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<document id="38E34F16AD658764196B81ADD248FB43" ID-CLB-Dataset="22458" ID-DOI="10.5852/ejt.2020.622" ID-GBIF-Dataset="9cb3ea96-0ee4-40b4-8006-19b5fed80b46" ID-Zenodo-Dep="3777282" ID-ZooBank="9CD3B2CD-F072-4994-8CA9-24145D343401" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1588217079841" checkinUser="valdenar" docAuthor="Darby, Michael" docDate="2020" docId="039987A13843FF9C2B28FC88FCBCFE20" docLanguage="en" docName="ejt-622_darby.pdf.imf" docOrigin="European Journal of Taxonomy 622" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Cissidium latum Sawada 2008" docType="treatment" docVersion="7" lastPageNumber="83" masterDocId="FFA0FFD93812FFCE2949FFEEFF9CFF98" masterDocTitle="A revision of Cissidium Motschulsky (Coleoptera: Ptiliidae) with seventy seven new species" masterLastPageNumber="188" masterPageNumber="1" pageNumber="82" updateTime="1698838121940" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="EBB2E6089335A6E8985FEB6F5E7A2617">A revision of Cissidium Motschulsky (Coleoptera: Ptiliidae) with seventy seven new species</mods:title>
</mods:titleInfo>
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<mods:roleTerm id="F5AD5DCD787502C596E316DA7CA53970">Author</mods:roleTerm>
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<mods:namePart id="B251656415299A3858BB323247E3DDC4">Darby, Michael</mods:namePart>
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<mods:title id="25E011C5CA54993B0A98BB1584629F24">European Journal of Taxonomy</mods:title>
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<mods:date id="3BDF14518712C23152A51E980F110CE5">2020</mods:date>
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<mods:number id="C29F3AD7906DC8E6AE006BDA442FF4FD">2020-04-02</mods:number>
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<mods:number id="ECE398743F2B62B2B4BA3174EA8D5F50">622</mods:number>
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<mods:classification id="CDDFC97FA0BBFD0DD67719C31C373829">journal article</mods:classification>
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<mods:identifier id="A2DDE4BA27D901C74ABBF4C39145DF86" type="DOI">10.5852/ejt.2020.622</mods:identifier>
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<treatment id="039987A13843FF9C2B28FC88FCBCFE20" ID-DOI="http://doi.org/10.5281/zenodo.3795584" ID-GBIF-Taxon="163877863" ID-Zenodo-Dep="3795584" LSID="urn:lsid:plazi:treatment:039987A13843FF9C2B28FC88FCBCFE20" httpUri="http://treatment.plazi.org/id/039987A13843FF9C2B28FC88FCBCFE20" lastPageId="82" lastPageNumber="83" pageId="81" pageNumber="82">
<subSubSection id="C32A653C3843FF9F2B28FC88FC4FFC18" box="[609,979,870,897]" pageId="81" pageNumber="82" type="nomenclature">
<paragraph id="8B8F36B73843FF9F2B28FC88FC4FFC18" blockId="81.[609,979,870,935]" box="[609,979,870,897]" pageId="81" pageNumber="82">
<heading id="D0C781DB3843FF9F2B28FC88FC4FFC18" box="[609,979,870,897]" centered="true" fontSize="11" level="2" pageId="81" pageNumber="82" reason="2">
