176 lines
21 KiB
XML
176 lines
21 KiB
XML
<document id="D833BB2A65E4B7CBB556C78D8A5C9B95" ID-CLB-Dataset="8783" ID-DOI="10.11646/zootaxa.4908.3.1" ID-GBIF-Dataset="6206d142-3b21-4c39-afca-75b602420619" ID-ISSN="1175-5326" ID-Zenodo-Dep="4441762" ID-ZooBank="F2774810-3C77-426E-A1BC-0F42F21E7F9B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1610699400433" checkinUser="plazi" docAuthor="Snegovaya, Nataly Yu. & Cokendolpher, James C." docDate="2021" docId="039C113F095DFFA598EBDDFBFE3FFAC3" docLanguage="en" docName="zootaxa.4908.3.1.pdf" docOrigin="Zootaxa 4908 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Homolophus milkoi Snegovaya & Cokendolpher 2021, sp. nov." docType="treatment" docVersion="7" lastPageNumber="338" masterDocId="FFA569470978FF80987CDD6CFFAEFFF1" masterDocTitle="Further studies on harvestman genus Homolophus (Opiliones: Phalangiidae), with descriptions of two new species" masterLastPageNumber="353" masterPageNumber="301" pageNumber="338" updateTime="1698924489785" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="00DF415435A626D8C2E4DD7A8F720C66">Further studies on harvestman genus Homolophus (Opiliones: Phalangiidae), with descriptions of two new species</mods:title>
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<mods:name id="B237D4679E88EAD259A5A3C23EE7661D" type="personal">
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<mods:roleTerm id="1A242ED6AC7C60258650468DCA2891E0">Author</mods:roleTerm>
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<mods:namePart id="C145EC2266CD28963D048F3C0F4022CE">Snegovaya, Nataly Yu.</mods:namePart>
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<mods:affiliation id="DBAF22E529AA0293F3D2DDF1E8433794">Institute of Zoology, Azerbaijan Academy of Sciences, pr. 1128, kv. 504, Baku AZ 1073, Azerbaijan.</mods:affiliation>
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<mods:namePart id="E5F47BBD3C5D231F08C765FD30DE9CF1">Cokendolpher, James C.</mods:namePart>
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<mods:affiliation id="347EE5FC2B8A4B8A0B2314A577E184CB">Invertebrate Zoology, Museum of Texas Tech University, Lubbock, Texas 79415, USA.</mods:affiliation>
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<mods:title id="FA84BEE30AC0CE84948649CD5F19384C">Zootaxa</mods:title>
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<mods:date id="060DAE3DB8FC20E307964CBC0D937DFE">2021</mods:date>
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<mods:number id="FB23706C8056138FCF23436F7281CD82">2021-01-15</mods:number>
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<mods:number id="CFBEDD7AC1089F80FE2390859F655D27">4908</mods:number>
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<treatment id="039C113F095DFFA598EBDDFBFE3FFAC3" ID-DOI="http://doi.org/10.5281/zenodo.4441766" ID-GBIF-Taxon="177009872" ID-Zenodo-Dep="4441766" LSID="urn:lsid:plazi:treatment:039C113F095DFFA598EBDDFBFE3FFAC3" httpUri="http://treatment.plazi.org/id/039C113F095DFFA598EBDDFBFE3FFAC3" lastPageNumber="338" pageId="37" pageNumber="338">
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<subSubSection id="C32FF3A2095DFFA598EBDDFBFE43FF43" box="[151,493,151,178]" pageId="37" pageNumber="338" type="nomenclature">
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<paragraph id="8B8AA029095DFFA598EBDDFBFE43FF43" blockId="37.[151,493,151,214]" box="[151,493,151,178]" pageId="37" pageNumber="338">
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<heading id="D0C21745095DFFA598EBDDFBFE43FF43" bold="true" box="[151,493,151,178]" fontSize="11" level="1" pageId="37" pageNumber="338" reason="1">
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<emphasis id="B9417C3B095DFFA598EBDDFBFE43FF43" bold="true" box="[151,493,151,178]" pageId="37" pageNumber="338">
