E94521342D5B16204F926" box="[1306,1389,1694,1714]" italics="true" pageId="8" pageNumber="208">Z. cisipa</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3550FE641E94521566D59364C1F945" box="[830,936,1724,1745]" class="Insecta" family="Formicidae" genus="Zatania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="208" phylum="Arthropoda" rank="species" species="gloriosa">
<emphasis id="B941F76F641E94521566D59364C1F945" box="[830,936,1724,1745]" italics="true" pageId="8" pageNumber="208">Z. gloriosa</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B8A2B7D641E94521566D5F563E1F8DD" pageId="8" pageNumber="208">
Measurements (
<emphasis id="B941F76F641E945215BAD5F56486F97B" bold="true" box="[994,1007,1754,1775]" italics="true" pageId="8" pageNumber="208">n</emphasis>
= 1): TL 2.85 mm; HW 0.68 mm; HL 0.62 mm; EL 0.29 mm; SL 0.98 mm; PW 0.51 mm; WL 1.28 mm; PrFL 1.01 mm; PrFW 0.15 mm; GL 0.95 mm. Indices: CI 101; REL 47; SI 145.
</paragraph>
</subSubSection>
<caption id="DF4A7BF5641F945316CFD0B066C1FC5A" pageId="9" pageNumber="209" targetBox="[159,749,174,892]" targetPageId="9">
<paragraph id="8B8A2B7D641F945316CFD0B066C1FC5A" pageId="9" pageNumber="209">
<emphasis id="B941F76F641F945316CFD0B06789FC27" bold="true" box="[151,224,927,947]" pageId="9" pageNumber="209">Fig. 30.</emphasis>
<taxonomicName id="4C3550FE641F945316B3D08E661AFC27" box="[235,371,929,947]" class="Insecta" family="Formicidae" genus="Zatania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="209" phylum="Arthropoda" rank="species" species="electra">
<emphasis id="B941F76F641F945316B3D08E661AFC27" box="[235,371,929,947]" italics="true" pageId="9" pageNumber="209">Zatania electra</emphasis>
</taxonomicName>
†: A, photograph of holotype worker; B, illustration of holotype worker.
</paragraph>
</caption>
<subSubSection id="C32F78F6641F945316F5D72E666FFBC5" pageId="9" pageNumber="209" type="etymology">
<paragraph id="8B8A2B7D641F945316F5D72E666FFBC5" pageId="9" pageNumber="209">
<emphasis id="B941F76F641F945316F5D72E6673FB82" box="[173,282,1025,1046]" italics="true" pageId="9" pageNumber="209">Etymology.</emphasis>
The species epithet is in reference to the fact that this is a fossil species found in amber (
<taxonomicName id="4C3550FE641F9453140CD73065FFFBA0" box="[596,662,1055,1076]" class="Insecta" family="Formicidae" genus="Zatania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="209" phylum="Arthropoda" rank="species" species="electra">
<emphasis id="B941F76F641F9453140CD73065FFFBA0" box="[596,662,1055,1076]" italics="true" pageId="9" pageNumber="209">electra</emphasis>
</taxonomicName>
is Greek for amber).
</paragraph>
</subSubSection>
<subSubSection id="C32F78F6641F945316F5D74A6570F9AD" pageId="9" pageNumber="209" type="discussion">
<paragraph id="8B8A2B7D641F945316F5D74A6570F9AD" pageId="9" pageNumber="209">
<emphasis id="B941F76F641F945316F5D74A6782FBEE" box="[173,235,1125,1146]" italics="true" pageId="9" pageNumber="209">Notes.</emphasis>
This is the only fossil species known in the genus. The workers, with respect to some morphological features, resemble the workers of
<taxonomicName id="4C3550FE641F945317F3D78E652DFB22" box="[427,580,1185,1206]" class="Insecta" family="Formicidae" genus="Zatania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="209" phylum="Arthropoda" rank="species" species="albimaculata">
<emphasis id="B941F76F641F945317F3D78E652DFB22" box="[427,580,1185,1206]" italics="true" pageId="9" pageNumber="209">Z. albimaculata</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3550FE641F945314D0D78E6586FB22" box="[648,751,1185,1206]" class="Insecta" family="Formicidae" genus="Zatania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="209" phylum="Arthropoda" rank="species" species="karstica">
<emphasis id="B941F76F641F945314D0D78E6586FB22" box="[648,751,1185,1206]" italics="true" pageId="9" pageNumber="209">Z. karstica</emphasis>
</taxonomicName>
, especially with regards to the relatively small size of the compound eyes and the constriction of the mesosoma immediately behind the pronotum. The dome-like propodeum is most similar to
<taxonomicName id="4C3550FE641F9453171ED63766D6FAB9" box="[326,447,1304,1325]" class="Insecta" family="Formicidae" genus="Prenolepis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="209" phylum="Arthropoda" rank="species" species="gibberosa">
<emphasis id="B941F76F641F9453171ED63766D6FAB9" box="[326,447,1304,1325]" italics="true" pageId="9" pageNumber="209">Z. gibberosa</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3550FE641F945317A8D6376531FAB9" box="[496,600,1304,1325]" class="Insecta" family="Formicidae" genus="Zatania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="209" phylum="Arthropoda" rank="species" species="karstica">
<emphasis id="B941F76F641F945317A8D6376531FAB9" box="[496,600,1304,1325]" italics="true" pageId="9" pageNumber="209">Z. karstica</emphasis>
</taxonomicName>
. The apparently uniform body colour of the fossil species is only shared with
<taxonomicName id="4C3550FE641F94531691D67B6658FAFC" box="[201,305,1364,1384]" class="Insecta" family="Formicidae" genus="Zatania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="209" phylum="Arthropoda" rank="species" species="karstica">
<emphasis id="B941F76F641F94531691D67B6658FAFC" box="[201,305,1364,1384]" italics="true" pageId="9" pageNumber="209">Z. karstica</emphasis>
</taxonomicName>
among extant species. Interestingly, however, despite the similarities of the workers to the three mentioned extant species, the narrow male parameres most closely resemble those of
<taxonomicName id="4C3550FE641F94531715D68266C8FA56" box="[333,417,1453,1474]" class="Insecta" family="Formicidae" genus="Paratrechina" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="209" phylum="Arthropoda" rank="species" species="cisipa">
<emphasis id="B941F76F641F94531715D68266C8FA56" box="[333,417,1453,1474]" italics="true" pageId="9" pageNumber="209">Z. cisipa</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3550FE641F9453178ED6826529FA56" box="[470,576,1453,1474]" class="Insecta" family="Formicidae" genus="Zatania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="209" phylum="Arthropoda" rank="species" species="gloriosa">
<emphasis id="B941F76F641F9453178ED6826529FA56" box="[470,576,1453,1474]" italics="true" pageId="9" pageNumber="209">Z. gloriosa</emphasis>
</taxonomicName>
. The narrow head of the
<taxonomicName id="4C3550FE641F94531684D6E46650FA4B" box="[220,313,1483,1503]" class="Insecta" family="Formicidae" genus="Zatania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="209" phylum="Arthropoda" rank="species" species="electra">
<emphasis id="B941F76F641F94531684D6E46650FA4B" box="[220,313,1483,1503]" italics="true" pageId="9" pageNumber="209">Z. electra</emphasis>
</taxonomicName>
worker is most similar to what is observed in
<taxonomicName id="4C3550FE641F945316EED6C76649FA69" box="[182,288,1512,1533]" class="Insecta" family="Formicidae" genus="Zatania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="209" phylum="Arthropoda" rank="species" species="gloriosa">
<emphasis id="B941F76F641F945316EED6C76649FA69" box="[182,288,1512,1533]" italics="true" pageId="9" pageNumber="209">Z. gloriosa</emphasis>
</taxonomicName>
†. It therefore appears that
<taxonomicName id="4C3550FE641F94531467D6C765F2FA69" box="[575,667,1512,1533]" class="Insecta" family="Formicidae" genus="Zatania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="209" phylum="Arthropoda" rank="species" species="electra">
<emphasis id="B941F76F641F94531467D6C765F2FA69" box="[575,667,1512,1533]" italics="true" pageId="9" pageNumber="209">Z. electra</emphasis>
</taxonomicName>
† has an interesting combination of morphological features observed among all of the known extant species.
</paragraph>
</subSubSection>
</treatment>
</document>