<taxonomicName id="4C304D343843FF9F2B28FC88FC4FFC18" ID-CoL="5Z94B" authority="Sawada, 2008" authorityName="Sawada" authorityYear="2008" box="[609,979,870,897]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="latum">
<emphasis id="B944EAA53843FF9F2B28FC88FCBBFC19" bold="true" box="[609,807,870,897]" italics="true" pageId="81" pageNumber="82">Cissidium latum</emphasis>
<bibRefCitation id="EFA14B463843FF9F2A67FC88FC4FFC18" author="Sawada Y." box="[814,979,870,896]" pageId="81" pageNumber="82" pagination="101 - 126" refId="ref59928" refString="Sawada Y. 2008. A Revision of the Genus Cissidium (Coleoptera: Ptiliidae) in Japan. Special Publication 2: 101 - 126. Japan Coelopterological Society, Osaka." type="journal article" year="2008">Sawada, 2008</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32A653C3843FF9F2BB9FC62FCDFFC3F" box="[752,835,908,935]" pageId="81" pageNumber="82" type="description">
<paragraph id="8B8F36B73843FF9F2BB9FC62FCDFFC3F" blockId="81.[609,979,870,935]" box="[752,835,908,935]" pageId="81" pageNumber="82">
<figureCitation id="130B2A323843FF9F2BB9FC62FCDFFC3F" box="[752,835,908,935]" captionStart="Fig" captionStartId="82.[189,232,1423,1449]" captionTargetBox="[191,1398,517,1380]" captionTargetId="figure@82.[189,1398,516,1380]" captionTargetPageId="82" captionText="Fig. 52. Cissidium latum Sawada, 2008. A. Habitus. B Pronotum, × 620. C. Mesoventrite showing median process of collar, mid-keel, keel and lateral margin, × 800." figureDoi="http://doi.org/10.5281/zenodo.3777388" httpUri="https://zenodo.org/record/3777388/files/figure.png" pageId="81" pageNumber="82">Fig. 52</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32A653C3843FF9C29F4FC3EFC6BFEBD" lastPageId="82" lastPageNumber="83" pageId="81" pageNumber="82" type="materials_examined">
<paragraph id="8B8F36B73843FF9F29F4FC3EFE36FC72" blockId="81.[189,426,976,1002]" box="[189,426,976,1002]" pageId="81" pageNumber="82">
<heading id="D0C781DB3843FF9F29F4FC3EFE36FC72" bold="true" box="[189,426,976,1002]" fontSize="11" level="3" pageId="81" pageNumber="82" reason="3">
<emphasis id="B944EAA53843FF9F29F4FC3EFE36FC72" bold="true" box="[189,426,976,1002]" pageId="81" pageNumber="82">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8B8F36B73843FF9F29A3FBECFEFBFB84" blockId="81.[189,1398,1026,1128]" box="[234,359,1026,1052]" pageId="81" pageNumber="82">
<materialsCitation id="3B583CEA3843FF9F29A3FBECFEFBFB84" ID-GBIF-Occurrence="2609493638" box="[234,359,1026,1052]" collectingDate="1987-10-21" collectionCode="BMNH" collectorName="Y. Nishikawa" country="Japan" elevation="290" location="Gaji" municipality="Ryukyu" pageId="81" pageNumber="82" specimenCount="1" specimenCount-male="1" stateProvince="Okinawa" typeStatus="paratype">
<emphasis id="B944EAA53843FF9F29A3FBECFEFBFB84" bold="true" box="[234,359,1026,1052]" pageId="81" pageNumber="82">
<typeStatus id="548B88153843FF9F29A3FBECFEFBFB84" box="[234,359,1026,1052]" pageId="81" pageNumber="82" type="paratype">Paratypes</typeStatus>
</emphasis>
</materialsCitation>
</paragraph>