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<taxonomicName id="4C35DBAA095DFFA598EBDDFBFE26FF43" authority="Snegovaya & Cokendolpher, 2021" authorityName="Snegovaya & Cokendolpher" authorityYear="2021" box="[151,392,151,178]" class="Arachnida" family="Phalangiidae" genus="Homolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="37" pageNumber="338" phylum="Arthropoda" rank="species" species="milkoi" status="sp. nov.">
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<emphasis id="B9417C3B095DFFA598EBDDFBFE26FF43" bold="true" box="[151,392,151,178]" italics="true" pageId="37" pageNumber="338">Homolophus milkoi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A272C140095DFFA599F3DDF4FE43FF43" box="[399,493,152,178]" pageId="37" pageNumber="338" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32FF3A2095DFFA598EBDDD0FEFAFF27" box="[151,340,188,214]" pageId="37" pageNumber="338" type="description">
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<paragraph id="8B8AA029095DFFA598EBDDD0FEFAFF27" blockId="37.[151,493,151,214]" box="[151,340,188,214]" pageId="37" pageNumber="338">
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<figureCitation id="130EBCAC095DFFA598EBDDD0FEACFF27" box="[151,258,188,214]" captionStart="FIGURE 7" captionStartId="15.[151,250,1630,1654]" captionTargetBox="[156,1428,709,1600]" captionTargetId="figure-289@15.[152,1436,705,1605]" captionTargetPageId="15" captionText="FIGURE 7. Distribution of some Homolophus species in the Palearctic: Central and Inner Asia. Data for pale colored symbols taken from literature; identifications not verified." figureDoi="http://doi.org/10.5281/zenodo.4441782" httpUri="https://zenodo.org/record/4441782/files/figure.png" pageId="37" pageNumber="338">Figures 7</figureCitation>
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,
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<figureCitation id="130EBCAC095DFFA59972DDD0FE82FF27" box="[270,300,188,214]" captionStart="FIGURE 24" captionStartId="34.[151,250,988,1012]" captionTargetBox="[170,1436,181,965]" captionTargetId="figure-22@34.[151,1436,181,965]" captionTargetPageId="34" captionText="FIGURE 24. Famous scientists: (left) Pyotr Kuz’mich Kozlov (1863–1935), a Russian traveler, geographer, ethnographer, archeologist, and explorer of Mongolia and Xîzàng (Tibet), for which Homolophus kozlovi sp. nov. is named; (right) Dmitry A. Milko, an entomologist, researcher, and zoologist from Kyrgyzstan for which Homolophus milkoi sp. nov. is named." figureDoi="http://doi.org/10.5281/zenodo.4441825" httpUri="https://zenodo.org/record/4441825/files/figure.png" pageId="37" pageNumber="338">24</figureCitation>
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,
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<figureCitation id="130EBCAC095DFFA59944DDD0FEFAFF27" box="[312,340,188,214]" captionStart="FIGURE 27" captionStartId="39.[151,250,1943,1967]" captionTargetBox="[193,1395,181,1909]" captionTargetId="figure-17@39.[189,1399,181,1919]" captionTargetPageId="39" captionText="FIGURE 27. Homolophus milkoi sp. nov., male: A, body, dorsal; B, ocularium, dorsal, C, right chelicera, ectal; D, right che- licera, mesal; E, right pedipalp, ectal; F, right pedipalp, mesal; G, femur I, dorsal; H, penis, lateral; I, penis glans, lateral; J, penis glans, dorsal; K, penis, dorsal." figureDoi="http://doi.org/10.5281/zenodo.4441833" httpUri="https://zenodo.org/record/4441833/files/figure.png" pageId="37" pageNumber="338">27</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32FF3A2095DFFA598EBDC6FFDA4FE06" pageId="37" pageNumber="338" type="diagnosis">
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<paragraph id="8B8AA029095DFFA598EBDC6FFDA4FE06" blockId="37.[151,1437,259,1330]" pageId="37" pageNumber="338">
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<emphasis id="B9417C3B095DFFA598EBDC6FFE4DFEEC" bold="true" box="[151,483,259,286]" pageId="37" pageNumber="338">Diagnosis and Comparisons</emphasis>
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. Body very dark brown with broad white belt on anterior free tergites and wide bright white areas on sides of body. Numerous darker spots in rows on white areas of anterior part of abdomen. Coloring pattern distinctive and only known from new species and
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<taxonomicName id="4C35DBAA095DFFA59B2ADC20FA32FE97" authority="Starega & Snegovaya, 2008" authorityName="Starega & Snegovaya" authorityYear="2008" box="[854,1436,332,358]" class="Arachnida" family="Phalangiidae" genus="Homolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="37" pageNumber="338" phylum="Arthropoda" rank="species" species="gricenkoi">
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<emphasis id="B9417C3B095DFFA59B2ADC20FBF9FE97" box="[854,1111,332,358]" italics="true" pageId="37" pageNumber="338">Homolophus gricenkoi</emphasis>
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<bibRefCitation id="EFA4DDD8095DFFA59C23DC20FA32FE97" author="Snegovaya, N. Yu. & Starega, W." box="[1119,1436,332,358]" pageId="37" pageNumber="338" pagination="5 - 7" refId="ref33152" refString="Snegovaya, N. Yu. & Starega, W. (2008) Redikorcevia platybunoides gen. & sp. n., a new harvestman from Kazakhstan, with establishment of a new tribe Scleropilionini trib. n. (Opiliones: Phalangiidae). Acta Arachnologica, 57, 5 - 7. https: // doi. org / 10.2476 / asjaa. 57.5" type="journal article" year="2008">Staręga & Snegovaya, 2008</bibRefCitation>
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</taxonomicName>
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(known only from NW
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<collectingCountry id="F322E0B9095DFFA599D8DC1CFDB6FE7A" box="[420,536,368,395]" name="Tajikistan" pageId="37" pageNumber="338">Tajikistan</collectingCountry>
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). New species differs by having different pattern of white on body (white areas larger around sides of body, ocularium lacking white stripe) and body dark brown rather than black. They also differ in structure of penis. New species with glans more rounded, versus glans noticeably laterally compressed in
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<taxonomicName id="4C35DBAA095DFFA59DFDDCD4FF73FE07" authorityName="Snegovaya & Cokendolpher" authorityYear="2021" class="Arachnida" family="Phalangiidae" genus="Homolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="37" pageNumber="338" phylum="Arthropoda" rank="species" species="milkoi">
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<emphasis id="B9417C3B095DFFA59DFDDCD4FF73FE07" italics="true" pageId="37" pageNumber="338">H. milkoi</emphasis>
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</taxonomicName>
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(best seen in dorsal view).
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C32FF3A2095DFFA598BBDF6CFC68FD93" pageId="37" pageNumber="338" type="materials_examined">
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<paragraph id="8B8AA029095DFFA598BBDF6CFF52FDCF" blockId="37.[151,1437,259,1330]" pageId="37" pageNumber="338">
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<materialsCitation id="3B5DAA74095DFFA598BBDF6CFF56FDCF" ID-GBIF-Occurrence="3019649318" collectorName="NW Kirgizia & Talas Mt. Ridge" country="Kyrgyzstan" elevation="2900" latitude="42.166668" location="Talas Mt." longLatPrecision="1152" longitude="71.55" municipality="Mt. Ridge" pageId="37" pageNumber="338" specimenCount="1" typeStatus="holotype">
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<emphasis id="B9417C3B095DFFA598BBDF6CFEC7FDE8" bold="true" box="[199,361,511,538]" pageId="37" pageNumber="338">
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<typeStatus id="548E1E8B095DFFA598BBDF6CFEACFDEB" box="[199,258,512,538]" pageId="37" pageNumber="338">Type</typeStatus>
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Locality
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</emphasis>
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.
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<collectingCountry id="F322E0B9095DFFA5990ADF6CFE59FDEB" box="[374,503,512,538]" name="Kyrgyzstan" pageId="37" pageNumber="338">Kyrgyzstan</collectingCountry>
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[=
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<collectorName id="26C0C5FF095DFFA59A66DF6CFD01FDEB" box="[538,687,511,538]" pageId="37" pageNumber="338">NW Kirgizia</collectorName>
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],
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<collectorName id="26C0C5FF095DFFA59AC2DF6CFCDCFDEB" box="[702,882,512,538]" pageId="37" pageNumber="338">
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<location id="8EEAF6F2095DFFA59AC2DF6CFC89FDEB" LSID="urn:lsid:plazi:treatment:039C113F095DFFA598EBDDFBFE3FFAC3:8EEAF6F2095DFFA59AC2DF6CFC89FDEB" box="[702,807,512,538]" country="Kyrgyzstan" latitude="42.166668" longLatPrecision="1152" longitude="71.55" municipality="Mt. Ridge" name="Talas Mt." pageId="37" pageNumber="338">
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Talas
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<collectingMunicipality id="6BEE3A53095DFFA59A83DF6CFC89FDEB" box="[767,807,512,538]" pageId="37" pageNumber="338">Mt.</collectingMunicipality>
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</location>
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Ridge
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</collectorName>
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, Chon-Kashka-Suu gorge,
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<elevation id="0018471A095DFFA59CE7DF6CFA87FDEB" box="[1179,1321,512,538]" metricMagnitude="3" metricUnit="m" metricValue="2.9" pageId="37" pageNumber="338" unit="m" value="2900.0">
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<quantity id="4CCD0DCC095DFFA59CE7DF6CFB41FDEB" box="[1179,1263,512,538]" metricMagnitude="3" metricUnit="m" metricValue="2.9" pageId="37" pageNumber="338" unit="m" value="2900.0">2900 m</quantity>
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a.s.l.
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</elevation>
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,
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<geoCoordinate id="EE01C6EE095DFFA59D48DF6CFA39FDEB" box="[1332,1431,512,538]" degrees="42" direction="north" minutes="10" orientation="latitude" pageId="37" pageNumber="338" precision="925" value="42.166668">42°10’N</geoCoordinate>
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,
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<geoCoordinate id="EE01C6EE095DFFA598EBDF48FF56FDCF" box="[151,248,548,574]" degrees="71" direction="east" minutes="33" orientation="longitude" pageId="37" pageNumber="338" precision="925" value="71.55">71°33’E</geoCoordinate>
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="8B8AA029095DFFA598BBDF24FC68FD93" blockId="37.[151,1437,259,1330]" box="[199,966,583,610]" pageId="37" pageNumber="338">
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<emphasis id="B9417C3B095DFFA598BBDF24FEACFD93" bold="true" box="[199,258,584,610]" pageId="37" pageNumber="338">Type</emphasis>
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||
<materialsCitation id="3B5DAA74095DFFA59975DF24FC6CFD93" ID-GBIF-Occurrence="3019649314" box="[265,962,583,610]" collectingDate="2017-08-09" collectionCode="ZMMU" collectorName="D. Milko" pageId="37" pageNumber="338" specimenCount="1" typeStatus="holotype">
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||
M
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<emphasis id="B9417C3B095DFFA5995EDF2BFEDEFD90" bold="true" box="[290,368,583,609]" pageId="37" pageNumber="338">aterial</emphasis>
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.
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||
<date id="FF8B86E9095DFFA59901DF24FE51FD93" box="[381,511,584,610]" pageId="37" pageNumber="338" value="2017-08-09">
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||
<collectingDate id="EFCF7F01095DFFA59901DF24FE51FD93" box="[381,511,584,610]" pageId="37" pageNumber="338" value="2017-08-09">08.09.2017</collectingDate>
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</date>
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||
, leg.
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||
<collectorName id="26C0C5FF095DFFA59A46DF24FD0EFD93" box="[570,672,584,610]" pageId="37" pageNumber="338">D. Milko</collectorName>
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(male
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||
<typeStatus id="548E1E8B095DFFA59A92DF24FCFDFD93" box="[750,851,584,610]" pageId="37" pageNumber="338" type="holotype">holotype</typeStatus>
|
||
,
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<collectionCode id="ED2438EC095DFFA59B22DF24FC14FD93" box="[862,954,584,610]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="37" pageNumber="338" type="Museum">ZMMU</collectionCode>
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||
)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32FF3A2095DFFA598BBDF07FC66FD77" box="[199,968,619,646]" pageId="37" pageNumber="338" type="distribution">
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<paragraph id="8B8AA029095DFFA598BBDF07FC66FD77" blockId="37.[151,1437,259,1330]" box="[199,968,619,646]" pageId="37" pageNumber="338">
|
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<emphasis id="B9417C3B095DFFA598BBDF07FEF7FD74" bold="true" box="[199,345,619,645]" pageId="37" pageNumber="338">Distribution</emphasis>
|
||
. Only known from
|
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<typeStatus id="548E1E8B095DFFA59A4BDF00FDC9FD77" box="[567,615,620,646]" pageId="37" pageNumber="338">type</typeStatus>
|
||
locality in
|
||
<collectingCountry id="F322E0B9095DFFA59A9ADF00FCC9FD77" box="[742,871,620,646]" name="Kyrgyzstan" pageId="37" pageNumber="338">Kyrgyzstan</collectingCountry>
|
||
(
|
||
<figureCitation id="130EBCAC095DFFA59B0ADF00FC14FD77" box="[886,954,620,646]" captionStart="FIGURE 7" captionStartId="15.[151,250,1630,1654]" captionTargetBox="[156,1428,709,1600]" captionTargetId="figure-289@15.[152,1436,705,1605]" captionTargetPageId="15" captionText="FIGURE 7. Distribution of some Homolophus species in the Palearctic: Central and Inner Asia. Data for pale colored symbols taken from literature; identifications not verified." figureDoi="http://doi.org/10.5281/zenodo.4441782" httpUri="https://zenodo.org/record/4441782/files/figure.png" pageId="37" pageNumber="338">Fig. 7</figureCitation>
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||
).
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C32FF3A2095DFFA598BBDFFCFA25FD3F" pageId="37" pageNumber="338" type="etymology">
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<paragraph id="8B8AA029095DFFA598BBDFFCFA25FD3F" blockId="37.[151,1437,259,1330]" pageId="37" pageNumber="338">
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||
<emphasis id="B9417C3B095DFFA598BBDFFCFEE6FD5B" bold="true" box="[199,328,656,682]" pageId="37" pageNumber="338">Etymology</emphasis>
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. The new species is named after famous researcher, zoologist, and entomologist Dmitry Anatolyevich Milko (
|
