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<document id="A3CA339A04F465201DD09722924BD8E7" ID-CLB-Dataset="22458" ID-DOI="10.5852/ejt.2020.622" ID-GBIF-Dataset="9cb3ea96-0ee4-40b4-8006-19b5fed80b46" ID-Zenodo-Dep="3777282" ID-ZooBank="9CD3B2CD-F072-4994-8CA9-24145D343401" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1588217079841" checkinUser="valdenar" docAuthor="Darby, Michael" docDate="2020" docId="039987A1386FFFB02BC3FB80FC79FD7E" docLanguage="en" docName="ejt-622_darby.pdf.imf" docOrigin="European Journal of Taxonomy 622" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Cissidium steeli Darby 2020, sp. nov." docType="treatment" docUuid="86C4969A-78B4-4DD6-A2BB-443588CDFCD0" docUuidSource="ZooBank" docVersion="7" lastPageNumber="127" masterDocId="FFA0FFD93812FFCE2949FFEEFF9CFF98" masterDocTitle="A revision of Cissidium Motschulsky (Coleoptera: Ptiliidae) with seventy seven new species" masterLastPageNumber="188" masterPageNumber="1" pageNumber="126" updateTime="1698838121940" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="25926611848751A2D3F1B5C2EF0E3CC9">
<mods:title id="D6A5C335A08ABC8202CC5FD9FC32FBCA">A revision of Cissidium Motschulsky (Coleoptera: Ptiliidae) with seventy seven new species</mods:title>
</mods:titleInfo>
<mods:name id="FE511194963D871646967A4A33DFAEB4" type="personal">
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<mods:namePart id="222F95C2A987BAC3861AAD06B631A3E1">Darby, Michael</mods:namePart>
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<mods:title id="7656AE831FEA25FF28E1C598B18546F0">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="1D0081ECF000366F63E3E62D4EEDB607">
<mods:date id="6782723E8A3AE09B3753AECCA1DC2662">2020</mods:date>
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<mods:number id="8764F0C7751C9A5E8305F6DE26DEB3C3">2020-04-02</mods:number>
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<mods:number id="3A7E05D7EA4D49EE7A27B3242C248A7A">622</mods:number>
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<mods:classification id="40362C7AC090816DBCB174E9F8CF4D84">journal article</mods:classification>
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<mods:identifier id="C8E4D771BDFB6AD9DFB881DCD36BB072" type="DOI">10.5852/ejt.2020.622</mods:identifier>
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<treatment id="039987A1386FFFB02BC3FB80FC79FD7E" ID-DOI="http://doi.org/10.5281/zenodo.3795646" ID-GBIF-Taxon="163877816" ID-Zenodo-Dep="3795646" LSID="urn:lsid:zoobank.org:act:86C4969A-78B4-4DD6-A2BB-443588CDFCD0" httpUri="http://treatment.plazi.org/id/039987A1386FFFB02BC3FB80FC79FD7E" lastPageId="126" lastPageNumber="127" pageId="125" pageNumber="126">
<subSubSection id="C32A653C386FFFB32BC3FB80FB51FB35" pageId="125" pageNumber="126" type="nomenclature">
<paragraph id="8B8F36B7386FFFB32BC3FB80FC36FB11" blockId="125.[358,1229,1134,1233]" box="[650,938,1134,1161]" pageId="125" pageNumber="126">
<heading id="D0C781DB386FFFB32BC3FB80FC36FB11" box="[650,938,1134,1161]" centered="true" fontSize="11" level="2" pageId="125" pageNumber="126" reason="2">
<taxonomicName id="4C304D34386FFFB32BC3FB80FCD4FB11" ID-CoL="8XXRR" authority="Darby, 2020" authorityName="Darby" authorityYear="2020" box="[650,840,1134,1161]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="125" pageNumber="126" phylum="Arthropoda" rank="species" species="steeli" status="sp. nov.">
<emphasis id="B944EAA5386FFFB32BC3FB80FCD4FB11" bold="true" box="[650,840,1134,1161]" italics="true" pageId="125" pageNumber="126">Cissidium steeli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757DE386FFFB32A06FB81FC36FB11" box="[847,938,1135,1161]" pageId="125" pageNumber="126" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="8B8F36B7386FFFB3282FFB7CFB51FB35" blockId="125.[358,1229,1134,1233]" box="[358,1229,1170,1197]" pageId="125" pageNumber="126">
<uri id="FFA13AB5386FFFB3282FFB7CFB51FB35" box="[358,1229,1170,1197]" pageId="125" pageNumber="126">
urn:lsid:zoobank.org:act:
<uuid id="FF960C62386FFFB32BCBFB7CFB51FB35" box="[642,1229,1170,1197]" pageId="125" pageNumber="126">86C4969A-78B4-4DD6-A2BB-443588CDFCD0</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C32A653C386FFFB32BB9FB59FCDFFB49" box="[752,835,1207,1233]" pageId="125" pageNumber="126" type="description">
<paragraph id="8B8F36B7386FFFB32BB9FB59FCDFFB49" blockId="125.[358,1229,1134,1233]" box="[752,835,1207,1233]" pageId="125" pageNumber="126">
<figureCitation id="130B2A32386FFFB32BB9FB59FCDFFB49" box="[752,835,1207,1233]" captionStart="Fig" captionStartId="126.[189,232,1797,1823]" captionTargetBox="[195,1392,848,1755]" captionTargetId="figure@126.[194,1395,847,1755]" captionTargetPageId="126" captionText="Fig. 80. Cissidium steeli sp. nov. A. Habitus. B. Pronotum, × 680. C. Mesoventrite showing median process of collar, mid-keel and keel, × 785." figureDoi="http://doi.org/10.5281/zenodo.3777444" httpUri="https://zenodo.org/record/3777444/files/figure.png" pageId="125" pageNumber="126">Fig. 80</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32A653C386FFFB329F4FB14FC4DFADC" box="[189,977,1274,1348]" pageId="125" pageNumber="126" type="etymology">
<paragraph id="8B8F36B7386FFFB329F4FB14FED8FA8C" blockId="125.[189,324,1274,1300]" box="[189,324,1274,1300]" pageId="125" pageNumber="126">
<heading id="D0C781DB386FFFB329F4FB14FED8FA8C" bold="true" box="[189,324,1274,1300]" fontSize="11" level="3" pageId="125" pageNumber="126" reason="3">
<emphasis id="B944EAA5386FFFB329F4FB14FED8FA8C" bold="true" box="[189,324,1274,1300]" pageId="125" pageNumber="126">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8B8F36B7386FFFB329F4FAC4FC4DFADC" blockId="125.[189,977,1322,1348]" box="[189,977,1322,1348]" pageId="125" pageNumber="126">
Johnson ms name, after W. Steel, the well-known
<collectingCountry id="F3277627386FFFB32A41FAC4FCAEFADC" box="[776,818,1322,1348]" name="United Kingdom" pageId="125" pageNumber="126">UK</collectingCountry>
Coleopterist.
</paragraph>
</subSubSection>
<subSubSection id="C32A653C386FFFB329F4FA82FBABFA43" pageId="125" pageNumber="126" type="materials_examined">
<paragraph id="8B8F36B7386FFFB329F4FA82FE36FA1E" blockId="125.[189,426,1388,1414]" box="[189,426,1388,1414]" pageId="125" pageNumber="126">
<heading id="D0C781DB386FFFB329F4FA82FE36FA1E" bold="true" box="[189,426,1388,1414]" fontSize="11" level="3" pageId="125" pageNumber="126" reason="3">
<emphasis id="B944EAA5386FFFB329F4FA82FE36FA1E" bold="true" box="[189,426,1388,1414]" pageId="125" pageNumber="126">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="3B583CEA386FFFB329A3FA73FBABFA43" ID-GBIF-Occurrence="2609493513" collectedFrom="leaf litter" collectingDate="1969-08" collectionCode="MMUE" collectorName="J. Darlington" country="Trinidad and Tobago" location="TRINIDAD" pageId="125" pageNumber="126" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<paragraph id="8B8F36B7386FFFB329A3FA73FEC0FA2F" blockId="125.[189,1079,1437,1499]" box="[234,348,1437,1463]" pageId="125" pageNumber="126">
<emphasis id="B944EAA5386FFFB329A3FA73FEC0FA2F" bold="true" box="[234,348,1437,1463]" pageId="125" pageNumber="126">
<typeStatus id="548B8815386FFFB329A3FA73FEC0FA2F" box="[234,348,1437,1463]" pageId="125" pageNumber="126" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8B8F36B7386FFFB329F4FA2FFBABFA43" blockId="125.[189,1079,1437,1499]" box="[189,1079,1473,1499]" pageId="125" pageNumber="126">
<collectingCountry id="F3277627386FFFB329F4FA2FFED7FA43" box="[189,331,1473,1499]" name="Trinidad and Tobago" pageId="125" pageNumber="126">TRINIDAD</collectingCountry>
•
<specimenCount id="9D36FD3E386FFFB3282DFA2FFEE7FA43" box="[356,379,1473,1499]" pageId="125" pageNumber="126" type="female">♀</specimenCount>
; Tamana;
<collectedFrom id="382C754A386FFFB328BCFA2FFDFFFA43" box="[501,611,1473,1499]" pageId="125" pageNumber="126">leaf litter</collectedFrom>
;
<date id="FF8E1077386FFFB32B24FA2FFD70FA43" box="[621,748,1473,1499]" pageId="125" pageNumber="126" value="1969-08">
<collectingDate id="EFCAE99F386FFFB32B24FA2FFD70FA43" box="[621,748,1473,1499]" pageId="125" pageNumber="126" value="1969-08">Aug. 1969</collectingDate>
</date>
;
<collectorName id="26C55361386FFFB32BB1FA2FFC0DFA43" box="[760,913,1473,1499]" pageId="125" pageNumber="126">J. Darlington</collectorName>
leg.;
<collectionCode id="ED21AE72386FFFB32A9DFA2FFBAFFA43" box="[980,1075,1473,1499]" collectionName="United Kingdom, Manchester, The University, Manchester Museum" country="United Kingdom" httpUri="http://grbio.org/cool/he6q-xsyw" name="Museum of Manchester University" pageId="125" pageNumber="126">MMUE</collectionCode>
.
</paragraph>
</materialsCitation>
</subSubSection>
<subSubSection id="C32A653C386FFFB029F4F9EDFCF1FDEE" lastPageId="126" lastPageNumber="127" pageId="125" pageNumber="126" type="description">
<paragraph id="8B8F36B7386FFFB329F4F9EDFED2F985" blockId="125.[189,334,1539,1565]" box="[189,334,1539,1565]" pageId="125" pageNumber="126">
<heading id="D0C781DB386FFFB329F4F9EDFED2F985" bold="true" box="[189,334,1539,1565]" fontSize="11" level="3" pageId="125" pageNumber="126" reason="3">
<emphasis id="B944EAA5386FFFB329F4F9EDFED2F985" bold="true" box="[189,334,1539,1565]" pageId="125" pageNumber="126">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8B8F36B7386FFFB329F4F9DAFD36F9D6" blockId="125.[189,682,1587,1614]" box="[189,682,1587,1614]" pageId="125" pageNumber="126">
SIZE. Habitus (
<figureCitation id="130B2A32386FFFB3282EF9DDFE48F9D6" box="[359,468,1587,1614]" captionStart="Fig" captionStartId="126.[189,232,1797,1823]" captionTargetBox="[195,1392,848,1755]" captionTargetId="figure@126.[194,1395,847,1755]" captionTargetPageId="126" captionText="Fig. 80. Cissidium steeli sp. nov. A. Habitus. B. Pronotum, × 680. C. Mesoventrite showing median process of collar, mid-keel and keel, × 785." figureDoi="http://doi.org/10.5281/zenodo.3777444" httpUri="https://zenodo.org/record/3777444/files/figure.png" pageId="125" pageNumber="126">Fig. 80A</figureCitation>
), length
<quantity id="4CC89B52386FFFB32B73F9DAFD39F9D6" box="[570,677,1588,1614]" metricMagnitude="-4" metricUnit="m" metricValue="7.1" pageId="125" pageNumber="126" unit="mm" value="0.71">0.71 mm</quantity>
.
</paragraph>
<paragraph id="8B8F36B7386FFFB329F4F998FBABF908" blockId="125.[189,1079,1654,1681]" box="[189,1079,1654,1681]" pageId="125" pageNumber="126">COLOUR. Dark brown, shining, pubescence, legs and antennae dusky yellow.</paragraph>
<paragraph id="8B8F36B7386FFFB329F4F954FAA9F960" blockId="125.[189,1398,1721,1784]" pageId="125" pageNumber="126">
HEAD. With a foveolate shallow linear depression behind the eyes, width across eyes
<quantity id="4CC89B52386FFFB32DD6F957FA96F94C" box="[1183,1290,1721,1748]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="125" pageNumber="126" unit="mm" value="0.2">0.20 mm</quantity>
; mentum as
<figureCitation id="130B2A32386FFFB32994F933FEA6F960" box="[221,314,1757,1784]" captionStart="Fig" captionStartId="5.[189,232,1927,1953]" captionTargetBox="[232,1334,1274,1884]" captionTargetId="figure@5.[232,1334,1274,1887]" captionTargetPageId="5" captionText="Fig. 4. Menta and submenta. A. Cissidium petri Darby, 2015, × 1380. B. Cissidium tigrum Darby, 2015, × 1220. C. Cissidium amboroense Darby, 2015, × 1260. D. Cissidium losbanos sp. nov., × 1100. E. Cissidium triangulum Darby, 2019, × 650. F. Cissidium pilosellum sp. nov., × 1420." figureDoi="http://doi.org/10.5281/zenodo.3777290" httpUri="https://zenodo.org/record/3777290/files/figure.png" pageId="125" pageNumber="126">Fig. 4A</figureCitation>
; antennomeres III–XI, length
<quantity id="4CC89B52386FFFB32BD3F930FC9AF960" box="[666,774,1758,1784]" metricMagnitude="-4" metricUnit="m" metricValue="3.1" pageId="125" pageNumber="126" unit="mm" value="0.31">0.31 mm</quantity>
, III–IX length
<quantity id="4CC89B52386FFFB32AFDF930FBBCF960" box="[948,1056,1758,1784]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="125" pageNumber="126" unit="mm" value="0.2">0.20 mm</quantity>
, X–XI length
<quantity id="4CC89B52386FFFB32D8FF930FAADF960" box="[1222,1329,1758,1784]" metricMagnitude="-4" metricUnit="m" metricValue="1.1" pageId="125" pageNumber="126" unit="mm" value="0.11">0.11 mm</quantity>
.
</paragraph>
<paragraph id="8B8F36B7386FFFB329F4F8CFFEA3F830" blockId="125.[189,1399,1824,1960]" pageId="125" pageNumber="126">
PRONOTUM.
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long,
<quantity id="4CC89B52386FFFB328B1F8CEFDFDF8A3" box="[504,609,1824,1851]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="125" pageNumber="126" unit="mm" value="0.3">0.30 mm</quantity>
wide, sparsely pubescent without either foveolae or foveae; lateral margins evenly rounded to slightly acute hind angles, narrowly bordered, the border running along the entire posterior margin though narrower in the sinuous medial emargination in front of the scutellum (
<figureCitation id="130B2A32386FFFB3298FF860FEADF830" box="[198,305,1934,1960]" captionStart="Fig" captionStartId="126.[189,232,1797,1823]" captionTargetBox="[195,1392,848,1755]" captionTargetId="figure@126.[194,1395,847,1755]" captionTargetPageId="126" captionText="Fig. 80. Cissidium steeli sp. nov. A. Habitus. B. Pronotum, × 680. C. Mesoventrite showing median process of collar, mid-keel and keel, × 785." figureDoi="http://doi.org/10.5281/zenodo.3777444" httpUri="https://zenodo.org/record/3777444/files/figure.png" pageId="125" pageNumber="126">Fig. 80B</figureCitation>
).
</paragraph>
<paragraph id="8B8F36B7386FFFB029F4F83FFA96FE17" blockId="125.[189,1173,2000,2027]" lastBlockId="126.[189,1399,267,399]" lastPageId="126" lastPageNumber="127" pageId="125" pageNumber="126">
ELYTRA.
<quantity id="4CC89B52386FFFB32869F83EFE15F873" box="[288,393,2000,2027]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="125" pageNumber="126" unit="mm" value="0.45">0.45 mm</quantity>
long,
<quantity id="4CC89B52386FFFB3289BF83EFDA7F873" box="[466,571,2000,2027]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="125" pageNumber="126" unit="mm" value="0.35">0.35 mm</quantity>
wide, pubescent as pronotum, shallowly foveolate. MESOVENTRITE. Medial extension of collar tapering posteriorly then parallel-sided; mid-keel broadest anteriorly then slightly concave to rounded posterior angles; keel with ± eight setae, long, parallel to pointed termination posterior to the interruption point of the mesocoxal margins; mesoventral lateral margins straight to rounded rectangular angles, without serrations; humeri toothed (
<figureCitation id="130B2A32386CFFB02DD8FE9BFB61FE17" box="[1169,1277,373,399]" captionStart="Fig" captionStartId="126.[189,232,1797,1823]" captionTargetBox="[195,1392,848,1755]" captionTargetId="figure@126.[194,1395,847,1755]" captionTargetPageId="126" captionText="Fig. 80. Cissidium steeli sp. nov. A. Habitus. B. Pronotum, × 680. C. Mesoventrite showing median process of collar, mid-keel and keel, × 785." figureDoi="http://doi.org/10.5281/zenodo.3777444" httpUri="https://zenodo.org/record/3777444/files/figure.png" pageId="126" pageNumber="127">Fig. 80C</figureCitation>
).
</paragraph>
<paragraph id="8B8F36B7386CFFB029F4FE59FD8AFE6C" blockId="126.[189,1398,438,500]" pageId="126" pageNumber="127">
METAVENTRITE. Sparsely pubescent, length
<quantity id="4CC89B52386CFFB02BE3FE58FC89FE49" box="[682,789,438,465]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="126" pageNumber="127" unit="mm" value="0.15">0.15 mm</quantity>
, disc simple, width across spines
<quantity id="4CC89B52386CFFB02DD0FE58FA9FFE49" box="[1177,1283,438,465]" metricMagnitude="-4" metricUnit="m" metricValue="1.2" pageId="126" pageNumber="127" unit="mm" value="0.12">0.12 mm</quantity>
; posterior margin of mesocoxae serrate.
</paragraph>
<paragraph id="8B8F36B7386CFFB029F4FDF2FE5EFDAE" blockId="126.[189,450,540,566]" box="[189,450,540,566]" pageId="126" pageNumber="127">WINGS. Macropterous.</paragraph>
<paragraph id="8B8F36B7386CFFB029F4FDB3FCF1FDEE" blockId="126.[189,877,604,631]" box="[189,877,604,631]" pageId="126" pageNumber="127">GENITALIA. Female spermatheca globular. Male not known.</paragraph>
</subSubSection>
<subSubSection id="C32A653C386CFFB029F4FD70FC79FD7E" box="[189,997,670,743]" pageId="126" pageNumber="127" type="discussion">
<paragraph id="8B8F36B7386CFFB029F4FD70FEB3FD20" blockId="126.[189,303,670,696]" box="[189,303,670,696]" pageId="126" pageNumber="127">
<heading id="D0C781DB386CFFB029F4FD70FEB3FD20" bold="true" box="[189,303,670,696]" fontSize="11" level="3" pageId="126" pageNumber="127" reason="3">
<emphasis id="B944EAA5386CFFB029F4FD70FEB3FD20" bold="true" box="[189,303,670,696]" pageId="126" pageNumber="127">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8B8F36B7386CFFB029F4FD22FC79FD7E" blockId="126.[189,997,716,743]" box="[189,997,716,743]" pageId="126" pageNumber="127">
The only species of
<taxonomicName id="4C304D34386CFFB028E0FD22FD82FD7E" box="[425,542,716,742]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="126" pageNumber="127" phylum="Arthropoda" rank="genus">
<emphasis id="B944EAA5386CFFB028E0FD22FD82FD7E" box="[425,542,716,742]" italics="true" pageId="126" pageNumber="127">Cissidium</emphasis>
</taxonomicName>
to have been described from
<collectingCountry id="F3277627386CFFB02A30FD22FC7DFD7E" box="[889,993,716,742]" name="Trinidad and Tobago" pageId="126" pageNumber="127">Trinidad</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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