184 lines
30 KiB
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184 lines
30 KiB
XML
<document id="6B65ABD43D80E089E8E2C55802773448" ID-DOI="10.14446/FI.2016.53" ID-ISSN="2533-4069" ID-Zenodo-Dep="4586149" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1614968823112" checkinUser="felipe" docAuthor="Rage, Jean-Claude" docDate="2016" docId="039987DFFFEBC7517FD4C198FC57FB6F" docLanguage="en" docName="FossilImprint.72.1-2.53-66.pdf" docOrigin="Fossil Imprint 72 (1 - 2)" docStyle="DocumentStyle:0EC54BF35E227CA4627F894F6BA30634.4:FossilImprint.2016-.journal_article" docStyleId="0EC54BF35E227CA4627F894F6BA30634" docStyleName="FossilImprint.2016-.journal_article" docStyleVersion="4" docTitle="Pelodytes BONAPARTE 1838" docType="treatment" docVersion="4" lastPageNumber="57" masterDocId="FFA0FFA7FFE8C7557C1EC22FFF80FFE1" masterDocTitle="Frogs (Amphibia, Anura) From The Eocene And Oligocene Of The Phosphorites Du Quercy (France). An Overview" masterLastPageNumber="66" masterPageNumber="53" pageNumber="56" updateTime="1698881274651" updateUser="plazi" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="28C2E4B61E917FA6503EC09F1A39EEFA">Frogs (Amphibia, Anura) From The Eocene And Oligocene Of The Phosphorites Du Quercy (France). An Overview</mods:title>
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<mods:namePart id="B6CB035DA8877D33C33C24E2EF18949C">Rage, Jean-Claude</mods:namePart>
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<mods:title id="132877E8BEEEC5B3DA3B3FE8CCFEF36C">Fossil Imprint</mods:title>
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<mods:part id="8318DC536FE7DBACB7893EF3C4BD81D6">
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<mods:date id="8FF35E47CDA4D333999C4D81BC400BD6">2016</mods:date>
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<mods:number id="7B2D85740C3BEA89E02A1D43F2FF8734">2016-08-15</mods:number>
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<treatment id="039987DFFFEBC7517FD4C198FC57FB6F" LSID="urn:lsid:plazi:treatment:039987DFFFEBC7517FD4C198FC57FB6F" httpUri="http://treatment.plazi.org/id/039987DFFFEBC7517FD4C198FC57FB6F" lastPageId="4" lastPageNumber="57" pageId="3" pageNumber="56">
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<subSubSection id="C32A6542FFEBC7567FD4C198FA9CFC2F" box="[970,1308,951,974]" pageId="3" pageNumber="56" type="nomenclature">
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<paragraph id="8B8F36C9FFEBC7567FD4C198FA9CFC2F" blockId="3.[970,1308,951,974]" box="[970,1308,951,974]" pageId="3" pageNumber="56">
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<heading id="D0C781A5FFEBC7567FD4C198FA9CFC2F" box="[970,1308,951,974]" centered="true" fontSize="8" level="2" pageId="3" pageNumber="56" reason="2">
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<emphasis id="B944EADBFFEBC7567FD4C198FA9CFC2F" bold="true" box="[970,1308,951,974]" pageId="3" pageNumber="56">
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cf.
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<taxonomicName id="4C304D4AFFEBC7567FF3C198FA9CFC2F" ID-CoL="6KCN" authority="BONAPARTE, 1838" authorityName="BONAPARTE" authorityYear="1838" box="[1005,1308,951,974]" class="Amphibia" family="Pelodytidae" genus="Pelodytes" kingdom="Animalia" order="Anura" pageId="3" pageNumber="56" phylum="Chordata" rank="genus">
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<emphasis id="B944EADBFFEBC7567FF3C198FBD5FC2F" bold="true" box="[1005,1109,951,974]" italics="true" pageId="3" pageNumber="56">Pelodytes</emphasis>
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<bibRefCitation id="EFA14B38FFEBC7567845C198FB5AFC2F" author="Bonaparte, C. L." box="[1115,1242,951,974]" pageId="3" pageNumber="56" refId="ref10439" refString="Bonaparte, C. L. (1838): Iconographia della Fauna Italica per le Quattro Classi degli Animali Vertebrati. II. Amfibi. Fascicolo 23 [Pictorial Book of the Italian Fauna for the Four Classes of Vertebrate Animals. II. Amphibians. Vol. 23]. - Salviucci, Roma, (not paginated). (in Italian)" type="book" year="1838">BONAPARTE</bibRefCitation>
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, 1838
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</taxonomicName>
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||
</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32A6542FFEBC7517828C1C8FC57FB6F" lastPageId="4" lastPageNumber="57" pageId="3" pageNumber="56" type="description">
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<paragraph id="8B8F36C9FFEBC7567828C1C8FB31FC1C" blockId="3.[1078,1201,999,1021]" box="[1078,1201,999,1021]" pageId="3" pageNumber="56">
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(
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<figureCitation id="130B2A4CFFEBC7567823C1C8FB29FC1C" box="[1085,1193,999,1021]" captionStart="Text-fig" captionStartId="4.[129,208,893,914]" captionTargetBox="[129,1482,152,852]" captionTargetId="figure-512@4.[138,1472,151,850]" captionTargetPageId="4" captionText="Text-fig. 3. Pelodytidae (cf. Pelodytes) from the Phosphorites du Quercy. 1: presacral vertebra UM-MAL 617, from the late Eocene of Malpérié (MP 17), in dorsal (a), anterior (b), and ventral (c) views. 2: posterior presacral vertebra UM-SNB 2406, from the late Eocene of Sainte-Néboule (MP 18), in anterior (a) and ventral (b) views. 3: right humerus UM-ECA 2556, from the late Eocene of Escamps (MP 19), in ventral view. 4: left ilium UM-ECA 2534, from the late Eocene of Escamps (MP 19), in lateral view. 5: right tibiale-fibulare UM-MAL 616, from the late Eocene (MP 17) of Malpérié (proximal section (a); dorsal view (b); and distal section (c)). Each scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.4586157" httpUri="https://zenodo.org/record/4586157/files/figure.png" pageId="3" pageNumber="56">Text-fig. 3</figureCitation>
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)
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</paragraph>
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<paragraph id="8B8F36C9FFEBC7567F44C63DFB0BFA1C" blockId="3.[820,1460,1042,2099]" pageId="3" pageNumber="56">
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In the Phosphorites, the taxon is represented by vertebrae, ilia, humeri and one fused tibiale-fibulare (
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<figureCitation id="130B2A4CFFEBC7567937C61EFA25FBA9" box="[1321,1445,1073,1096]" captionStart="Text-fig" captionStartId="4.[129,208,893,914]" captionTargetBox="[129,1482,152,852]" captionTargetId="figure-512@4.[138,1472,151,850]" captionTargetPageId="4" captionText="Text-fig. 3. Pelodytidae (cf. Pelodytes) from the Phosphorites du Quercy. 1: presacral vertebra UM-MAL 617, from the late Eocene of Malpérié (MP 17), in dorsal (a), anterior (b), and ventral (c) views. 2: posterior presacral vertebra UM-SNB 2406, from the late Eocene of Sainte-Néboule (MP 18), in anterior (a) and ventral (b) views. 3: right humerus UM-ECA 2556, from the late Eocene of Escamps (MP 19), in ventral view. 4: left ilium UM-ECA 2534, from the late Eocene of Escamps (MP 19), in lateral view. 5: right tibiale-fibulare UM-MAL 616, from the late Eocene (MP 17) of Malpérié (proximal section (a); dorsal view (b); and distal section (c)). Each scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.4586157" httpUri="https://zenodo.org/record/4586157/files/figure.png" pageId="3" pageNumber="56">Text-fig. 3</figureCitation>
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). Vertebrae show a combination of features that is characteristic of
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<taxonomicName id="4C304D4AFFEBC7567F8FC65FFC79FB66" authorityName="BONAPARTE" authorityYear="1838" box="[913,1017,1136,1159]" class="Amphibia" family="Pelodytidae" genus="Pelodytes" kingdom="Animalia" order="Anura" pageId="3" pageNumber="56" phylum="Chordata" rank="genus">
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<emphasis id="B944EADBFFEBC7567F8FC65FFC79FB66" box="[913,1017,1136,1159]" italics="true" pageId="3" pageNumber="56">Pelodytes</emphasis>
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</taxonomicName>
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(
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<figureCitation id="130B2A4CFFEBC7567817C65FFB00FB66" box="[1033,1152,1136,1159]" captionStart="Text-fig" captionStartId="4.[129,208,893,914]" captionTargetBox="[129,1482,152,852]" captionTargetId="figure-512@4.[138,1472,151,850]" captionTargetPageId="4" captionText="Text-fig. 3. Pelodytidae (cf. Pelodytes) from the Phosphorites du Quercy. 1: presacral vertebra UM-MAL 617, from the late Eocene of Malpérié (MP 17), in dorsal (a), anterior (b), and ventral (c) views. 2: posterior presacral vertebra UM-SNB 2406, from the late Eocene of Sainte-Néboule (MP 18), in anterior (a) and ventral (b) views. 3: right humerus UM-ECA 2556, from the late Eocene of Escamps (MP 19), in ventral view. 4: left ilium UM-ECA 2534, from the late Eocene of Escamps (MP 19), in lateral view. 5: right tibiale-fibulare UM-MAL 616, from the late Eocene (MP 17) of Malpérié (proximal section (a); dorsal view (b); and distal section (c)). Each scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.4586157" httpUri="https://zenodo.org/record/4586157/files/figure.png" pageId="3" pageNumber="56">Text-fig. 3</figureCitation>
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: 1, 2). They are proceolous, lightly built and the centrum, cotyle and condyle all are markedly depressed dorsoventrally. In ventral aspect, the ventral parts of the lateral walls typically extend laterally on either side of the centrum. The neural arch is of the imbricate
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<typeStatus id="548B886BFFEBC7567F2AC723FCE4FAC2" box="[820,868,1292,1315]" pageId="3" pageNumber="56">type</typeStatus>
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, i.e. it is anteroposteriorly long. The posterior projection of the neural spine is relatively weak. In posterior presacral vertebrae, the bases of the transverse processes originate approximately below the prezygapophyses and they project markedly anterolaterally. One incomplete sacral vertebra may belong to this taxon; however, it is regarded as belonging to an indeterminate anuran because it bears postzygapophyses (see below: ‘Problematic taxa’).
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</paragraph>
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<paragraph id="8B8F36C9FFEBC7567F44C42AFAF9F978" blockId="3.[820,1460,1042,2099]" pageId="3" pageNumber="56">
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The ilia lack both a dorsal crest and a tuber superius, as do those of pelobatids (
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<figureCitation id="130B2A4CFFEBC756782FC40BFB26F9DA" box="[1073,1190,1572,1595]" captionStart="Text-fig" captionStartId="4.[129,208,893,914]" captionTargetBox="[129,1482,152,852]" captionTargetId="figure-512@4.[138,1472,151,850]" captionTargetPageId="4" captionText="Text-fig. 3. Pelodytidae (cf. Pelodytes) from the Phosphorites du Quercy. 1: presacral vertebra UM-MAL 617, from the late Eocene of Malpérié (MP 17), in dorsal (a), anterior (b), and ventral (c) views. 2: posterior presacral vertebra UM-SNB 2406, from the late Eocene of Sainte-Néboule (MP 18), in anterior (a) and ventral (b) views. 3: right humerus UM-ECA 2556, from the late Eocene of Escamps (MP 19), in ventral view. 4: left ilium UM-ECA 2534, from the late Eocene of Escamps (MP 19), in lateral view. 5: right tibiale-fibulare UM-MAL 616, from the late Eocene (MP 17) of Malpérié (proximal section (a); dorsal view (b); and distal section (c)). Each scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.4586157" httpUri="https://zenodo.org/record/4586157/files/figure.png" pageId="3" pageNumber="56">Text-fig. 3</figureCitation>
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: 4). However, their shaft is markedly curved in lateral view and the dorsal border of their pars ascendens (ischiatic process) is slightly curved dorsally, which distinguishes them from pelobatid ilia.
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</paragraph>
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<paragraph id="8B8F36C9FFEBC7567F44C48EFB6DF872" blockId="3.[820,1460,1042,2099]" pageId="3" pageNumber="56">
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Humeri are very similar to those of
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<taxonomicName id="4C304D4AFFEBC75678C0C48EFAC6F959" authorityName="BONAPARTE" authorityYear="1838" box="[1246,1350,1697,1720]" class="Amphibia" family="Pelodytidae" genus="Pelodytes" kingdom="Animalia" order="Anura" pageId="3" pageNumber="56" phylum="Chordata" rank="genus">
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<emphasis id="B944EADBFFEBC75678C0C48EFAC6F959" box="[1246,1350,1697,1720]" italics="true" pageId="3" pageNumber="56">Pelodytes</emphasis>
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</taxonomicName>
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(
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<figureCitation id="130B2A4CFFEBC756794BC48EFCC4F936" captionStart="Text-fig" captionStartId="4.[129,208,893,914]" captionTargetBox="[129,1482,152,852]" captionTargetId="figure-512@4.[138,1472,151,850]" captionTargetPageId="4" captionText="Text-fig. 3. Pelodytidae (cf. Pelodytes) from the Phosphorites du Quercy. 1: presacral vertebra UM-MAL 617, from the late Eocene of Malpérié (MP 17), in dorsal (a), anterior (b), and ventral (c) views. 2: posterior presacral vertebra UM-SNB 2406, from the late Eocene of Sainte-Néboule (MP 18), in anterior (a) and ventral (b) views. 3: right humerus UM-ECA 2556, from the late Eocene of Escamps (MP 19), in ventral view. 4: left ilium UM-ECA 2534, from the late Eocene of Escamps (MP 19), in lateral view. 5: right tibiale-fibulare UM-MAL 616, from the late Eocene (MP 17) of Malpérié (proximal section (a); dorsal view (b); and distal section (c)). Each scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.4586157" httpUri="https://zenodo.org/record/4586157/files/figure.png" pageId="3" pageNumber="56">Text-fig. 3</figureCitation>
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: 3). They have a slender and almost straight diaphysis that bears ventral and paraventral crests. The articular ball is well defined, comparatively small and scarcely shifted laterally. The distal part of the lateral border of the bone (crista lateralis not taken into account), close to the articular ball, forms a gentle curve that is slightly convex laterally; this part of the humerus is generally concave in anurans.
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</paragraph>
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<paragraph id="8B8F36C9FFEBC7517F44C5B3FE82F8D2" blockId="3.[820,1460,1042,2099]" lastBlockId="4.[128,768,1111,2099]" lastPageId="4" lastPageNumber="57" pageId="3" pageNumber="56">
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A fused tibiale-fibulare (
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<figureCitation id="130B2A4CFFEBC756786AC5B3FB6FF852" box="[1140,1263,1948,1971]" captionStart="Text-fig" captionStartId="4.[129,208,893,914]" captionTargetBox="[129,1482,152,852]" captionTargetId="figure-512@4.[138,1472,151,850]" captionTargetPageId="4" captionText="Text-fig. 3. Pelodytidae (cf. Pelodytes) from the Phosphorites du Quercy. 1: presacral vertebra UM-MAL 617, from the late Eocene of Malpérié (MP 17), in dorsal (a), anterior (b), and ventral (c) views. 2: posterior presacral vertebra UM-SNB 2406, from the late Eocene of Sainte-Néboule (MP 18), in anterior (a) and ventral (b) views. 3: right humerus UM-ECA 2556, from the late Eocene of Escamps (MP 19), in ventral view. 4: left ilium UM-ECA 2534, from the late Eocene of Escamps (MP 19), in lateral view. 5: right tibiale-fibulare UM-MAL 616, from the late Eocene (MP 17) of Malpérié (proximal section (a); dorsal view (b); and distal section (c)). Each scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.4586157" httpUri="https://zenodo.org/record/4586157/files/figure.png" pageId="3" pageNumber="56">Text-fig. 3</figureCitation>
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: 5) belonging to a pelodytid was recovered among the specimens of anurans from the late Eocene of Malpérié (MP 17) by
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<bibRefCitation id="EFA14B38FFEBC7567912C5F3FA30F812" author="Duffaud, S." box="[1292,1456,2012,2035]" pageId="3" pageNumber="56" refId="ref11043" refString="Duffaud, S. (2000): Les faunes d'amphibiens du Cretace superieur a l'Oligocene inferieur en Europe: paleobiodiversite, evolution, mise en place; PhD Thesis. - MS, Museum national d'Histoire naturelle, Paris, France, 221 pp. (copy in author's private library)." type="book" year="2000">Duffaud (2000)</bibRefCitation>
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. In almost all anurans, the tibiale and fibulare are elongate and fused only at their extremities. However, in pelodytids, centrolenids (an extant Central and South American family) and
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<taxonomicName id="4C304D4AFFECC7517C9FC658FE1EFB6F" authority="HENRICI, 1994" authorityName="HENRICI" authorityYear="1994" box="[129,414,1143,1166]" class="Amphibia" family="Pelodytidae" genus="Tephrodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="4" pageNumber="57" phylum="Chordata" rank="genus">
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<emphasis id="B944EADBFFECC7517C9FC658FF7FFB6F" box="[129,255,1143,1166]" italics="true" pageId="4" pageNumber="57">Tephrodytes</emphasis>
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<bibRefCitation id="EFA14B38FFECC7517D18C658FE1EFB6F" author="Henrici, A. C." box="[262,414,1143,1166]" pageId="4" pageNumber="57" pagination="155 - 183" refId="ref12286" refString="Henrici, A. C. (1994): Tephrodytes brassicarvalis, new genus and species (Anura: Pelodytidae), from the Arikareean cabbage patch beds of Montana, USA, and Pelodytid- Pelobatid relationships. - Annals of Carnegie Museum, 63: 155 - 183." type="journal article" year="1994">HENRICI, 1994</bibRefCitation>
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</taxonomicName>
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(but not in rhinophrynids, contra
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<bibRefCitation id="EFA14B38FFECC7517C9FC6B8FF7CFB4F" author="Trueb, L." box="[129,252,1175,1198]" pageId="4" pageNumber="57" pagination="65 - 132" refId="ref15413" refString="Trueb, L. (1973): Bones, frogs, and evolution. - In: Vial, J. L. (ed.), Evolutionary biology of the anurans, University of Missouri Press, Columbia, pp. 65 - 132." type="book chapter" year="1973">Trueb 1973</bibRefCitation>
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), they are fused throughout their length and form a single element. This element thus resembles the anuran tibiofibula; however, it is shorter and its extremities are more expanded. In addition the proximal ends of the tibiale and fibulare are approximately circular, whereas the distal extremities are clearly flattened. Traces of the fusion remain as grooves, except in the central portion of the bone. This tibiale-fibulare from Malpérié clearly displays this morphology, which is characteristic of
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<taxonomicName id="4C304D4AFFECC7517D93C7B6FD1BFA51" authority="(Sanchiz 1978)" baseAuthorityName="Sanchiz" baseAuthorityYear="1978" box="[397,667,1433,1456]" class="Amphibia" family="Pelodytidae" genus="Pelodytes" kingdom="Animalia" order="Anura" pageId="4" pageNumber="57" phylum="Chordata" rank="genus">
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<emphasis id="B944EADBFFECC7517D93C7B6FE71FA51" box="[397,497,1433,1456]" italics="true" pageId="4" pageNumber="57">Pelodytes</emphasis>
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(
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<bibRefCitation id="EFA14B38FFECC7517E1EC7B6FD13FA51" author="Sanchiz, B." box="[512,659,1433,1456]" pageId="4" pageNumber="57" pagination="9 - 27" refId="ref14865" refString="Sanchiz, B. (1978): Nuevos restos fosiles de la familia Pelodytidae (Amphibia, Anura) [New fossil remains of the family Pelodytidae (Amphibia, Anura)]. - Estudios Geologicos, 34: 9 - 27. (in Spanish)" type="journal article" year="1978">Sanchiz 1978</bibRefCitation>
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)
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</taxonomicName>
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. It differs from that of
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<taxonomicName id="4C304D4AFFECC7517CE3C796FD99FA31" authority="TAYLOR, 1941" authorityName="TAYLOR" authorityYear="1941" box="[253,537,1465,1488]" class="Amphibia" family="Pelodytidae" genus="Miopelodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="4" pageNumber="57" phylum="Chordata" rank="genus">
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<emphasis id="B944EADBFFECC7517CE3C796FE07FA31" box="[253,391,1465,1488]" italics="true" pageId="4" pageNumber="57">Miopelodytes</emphasis>
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<bibRefCitation id="EFA14B38FFECC7517D92C796FD99FA31" author="Taylor, E. H." box="[396,537,1465,1488]" pageId="4" pageNumber="57" pagination="61 - 69" refId="ref15341" refString="Taylor, E. H. (1941): A new anuran from the middle Miocene of Nevada. - University of Kansas Science Bulletin, 27: 61 - 69." type="journal article" year="1941">
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TAYLOR,
|
||
<quantity id="4CC89B2CFFECC7517DFBC796FD99FA31" box="[485,537,1465,1488]" metricMagnitude="1" metricUnit="m" metricValue="4.93014" pageId="4" pageNumber="57" unit="in" value="1941.0">1941</quantity>
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
in being more slender; more specifically, its extremities are less expanded. The fused tibiale-fibulare of centrolenids is markedly more elongate and slender (
|
||
<bibRefCitation id="EFA14B38FFECC7517CC2C435FE5EF9D0" author="Guayasamin, J. M. & Castroviejo-Fischer, S. & Trueb, L. & Ayarzaguena, J. & Rada, M. & Vila, C." box="[220,478,1562,1585]" pageId="4" pageNumber="57" pagination="1 - 97" refId="ref12161" refString="Guayasamin, J. M., Castroviejo-Fischer, S., Trueb, L., Ayarzaguena, J., Rada, M., Vila, C. (2009): Phylogenetic systematics of glassfrogs (Amphibia: Centrolenidae) and their sister taxon Allophryne ruthveni. - Zootaxa, 2100: 1 - 97." type="journal article" year="2009">Guayasamin et al. 2009</bibRefCitation>
|
||
). The fused tibiale-fibulare from Malpérié cannot be distinguished from the tibiale-fibulare of extant
|
||
<taxonomicName id="4C304D4AFFECC7517CF0C474FEDBF993" baseAuthorityName="Sanchiz" baseAuthorityYear="1978" box="[238,347,1627,1650]" class="Amphibia" family="Pelodytidae" genus="Pelodytes" kingdom="Animalia" order="Anura" pageId="4" pageNumber="57" phylum="Chordata" rank="genus">
|
||
<emphasis id="B944EADBFFECC7517CF0C474FEDBF993" box="[238,347,1627,1650]" italics="true" pageId="4" pageNumber="57">Pelodytes</emphasis>
|
||
</taxonomicName>
|
||
and it also closely resembles that of
|
||
<taxonomicName id="4C304D4AFFECC7517C9FC454FE88F973" authorityName="HENRICI" authorityYear="1994" box="[129,264,1659,1682]" class="Amphibia" family="Pelodytidae" genus="Tephrodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="4" pageNumber="57" phylum="Chordata" rank="genus">
|
||
<emphasis id="B944EADBFFECC7517C9FC454FE88F973" box="[129,264,1659,1682]" italics="true" pageId="4" pageNumber="57">Tephrodytes</emphasis>
|
||
</taxonomicName>
|
||
(Oligocene-Miocene transition of the
|
||
<collectingCountry id="F3277659FFECC7517EDFC454FD7BF973" box="[705,763,1659,1682]" name="United States of America" pageId="4" pageNumber="57">USA</collectingCountry>
|
||
;
|
||
<bibRefCitation id="EFA14B38FFECC7517C9FC4B4FE89F953" author="Henrici, A. C." box="[129,265,1691,1714]" pageId="4" pageNumber="57" pagination="155 - 183" refId="ref12286" refString="Henrici, A. C. (1994): Tephrodytes brassicarvalis, new genus and species (Anura: Pelodytidae), from the Arikareean cabbage patch beds of Montana, USA, and Pelodytid- Pelobatid relationships. - Annals of Carnegie Museum, 63: 155 - 183." type="journal article" year="1994">Henrici 1994</bibRefCitation>
|
||
).
|
||
<taxonomicName id="4C304D4AFFECC7517D07C4B4FE14F953" authorityName="HENRICI" authorityYear="1994" box="[281,404,1691,1714]" class="Amphibia" family="Pelodytidae" genus="Tephrodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="4" pageNumber="57" phylum="Chordata" rank="genus">
|
||
<emphasis id="B944EADBFFECC7517D07C4B4FE14F953" box="[281,404,1691,1714]" italics="true" pageId="4" pageNumber="57">Tephrodytes</emphasis>
|
||
</taxonomicName>
|
||
was first referred to the
|
||
<taxonomicName id="4C304D4AFFECC7517E99C4B4FEA1F933" authority="(Henrici 1994)" baseAuthorityName="Henrici" baseAuthorityYear="1994" class="Amphibia" family="Pelodytidae" kingdom="Animalia" order="Anura" pageId="4" pageNumber="57" phylum="Chordata" rank="family">
|
||
Pelodytidae (
|
||
<bibRefCitation id="EFA14B38FFECC7517C96C494FE9AF933" author="Henrici, A. C." box="[136,282,1723,1746]" pageId="4" pageNumber="57" pagination="155 - 183" refId="ref12286" refString="Henrici, A. C. (1994): Tephrodytes brassicarvalis, new genus and species (Anura: Pelodytidae), from the Arikareean cabbage patch beds of Montana, USA, and Pelodytid- Pelobatid relationships. - Annals of Carnegie Museum, 63: 155 - 183." type="journal article" year="1994">Henrici 1994</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
but
|
||
<bibRefCitation id="EFA14B38FFECC7517D45C494FDC0F933" author="Henrici, A. C. & Baez, A. M. & Grande, L." box="[347,576,1723,1746]" pageId="4" pageNumber="57" pagination="295 - 309" refId="ref12337" refString="Henrici, A. C., Baez, A. M., Grande, L. (2013): Aerugoamnis paulus, new genus and new species (Anura: Anomocoela): first reported anuran from the early Eocene (Wasatchian) Fossil Butte Member of the Green River Formation, Wyoming. - Annals of Carnegie Museum, 81: 295 - 309. http: // dx. doi. org / 10.2992 / 007.081.0402" type="journal article" year="2013">Henrici et al. (2013)</bibRefCitation>
|
||
placed it among pelobatids s.l. However, the morphology of various bones suggests that
|
||
<taxonomicName id="4C304D4AFFECC7517D10C4D3FE0BF8F2" authorityName="HENRICI" authorityYear="1994" box="[270,395,1788,1811]" class="Amphibia" family="Pelodytidae" genus="Tephrodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="4" pageNumber="57" phylum="Chordata" rank="genus">
|
||
<emphasis id="B944EADBFFECC7517D10C4D3FE0BF8F2" box="[270,395,1788,1811]" italics="true" pageId="4" pageNumber="57">Tephrodytes</emphasis>
|
||
</taxonomicName>
|
||
may be more closely related to the
|
||
<taxonomicName id="4C304D4AFFECC7517C9EC533FF7DF8D2" baseAuthorityName="Henrici" baseAuthorityYear="1994" box="[128,253,1820,1843]" class="Amphibia" family="Pelodytidae" kingdom="Animalia" order="Anura" pageId="4" pageNumber="57" phylum="Chordata" rank="family">Pelodytidae</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF4F6641FFECC7517C9FC152FA1AFBE4" ID-DOI="http://doi.org/10.5281/zenodo.4586157" ID-Zenodo-Dep="4586157" httpUri="https://zenodo.org/record/4586157/files/figure.png" pageId="4" pageNumber="57" startId="4.[129,208,893,914]" targetBox="[129,1482,152,852]" targetPageId="4">
|
||
<paragraph id="8B8F36C9FFECC7517C9FC152FA1AFBE4" blockId="4.[128,1483,893,1029]" pageId="4" pageNumber="57">
|
||
Text-fig. 3.
|
||
<taxonomicName id="4C304D4AFFECC7517CEAC152FEEAFC73" baseAuthorityName="Henrici" baseAuthorityYear="1994" box="[244,362,893,914]" class="Amphibia" family="Pelodytidae" kingdom="Animalia" order="Anura" pageId="4" pageNumber="57" phylum="Chordata" rank="family">Pelodytidae</taxonomicName>
|
||
(cf.
|
||
<taxonomicName id="4C304D4AFFECC7517D89C152FE72FC73" baseAuthorityName="Sanchiz" baseAuthorityYear="1978" box="[407,498,893,914]" class="Amphibia" family="Pelodytidae" genus="Pelodytes" kingdom="Animalia" order="Anura" pageId="4" pageNumber="57" phylum="Chordata" rank="genus">
|
||
<emphasis id="B944EADBFFECC7517D89C152FE72FC73" bold="true" box="[407,498,893,914]" italics="true" pageId="4" pageNumber="57">Pelodytes</emphasis>
|
||
</taxonomicName>
|
||
) from the Phosphorites du Quercy. 1: presacral vertebra UM-MAL 617, from the late Eocene of Malpérié (MP 17), in dorsal (a), anterior (b), and ventral (c) views. 2: posterior presacral vertebra UM-SNB 2406, from the late Eocene of Sainte-Néboule (MP 18), in anterior (a) and ventral (b) views. 3: right humerus UM-ECA 2556, from the late Eocene of Escamps (MP 19), in ventral view. 4: left ilium UM-ECA 2534, from the late Eocene of Escamps (MP 19), in lateral view. 5: right tibiale-fibulare UM-MAL 616, from the late Eocene (MP 17) of Malpérié (proximal section (a); dorsal view (b); and distal section (c)). Each scale bar = 3 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8F36C9FFECC7517CB8C513FC57FB6F" blockId="4.[128,768,1111,2099]" lastBlockId="4.[843,1482,1111,1166]" pageId="4" pageNumber="57">
|
||
In conclusion, the specimens from the Phosphorites allocated to
|
||
<taxonomicName id="4C304D4AFFECC7517D1BC573FE02F892" baseAuthorityName="Henrici" baseAuthorityYear="1994" box="[261,386,1884,1907]" class="Amphibia" family="Pelodytidae" kingdom="Animalia" order="Anura" pageId="4" pageNumber="57" phylum="Chordata" rank="family">Pelodytidae</taxonomicName>
|
||
are quite similar to those of extant
|
||
<taxonomicName id="4C304D4AFFECC7517C9EC553FF64F872" baseAuthorityName="Sanchiz" baseAuthorityYear="1978" box="[128,228,1916,1939]" class="Amphibia" family="Pelodytidae" genus="Pelodytes" kingdom="Animalia" order="Anura" pageId="4" pageNumber="57" phylum="Chordata" rank="genus">
|
||
<emphasis id="B944EADBFFECC7517C9EC553FF64F872" box="[128,228,1916,1939]" italics="true" pageId="4" pageNumber="57">Pelodytes</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation id="EFA14B38FFECC7517CEEC553FE71F872" author="Henrici, A. C. & Baez, A. M. & Grande, L." box="[240,497,1916,1939]" pageId="4" pageNumber="57" pagination="295 - 309" refId="ref12337" refString="Henrici, A. C., Baez, A. M., Grande, L. (2013): Aerugoamnis paulus, new genus and new species (Anura: Anomocoela): first reported anuran from the early Eocene (Wasatchian) Fossil Butte Member of the Green River Formation, Wyoming. - Annals of Carnegie Museum, 81: 295 - 309. http: // dx. doi. org / 10.2992 / 007.081.0402" type="journal article" year="2013">Henrici et al. (2013: 304)</bibRefCitation>
|
||
noted that reliable referral of the remains from the Eocene of Europe to pelodytids (as cf.
|
||
<taxonomicName id="4C304D4AFFECC7517C9EC593FF64F832" baseAuthorityName="Sanchiz" baseAuthorityYear="1978" box="[128,228,1980,2003]" class="Amphibia" family="Pelodytidae" genus="Pelodytes" kingdom="Animalia" order="Anura" pageId="4" pageNumber="57" phylum="Chordata" rank="genus">
|
||
<emphasis id="B944EADBFFECC7517C9EC593FF64F832" box="[128,228,1980,2003]" italics="true" pageId="4" pageNumber="57">Pelodytes</emphasis>
|
||
</taxonomicName>
|
||
) would require finding of a fused tibiale-fibulare in the same localities. The recovery of such a tibiale-fibulare confirms that a fossil taxon closely related to
|
||
<taxonomicName id="4C304D4AFFECC7517E49C5D3FD3BF7F2" baseAuthorityName="Sanchiz" baseAuthorityYear="1978" box="[599,699,2044,2067]" class="Amphibia" family="Pelodytidae" genus="Pelodytes" kingdom="Animalia" order="Anura" pageId="4" pageNumber="57" phylum="Chordata" rank="genus">
|
||
<emphasis id="B944EADBFFECC7517E49C5D3FD3BF7F2" box="[599,699,2044,2067]" italics="true" pageId="4" pageNumber="57">Pelodytes</emphasis>
|
||
</taxonomicName>
|
||
, if not
|
||
<taxonomicName id="4C304D4AFFECC7517C9ECA33FF64F7D2" baseAuthorityName="Sanchiz" baseAuthorityYear="1978" box="[128,228,2076,2099]" class="Amphibia" family="Pelodytidae" genus="Pelodytes" kingdom="Animalia" order="Anura" pageId="4" pageNumber="57" phylum="Chordata" rank="genus">
|
||
<emphasis id="B944EADBFFECC7517C9ECA33FF64F7D2" box="[128,228,2076,2099]" italics="true" pageId="4" pageNumber="57">Pelodytes</emphasis>
|
||
</taxonomicName>
|
||
, is present in the late middle and late Eocene (from MP 16 to MP 19) of western Europe, more specifically in the Phosphorites.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |