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<document id="0201F14BBD1D71AC5391DD3FCC4262F4" ID-CLB-Dataset="22458" ID-DOI="10.5852/ejt.2020.622" ID-GBIF-Dataset="9cb3ea96-0ee4-40b4-8006-19b5fed80b46" ID-Zenodo-Dep="3777282" ID-ZooBank="9CD3B2CD-F072-4994-8CA9-24145D343401" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1588217079841" checkinUser="valdenar" docAuthor="Darby, Michael" docDate="2020" docId="039987A13847FF982B03FA85FEFEFC09" docLanguage="en" docName="ejt-622_darby.pdf.imf" docOrigin="European Journal of Taxonomy 622" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Cissidium marshallae Darby 2019" docType="treatment" docVersion="7" lastPageNumber="87" masterDocId="FFA0FFD93812FFCE2949FFEEFF9CFF98" masterDocTitle="A revision of Cissidium Motschulsky (Coleoptera: Ptiliidae) with seventy seven new species" masterLastPageNumber="188" masterPageNumber="1" pageNumber="86" updateTime="1698838121940" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="04B1CBFCED5A9DBF112C8E4DFBA4CC38">A revision of Cissidium Motschulsky (Coleoptera: Ptiliidae) with seventy seven new species</mods:title>
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<mods:namePart id="E4D47E93675201733932DFCD2EFE1D02">Darby, Michael</mods:namePart>
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<mods:title id="4162BAA7B64B6F27C4EDA08746ED3085">European Journal of Taxonomy</mods:title>
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<mods:date id="65F538FDA5356DF797E0E99650F50A91">2020</mods:date>
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<treatment id="039987A13847FF982B03FA85FEFEFC09" ID-DOI="http://doi.org/10.5281/zenodo.3795590" ID-GBIF-Taxon="163877866" ID-Zenodo-Dep="3795590" LSID="urn:lsid:plazi:treatment:039987A13847FF982B03FA85FEFEFC09" httpUri="http://treatment.plazi.org/id/039987A13847FF982B03FA85FEFEFC09" lastPageId="86" lastPageNumber="87" pageId="85" pageNumber="86">
<subSubSection id="C32A653C3847FF9B2B03FA85FC76FA1D" box="[586,1002,1387,1414]" pageId="85" pageNumber="86" type="nomenclature">
<paragraph id="8B8F36B73847FF9B2B03FA85FC76FA1D" blockId="85.[586,1002,1387,1451]" box="[586,1002,1387,1414]" pageId="85" pageNumber="86">
<heading id="D0C781DB3847FF9B2B03FA85FC76FA1D" box="[586,1002,1387,1414]" centered="true" fontSize="11" level="2" pageId="85" pageNumber="86" reason="2">
<taxonomicName id="4C304D343847FF9B2B03FA85FC76FA1D" ID-CoL="8XXQR" authority="Darby, 2019" authorityName="Darby" authorityYear="2019" box="[586,1002,1387,1414]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="marshallae">
<emphasis id="B944EAA53847FF9B2B03FA85FCCDFA1E" bold="true" box="[586,849,1387,1414]" italics="true" pageId="85" pageNumber="86">Cissidium marshallae</emphasis>
<bibRefCitation id="EFA14B463847FF9B2A10FA85FC76FA1D" author="Darby M." box="[857,1002,1387,1413]" pageId="85" pageNumber="86" pagination="1 - 50" refId="ref59424" refString="Darby M. 2019. New Ptiliidae (Coleoptera) from Sarawak in the spirit collection of the Natural History Museum, London. European Journal of Taxonomy 512: 1 - 50. https: // doi. org / 10.5852 / ejt. 2019.512" type="journal article" year="2019">Darby, 2019</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32A653C3847FF9B2BB9FA7EFCDFFA33" box="[752,835,1424,1451]" pageId="85" pageNumber="86" type="description">
<paragraph id="8B8F36B73847FF9B2BB9FA7EFCDFFA33" blockId="85.[586,1002,1387,1451]" box="[752,835,1424,1451]" pageId="85" pageNumber="86">
<figureCitation id="130B2A323847FF9B2BB9FA7EFCDFFA33" box="[752,835,1424,1451]" captionStart="Fig" captionStartId="86.[189,232,1964,1990]" captionTargetBox="[194,1398,1012,1922]" captionTargetId="figure@86.[189,1398,1012,1922]" captionTargetPageId="86" captionText="Fig. 55. Cissidium marshallae Darby, 2019. A. Habitus. B. Pronotum, × 640. C. Mesoventrite showing median process of collar, mid-keel, keel and lateral margin, × 750." figureDoi="http://doi.org/10.5281/zenodo.3777394" httpUri="https://zenodo.org/record/3777394/files/figure.png" pageId="85" pageNumber="86">Fig. 55</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32A653C3847FF9829F4FA3DFCDCFD66" lastPageId="86" lastPageNumber="87" pageId="85" pageNumber="86" type="materials_examined">
<paragraph id="8B8F36B73847FF9B29F4FA3DFE36FA75" blockId="85.[189,426,1491,1517]" box="[189,426,1491,1517]" pageId="85" pageNumber="86">
<heading id="D0C781DB3847FF9B29F4FA3DFE36FA75" bold="true" box="[189,426,1491,1517]" fontSize="11" level="3" pageId="85" pageNumber="86" reason="3">
<emphasis id="B944EAA53847FF9B29F4FA3DFE36FA75" bold="true" box="[189,426,1491,1517]" pageId="85" pageNumber="86">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="3B583CEA3847FF9B29A3F9EDFC0FF9FF" ID-GBIF-Occurrence="2609493702" collectedFrom="alluvial forest litter" collectionCode="BMNH" collectorName="P. M. Hammond &amp; J. E. Marshall leg." country="Malaysia" location="Sarawak" pageId="85" pageNumber="86" specimenCount="1" specimenCount-female="1" stateProvince="Sarawak" typeStatus="holotype">
<paragraph id="8B8F36B73847FF9B29A3F9EDFEC0F985" blockId="85.[189,1398,1539,1639]" box="[234,348,1539,1565]" pageId="85" pageNumber="86">
<emphasis id="B944EAA53847FF9B29A3F9EDFEC0F985" bold="true" box="[234,348,1539,1565]" pageId="85" pageNumber="86">
<typeStatus id="548B88153847FF9B29A3F9EDFEC0F985" box="[234,348,1539,1565]" pageId="85" pageNumber="86" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8B8F36B73847FF9B29F4F9C6FC0FF9FF" blockId="85.[189,1398,1539,1639]" pageId="85" pageNumber="86">
<collectingCountry id="F32776273847FF9B29F4F9C6FECFF9DA" box="[189,339,1576,1602]" name="Malaysia" pageId="85" pageNumber="86">MALAYSIA</collectingCountry>
<specimenCount id="9D36FD3E3847FF9B2838F9C6FE14F9DA" box="[369,392,1576,1602]" pageId="85" pageNumber="86" type="female"></specimenCount>
;
<collectingRegion id="49F4F8553847FF9B28D1F9C6FD9CF9DA" box="[408,512,1576,1602]" country="Malaysia" name="Sarawak" pageId="85" pageNumber="86">Sarawak</collectingRegion>
, 4
<superScript id="7C459BFF3847FF9B2B54F9C8FDB6F9AC" attach="left" box="[541,554,1574,1588]" fontSize="6" pageId="85" pageNumber="86">th</superScript>
division, on Mulu NP, Camp 5; MayAug. 1978;
<collectedFrom id="382C754A3847FF9B2DC0F9C6FAEEF9DA" box="[1161,1394,1576,1602]" pageId="85" pageNumber="86">alluvial forest litter</collectedFrom>
;
<collectorName id="26C553613847FF9B29F4F9A3FEEAF9FF" box="[189,374,1613,1639]" pageId="85" pageNumber="86">P.M. Hammond</collectorName>
and
<collectorName id="26C553613847FF9B28E6F9A3FD1CF9FF" box="[431,640,1613,1639]" pageId="85" pageNumber="86">J.E. Marshall leg.</collectorName>
; BM 1978-49;
<collectionCode id="ED21AE723847FF9B2A7AF9A3FC13F9FF" box="[819,911,1613,1639]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="85" pageNumber="86">BMNH</collectionCode>
.
</paragraph>
</materialsCitation>
<paragraph id="8B8F36B73847FF9B29A3F97EFEC0F932" blockId="85.[189,1369,1680,1743]" box="[234,348,1680,1706]" pageId="85" pageNumber="86">
<materialsCitation id="3B583CEA3847FF9B29A3F97EFEC0F932" ID-GBIF-Occurrence="2609493716" box="[234,348,1680,1706]" collectedFrom="alluvial forest litter" collectionCode="BMNH" collectorName="Mulu NP &amp; Camp &amp; May-Aug. &amp; P. M. Hammond &amp; J. E. Marshall" country="Malaysia" location="MALAYSIA" pageId="85" pageNumber="86" specimenCount="1" stateProvince="Sarawak" typeStatus="paratype">
<emphasis id="B944EAA53847FF9B29A3F97EFEC0F932" bold="true" box="[234,348,1680,1706]" pageId="85" pageNumber="86">
<typeStatus id="548B88153847FF9B29A3F97EFEC0F932" box="[234,348,1680,1706]" pageId="85" pageNumber="86" type="paratype">Paratype</typeStatus>
</emphasis>
</materialsCitation>
</paragraph>
<paragraph id="8B8F36B73847FF9B29F4F95BFAC5F957" blockId="85.[189,1369,1680,1743]" box="[189,1369,1716,1743]" pageId="85" pageNumber="86">
<collectingCountry id="F32776273847FF9B29F4F95BFECFF957" box="[189,339,1717,1743]" name="Malaysia" pageId="85" pageNumber="86">MALAYSIA</collectingCountry>
<specimenCount id="9D36FD3E3847FF9B2825F95BFE34F957" box="[364,424,1717,1743]" pageId="85" pageNumber="86" type="generic">1 ex.</specimenCount>
; same collection data as for holotype; mounted as a disarticulated slide;
<collectionCode id="ED21AE723847FF9B2DB3F95BFACAF957" box="[1274,1366,1717,1743]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="85" pageNumber="86">BMNH</collectionCode>
.
</paragraph>
<paragraph id="8B8F36B73847FF9B29F4F919FD8DF889" blockId="85.[189,529,1783,1809]" box="[189,529,1783,1809]" pageId="85" pageNumber="86">
<heading id="D0C781DB3847FF9B29F4F919FD8DF889" bold="true" box="[189,529,1783,1809]" fontSize="11" level="3" pageId="85" pageNumber="86" reason="3">
<emphasis id="B944EAA53847FF9B29F4F919FD8DF889" bold="true" box="[189,529,1783,1809]" pageId="85" pageNumber="86">Supplementary description</emphasis>
</heading>
</paragraph>
<paragraph id="8B8F36B73847FF9B29F4F8C6FA82F8FF" blockId="85.[189,1398,1832,1895]" pageId="85" pageNumber="86">
The following description amplifies the
<typeStatus id="548B88153847FF9B2BDCF8C6FD54F8DA" box="[661,712,1832,1858]" pageId="85" pageNumber="86">type</typeStatus>
description of
<bibRefCitation id="EFA14B463847FF9B2A30F8C6FB86F8DA" author="Darby M." box="[889,1050,1832,1858]" pageId="85" pageNumber="86" pagination="1 - 50" refId="ref59424" refString="Darby M. 2019. New Ptiliidae (Coleoptera) from Sarawak in the spirit collection of the Natural History Museum, London. European Journal of Taxonomy 512: 1 - 50. https: // doi. org / 10.5852 / ejt. 2019.512" type="journal article" year="2019">Darby (2019)</bibRefCitation>
, in particular by adding more details of the meso- and metaventral characters to conform with the entries in the present paper.
</paragraph>
<paragraph id="8B8F36B73847FF9B29F4F87EFD36F832" blockId="85.[189,682,1935,1962]" box="[189,682,1935,1962]" pageId="85" pageNumber="86">
SIZE. Habitus (
<figureCitation id="130B2A323847FF9B282EF861FE48F832" box="[359,468,1935,1962]" captionStart="Fig" captionStartId="86.[189,232,1964,1990]" captionTargetBox="[194,1398,1012,1922]" captionTargetId="figure@86.[189,1398,1012,1922]" captionTargetPageId="86" captionText="Fig. 55. Cissidium marshallae Darby, 2019. A. Habitus. B. Pronotum, × 640. C. Mesoventrite showing median process of collar, mid-keel, keel and lateral margin, × 750." figureDoi="http://doi.org/10.5281/zenodo.3777394" httpUri="https://zenodo.org/record/3777394/files/figure.png" pageId="85" pageNumber="86">Fig. 55A</figureCitation>
), length
<quantity id="4CC89B523847FF9B2B73F861FD39F832" box="[570,677,1935,1962]" metricMagnitude="-4" metricUnit="m" metricValue="5.8" pageId="85" pageNumber="86" unit="mm" value="0.58">0.58 mm</quantity>
.
</paragraph>
<paragraph id="8B8F36B73847FF9829F4F83DFAAEFEBD" blockId="85.[189,901,2002,2029]" lastBlockId="86.[189,1330,267,294]" lastPageId="86" lastPageNumber="87" pageId="85" pageNumber="86">
COLOUR. Dark brown, antennae, legs and pubescence yellow. HEAD. Distance across eyes
<quantity id="4CC89B523844FF982B4CFEE5FDEDFEBE" box="[517,625,267,294]" metricMagnitude="-4" metricUnit="m" metricValue="1.9" pageId="86" pageNumber="87" unit="mm" value="0.19">0.19 mm</quantity>
; antennomeres IIIXI, length
<quantity id="4CC89B523844FF982A98FEE5FBA1FEBE" box="[977,1085,267,294]" metricMagnitude="-4" metricUnit="m" metricValue="2.9" pageId="86" pageNumber="87" unit="mm" value="0.29">0.29 mm</quantity>
; mentum as
<figureCitation id="130B2A323844FF982D99FEE5FAAEFEBD" box="[1232,1330,267,293]" captionStart="Fig" captionStartId="5.[189,232,1927,1953]" captionTargetBox="[232,1334,1274,1884]" captionTargetId="figure@5.[232,1334,1274,1887]" captionTargetPageId="5" captionText="Fig. 4. Menta and submenta. A. Cissidium petri Darby, 2015, × 1380. B. Cissidium tigrum Darby, 2015, × 1220. C. Cissidium amboroense Darby, 2015, × 1260. D. Cissidium losbanos sp. nov., × 1100. E. Cissidium triangulum Darby, 2019, × 650. F. Cissidium pilosellum sp. nov., × 1420." figureDoi="http://doi.org/10.5281/zenodo.3777290" httpUri="https://zenodo.org/record/3777290/files/figure.png" pageId="86" pageNumber="87">Fig. 4A.</figureCitation>
</paragraph>
<paragraph id="8B8F36B73844FF9829F4FEA3FDD2FE49" blockId="86.[189,1398,333,465]" pageId="86" pageNumber="87">
PRONOTUM. Length
<quantity id="4CC89B523844FF9828D1FEA3FD9DFEFF" box="[408,513,333,359]" metricMagnitude="-4" metricUnit="m" metricValue="1.6" pageId="86" pageNumber="87" unit="mm" value="0.16">0.16 mm</quantity>
, width
<quantity id="4CC89B523844FF982B18FEA3FD25FEFF" box="[593,697,333,359]" metricMagnitude="-4" metricUnit="m" metricValue="2.6" pageId="86" pageNumber="87" unit="mm" value="0.26">0.26 mm</quantity>
, pubescent and foveolate throughout, foveolae forming short, obscure, longitudinal depressions at base; side margins sharply angled, thence concave to rectangular hind angles, bordered, the border continuing along the posterior margin to the medial emargination opposite the scutellum (
<figureCitation id="130B2A323844FF98289CFE58FDDDFE49" box="[469,577,438,465]" captionStart="Fig" captionStartId="86.[189,232,1964,1990]" captionTargetBox="[194,1398,1012,1922]" captionTargetId="figure@86.[189,1398,1012,1922]" captionTargetPageId="86" captionText="Fig. 55. Cissidium marshallae Darby, 2019. A. Habitus. B. Pronotum, × 640. C. Mesoventrite showing median process of collar, mid-keel, keel and lateral margin, × 750." figureDoi="http://doi.org/10.5281/zenodo.3777394" httpUri="https://zenodo.org/record/3777394/files/figure.png" pageId="86" pageNumber="87">Fig. 55B</figureCitation>
).
</paragraph>
<paragraph id="8B8F36B73844FF9829F4FE16FC24FD8A" blockId="86.[189,952,504,531]" box="[189,952,504,531]" pageId="86" pageNumber="87">
ELYTRA. Length
<quantity id="4CC89B523844FF982833FE16FE7AFD8A" box="[378,486,504,530]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="86" pageNumber="87" unit="mm" value="0.4">0.40 mm</quantity>
, width
<quantity id="4CC89B523844FF982B72FE16FD3BFD8A" box="[571,679,504,531]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="86" pageNumber="87" unit="mm" value="0.35">0.35 mm</quantity>
, pubescent throughout.
</paragraph>
<paragraph id="8B8F36B73844FF9829F4FDD4FB1BFD26" blockId="86.[189,1399,569,702]" pageId="86" pageNumber="87">
MESOVENTRITE. Medial extension of collar narrow, parallel-sided, slightly widened posteriorly; midkeel parallel-sided, hind angles not reaching mesocoxal anterior borders; keel foveolate with ± seven setae, broad basally, tapering to parallel-sided with rounded termination past interruption of mesocoxal borders; mesoventral lateral margins with ¾ serrations; humeri toothed (
<figureCitation id="130B2A323844FF982D47FD4DFBE6FD26" box="[1038,1146,675,702]" captionStart="Fig" captionStartId="86.[189,232,1964,1990]" captionTargetBox="[194,1398,1012,1922]" captionTargetId="figure@86.[189,1398,1012,1922]" captionTargetPageId="86" captionText="Fig. 55. Cissidium marshallae Darby, 2019. A. Habitus. B. Pronotum, × 640. C. Mesoventrite showing median process of collar, mid-keel, keel and lateral margin, × 750." figureDoi="http://doi.org/10.5281/zenodo.3777394" httpUri="https://zenodo.org/record/3777394/files/figure.png" pageId="86" pageNumber="87">Fig. 55C</figureCitation>
).
</paragraph>
<paragraph id="8B8F36B73844FF9829F4FD0BFCDCFD66" blockId="86.[189,832,740,767]" box="[189,832,740,767]" pageId="86" pageNumber="87">METAVENTRITE. With scattered pubescence, disc simple.</paragraph>
</subSubSection>
<subSubSection id="C32A653C3844FF9829F4FCC8FEFEFC09" pageId="86" pageNumber="87" type="discussion">
<paragraph id="8B8F36B73844FF9829F4FCC8FEB3FCD8" blockId="86.[189,303,806,832]" box="[189,303,806,832]" pageId="86" pageNumber="87">
<heading id="D0C781DB3844FF9829F4FCC8FEB3FCD8" bold="true" box="[189,303,806,832]" fontSize="11" level="3" pageId="86" pageNumber="87" reason="3">
<emphasis id="B944EAA53844FF9829F4FCC8FEB3FCD8" bold="true" box="[189,303,806,832]" pageId="86" pageNumber="87">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8B8F36B73844FF9829F4FCBAFEFEFC09" blockId="86.[189,1399,852,913]" pageId="86" pageNumber="87">Distinguished from the other four Peruvian species in this group by the pair of linear depressions at the pronotal base.</paragraph>
</subSubSection>
</treatment>
</document>