<paragraph id="8B8F36B73843FF9F29F4FBC6FB7CFBF0" blockId="81.[189,1398,1026,1128]" pageId="81" pageNumber="82">
<collectingCountry id="F32776273843FF9F29F4FBC6FE8AFBDA" box="[189,278,1064,1090]" name="Japan" pageId="81" pageNumber="82">JAPAN</collectingCountry>
•
<specimenCount id="9D36FD3E3843FF9F2863FBC6FECAFBDA" box="[298,342,1064,1090]" pageId="81" pageNumber="82" type="male">1 ♂</specimenCount>
;
<collectingRegion id="49F4F8553843FF9F2828FBC6FE52FBDA" box="[353,462,1064,1090]" country="Japan" name="Okinawa" pageId="81" pageNumber="82">Okinawa</collectingRegion>
,
<collectingMunicipality id="6BEBACCD3843FF9F2891FBC6FDABFBDA" box="[472,567,1064,1090]" pageId="81" pageNumber="82">Ryukyu</collectingMunicipality>
,
<location id="8EEF606C3843FF9F2B09FBC6FDEFFBDA" LSID="urn:lsid:plazi:treatment:039987A13843FF9C2B28FC88FCBCFE20:8EEF606C3843FF9F2B09FBC6FDEFFBDA" box="[576,627,1064,1090]" country="Japan" municipality="Ryukyu" name="Gaji" pageId="81" pageNumber="82" stateProvince="Okinawa">Gaji</location>
rindo,
<locationDeviation id="9A3D2F953843FF9F2B8BFBC6FC3CFBDA" box="[706,928,1064,1090]" location="Mt Terukubi" pageId="81" pageNumber="82">NE of Mt Terukubi</locationDeviation>
;
<elevation id="001DD1843843FF9F2AE3FBC6FBB3FBDA" box="[938,1071,1064,1090]" metricMagnitude="2" metricUnit="m" metricValue="2.9" pageId="81" pageNumber="82" unit="m" value="290.0">
<quantity id="4CC89B523843FF9F2AE3FBC6FC6EFBDA" box="[938,1010,1064,1090]" metricMagnitude="2" metricUnit="m" metricValue="2.9" pageId="81" pageNumber="82" unit="m" value="290.0">290 m</quantity>
a.s.l.
</elevation>
;
<date id="FF8E10773843FF9F2D72FBC6FB4EFBDA" box="[1083,1234,1064,1090]" pageId="81" pageNumber="82" value="1987-10-21">
<collectingDate id="EFCAE99F3843FF9F2D72FBC6FB4EFBDA" box="[1083,1234,1064,1090]" pageId="81" pageNumber="82" value="1987-10-21">21 Oct. 1987</collectingDate>
</date>
;
<collectorName id="26C553613843FF9F2D92FBC6FAEAFBDA" box="[1243,1398,1064,1090]" pageId="81" pageNumber="82">Y. Nishikawa</collectorName>
leg.;
<collectionCode id="ED21AE723843FF9F29B1FBA0FECCFBF0" box="[248,336,1102,1128]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="81" pageNumber="82">BMNH</collectionCode>
•
<materialsCitation id="3B583CEA3843FF9F2820FBA0FB40FBF0" ID-GBIF-Occurrence="2609493535" box="[361,1244,1102,1128]" collectingDate="1997-04-12" collectionCode="YS" collectorName="Y. Sawada" country="Japan" location="Kunigami-son" municipality="Hiji" pageId="81" pageNumber="82" specimenCount="1" specimenCount-male="1" stateProvince="Okinawa Pref." typeStatus="paratype">
<specimenCount id="9D36FD3E3843FF9F2820FBA0FE0BFBF0" box="[361,407,1102,1128]" pageId="81" pageNumber="82" type="male">1 ♂</specimenCount>
;
<collectingMunicipality id="6BEBACCD3843FF9F28EDFBA0FE49FBF0" box="[420,469,1102,1128]" pageId="81" pageNumber="82">Hiji</collectingMunicipality>
,
<location id="8EEF606C3843FF9F28A9FBA0FD17FBF0" LSID="urn:lsid:plazi:treatment:039987A13843FF9C2B28FC88FCBCFE20:8EEF606C3843FF9F28A9FBA0FD17FBF0" box="[480,651,1102,1128]" country="Japan" municipality="Hiji" name="Kunigami-son" pageId="81" pageNumber="82" stateProvince="Okinawa Pref.">Kunigami-son</location>
,
<collectingRegion id="49F4F8553843FF9F2BDEFBA0FCD8FBF0" box="[663,836,1102,1128]" country="Japan" name="Okinawa" pageId="81" pageNumber="82">Okinawa Pref.</collectingRegion>
;
<date id="FF8E10773843FF9F2A19FBA0FC77FBF0" box="[848,1003,1102,1128]" pageId="81" pageNumber="82" value="1997-04-12">
<collectingDate id="EFCAE99F3843FF9F2A19FBA0FC77FBF0" box="[848,1003,1102,1128]" pageId="81" pageNumber="82" value="1997-04-12">12 Apr. 1997</collectingDate>
</date>
;
<collectorName id="26C553613843FF9F2ABEFBA0FBEEFBF0" box="[1015,1138,1102,1128]" pageId="81" pageNumber="82">Y. Sawada</collectorName>
leg.;
<collectionCode id="ED21AE723843FF9F2DFAFBA0FB40FBF0" box="[1203,1244,1102,1128]" pageId="81" pageNumber="82">YS</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8B8F36B73843FF9F29F4FB7CFD8DFB34" blockId="81.[189,529,1170,1196]" box="[189,529,1170,1196]" pageId="81" pageNumber="82">
<heading id="D0C781DB3843FF9F29F4FB7CFD8DFB34" bold="true" box="[189,529,1170,1196]" fontSize="11" level="3" pageId="81" pageNumber="82" reason="3">
<emphasis id="B944EAA53843FF9F29F4FB7CFD8DFB34" bold="true" box="[189,529,1170,1196]" pageId="81" pageNumber="82">Supplementary description</emphasis>
</heading>
</paragraph>
<paragraph id="8B8F36B73843FF9F29F4FB2DFE36FA9C" blockId="81.[189,1398,1219,1284]" pageId="81" pageNumber="82">
The following description amplifies the
<typeStatus id="548B88153843FF9F2BDEFB2AFD56FB46" box="[663,714,1220,1246]" pageId="81" pageNumber="82">type</typeStatus>
description of
<bibRefCitation id="EFA14B463843FF9F2A35FB2DFBB1FB45" author="Sawada Y." box="[892,1069,1219,1245]" pageId="81" pageNumber="82" pagination="101 - 126" refId="ref59928" refString="Sawada Y. 2008. A Revision of the Genus Cissidium (Coleoptera: Ptiliidae) in Japan. Special Publication 2: 101 - 126. Japan Coelopterological Society, Osaka." type="journal article" year="2008">Sawada (2008)</bibRefCitation>
to conform with the entries in the present paper.
</paragraph>
<paragraph id="8B8F36B73843FF9F29F4FAC0FD36FAD0" blockId="81.[189,682,1325,1352]" box="[189,682,1325,1352]" pageId="81" pageNumber="82">
SIZE. Habitus (
<figureCitation id="130B2A323843FF9F282EFAC3FE48FAD0" box="[359,468,1325,1352]" captionStart="Fig" captionStartId="82.[189,232,1423,1449]" captionTargetBox="[191,1398,517,1380]" captionTargetId="figure@82.[189,1398,516,1380]" captionTargetPageId="82" captionText="Fig. 52. Cissidium latum Sawada, 2008. A. Habitus. B Pronotum, × 620. C. Mesoventrite showing median process of collar, mid-keel, keel and lateral margin, × 800." figureDoi="http://doi.org/10.5281/zenodo.3777388" httpUri="https://zenodo.org/record/3777388/files/figure.png" pageId="81" pageNumber="82">Fig. 52A</figureCitation>
), length
<quantity id="4CC89B523843FF9F2B73FAC3FD39FAD0" box="[570,677,1325,1352]" metricMagnitude="-4" metricUnit="m" metricValue="7.6" pageId="81" pageNumber="82" unit="mm" value="0.76">0.76 mm</quantity>
.
</paragraph>
<paragraph id="8B8F36B73843FF9F29F4FA9CFBDEFA14" blockId="81.[189,1090,1394,1420]" box="[189,1090,1394,1420]" pageId="81" pageNumber="82">COLOUR. Reddish brown to dark brown, antennae, legs and pubescence paler.</paragraph>
<paragraph id="8B8F36B73843FF9F29F4FA58FBCAFA48" blockId="81.[189,1110,1462,1488]" box="[189,1110,1462,1488]" pageId="81" pageNumber="82">HEAD. With a transverse row of indistinctly formed foveolae between the eyes.</paragraph>
<paragraph id="8B8F36B73843FF9F29F4FA15FE24F91F" blockId="81.[189,1399,1530,1671]" pageId="81" pageNumber="82">
PRONOTUM. Length
<quantity id="4CC89B523843FF9F28F8FA14FDBAF98D" box="[433,550,1530,1557]" metricMagnitude="-4" metricUnit="m" metricValue="2.1" pageId="81" pageNumber="82" unit="mm" value="0.21">0.21 mm</quantity>
, width
<quantity id="4CC89B523843FF9F2BC7FA14FC9EF98D" box="[654,770,1530,1557]" metricMagnitude="-4" metricUnit="m" metricValue="3.7" pageId="81" pageNumber="82" unit="mm" value="0.37">0.37 mm</quantity>
, densely pubescent and very shallowly foveolate throughout; side margins rounded to concave before rectangular hind angles, narrowly bordered, the border continuing along the entire posterior margin, which is without an emargination opposite the scutellum (
<figureCitation id="130B2A323843FF9F2876F982FE37F91F" box="[319,427,1644,1671]" captionStart="Fig" captionStartId="82.[189,232,1423,1449]" captionTargetBox="[191,1398,517,1380]" captionTargetId="figure@82.[189,1398,516,1380]" captionTargetPageId="82" captionText="Fig. 52. Cissidium latum Sawada, 2008. A. Habitus. B Pronotum, × 620. C. Mesoventrite showing median process of collar, mid-keel, keel and lateral margin, × 800." figureDoi="http://doi.org/10.5281/zenodo.3777388" httpUri="https://zenodo.org/record/3777388/files/figure.png" pageId="81" pageNumber="82">Fig. 52B</figureCitation>
).
</paragraph>
<paragraph id="8B8F36B73843FF9F29F4F95FFA9FF953" blockId="81.[189,1283,1712,1739]" box="[189,1283,1712,1739]" pageId="81" pageNumber="82">
ELYTRA. Length
<quantity id="4CC89B523843FF9F2833F95EFE7AF953" box="[378,486,1712,1739]" metricMagnitude="-4" metricUnit="m" metricValue="4.699999999999999" pageId="81" pageNumber="82" unit="mm" value="0.47">0.47 mm</quantity>
, width
<quantity id="4CC89B523843FF9F2B72F95EFD3BF953" box="[571,679,1712,1739]" metricMagnitude="-4" metricUnit="m" metricValue="4.2" pageId="81" pageNumber="82" unit="mm" value="0.42">0.42 mm</quantity>
, pubescent more widely distributed than pronotum.
</paragraph>
<paragraph id="8B8F36B73843FF9F29F4F91BFADAF819" blockId="81.[189,1399,1780,1921]" pageId="81" pageNumber="82">
MESOVENTRITE. Collar with 3 projecting setae along the posterior margin opposite the mid-keel, median extension widest anteriorly then slightly tapering posteriorly, with very short setae; mid-keel strongly sculpted and pubescent, widest medially; keel long, pointed, terminating past the mid-point of the mesocoxae; lateral margins very shallowly rounded, without serrations; humeri toothed (
<figureCitation id="130B2A323843FF9F2D84F888FAA6F819" box="[1229,1338,1894,1921]" captionStart="Fig" captionStartId="82.[189,232,1423,1449]" captionTargetBox="[191,1398,517,1380]" captionTargetId="figure@82.[189,1398,516,1380]" captionTargetPageId="82" captionText="Fig. 52. Cissidium latum Sawada, 2008. A. Habitus. B Pronotum, × 620. C. Mesoventrite showing median process of collar, mid-keel, keel and lateral margin, × 800." figureDoi="http://doi.org/10.5281/zenodo.3777388" httpUri="https://zenodo.org/record/3777388/files/figure.png" pageId="81" pageNumber="82">Fig. 52C</figureCitation>
).
</paragraph>
<paragraph id="8B8F36B73843FF9F29F4F845FB9CF873" blockId="81.[189,1398,1962,2027]" pageId="81" pageNumber="82">
METAVENTRITE.
<quantity id="4CC89B523843FF9F283DF844FE78F85D" box="[372,484,1962,1989]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="81" pageNumber="82" unit="mm" value="0.15">0.15 mm</quantity>
long, reticulate towards lateral margins, sparsely pubescent, disc simple, distance between spines
<quantity id="4CC89B523843FF9F2897F83FFDD6F873" box="[478,586,2001,2027]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="81" pageNumber="82" unit="mm" value="0.1">0.10 mm</quantity>
; mesocoxal posterior borders serrate.
</paragraph>
<paragraph id="8B8F36B73840FF9C29F4FEE5FC6BFEBD" blockId="82.[189,1015,267,294]" box="[189,1015,267,294]" pageId="82" pageNumber="83">
GENITALIA. Male aedeagus as
<figureCitation id="130B2A323840FF9C2B5CFEE5FDC4FEBD" box="[533,600,267,293]" captionStart="Fig" captionStartId="7.[189,232,1825,1851]" captionTargetBox="[202,1398,769,1790]" captionTargetId="figure@7.[189,1398,769,1799]" captionTargetPageId="7" captionText="Fig. 6. Figures to show genitalia types. – Spermathecae. A. Cissidium bilineatum sp. nov. B. Cissidium amboroense Darby, 2015. C. Cissidium sawadai Darby, 2015. D. Cissidium arcuatum sp. nov. E. Cissidium murphyi sp. nov. – Aedeagi. F. Cissidium petri Darby, 2015. G. Cissidium clareae sp. nov. H. Cissidium fraternum sp. nov. I. Cissidium tigrum Darby, 2015. J. Cissidium apterum sp. nov. K. Cissidium brachypterum sp. nov. L. Cissidium greensladei sp. nov. M. Cissidium kolombangaricum sp. nov. N. Cissidium mindanao sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3777294" httpUri="https://zenodo.org/record/3777294/files/figure.png" pageId="82" pageNumber="83">Fig. 6</figureCitation>
Fa–b but beak not so strongly bent.
</paragraph>
</subSubSection>
<subSubSection id="C32A653C3840FF9C29F4FEA3FCBCFE20" pageId="82" pageNumber="83" type="discussion">
<paragraph id="8B8F36B73840FF9C29F4FEA3FEB3FEFF" blockId="82.[189,303,333,359]" box="[189,303,333,359]" pageId="82" pageNumber="83">
<heading id="D0C781DB3840FF9C29F4FEA3FEB3FEFF" bold="true" box="[189,303,333,359]" fontSize="11" level="3" pageId="82" pageNumber="83" reason="3">
<emphasis id="B944EAA53840FF9C29F4FEA3FEB3FEFF" bold="true" box="[189,303,333,359]" pageId="82" pageNumber="83">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8B8F36B73840FF9C29F4FE95FCBCFE20" blockId="82.[189,1398,379,441]" pageId="82" pageNumber="83">
This is one of three species in this group from
<collectingCountry id="F32776273840FF9C2BA8FE92FCB8FE0E" box="[737,804,380,406]" name="Japan" pageId="82" pageNumber="83">Japan</collectingCountry>
and differs from them and from all other Japanese
<taxonomicName id="4C304D343840FF9C29F4FE70FECDFE20" authority="in" authorityName="In" box="[189,337,414,440]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B944EAA53840FF9C29F4FE70FEAEFE20" box="[189,306,414,440]" italics="true" pageId="82" pageNumber="83">Cissidium</emphasis>
in
</taxonomicName>
having wider and more rounded elytra.
</paragraph>
</subSubSection>
</treatment>
</document>
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