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<figureCitation id="130EBCAC095DFFA59968DFD8FEC8FD3F" box="[276,358,692,718]" captionStart="FIGURE 24" captionStartId="34.[151,250,988,1012]" captionTargetBox="[170,1436,181,965]" captionTargetId="figure-22@34.[151,1436,181,965]" captionTargetPageId="34" captionText="FIGURE 24. Famous scientists: (left) Pyotr Kuz’mich Kozlov (1863–1935), a Russian traveler, geographer, ethnographer, archeologist, and explorer of Mongolia and Xîzàng (Tibet), for which Homolophus kozlovi sp. nov. is named; (right) Dmitry A. Milko, an entomologist, researcher, and zoologist from Kyrgyzstan for which Homolophus milkoi sp. nov. is named." figureDoi="http://doi.org/10.5281/zenodo.4441825" httpUri="https://zenodo.org/record/4441825/files/figure.png" pageId="37" pageNumber="338">Fig. 24</figureCitation>
|
||
) of the Biological and Soil Institute of National Academy of Sciences of the
|
||
<collectingCountry id="F322E0B9095DFFA59CB8DFD8FA29FD3F" box="[1220,1415,692,718]" name="Kyrgyzstan" pageId="37" pageNumber="338">Kyrgyz Republic</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32FF3A2095DFFA598BBDFBBFE3FFAC3" pageId="37" pageNumber="338" type="description">
|
||
<paragraph id="8B8AA029095DFFA598BBDFBBFD8DFAFF" blockId="37.[151,1437,259,1330]" pageId="37" pageNumber="338">
|
||
<emphasis id="B9417C3B095DFFA598BBDFBBFEFFFD00" bold="true" box="[199,337,727,753]" pageId="37" pageNumber="338">Description</emphasis>
|
||
. Medium-sized harvestmen, male body rectangular, with rounded corners; medium sclerotization of body cuticle, less slender, 1.44 times longer than wide; length
|
||
<quantity id="4CCD0DCC095DFFA59BF9DF90FC71FCE7" box="[901,991,764,790]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="37" pageNumber="338" unit="mm" value="4.8">4.8 mm</quantity>
|
||
, width 3.0 mm; body dark-brown with broad white belt across pro- and metapeltidium and wide bright white splotches, dots on sides of body which grow into entire regions anteriorly on anterolateral sides of abdomen and cephalothorax; preocular area and anterior edge of cephalothorax also with extensive white coloring. Body with transverse rows of small black-tipped denticles on borders of tergites. Venter, genital operculum, and coxae with only setae. Ocularium low, slightly wider than long, small, 1.5 times its length from anterior margin of cephalothorax with 6–8 black-tipped denticles (with subterminal short erect setae on each denticle) around both center dark lines (dark markings showing muscle attachments sites). Legs long, slender, cylindrical in cross-section, Fe not thickened. Leg lengths (mm): I 5.2 + 1.4 + 4.6 + 5.5 + 8.2 = 24.9, II 11.5 + 1.8 + 10.0 + 8.0 + 21.0 = 52.3, III 4.0 + 1.2 + 4.6 + 6.0 + 9.0 = 24.8, IV 6.5 + 1.2 + 5.6 + 8.0 + 13.2 = 34.5. Venter yellow, coxae light brown. Pedipalps not thickened, Fe only dorsally with some small denticles, other segments only ventrally with microdenticles. Pedipalp segment lengths (mm): 1.05 + 0.6 + 1.25 + 1.6 = 4.5. Chelicerae small, slender, with some tabby marks, without denticles. Cheliceral basal segment ventrally smooth without bulge, length
|
||
<quantity id="4CCD0DCC095DFFA59949D9C7FE33FB37" box="[309,413,1195,1222]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" pageId="37" pageNumber="338" unit="mm" value="1.45">1.45 mm</quantity>
|
||
, distal segment length
|
||
<quantity id="4CCD0DCC095DFFA59ADAD9C7FCA1FB37" box="[678,783,1195,1222]" metricMagnitude="-3" metricUnit="m" metricValue="1.65" pageId="37" pageNumber="338" unit="mm" value="1.65">1.65 mm</quantity>
|
||
. Penis short, truncus dorsoventrally flattened, bowed, recurved in lateral view, wide at base, tapering to glans; glans boat-shaped in lateral view and narrow; length (mm): truncus 2.13, glans 0.38, stylus 0.1.
|
||
</paragraph>
|
||
<paragraph id="8B8AA029095DFFA598BBD874FE3FFAC3" blockId="37.[151,1437,259,1330]" box="[199,401,1304,1330]" pageId="37" pageNumber="338">Female unknown.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |