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<document ID-DOI="http://doi.org/10.5281/zenodo.4761990" ID-GBIF-Dataset="28228da7-1a79-45b8-bb9c-698fe9764316" ID-GBIF-Taxon="182070773" ID-ISSN="1854-0392" ID-Zenodo-Dep="4761990" approvalRequired="1" approvalRequired_for_textStreams="1" checkinTime="1620933921553" checkinUser="felipe" docAuthor="McLellan, Ian D. &amp; Zwick, Peter" docDate="2007" docId="0399AB12FFBBFFCFE91150695246FBC2" docLanguage="en" docName="Illiesia.3.4.20-42.pdf" docOrigin="Illiesia 3 (4)" docStyle="DocumentStyle:1DEF1196F206C1C297CF0540A5C6768F.1:Illesia.2005-2008.journal_article" docStyleId="1DEF1196F206C1C297CF0540A5C6768F" docStyleName="Illesia.2005-2008.journal_article" docStyleVersion="1" docTitle="Uncicauda pirata McLellan &amp; Zwick 2007, gen. n., sp. n." docType="treatment" docVersion="6" lastPageId="4" lastPageNumber="24" masterDocId="FFA0D36AFFB9FFCBE82B5344536CFF9B" masterDocTitle="New Species Of And Keys To South American Gripopterygidae (Plecoptera)" masterLastPageNumber="42" masterPageNumber="20" pageId="2" pageNumber="22" updateTime="1632015891772" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>New Species Of And Keys To South American Gripopterygidae (Plecoptera)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>McLellan, Ian D.</mods:namePart>
<mods:affiliation>and Peter Zwick &amp; P. O. Box 95, Westport, New Zealand E-mail: mclellan @ xtra. co. nz</mods:affiliation>
<mods:nameIdentifier type="email">mclellan@xtra.co.nz</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zwick, Peter</mods:namePart>
<mods:affiliation>Limnologische Fluss-Station des Max-Planck-Instituts für Limnologie, Schlitz, Germany Correspondence: Peter Zwick, Schwarzer Stock 9, D- 36110 Schlitz, Germany E-mail: pleco-p. zwick @ t-online. de</mods:affiliation>
<mods:nameIdentifier type="email">pleco-p.zwick@t-online.de</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Illiesia</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2007</mods:date>
<mods:detail type="pubDate">
<mods:number>2007-06-26</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>3</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>4</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>20</mods:start>
<mods:end>42</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.4758655</mods:identifier>
<mods:identifier type="GBIF-Dataset">28228da7-1a79-45b8-bb9c-698fe9764316</mods:identifier>
<mods:identifier type="ISSN">1854-0392</mods:identifier>
<mods:identifier type="Zenodo-Dep">4758655</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.4761990" ID-GBIF-Taxon="182070773" ID-Zenodo-Dep="4761990" LSID="urn:lsid:plazi:treatment:0399AB12FFBBFFCFE91150695246FBC2" httpUri="http://treatment.plazi.org/id/0399AB12FFBBFFCFE91150695246FBC2" lastPageId="4" lastPageNumber="24" pageId="2" pageNumber="22">
<subSubSection box="[314,682,813,838]" pageId="2" pageNumber="22" type="nomenclature">
<paragraph blockId="2.[314,682,813,838]" box="[314,682,813,838]" pageId="2" pageNumber="22">
<heading bold="true" box="[314,682,813,838]" fontSize="10" level="2" pageId="2" pageNumber="22" reason="2">
<emphasis bold="true" box="[314,682,813,838]" pageId="2" pageNumber="22">
<taxonomicName authority="McLellan &amp; Zwick, 2007" authorityName="McLellan &amp; Zwick" authorityYear="2007" box="[314,516,813,837]" class="Insecta" family="Gripopterygidae" genus="Uncicauda" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="22" phylum="Arthropoda" rank="species" species="pirata" status="gen. n., sp. n.">
<emphasis bold="true" box="[314,516,813,837]" italics="true" pageId="2" pageNumber="22">Uncicauda pirata</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[528,682,814,838]" pageId="2" pageNumber="22" rank="species">gen. n., sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[442,554,848,874]" pageId="2" pageNumber="22" type="description">
<paragraph blockId="2.[442,554,848,874]" box="[442,554,848,874]" pageId="2" pageNumber="22">
(
<figureCitation box="[450,545,848,874]" captionStart="Figs" captionStartId="3.[184,234,1780,1804]" captionTargetBox="[299,1285,305,1716]" captionTargetId="figure-108@3.[279,1308,285,1728]" captionTargetPageId="3" captionText="Figs. 1‐5. Uncicauda pirata sp. n., adult. 1, pair of wings; 2, male abdominal tip, lateral; 3, same, ventral; 4, same, dorsal; 5, female abdominal tip, ventral. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4758657" httpUri="https://zenodo.org/record/4758657/files/figure.png" pageId="2" pageNumber="22">
Figs. 1
<emphasis bold="true" box="[524,533,849,874]" pageId="2" pageNumber="22">‐</emphasis>
5
</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="22" type="materials_examined">
<paragraph blockId="2.[184,776,917,1978]" pageId="2" pageNumber="22">
<emphasis bold="true" box="[184,413,917,941]" pageId="2" pageNumber="22">Material examined.</emphasis>
<materialsCitation ID-GBIF-Occurrence="3111445317" collectingDate="1993-12-15" collectingDateMax="1993-12-17" collectingDateMin="1993-12-15" collectionCode="CMF, OSF, USNM" country="Chile" county="Pino Hachado" elevation="1480" latitude="-38.65" location="Nahuelbuta" longLatPrecision="1173" longitude="-70.96667" municipality="Malleco" pageId="2" pageNumber="22" specimenCount="3" specimenCount-female="1" specimenCount-male="2" typeStatus="holotype">
<typeStatus box="[420,528,917,941]" pageId="2" pageNumber="22">Holotype</typeStatus>
<specimenCount box="[535,554,917,941]" pageId="2" pageNumber="22" type="male">♂</specimenCount>
,
<typeStatus box="[569,671,918,942]" pageId="2" pageNumber="22">paratype</typeStatus>
<specimenCount box="[678,697,917,941]" pageId="2" pageNumber="22" type="male">♂</specimenCount>
and
<specimenCount box="[756,775,917,941]" pageId="2" pageNumber="22" type="female">♀</specimenCount>
<collectingCountry box="[184,266,952,976]" name="Chile" pageId="2" pageNumber="22">CHILE</collectingCountry>
,
<collectingMunicipality box="[278,371,952,976]" country="Chile" county="Pino Hachado" latitude="-38.65" longLatPrecision="1173" longitude="-70.96667" municipality="Malleco" name="Malleco" pageId="2" pageNumber="22">Malleco</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:0399AB12FFBBFFCFE91150695246FBC2:8EEF4CDFFFBBFFC9E95450FC5165FC4B" box="[383,521,952,976]" country="Chile" county="Pino Hachado" latitude="-38.65" longLatPrecision="1173" longitude="-70.96667" municipality="Malleco" name="Nahuelbuta" pageId="2" pageNumber="22">Nahuelbuta</location>
,
<location LSID="urn:lsid:plazi:treatment:0399AB12FFBBFFCFE91150695246FBC2:8EEF4CDFFFBBFFC9EA3E50FD51C8FC4B" box="[533,676,952,976]" country="Chile" county="Pino Hachado" latitude="-38.65" longLatPrecision="1173" longitude="-70.96667" municipality="Malleco" name="Los Gringos" pageId="2" pageNumber="22">Los Gringos</location>
,
<geoCoordinate box="[688,770,952,977]" degrees="37" direction="south" minutes="48" orientation="latitude" pageId="2" pageNumber="22" precision="925" value="-37.8">
37°48
<emphasis bold="true" box="[750,755,952,977]" pageId="2" pageNumber="22">ʹ</emphasis>
S
</geoCoordinate>
;
<geoCoordinate box="[184,279,987,1012]" degrees="73" direction="west" minutes="01" orientation="longitude" pageId="2" pageNumber="22" precision="925" value="-73.01667">
73°01
<emphasis bold="true" box="[246,251,987,1012]" pageId="2" pageNumber="22">ʹ</emphasis>
W
</geoCoordinate>
,
<date box="[289,531,986,1012]" pageId="2" pageNumber="22" value="1993-12-15" valueMax="1993-12-17" valueMin="1993-12-15">
<collectingDate box="[289,531,986,1012]" pageId="2" pageNumber="22" value="1993-12-15" valueMax="1993-12-17" valueMin="1993-12-15">
15
<emphasis bold="true" box="[315,324,987,1012]" pageId="2" pageNumber="22">‐</emphasis>
17 December 1993
</collectingDate>
</date>
,
<collectionCode box="[541,598,987,1011]" pageId="2" pageNumber="22">CMF</collectionCode>
&amp;
<collectionCode box="[618,667,987,1011]" pageId="2" pageNumber="22">OSF</collectionCode>
(
<collectionCode box="[680,764,987,1011]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="2" pageNumber="22" type="Museum">USNM</collectionCode>
).
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3111445323" collectingDate="1988-02-04" collectingDateMax="1994-01-09" collectingDateMin="1988-02-04" collectionCode="CMF, OSF" country="Chile" county="Pino Hachado" elevation="1480" latitude="-38.65" location="Chile" longLatPrecision="1173" longitude="-70.96667" municipality="Malleco" pageId="2" pageNumber="22" specimenCount="4" specimenCount-female="1" specimenCount-male="3" typeStatus="paratype">
Additional
<typeStatus box="[314,430,1022,1046]" pageId="2" pageNumber="22">paratypes</typeStatus>
:
<specimenCount box="[439,471,1021,1045]" pageId="2" pageNumber="22" type="male">1♂</specimenCount>
,
<specimenCount box="[484,516,1021,1045]" pageId="2" pageNumber="22" type="female">1♀</specimenCount>
,
<collectingCountry box="[528,610,1021,1045]" name="Chile" pageId="2" pageNumber="22">CHILE</collectingCountry>
,
<collectingMunicipality box="[620,713,1021,1045]" pageId="2" pageNumber="22">Malleco</collectingMunicipality>
,
<collectingCounty pageId="2" pageNumber="22">Pino Hachado</collectingCounty>
,
<quantity box="[300,381,1056,1080]" metricMagnitude="3" metricUnit="m" metricValue="1.48" pageId="2" pageNumber="22" unit="m" value="1480.0">
<elevation box="[300,381,1056,1080]" metricMagnitude="3" metricUnit="m" metricValue="1.48" pageId="2" pageNumber="22" unit="m" value="1480.0">1480 m</elevation>
</quantity>
,
<geoCoordinate box="[391,474,1056,1081]" degrees="38" direction="south" minutes="39" orientation="latitude" pageId="2" pageNumber="22" precision="925" value="-38.65">
38°39
<emphasis bold="true" box="[453,458,1056,1081]" pageId="2" pageNumber="22">ʹ</emphasis>
S
</geoCoordinate>
;
<geoCoordinate box="[484,579,1056,1081]" degrees="70" direction="west" minutes="58" orientation="longitude" pageId="2" pageNumber="22" precision="925" value="-70.96667">
70°58
<emphasis bold="true" box="[546,551,1056,1081]" pageId="2" pageNumber="22">ʹ</emphasis>
W
</geoCoordinate>
,
<date box="[589,776,1056,1081]" pageId="2" pageNumber="22" value="1994-01-08" valueMax="1994-01-09" valueMin="1994-01-08">
<collectingDate box="[589,776,1056,1081]" pageId="2" pageNumber="22" value="1994-01-08" valueMax="1994-01-09" valueMin="1994-01-08">
8
<emphasis bold="true" box="[602,610,1056,1081]" pageId="2" pageNumber="22">‐</emphasis>
9 January 1994
</collectingDate>
</date>
<collectionCode box="[184,242,1090,1114]" pageId="2" pageNumber="22">CMF</collectionCode>
&amp;
<collectionCode box="[261,310,1090,1114]" pageId="2" pageNumber="22">OSF</collectionCode>
;
<specimenCount box="[319,351,1090,1114]" pageId="2" pageNumber="22" type="male">2♂</specimenCount>
,
<location LSID="urn:lsid:plazi:treatment:0399AB12FFBBFFCFE91150695246FBC2:8EEF4CDFFFBBFFC9E940570652C5FBC1" box="[363,425,1090,1114]" country="Chile" county="Pino Hachado" latitude="-38.65" longLatPrecision="1173" longitude="-70.96667" municipality="Malleco" name="Chile" pageId="2" pageNumber="22">Chile</location>
,
<collectorName box="[434,513,1090,1114]" pageId="2" pageNumber="22">Cautin</collectorName>
,
<collectorName box="[522,644,1090,1114]" pageId="2" pageNumber="22">Conquillio</collectorName>
,
<quantity box="[653,733,1090,1114]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="22" unit="m" value="1200.0">
<elevation box="[653,733,1090,1114]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="22" unit="m" value="1200.0">1200 m</elevation>
</quantity>
,
<date pageId="2" pageNumber="22" value="1988-02-04" valueMax="1988-02-08" valueMin="1988-02-04">
<collectingDate pageId="2" pageNumber="22" value="1988-02-04" valueMax="1988-02-08" valueMin="1988-02-04">
4
<emphasis bold="true" box="[754,762,1091,1116]" pageId="2" pageNumber="22">‐</emphasis>
8 February 1988
</collectingDate>
</date>
,
<collectorName box="[360,480,1125,1148]" pageId="2" pageNumber="22">L. E. Peña.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="22" type="description">
<paragraph blockId="2.[184,776,917,1978]" pageId="2" pageNumber="22">
<emphasis bold="true" box="[184,332,1159,1183]" pageId="2" pageNumber="22">Dimensions.</emphasis>
Male: body length 6.0
<emphasis bold="true" box="[576,584,1160,1185]" pageId="2" pageNumber="22">‐</emphasis>
7.0, forewing 7.5
<emphasis bold="true" box="[767,776,1160,1185]" pageId="2" pageNumber="22">‐</emphasis>
8.0; antenna 4.5. Female: body length 8.0; forewing 8.0
<emphasis bold="true" box="[216,224,1229,1254]" pageId="2" pageNumber="22">‐</emphasis>
9.0; antenna 5.0.
</paragraph>
<paragraph blockId="2.[184,776,917,1978]" pageId="2" pageNumber="22">
<emphasis bold="true" box="[184,259,1263,1287]" pageId="2" pageNumber="22">Adult.</emphasis>
Small slender light brown animals. Pronotum rectangular, with angles rounded. Head a little paler along sutures, muscle attachments on head and thorax a little darker, no prominent pattern. Ocelli present. Antenna brown, long, slender, individual segments distinctly expanded apically. Palps short, brown. Row of setae runs across apex of lacinia, in front of broad terminal teeth (see Fig. 68). Legs slender, essentially brown, base of femur a little paler, especially in front leg. Front and middle tibiae paler towards apex, hind tibiae with base to subgenual organ dark brown, remainder yellowish. Tibial spurs present.
</paragraph>
<paragraph blockId="2.[184,776,917,1978]" lastBlockId="2.[812,1404,260,1943]" pageId="2" pageNumber="22">
Wings (
<figureCitation box="[316,384,1713,1737]" captionStart="Figs" captionStartId="3.[184,234,1780,1804]" captionTargetBox="[299,1285,305,1716]" captionTargetId="figure-108@3.[279,1308,285,1728]" captionTargetPageId="3" captionText="Figs. 1‐5. Uncicauda pirata sp. n., adult. 1, pair of wings; 2, male abdominal tip, lateral; 3, same, ventral; 4, same, dorsal; 5, female abdominal tip, ventral. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4758657" httpUri="https://zenodo.org/record/4758657/files/figure.png" pageId="2" pageNumber="22">Fig. 1</figureCitation>
): Venation typical of the genus except for a very short fork of Rs without crossvein. The hind wing of specimens from Los Gringos (including the
<typeStatus box="[352,454,1816,1840]" pageId="2" pageNumber="22">holotype</typeStatus>
) has the posterior branch of M coalesced with Cu1 for a short distance. In other specimens the veins are separate but connected by a short crossvein. Veins pale brown and membrane with light brown tinge. Forewing with a darker tint around crossveins and in some specimens a purplish pigment from C to M but without speckles in cells.
</paragraph>
<paragraph blockId="2.[812,1404,260,1943]" pageId="2" pageNumber="22">
Abdomen sexually dimorphic. Cerci yellowish, short, downcurved with 8
<emphasis bold="true" box="[1106,1114,364,389]" pageId="2" pageNumber="22">‐</emphasis>
10 segments in males and
<quantity box="[812,889,398,423]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" metricValueMax="2.794" metricValueMin="2.286" pageId="2" pageNumber="22" unit="in" value="10.0" valueMax="11.0" valueMin="9.0">
9
<emphasis bold="true" box="[825,834,398,423]" pageId="2" pageNumber="22">‐</emphasis>
11 in
</quantity>
females. Pilosity inconspicuous.
</paragraph>
<paragraph blockId="2.[812,1404,260,1943]" pageId="2" pageNumber="22">
<emphasis bold="true" box="[812,992,432,457]" pageId="2" pageNumber="22">Male genitalia.</emphasis>
(
<figureCitation box="[1011,1113,433,458]" captionStart="Figs" captionStartId="3.[184,234,1780,1804]" captionTargetBox="[299,1285,305,1716]" captionTargetId="figure-108@3.[279,1308,285,1728]" captionTargetPageId="3" captionText="Figs. 1‐5. Uncicauda pirata sp. n., adult. 1, pair of wings; 2, male abdominal tip, lateral; 3, same, ventral; 4, same, dorsal; 5, female abdominal tip, ventral. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4758657" httpUri="https://zenodo.org/record/4758657/files/figure.png" pageId="2" pageNumber="22">
Figs. 2
<emphasis bold="true" box="[1088,1096,433,458]" pageId="2" pageNumber="22">‐</emphasis>
4
</figureCitation>
): Abdomen brownish or purplish, only slightly sclerotised. Each tergite with an anterior sclerite band that is medially divided on anterior tergites but entire on tergites 9
<emphasis bold="true" box="[1254,1263,537,562]" pageId="2" pageNumber="22">‐</emphasis>
10, although it is deeply incised from behind on segment 9. Sternites incompletely sclerotised, segments 2
<emphasis bold="true" box="[1326,1334,606,631]" pageId="2" pageNumber="22">‐</emphasis>
7 with widely separate narrow transverse anterior sclerites plus a continuous sclerite band across posterior third of segment; sternite 8 completely sclerotised. Subgenital plate a broadly rounded transversely oval sclerite almost truncate posteriorly. Tergite 10 well sclerotised, except for a small notch in anterior margin and a median elongate pale band in front of a sharply pointed downcurved posterior sclerite (
<bibRefCitation author="McLellan, I. D." box="[819,994,916,941]" pageId="2" pageNumber="22" pagination="1 - 79" refId="ref11883" refString="McLellan, I. D. 1971. A revision of Australian Gripopterygidae (Insecta: Plecoptera). Australian Journal of Zoology, Supplementary Series 2: 1 - 79." type="book chapter" year="1971">McLellan 1971</bibRefCitation>
). No epiproct. Paraprocts laterally dark brown with pale tips. The long slender sclerotised struts in side view are regularly curved and apically spatulate. Dorsoventrally the paraprocts have a narrow membranous triangular base that tapers until the sclerotised struts remain with their flat apices sometimes against each other. Penis structure not studied.
</paragraph>
<paragraph blockId="2.[812,1404,260,1943]" pageId="2" pageNumber="22">
<emphasis bold="true" box="[812,1023,1193,1217]" pageId="2" pageNumber="22">Female genitalia.</emphasis>
(
<figureCitation box="[1048,1123,1193,1217]" captionStart="Figs" captionStartId="3.[184,234,1780,1804]" captionTargetBox="[299,1285,305,1716]" captionTargetId="figure-108@3.[279,1308,285,1728]" captionTargetPageId="3" captionText="Figs. 1‐5. Uncicauda pirata sp. n., adult. 1, pair of wings; 2, male abdominal tip, lateral; 3, same, ventral; 4, same, dorsal; 5, female abdominal tip, ventral. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4758657" httpUri="https://zenodo.org/record/4758657/files/figure.png" pageId="2" pageNumber="22">Fig. 5</figureCitation>
): Abdomen pale, soft, dorsally unsclerotised except tergite 10, which is rounded apically. Sternites resemble those in males but anterior sclerites narrower and on sternite 7 posterior band shaped sclerite split into separate lateral sclerites that are slightly raised. Sternite 8 forming a sclerotised subgenital plate covering the posterior segments. Sides of subgenital plate parallel to slightly diverging posteriorly; distal margin with a broad, shallow excision. Subanal lobes triangular. The posterior sclerites on sternite 7, subgenital plate and ventrolateral face of the subanal lobes with a thick coat of stout curved colourless hairs which form a dense brush along the posterior margin of the subgenital plate.
</paragraph>
<paragraph blockId="2.[812,1404,260,1943]" box="[812,1013,1711,1735]" pageId="2" pageNumber="22">
<emphasis bold="true" box="[812,885,1711,1735]" pageId="2" pageNumber="22">Larva.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection lastPageId="4" lastPageNumber="24" pageId="2" pageNumber="22" type="discussion">
<paragraph blockId="2.[812,1404,260,1943]" pageId="2" pageNumber="22">
<emphasis bold="true" box="[812,924,1746,1770]" pageId="2" pageNumber="22">Remarks.</emphasis>
Sternite 9 of the female
<typeStatus box="[1265,1367,1747,1771]" pageId="2" pageNumber="22">paratype</typeStatus>
is completely telescoped under sternite 8. The generic name describes the hooked (Latin: uncus, hook; cauda, tail) caudal end of the male and the pirate
<emphasis bold="true" box="[1354,1362,1850,1875]" pageId="2" pageNumber="22">‐</emphasis>
like hook of tergite 10 posterior sclerite of the male suggested the specific name.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4758657" ID-Zenodo-Dep="4758657" httpUri="https://zenodo.org/record/4758657/files/figure.png" pageId="3" pageNumber="23" startId="3.[184,234,1780,1804]" targetBox="[299,1285,305,1716]" targetPageId="3">
<paragraph blockId="3.[184,1403,1779,1838]" pageId="3" pageNumber="23">
Figs. 1
<emphasis bold="true" box="[255,263,1780,1805]" pageId="3" pageNumber="23">‐</emphasis>
5.
<taxonomicName authority="McLellan &amp; Zwick, 2007" authorityName="McLellan &amp; Zwick" authorityYear="2007" box="[288,468,1780,1804]" class="Insecta" family="Gripopterygidae" genus="Uncicauda" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="23" phylum="Arthropoda" rank="species" species="pirata" status="sp. n.">
<emphasis box="[288,468,1780,1804]" italics="true" pageId="3" pageNumber="23">Uncicauda pirata</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[473,533,1780,1803]" pageId="3" pageNumber="23" rank="species">sp. n.</taxonomicNameLabel>
, adult. 1, pair of wings; 2, male abdominal tip, lateral; 3, same, ventral; 4, same, dorsal; 5, female abdominal tip, ventral. Not to scale.
</paragraph>
</caption>
<paragraph blockId="3.[184,775,1918,1977]" lastBlockId="4.[184,776,260,1113]" lastPageId="4" lastPageNumber="24" pageId="3" pageNumber="23">
<emphasis bold="true" box="[184,305,1918,1942]" pageId="3" pageNumber="23">Affinities.</emphasis>
The new taxon is a member of
<taxonomicName authorityName="Enderlein" authorityYear="1909" box="[184,374,1952,1976]" class="Insecta" family="Gripopterygidae" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="23" phylum="Arthropoda" rank="subFamily" subFamily="Gripopteryginae">Gripopteryginae</taxonomicName>
in the sense of
<bibRefCitation author="McLellan, I. D." box="[542,724,1952,1977]" pageId="3" pageNumber="23" pagination="119 - 147" refId="ref11915" refString="McLellan, I. D. 1977. New alpine and southern Plecoptera from New Zealand, and a new classification of the Gripopterygidae. New Zealand Journal of Zoology 4 (2): 119 - 147." type="journal article" year="1977">McLellan (1977)</bibRefCitation>
. The insect has no striking autapomorphies but at the same time fits into none of the existing genera. Analysis of its affinities is hampered by lack of its larva. Therefore, generic rank as well as placement are somewhat provisional.
</paragraph>
<paragraph blockId="4.[184,776,260,1113]" pageId="4" pageNumber="24">
The known apomorphic characters of
<taxonomicName authorityName="McLellan &amp; Zwick" authorityYear="2007" box="[662,775,363,387]" class="Insecta" family="Gripopterygidae" genus="Uncicauda" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="24" phylum="Arthropoda" rank="genus">
<emphasis box="[662,775,363,387]" italics="true" pageId="4" pageNumber="24">Uncicauda</emphasis>
</taxonomicName>
suggest relations with some of the smaller genera which
<bibRefCitation author="Illies, J." box="[260,388,432,456]" pageId="4" pageNumber="24" pagination="145 - 248" refId="ref11858" refString="Illies, J. 1963: Revision der sudamerikanischen Gripopterygidae (Plecoptera). Mitteilungen der Schweizerischen Entomologischen Gesellschaft 36: 145 - 248." type="journal article" year="1963">Illies (1963)</bibRefCitation>
classified as
<taxonomicName authorityName="Illies" authorityYear="1963" box="[532,772,432,456]" class="Insecta" family="Gripopterygidae" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="24" phylum="Arthropoda" rank="subFamily" subFamily="Paragripopteryginae">Paragripopteryginae</taxonomicName>
. However, all of the apomorphies in question represent reductions that may have occurred repeatedly and independently, especially shortening of the fork of Rs and of the cerci. Even loss of the male epiproct, although very unusual and known only among South American
<taxonomicName authorityName="Enderlein" authorityYear="1909" box="[530,723,640,664]" class="Insecta" family="Gripopterygidae" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="24" phylum="Arthropoda" rank="family">Gripopterygidae</taxonomicName>
, has occurred independently. Within
<taxonomicName box="[562,690,675,698]" class="Insecta" family="Gripopterygidae" genus="Gripopteryx" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="24" phylum="Arthropoda" rank="genus">
<emphasis box="[562,690,675,698]" italics="true" pageId="4" pageNumber="24">Gripopteryx</emphasis>
</taxonomicName>
which, by its bizarre larvae, is undoubtedly monophyletic several species have very small epiprocts, and
<taxonomicName authority="Froehlich" authorityName="Froehlich" authorityYear="1993" class="Insecta" family="Gripopterygidae" genus="Gripopteryx" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="24" phylum="Arthropoda" rank="species" species="serrensis">
<emphasis italics="true" pageId="4" pageNumber="24">G. serrensis</emphasis>
Froehlich
</taxonomicName>
lacks it completely (
<bibRefCitation author="Froehlich, C. G." pageId="4" pageNumber="24" pagination="21 - 38" refId="ref11527" refString="Froehlich, C. G. 1993. Brazilian Plecoptera: 7. Old and new species of Gripopteryx (Gripopterygidae). - Aquatic Insects 15 (1): 21 - 38." type="journal article" year="1993">Froehlich 1993</bibRefCitation>
). The remaining genera without male epiproct (
<taxonomicName authorityName="Enderlein" authorityYear="1909" box="[192,361,848,871]" class="Insecta" family="Gripopterygidae" genus="Paragripopteryx" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="24" phylum="Arthropoda" rank="genus">
<emphasis box="[192,361,848,871]" italics="true" pageId="4" pageNumber="24">Paragripopteryx</emphasis>
</taxonomicName>
,
<taxonomicName box="[372,472,847,871]" class="Insecta" family="Gripopterygidae" genus="Tupiperla" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="24" phylum="Arthropoda" rank="genus">
<emphasis box="[372,472,847,871]" italics="true" pageId="4" pageNumber="24">Tupiperla</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Froehlich" authorityYear="2001" box="[483,625,847,871]" class="Insecta" family="Gripopterygidae" genus="Guaranyperla" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="24" phylum="Arthropoda" rank="genus">
<emphasis box="[483,625,847,871]" italics="true" pageId="4" pageNumber="24">Guaranyperla</emphasis>
</taxonomicName>
(all NE South American) and
<taxonomicName authority="(Andean))" baseAuthorityName="Andean" box="[359,612,881,905]" class="Insecta" family="Gripopterygidae" genus="Potamoperla" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="24" phylum="Arthropoda" rank="genus">
<emphasis box="[359,489,881,905]" italics="true" pageId="4" pageNumber="24">Potamoperla</emphasis>
(Andean))
</taxonomicName>
have a double point of male tergite 10 which is unique in the subfamily and suggests these genera represent a monophylum. We believe the new genus
<taxonomicName authorityName="McLellan &amp; Zwick" authorityYear="2007" box="[662,775,985,1009]" class="Insecta" family="Gripopterygidae" genus="Uncicauda" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="24" phylum="Arthropoda" rank="genus">
<emphasis box="[662,775,985,1009]" italics="true" pageId="4" pageNumber="24">Uncicauda</emphasis>
</taxonomicName>
with a sharp single point of tergite 10 may be an early member or the sister taxon of this small group of genera.
</paragraph>
</subSubSection>
</treatment>
</document>
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