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27 KiB
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204 lines
27 KiB
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<mods:title id="D499BB04F4B84C062B250E2C945D281B">The known species of the genus Achagua Rindge, 1983, with the description of three new species from the Neotropics (Geometridae: Ennominae)</mods:title>
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<mods:namePart id="0ED8264E08C0F4A3915A8A0492F60A5D">Matson, Tanner A.</mods:namePart>
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<mods:date id="5E67D8D8FD699FCAB3184CD08E825630">2023</mods:date>
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<paragraph id="8B8F36BBFFC6E029D398FC51FE26FBB1" blockId="1.[151,431,1014,1041]" box="[151,431,1014,1041]" pageId="1" pageNumber="566">
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<taxonomicName id="4C304D38FFC6E029D398FC51FE26FBB1" ID-CoL="8ZYLT" authority="Rindge, 1983" authorityName="Rindge" authorityYear="1983" box="[151,431,1014,1041]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="566" phylum="Arthropoda" rank="genus">
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<emphasis id="B944EAA9FFC6E029D398FC51FE26FBB1" bold="true" box="[151,431,1014,1041]" pageId="1" pageNumber="566">
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<emphasis id="B944EAA9FFC6E029D398FC51FE8DFBB0" bold="true" box="[151,260,1014,1041]" italics="true" pageId="1" pageNumber="566">Achagua</emphasis>
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<bibRefCitation id="EFA14B4AFFC6E029D203FC51FE26FBB1" author="Rindge, F. H." box="[268,431,1014,1040]" pageId="1" pageNumber="566" pagination="147 - 262" refId="ref7036" refString="Rindge, F. H. (1983) A generic revision of the New World Nacophorini (Lepidoptera: Geometridae). Bulletin of the American Museum of Natural History, 175, 147 - 262." type="journal article" year="1983">Rindge, 1983</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<subSubSection id="C32A6530FFC6E02AD398FB98FD9FFEBC" lastPageId="2" lastPageNumber="567" pageId="1" pageNumber="566" type="type_taxon">
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<paragraph id="8B8F36BBFFC6E029D398FB98FC52FBF9" blockId="1.[151,987,1086,1113]" box="[151,987,1086,1113]" pageId="1" pageNumber="566">
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<heading id="D0C781D7FFC6E029D398FB98FC52FBF9" bold="true" box="[151,987,1086,1113]" fontSize="11" level="1" pageId="1" pageNumber="566" reason="1">
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<emphasis id="B944EAA9FFC6E029D398FB98FC52FBF9" bold="true" box="[151,987,1086,1113]" pageId="1" pageNumber="566">
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<typeStatus id="548B8819FFC6E029D398FB98FF5DFBF8" box="[151,212,1087,1113]" pageId="1" pageNumber="566">Type</typeStatus>
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Species:
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<taxonomicName id="4C304D38FFC6E029D247FB99FD47FBF9" authority="Rindge, 1983" authorityName="Rindge" authorityYear="1983" box="[328,718,1086,1113]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="566" phylum="Arthropoda" rank="species" species="obsoleta">
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<emphasis id="B944EAA9FFC6E029D247FB99FDA9FBF8" bold="true" box="[328,544,1086,1113]" italics="true" pageId="1" pageNumber="566">Achagua obsoleta</emphasis>
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<bibRefCitation id="EFA14B4AFFC6E029D127FB99FD47FBF9" author="Rindge, F. H." box="[552,718,1086,1112]" pageId="1" pageNumber="566" pagination="147 - 262" refId="ref7036" refString="Rindge, F. H. (1983) A generic revision of the New World Nacophorini (Lepidoptera: Geometridae). Bulletin of the American Museum of Natural History, 175, 147 - 262." type="journal article" year="1983">Rindge, 1983</bibRefCitation>
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</taxonomicName>
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; original designation.
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B8F36BBFFC6E029D398FB21FEB4F9F0" blockId="1.[151,1437,1158,2013]" pageId="1" pageNumber="566">
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<emphasis id="B944EAA9FFC6E029D398FB21FEBEFB01" bold="true" box="[151,311,1158,1184]" pageId="1" pageNumber="566">Re-diagnosis.</emphasis>
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Both
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<bibRefCitation id="EFA14B4AFFC6E029D28DFB20FDA1FB00" author="Rindge, F. H." box="[386,552,1159,1185]" pageId="1" pageNumber="566" pagination="147 - 262" refId="ref7036" refString="Rindge, F. H. (1983) A generic revision of the New World Nacophorini (Lepidoptera: Geometridae). Bulletin of the American Museum of Natural History, 175, 147 - 262." type="journal article" year="1983">Rindge (1983)</bibRefCitation>
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and
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<bibRefCitation id="EFA14B4AFFC6E029D16AFB20FD74FB00" author="Pitkin, L." box="[613,765,1159,1185]" pageId="1" pageNumber="566" pagination="121 - 401" refId="ref6844" refString="Pitkin, L. (2002) Neotropical Ennomine moths: a review of the genera (Lepidoptera: Geometridae). Zoological Journal of the Linnean Society, 135 (2 - 3), 121 - 401. https: // doi. org / 10.1046 / j. 1096 - 3642.2002.01200. x" type="journal article" year="2002">Pitkin (2002)</bibRefCitation>
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provided diagnostic comments for
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<taxonomicName id="4C304D38FFC6E029D794FB20FB76FB00" authorityName="Rindge" authorityYear="1983" box="[1179,1279,1159,1185]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="566" phylum="Arthropoda" rank="genus">
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<emphasis id="B944EAA9FFC6E029D794FB20FB76FB00" box="[1179,1279,1159,1185]" italics="true" pageId="1" pageNumber="566">Achagua</emphasis>
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</taxonomicName>
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, but with the discovery of three undescribed species, it is necessary to revisit these diagnoses. The mostly bipectinate antennae of male
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<taxonomicName id="4C304D38FFC6E029D3F7FB68FED5FB48" authorityName="Rindge" authorityYear="1983" box="[248,348,1231,1257]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="566" phylum="Arthropoda" rank="genus">
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<emphasis id="B944EAA9FFC6E029D3F7FB68FED5FB48" box="[248,348,1231,1257]" italics="true" pageId="1" pageNumber="566">Achagua</emphasis>
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</taxonomicName>
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, with the apical third to quarter being simple filiform, is a shared characteristic among related nacophorine genera, e.g.,
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<emphasis id="B944EAA9FFC6E029D2B2FB54FDA8FAAC" box="[445,545,1267,1293]" italics="true" pageId="1" pageNumber="566">Cargolia</emphasis>
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and
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<emphasis id="B944EAA9FFC6E029D154FB54FD4BFAAC" box="[603,706,1267,1293]" italics="true" pageId="1" pageNumber="566">Gabriola</emphasis>
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, and thus, does not disambiguate closely related taxa. In males, the two processes of the uncus in conjunction with the subtriangular anellar processes appear to be the most reliable diagnostic features (
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<figureCitation id="130B2A3EFFC6E029D2A0FA9CFD0DFAF4" box="[431,644,1338,1365]" captionStart="FIGURES 5–8" captionStartId="4.[152,270,1869,1893]" captionTargetBox="[158,1432,190,1832]" captionTargetId="figure-25@4.[151,1436,181,1845]" captionTargetPageId="4" captionText="FIGURES 5–8. Achagua male genitalia. (a) genital capsule, (b) phallus. (5) A. obsoleta Rindge, TAM-2023-270, AMNH_IZC 00353009, paratype, AMNH; (6) A. magna n. sp., TAM-2023-268, AMNH_IZC 00353023, paratype. AMNH; (7) A. cooperae n. sp., TAM-2023-251,AMNH_IZC 00353028, paratype, AMNH; (8) A. velata n. sp., TAM-2023-289, holotype, MNHN. Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.8426642" httpUri="https://zenodo.org/record/8426642/files/figure.png" pageId="1" pageNumber="566">Figs. 5a, 6a, 7a, 8a</figureCitation>
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).
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<bibRefCitation id="EFA14B4AFFC6E029D196FA9CFCA4FAF4" author="Pitkin, L." box="[665,813,1339,1365]" pageId="1" pageNumber="566" pagination="121 - 401" refId="ref6844" refString="Pitkin, L. (2002) Neotropical Ennomine moths: a review of the genera (Lepidoptera: Geometridae). Zoological Journal of the Linnean Society, 135 (2 - 3), 121 - 401. https: // doi. org / 10.1046 / j. 1096 - 3642.2002.01200. x" type="journal article" year="2002">Pitkin (2002)</bibRefCitation>
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listed the strongly spatulate (or capitate) dorsal process of the uncus as an apomorphy for the genus, but two of the new species described herein possess a non-spatulate dorsal process (
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<figureCitation id="130B2A3EFFC6E029D24BFA24FE4EFA3C" box="[324,455,1411,1437]" captionStart="FIGURES 5–8" captionStartId="4.[152,270,1869,1893]" captionTargetBox="[158,1432,190,1832]" captionTargetId="figure-25@4.[151,1436,181,1845]" captionTargetPageId="4" captionText="FIGURES 5–8. Achagua male genitalia. (a) genital capsule, (b) phallus. (5) A. obsoleta Rindge, TAM-2023-270, AMNH_IZC 00353009, paratype, AMNH; (6) A. magna n. sp., TAM-2023-268, AMNH_IZC 00353023, paratype. AMNH; (7) A. cooperae n. sp., TAM-2023-251,AMNH_IZC 00353028, paratype, AMNH; (8) A. velata n. sp., TAM-2023-289, holotype, MNHN. Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.8426642" httpUri="https://zenodo.org/record/8426642/files/figure.png" pageId="1" pageNumber="566">Figs. 7a, 8a</figureCitation>
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). In females, the large, elongate, apically pointed, denticulate signum may be diagnostic, but only a single preparation of
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<taxonomicName id="4C304D38FFC6E029D10BFA00FD0AFA60" box="[516,643,1447,1473]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="566" phylum="Arthropoda" rank="species" species="obsoleta">
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<emphasis id="B944EAA9FFC6E029D10BFA00FD0AFA60" box="[516,643,1447,1473]" italics="true" pageId="1" pageNumber="566">A. obsoleta</emphasis>
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</taxonomicName>
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(
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<figureCitation id="130B2A3EFFC6E029D19CFA00FD51FA60" box="[659,728,1447,1473]" captionStart="FIGURES 9–10" captionStartId="5.[152,270,1436,1460]" captionTargetBox="[256,1331,198,1417]" captionTargetId="figure-16@5.[244,1343,198,1429]" captionTargetPageId="5" captionText="FIGURES 9–10. Achagua female genitalia. (9) A. obsoleta Rindge, No. 20288, NHMUK010293435, NHMUK; photo credit: Louise Berridge (NHMUK); (10) A. cooperae n. sp., TAM-2023-276, USNMENT01771269, paratype, USNM. Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.8426644" httpUri="https://zenodo.org/record/8426644/files/figure.png" pageId="1" pageNumber="566">Fig. 9</figureCitation>
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) and two preparations of
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<taxonomicName id="4C304D38FFC6E029D0F2FA00FB0CFA61" authorityName="Matson" authorityYear="2023" box="[1021,1157,1447,1473]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="566" phylum="Arthropoda" rank="species" species="cooperae">
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<emphasis id="B944EAA9FFC6E029D0F2FA00FB0CFA61" box="[1021,1157,1447,1473]" italics="true" pageId="1" pageNumber="566">A. cooperae</emphasis>
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</taxonomicName>
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(
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<figureCitation id="130B2A3EFFC6E029D799FA00FB61FA60" box="[1174,1256,1447,1473]" captionStart="FIGURES 9–10" captionStartId="5.[152,270,1436,1460]" captionTargetBox="[256,1331,198,1417]" captionTargetId="figure-16@5.[244,1343,198,1429]" captionTargetPageId="5" captionText="FIGURES 9–10. Achagua female genitalia. (9) A. obsoleta Rindge, No. 20288, NHMUK010293435, NHMUK; photo credit: Louise Berridge (NHMUK); (10) A. cooperae n. sp., TAM-2023-276, USNMENT01771269, paratype, USNM. Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.8426644" httpUri="https://zenodo.org/record/8426644/files/figure.png" pageId="1" pageNumber="566">Fig. 10</figureCitation>
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) were available for study. Superficially, the pearly-white ground color, large size, brown to black wing margins, and lack of welldefined antemedial and postmedial lines seem to also define
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<taxonomicName id="4C304D38FFC6E029D032FA48FC28F9A8" authorityName="Rindge" authorityYear="1983" box="[829,929,1519,1545]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="566" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B944EAA9FFC6E029D032FA48FC28F9A8" box="[829,929,1519,1545]" italics="true" pageId="1" pageNumber="566">Achagua</emphasis>
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||
</taxonomicName>
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||
. Further, the multi-locus phylogenetic results of
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<bibRefCitation id="EFA14B4AFFC6E029D3B8F9B4FE70F98C" author="Murillo-Ramos, Ld. C. & Brehm, G. & Sihvonen, P. & Hausmann, A. & Holm, S. & Ghanavi, H. R. & Ounap, E. & Truuverk, A. & Staude, H. & Friedrich, E. & Tammaru, T. & Wahlberg, N." box="[183,505,1555,1581]" pageId="1" pageNumber="566" pagination="7386" refId="ref6742" refString="Murillo-Ramos, Ld. C., Brehm, G., Sihvonen, P., Hausmann, A., Holm, S., Ghanavi, H. R., Ounap, E., Truuverk, A., Staude, H., Friedrich, E., Tammaru, T. & Wahlberg, N. (2019) A comprehensive molecular phylogeny of Geometridae (Lepidoptera) with a focus on enigmatic small subfamilies. PeerJ, 7, e 7386. https: // doi. org / 10.7717 / peerj. 7386" type="journal article" year="2019">
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||
Murillo-Ramos
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||
<emphasis id="B944EAA9FFC6E029D263F9B4FE2FF98C" box="[364,422,1555,1581]" italics="true" pageId="1" pageNumber="566">et al.</emphasis>
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||
(2019)
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||
</bibRefCitation>
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||
and
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||
<bibRefCitation id="EFA14B4AFFC6E029D13DF9B4FC9AF98C" author="Brehm, G. & Murillo-Ramos, L. & Sihvonen, P. & Hausmann, A. & Schmidt, B. C. & Ounap, E. & Moser A. & Mortter, R. & Bolt, D. & Bodner, F. & Lindt, A. & Parra, L. E. & Wahlberg, N." box="[562,787,1555,1581]" pageId="1" pageNumber="566" pagination="457 - 486" refId="ref6406" refString="Brehm, G., Murillo-Ramos, L., Sihvonen, P., Hausmann, A., Schmidt, B. C., Ounap, E., Moser A., Mortter, R., Bolt, D., Bodner, F., Lindt, A., Parra, L. E. & Wahlberg, N. (2019) New World geometrid moths (Lepidoptera: Geometridae): Molecular phylogeny, biogeography, taxonomic updates and description of 11 new tribes. Arthropod Systematics and Phylogeny, 77 (3), 457 - 486. https: // doi. org / 10.26049 / ASP 77 - 3 - 2019 - 5" type="journal article" year="2019">
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||
Brehm
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<emphasis id="B944EAA9FFC6E029D189F9B4FD49F98C" box="[646,704,1555,1581]" italics="true" pageId="1" pageNumber="566">et al.</emphasis>
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||
(2019)
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||
</bibRefCitation>
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||
also support the recognition of
|
||
<taxonomicName id="4C304D38FFC6E029D770F9B4FB6AF98C" authorityName="Rindge" authorityYear="1983" box="[1151,1251,1555,1581]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="566" phylum="Arthropoda" rank="genus">
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<emphasis id="B944EAA9FFC6E029D770F9B4FB6AF98C" box="[1151,1251,1555,1581]" italics="true" pageId="1" pageNumber="566">Achagua</emphasis>
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</taxonomicName>
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||
as distinct from related genera.
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||
</paragraph>
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<paragraph id="8B8F36BBFFC6E029D3C8F9FDFE95F940" blockId="1.[151,1437,1158,2013]" pageId="1" pageNumber="566">
|
||
<emphasis id="B944EAA9FFC6E029D3C8F9FDFED0F9D5" bold="true" box="[199,345,1626,1652]" pageId="1" pageNumber="566">Distribution</emphasis>
|
||
(
|
||
<figureCitation id="130B2A3EFFC6E029D263F9FCFE36F9D4" box="[364,447,1627,1653]" captionStart="FIGURE 11" captionStartId="9.[152,255,1039,1063]" captionTargetBox="[151,1436,181,1015]" captionTargetId="figure-16@9.[151,1436,181,1015]" captionTargetPageId="9" captionText="FIGURE 11. Geographic distribution of Achagua: A. obsoleta Rindge (star), A. magna n. sp. (triangle), A. cooperae n. sp. (circle), and A. velata n. sp. (square). Single dots may represent>1 individual. Scale bar = 600 km." figureDoi="http://doi.org/10.5281/zenodo.8426646" httpUri="https://zenodo.org/record/8426646/files/figure.png" pageId="1" pageNumber="566">Fig. 11</figureCitation>
|
||
).
|
||
<taxonomicName id="4C304D38FFC6E029D2D6F9FCFDB4F9D4" authorityName="Rindge" authorityYear="1983" box="[473,573,1627,1653]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="566" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B944EAA9FFC6E029D2D6F9FCFDB4F9D4" box="[473,573,1627,1653]" italics="true" pageId="1" pageNumber="566">Achagua</emphasis>
|
||
</taxonomicName>
|
||
inhabit the tropical and subtropical moist broadleaf forests of Central and South America from the Sierra de los Tuxtlas of
|
||
<collectingRegion id="49F4F859FFC6E029D1B7F9D8FC96F938" box="[696,799,1663,1689]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="1" pageNumber="566">Veracruz</collectingRegion>
|
||
,
|
||
<collectingCountry id="F327762BFFC6E029D025F9D8FC09F938" box="[810,896,1663,1689]" name="Mexico" pageId="1" pageNumber="566">Mexico</collectingCountry>
|
||
(
|
||
<taxonomicName id="4C304D38FFC6E029D09FF9D8FB9EF939" authorityName="Matson" authorityYear="2023" box="[912,1047,1663,1689]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="566" phylum="Arthropoda" rank="species" species="cooperae">
|
||
<emphasis id="B944EAA9FFC6E029D09FF9D8FB9EF939" box="[912,1047,1663,1689]" italics="true" pageId="1" pageNumber="566">A. cooperae</emphasis>
|
||
</taxonomicName>
|
||
), east to the Guiana Highlands (
|
||
<taxonomicName id="4C304D38FFC6E029D68BF9D8FF52F91C" authorityName="Matson" authorityYear="2023" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="566" phylum="Arthropoda" rank="species" species="velata">
|
||
<emphasis id="B944EAA9FFC6E029D68BF9D8FF52F91C" italics="true" pageId="1" pageNumber="566">A. velata</emphasis>
|
||
</taxonomicName>
|
||
), and southward through the eastern Andes of
|
||
<collectingCountry id="F327762BFFC6E029D1E0F904FCE9F91C" box="[751,864,1699,1725]" name="Colombia" pageId="1" pageNumber="566">Colombia</collectingCountry>
|
||
,
|
||
<collectingCountry id="F327762BFFC6E029D063F904FC42F91C" box="[876,971,1699,1725]" name="Ecuador" pageId="1" pageNumber="566">Ecuador</collectingCountry>
|
||
, and
|
||
<collectingCountry id="F327762BFFC6E029D707F904FBB2F91C" box="[1032,1083,1699,1725]" name="Peru" pageId="1" pageNumber="566">Peru</collectingCountry>
|
||
to at least the Bolivian Yungas (
|
||
<taxonomicName id="4C304D38FFC6E029D3AEF960FE85F941" authorityName="Matson" authorityYear="2023" box="[161,268,1735,1761]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="566" phylum="Arthropoda" rank="species" species="magna">
|
||
<emphasis id="B944EAA9FFC6E029D3AEF960FE85F941" box="[161,268,1735,1761]" italics="true" pageId="1" pageNumber="566">A. magna</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8F36BBFFC6E029D3C8F94DFF5BF888" blockId="1.[151,1437,1158,2013]" pageId="1" pageNumber="566">
|
||
<emphasis id="B944EAA9FFC6E029D3C8F94DFEAEF8A5" bold="true" box="[199,295,1770,1796]" pageId="1" pageNumber="566">Biology.</emphasis>
|
||
Thus far, no life history information is known for the genus. Adults are infrequently collected at light.
|
||
</paragraph>
|
||
<paragraph id="8B8F36BBFFC6E029D3C8F894FEA1F87C" blockId="1.[151,1437,1158,2013]" pageId="1" pageNumber="566">
|
||
<emphasis id="B944EAA9FFC6E029D3C8F894FEB2F8EC" bold="true" box="[199,315,1843,1869]" pageId="1" pageNumber="566">Remarks.</emphasis>
|
||
In
|
||
<taxonomicName id="4C304D38FFC6E029D26EF894FE4CF8EC" authorityName="Rindge" authorityYear="1983" box="[353,453,1843,1869]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="566" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B944EAA9FFC6E029D26EF894FE4CF8EC" box="[353,453,1843,1869]" italics="true" pageId="1" pageNumber="566">Achagua</emphasis>
|
||
</taxonomicName>
|
||
, the uncus is characterized by two distinct processes. However, there is some disagreement regarding the nomenclature of these processes.
|
||
<bibRefCitation id="EFA14B4AFFC6E029D1FBF8F0FC11F8D0" author="Rindge, F. H." box="[756,920,1879,1905]" pageId="1" pageNumber="566" pagination="147 - 262" refId="ref7036" refString="Rindge, F. H. (1983) A generic revision of the New World Nacophorini (Lepidoptera: Geometridae). Bulletin of the American Museum of Natural History, 175, 147 - 262." type="journal article" year="1983">Rindge (1983)</bibRefCitation>
|
||
referred to the much larger dorsal process as the pseudouncus, but it is unclear how he arrived at this conclusion. Alternatively, I adopt the terminology used by
|
||
<bibRefCitation id="EFA14B4AFFC6E029D398F838FEA4F818" author="Pitkin, L." box="[151,301,1951,1977]" pageId="1" pageNumber="566" pagination="121 - 401" refId="ref6844" refString="Pitkin, L. (2002) Neotropical Ennomine moths: a review of the genera (Lepidoptera: Geometridae). Zoological Journal of the Linnean Society, 135 (2 - 3), 121 - 401. https: // doi. org / 10.1046 / j. 1096 - 3642.2002.01200. x" type="journal article" year="2002">Pitkin (2002)</bibRefCitation>
|
||
and refer to these processes based on their orientation, specifically as the dorsal or ventral processes of the uncus.
|
||
</paragraph>
|
||
<paragraph id="8B8F36BBFFC5E02AD3C8FF30FD9FFEBC" blockId="2.[151,1437,151,285]" pageId="2" pageNumber="567">
|
||
<bibRefCitation id="EFA14B4AFFC5E02AD3C8FF30FEFAFF10" author="Campos, M. R." box="[199,371,151,177]" pageId="2" pageNumber="567" pagination="938 - 943" refId="ref6531" refString="Campos, M. R. (2001) A new genus and species of freshwater crab from Colombia (Crustacea: Decapoda: Pseudothelphusidae). Proceedings of the Biological Society of Washington 114 (4), 938 - 943." type="journal article" year="2001">Campos (2001)</bibRefCitation>
|
||
described the genus “
|
||
<taxonomicName id="4C304D38FFC5E02AD169FF30FD43FF10" authorityName="Rindge" authorityYear="1983" box="[614,714,151,177]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="567" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B944EAA9FFC5E02AD169FF30FD43FF10" box="[614,714,151,177]" italics="true" pageId="2" pageNumber="567">Achagua</emphasis>
|
||
</taxonomicName>
|
||
” for a group of freshwater crabs. As the name was already preoccupied by
|
||
<taxonomicName id="4C304D38FFC5E02AD206FF1CFE4DFF74" authority="Rindge" authorityName="Rindge" authorityYear="1983" box="[265,452,187,213]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="567" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B944EAA9FFC5E02AD206FF1CFEE4FF74" box="[265,365,187,213]" italics="true" pageId="2" pageNumber="567">Achagua</emphasis>
|
||
Rindge
|
||
</taxonomicName>
|
||
(the subject of this work), “
|
||
<taxonomicName id="4C304D38FFC5E02AD1F8FF1CFCD2FF74" authority="Campos" authorityName="Rindge" authorityYear="1983" box="[759,859,187,213]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="567" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B944EAA9FFC5E02AD1F8FF1CFCD2FF74" box="[759,859,187,213]" italics="true" pageId="2" pageNumber="567">Achagua</emphasis>
|
||
</taxonomicName>
|
||
” Campos was rendered a junior homonym. Campos and Magalh„es (2004) synonymized “
|
||
<taxonomicName id="4C304D38FFC5E02AD132FF78FD28FF58" authority="Campos" authorityName="Rindge" authorityYear="1983" box="[573,673,223,249]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="567" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B944EAA9FFC5E02AD132FF78FD28FF58" box="[573,673,223,249]" italics="true" pageId="2" pageNumber="567">Achagua</emphasis>
|
||
</taxonomicName>
|
||
” Campos with
|
||
<taxonomicName id="4C304D38FFC5E02AD043FF78FBC8FF58" authority="Pretzmann" authorityName="Pretzmann" authorityYear="1971" box="[844,1089,223,249]" class="Malacostraca" family="Pseudothelphusidae" genus="Eudaniela" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="567" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B944EAA9FFC5E02AD043FF78FC36FF58" box="[844,959,223,249]" italics="true" pageId="2" pageNumber="567">Eudaniela</emphasis>
|
||
Pretzmann
|
||
</taxonomicName>
|
||
, such that this junior homonym has not been in contemporary use.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |