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<document id="3CCD89D763A9870A76F087A24B848D3B" ID-DOI="10.5281/zenodo.158689" ID-GBIF-Dataset="999b5ab0-d412-44b7-a113-70ddad24a49e" ID-ISSN="1175­5326" ID-Zenodo-Dep="158689" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461024304210" checkinUser="plazi" docAuthor="Caterino, Michael S." docDate="2004" docId="0399879AFF98FFD7FE98FA17FB54009F" docLanguage="en" docName="zt00601.pdf" docOrigin="Zootaxa 601" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Hister laevimargo Lewis" docType="treatment" docVersion="9" lastPageNumber="9" masterDocId="FFA0FFE2FF9EFFDEFF90FFF6FFBA0149" masterDocTitle="Taxonomy of the Hister criticus group, with description of a new species (Coleoptera: Histeridae)" masterLastPageNumber="16" masterPageNumber="1" pageNumber="7" updateTime="1698571502456" updateUser="plazi">
<mods:mods id="FE3C1B0D3047D67DC4F8394718E9532E" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="D3D0BCFECA426C52F4CC6555FF0B6B90">Taxonomy of the Hister criticus group, with description of a new species (Coleoptera: Histeridae)</mods:title>
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<mods:namePart id="CD59482172CE9B8AAFFAC286E60679F1">Caterino, Michael S.</mods:namePart>
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<mods:date id="D29F66A9D95098CFE3D55022E3591A86">2004</mods:date>
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<treatment id="0399879AFF98FFD7FE98FA17FB54009F" ID-DOI="http://doi.org/10.5281/zenodo.6271826" ID-GBIF-Taxon="119605797" ID-Zenodo-Dep="6271826" LSID="urn:lsid:plazi:treatment:0399879AFF98FFD7FE98FA17FB54009F" httpUri="http://treatment.plazi.org/id/0399879AFF98FFD7FE98FA17FB54009F" lastPageId="9" lastPageNumber="9" pageId="6" pageNumber="7">
<subSubSection id="C32A6507FF98FFD8FE98FA17FCBA04B3" box="[264,768,1504,1531]" pageId="6" pageNumber="7" type="nomenclature">
<paragraph id="8B8F368CFF98FFD8FE98FA17FCBA04B3" blockId="6.[264,768,1504,1531]" box="[264,768,1504,1531]" pageId="6" pageNumber="7">
<heading id="D0C781E0FF98FFD8FE98FA17FCBA04B3" bold="true" box="[264,768,1504,1531]" fontSize="11" level="1" pageId="6" pageNumber="7" reason="1">
<emphasis id="B944EA9EFF98FFD8FE98FA17FCBA04B3" bold="true" box="[264,768,1504,1531]" pageId="6" pageNumber="7">
<taxonomicName id="4C304D0FFF98FFD8FE98FA17FD8804B3" ID-CoL="3M57P" authority="Lewis" authorityName="Lewis" box="[264,562,1504,1531]" class="Insecta" family="Histeridae" genus="Hister" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="laevimargo">
<emphasis id="B944EA9EFF98FFD8FE98FA17FE5B04B2" bold="true" box="[264,481,1505,1531]" italics="true" pageId="6" pageNumber="7">Hister laevimargo</emphasis>
Lewis
</taxonomicName>
(
<figureCitation id="130B2A09FF98FFD8FDD1FA16FD2E04B3" box="[577,660,1504,1530]" captionStart="FIGURE 1" captionStartId="1.[264,368,1201,1225]" captionTargetBox="[369,1221,573,1164]" captionTargetId="figure@1.[275,1315,531,1197]" captionTargetPageId="1" captionText="FIGURE 1. Map of specimen records for Hister criticus, H. laevimargo, and H. latimargo." httpUri="https://zenodo.org/record/158690/files/figure.png" pageId="6" pageNumber="7">Figs. 1</figureCitation>
,
<figureCitation id="130B2A09FF98FFD8FD33FA16FD0804B3" box="[675,690,1504,1530]" captionStart="FIGURE 3" captionStartId="5.[264,368,1741,1765]" captionTargetBox="[384,1203,489,1713]" captionTargetId="figure@5.[384,1203,489,1715]" captionTargetPageId="5" captionText="FIGURE 3. Dorsal habitus photographs of Hister spp. A. Hister criticus; B. H. laevimargo; C. H. latimargo." httpUri="https://zenodo.org/record/169295/files/figure.png" pageId="6" pageNumber="7">3</figureCitation>
B, 4B)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32A6507FF98FFD8FE98F9DAFE4807AA" pageId="6" pageNumber="7" type="reference_group">
<paragraph id="8B8F368CFF98FFD8FE98F9DAFC8407CA" blockId="6.[264,1323,1580,1763]" pageId="6" pageNumber="7">
<treatmentCitationGroup id="AB2011A2FF98FFD8FE98F9DAFC8407CA" infered="true" pageId="6" pageNumber="7">
<taxonomicName id="4C304D0FFF98FFD8FE98F9DAFD36070A" ID-CoL="3M57P" authority="Lewis, 1900: 246" authorityName="Lewis" authorityPageNumber="246" authorityYear="1900" box="[264,652,1580,1603]" class="Insecta" family="Histeridae" genus="Hister" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="laevimargo">
<emphasis id="B944EA9EFF98FFD8FE98F9DAFE73070A" box="[264,457,1580,1603]" italics="true" pageId="6" pageNumber="7">Hister laevimargo</emphasis>
<treatmentCitation id="0A91109DFF98FFD8FE40F9DAFD36070A" author="Lewis" box="[464,652,1580,1603]" page="246" pageId="6" pageNumber="7" year="1900">
<bibRefCitation id="EFA14B7DFF98FFD8FE40F9DAFDED070A" author="Lewis" box="[464,599,1580,1603]" pageId="6" pageNumber="16" refString="Lewis, G. (1900) On new species of Histeridae and notices of others. Annals and Magazine of Natural History (7), 5, 246 - 254." type="journal article" year="1900">Lewis, 1900</bibRefCitation>
: 246
</treatmentCitation>
</taxonomicName>
;
<typeStatus id="548B882EFF98FFD8FD07F9DAFD43070A" box="[663,761,1580,1603]" pageId="6" pageNumber="7" type="lectotype">lectotype</typeStatus>
female, hereby designated: “
<collectingCountry id="F327761CFF98FFD8FBBDF9DAFB1C070A" box="[1069,1190,1580,1603]" name="Costa Rica" pageId="6" pageNumber="7">Costa Rica</collectingCountry>
, P. Biolley”/ “G.Lewis Coll., B.M.1926­369.”/ “
<taxonomicName id="4C304D0FFF98FFD8FD2DF9BAFC68072A" ID-CoL="3M57P" authority="Lewis" authorityName="Lewis" box="[701,978,1612,1635]" class="Insecta" family="Histeridae" genus="Hister" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="laevimargo">Hister laevimargo Lewis</taxonomicName>
Co­type”/ “Barba. Esmeralda,
<date id="FF8E104CFF98FFD8FEA8F99AFED107CA" box="[312,363,1644,1667]" pageId="6" pageNumber="7" value="1990-05">V.90</date>
, Troncs.”, BMNH; examined by the author.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B8F368CFF98FFD8FE98F97AFE4807AA" blockId="6.[264,1323,1580,1763]" pageId="6" pageNumber="7">
<treatmentCitationGroup id="AB2011A2FF98FFD8FE98F97AFE4807AA" infered="true" pageId="6" pageNumber="7">
<taxonomicName id="4C304D0FFF98FFD8FE98F97AFD1F07EA" ID-CoL="9TM2D" authority="Lewis, 1900: 247" authorityName="Lewis" authorityPageNumber="247" authorityYear="1900" box="[264,677,1676,1699]" class="Insecta" family="Histeridae" genus="Hister" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="planimargo">
<emphasis id="B944EA9EFF98FFD8FE98F97AFE6807EA" box="[264,466,1676,1699]" italics="true" pageId="6" pageNumber="7">Hister planimargo</emphasis>
<treatmentCitation id="0A91109DFF98FFD8FE4EF97AFD1F07EA" author="Lewis" box="[478,677,1676,1699]" page="247" pageId="6" pageNumber="7" year="1900">
<bibRefCitation id="EFA14B7DFF98FFD8FE4EF97AFDD107EA" author="Lewis" box="[478,619,1676,1699]" pageId="6" pageNumber="16" refString="Lewis, G. (1900) On new species of Histeridae and notices of others. Annals and Magazine of Natural History (7), 5, 246 - 254." type="journal article" year="1900">Lewis, 1900</bibRefCitation>
: 247
</treatmentCitation>
</taxonomicName>
;
<typeStatus id="548B882EFF98FFD8FD25F97AFCA007EA" box="[693,794,1676,1699]" pageId="6" pageNumber="7" type="lectotype">lectotype</typeStatus>
, hereby designated: “
<collectingCountry id="F327761CFF98FFD8FB9DF97AFB3607EA" box="[1037,1164,1676,1699]" name="Costa Rica" pageId="6" pageNumber="7">Costa Rica</collectingCountry>
, (Donckier).”/ “G.Lewis Coll., B.M.1926­369.”/ “
<taxonomicName id="4C304D0FFF98FFD8FD3FF95AFC07078A" ID-CoL="9TM2D" authority="Lewis" authorityName="Lewis" box="[687,957,1708,1731]" class="Insecta" family="Histeridae" genus="Hister" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="planimargo">Hister planimargo Lewis</taxonomicName>
<typeStatus id="548B882EFF98FFD8FC56F95AFC41078A" box="[966,1019,1708,1731]" pageId="6" pageNumber="7">Type</typeStatus>
”, MNHN; examined by the author;
<emphasis id="B944EA9EFF98FFD8FE1AF93DFE4807AA" bold="true" box="[394,498,1739,1763]" pageId="6" pageNumber="7">Syn. nov.</emphasis>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C32A6507FF98FFD9FE98F8E2FD3E00E7" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="diagnosis">
<paragraph id="8B8F368CFF98FFD9FE98F8E2FD3E00E7" blockId="6.[264,1323,1812,1878]" lastBlockId="7.[264,1323,284,710]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
<emphasis id="B944EA9EFF98FFD8FE98F8E2FE330664" box="[264,393,1812,1837]" italics="true" pageId="6" pageNumber="7">Diagnosis:</emphasis>
This species shares with
<taxonomicName id="4C304D0FFF98FFD8FD27F8E2FC890664" box="[695,819,1812,1837]" class="Insecta" family="Histeridae" genus="Hister" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="criticus">
<emphasis id="B944EA9EFF98FFD8FD27F8E2FC890664" box="[695,819,1812,1837]" italics="true" pageId="6" pageNumber="7">H. criticus</emphasis>
</taxonomicName>
the relatively narrow lateral pronotal marginal bead (between the single lateral stria and the margin), and the presence of prosternal striae. However, elytral stria 4, interrupted or absent in
<taxonomicName id="4C304D0FFF99FFD9FC1FFEEAFBB1007C" box="[911,1035,284,309]" class="Insecta" family="Histeridae" genus="Hister" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="criticus">
<emphasis id="B944EA9EFF99FFD9FC1FFEEAFBB1007C" box="[911,1035,284,309]" italics="true" pageId="7" pageNumber="8">H. criticus</emphasis>
</taxonomicName>
, is complete in
<taxonomicName id="4C304D0FFF99FFD9FB54FEEAFEE90014" class="Insecta" family="Histeridae" genus="Hister" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="laevimargo">
<emphasis id="B944EA9EFF99FFD9FB54FEEAFEE90014" italics="true" pageId="7" pageNumber="8">H. laevimargo</emphasis>
</taxonomicName>
. The propygidial and pygidial punctation also differs substantially, with
<taxonomicName id="4C304D0FFF99FFD9FB31FEB2FE9400CC" class="Insecta" family="Histeridae" genus="Hister" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="laevimargo">
<emphasis id="B944EA9EFF99FFD9FB31FEB2FE9400CC" italics="true" pageId="7" pageNumber="8">H. laevimargo</emphasis>
</taxonomicName>
having relatively small, and very densely set punctures, most separated by one­third their diameters or less (
<figureCitation id="130B2A09FF99FFD9FD8AFE62FDD800E7" box="[538,610,404,430]" captionStart="FIGURE 4" captionStartId="6.[264,367,1275,1299]" captionTargetBox="[364,1223,330,1248]" captionTargetId="figure@6.[364,1223,329,1249]" captionTargetPageId="6" captionText="FIGURE 4. Photographs of propygidium and pygidium of Hister spp. A. Hister criticus; B. H. laevimargo; C. H. latimargo; D. H. guanacaste; E. H. foveipygus." httpUri="https://zenodo.org/record/169296/files/figure.png" pageId="7" pageNumber="8">Fig. 4</figureCitation>
B).
</paragraph>
</subSubSection>
<subSubSection id="C32A6507FF99FFD9FEA8FE4AFD2C038F" pageId="7" pageNumber="8" type="description">
<paragraph id="8B8F368CFF99FFD9FEA8FE4AFC850307" blockId="7.[264,1323,284,710]" pageId="7" pageNumber="8">
<emphasis id="B944EA9EFF99FFD9FEA8FE4AFE76009C" box="[312,460,444,469]" italics="true" pageId="7" pageNumber="8">Description:</emphasis>
Average length:
<quantity id="4CC89B69FF99FFD9FD02FE4AFD56009F" box="[658,748,444,470]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="7" pageNumber="8" unit="mm" value="3.5">3.5 mm</quantity>
(range 3.2–3.8); Avg. width:
<quantity id="4CC89B69FF99FFD9FBD5FE4AFB25009F" box="[1093,1183,444,470]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="7" pageNumber="8" unit="mm" value="3.3">3.3 mm</quantity>
(range 3.0– 3.6). This species is extremely similar to
<taxonomicName id="4C304D0FFF99FFD9FC97FE12FC3D00B4" box="[775,903,484,509]" class="Insecta" family="Histeridae" genus="Hister" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="criticus">
<emphasis id="B944EA9EFF99FFD9FC97FE12FC3D00B4" box="[775,903,484,509]" italics="true" pageId="7" pageNumber="8">H. criticus</emphasis>
</taxonomicName>
, externally differing only in those characters given in the diagnosis. The aedeagus has not been examined because the species is only known from
<typeStatus id="548B882EFF99FFD9FE66FDC2FD8E0307" box="[502,564,564,590]" pageId="7" pageNumber="8">types</typeStatus>
in marginal condition.
</paragraph>
<paragraph id="8B8F368CFF99FFD9FEA8FDAAFD2C038F" blockId="7.[264,1323,284,710]" pageId="7" pageNumber="8">
<emphasis id="B944EA9EFF99FFD9FEA8FDAAFE2D033C" box="[312,407,604,629]" italics="true" pageId="7" pageNumber="8">Records</emphasis>
(
<figureCitation id="130B2A09FF99FFD9FE38FDAAFE54033F" box="[424,494,604,630]" captionStart="FIGURE 1" captionStartId="1.[264,368,1201,1225]" captionTargetBox="[369,1221,573,1164]" captionTargetId="figure@1.[275,1315,531,1197]" captionTargetPageId="1" captionText="FIGURE 1. Map of specimen records for Hister criticus, H. laevimargo, and H. latimargo." httpUri="https://zenodo.org/record/158690/files/figure.png" pageId="7" pageNumber="8">Fig. 1</figureCitation>
): This species is known only from the
<typeStatus id="548B882EFF99FFD9FC51FDAAFBB9033F" box="[961,1027,604,630]" pageId="7" pageNumber="8">types</typeStatus>
. The
<typeStatus id="548B882EFF99FFD9FBD4FDAAFBCD033F" box="[1092,1143,604,630]" pageId="7" pageNumber="8">type</typeStatus>
locality, Barba Esmeralda, apparently refers to a site on the slope of Volcán Barva in Heredia, about
<quantity id="4CC89B69FF99FFD9FA9DFD72FE97038F" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="8" unit="km" value="20.0">20 km</quantity>
north of San José,
<collectingCountry id="F327761CFF99FFD9FD9EFD5AFD28038F" box="[526,658,684,710]" name="Costa Rica" pageId="7" pageNumber="8">Costa Rica</collectingCountry>
.
</paragraph>
</subSubSection>
<caption id="DF4F6604FF99FFD9FE98F8E9FDF00610" httpUri="https://zenodo.org/record/158694/files/figure.png" pageId="7" pageNumber="8" targetBox="[510,1087,776,1806]" targetPageId="7">
<paragraph id="8B8F368CFF99FFD9FE98F8E9FDF00610" blockId="7.[264,1323,1823,1881]" pageId="7" pageNumber="8">
<emphasis id="B944EA9EFF99FFD9FE98F8E9FE36067E" bold="true" box="[264,396,1823,1847]" pageId="7" pageNumber="8">FIGURE 5.</emphasis>
Posterior view of protibiae of
<taxonomicName id="4C304D0FFF99FFD9FD73F8E9FC9F067F" box="[739,805,1823,1846]" class="Insecta" family="Histeridae" genus="Hister" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B944EA9EFF99FFD9FD73F8E9FC9F067F" box="[739,805,1823,1846]" italics="true" pageId="7" pageNumber="8">Hister</emphasis>
</taxonomicName>
spp. A.
<taxonomicName id="4C304D0FFF99FFD9FC16F8E9FB98067F" box="[902,1058,1823,1846]" class="Insecta" family="Histeridae" genus="Hister" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="criticus">
<emphasis id="B944EA9EFF99FFD9FC16F8E9FB98067F" box="[902,1058,1823,1846]" italics="true" pageId="7" pageNumber="8">Hister criticus</emphasis>
</taxonomicName>
; B.
<taxonomicName id="4C304D0FFF99FFD9FBC4F8E9FB64067F" box="[1108,1246,1823,1846]" class="Insecta" family="Histeridae" genus="Hister" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="latimargo">
<emphasis id="B944EA9EFF99FFD9FBC4F8E9FB64067F" box="[1108,1246,1823,1846]" italics="true" pageId="7" pageNumber="8">H. latimargo</emphasis>
</taxonomicName>
; C.
<taxonomicName id="4C304D0FFF99FFD9FA81F8E9FE3B0610" class="Insecta" family="Histeridae" genus="Hister" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="guanacaste">
<emphasis id="B944EA9EFF99FFD9FA81F8E9FE3B0610" italics="true" pageId="7" pageNumber="8">H. guanacaste</emphasis>
</taxonomicName>
; D.
<taxonomicName id="4C304D0FFF99FFD9FE20F8B4FDF90610" box="[432,579,1858,1881]" class="Insecta" family="Histeridae" genus="Hister" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="foveipygus">
<emphasis id="B944EA9EFF99FFD9FE20F8B4FDF90610" box="[432,579,1858,1881]" italics="true" pageId="7" pageNumber="8">H. foveipygus</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<caption id="DF4F6604FF96FFD6FE98FB68FB7C0591" httpUri="https://zenodo.org/record/158695/files/figure.png" pageId="8" pageNumber="9" targetBox="[450,1127,317,1146]" targetPageId="8">
<paragraph id="8B8F368CFF96FFD6FE98FB68FB7C0591" blockId="8.[264,1323,1182,1241]" pageId="8" pageNumber="9">
<emphasis id="B944EA9EFF96FFD6FE98FB68FE3305FF" bold="true" box="[264,393,1182,1206]" pageId="8" pageNumber="9">FIGURE 6.</emphasis>
Dorsal view of components of aedeagus of
<taxonomicName id="4C304D0FFF96FFD6FCCAFB69FC2705FF" box="[858,925,1183,1206]" class="Insecta" family="Histeridae" genus="Hister" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B944EA9EFF96FFD6FCCAFB69FC2705FF" box="[858,925,1183,1206]" italics="true" pageId="8" pageNumber="9">Hister</emphasis>
</taxonomicName>
spp. A. Tegmen of
<taxonomicName id="4C304D0FFF96FFD6FBFCFB69FABF05FF" box="[1132,1285,1183,1206]" class="Insecta" family="Histeridae" genus="Hister" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="criticus">
<emphasis id="B944EA9EFF96FFD6FBFCFB69FABF05FF" box="[1132,1285,1183,1206]" italics="true" pageId="8" pageNumber="9">Hister criticus</emphasis>
</taxonomicName>
; B. Tegmen of
<taxonomicName id="4C304D0FFF96FFD6FE11FB34FDA10590" box="[385,539,1218,1241]" class="Insecta" family="Histeridae" genus="Hister" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="guanacaste">
<emphasis id="B944EA9EFF96FFD6FE11FB34FDA10590" box="[385,539,1218,1241]" italics="true" pageId="8" pageNumber="9">H. guanacaste</emphasis>
</taxonomicName>
; C. Tegmen of
<taxonomicName id="4C304D0FFF96FFD6FD51FB34FCEE0590" box="[705,852,1218,1241]" class="Insecta" family="Histeridae" genus="Hister" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="foveipygus">
<emphasis id="B944EA9EFF96FFD6FD51FB34FCEE0590" box="[705,852,1218,1241]" italics="true" pageId="8" pageNumber="9">H. foveipygus</emphasis>
</taxonomicName>
; D. Median lobe of
<taxonomicName id="4C304D0FFF96FFD6FBBCFB34FB050590" box="[1068,1215,1218,1241]" class="Insecta" family="Histeridae" genus="Hister" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="foveipygus">
<emphasis id="B944EA9EFF96FFD6FBBCFB34FB050590" box="[1068,1215,1218,1241]" italics="true" pageId="8" pageNumber="9">H. foveipygus</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<subSubSection id="C32A6507FF96FFD7FEA8FAC2FB54009F" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9" type="discussion">
<paragraph id="8B8F368CFF96FFD6FEA8FAC2FB04064F" blockId="8.[264,1323,1332,1878]" pageId="8" pageNumber="9">
<emphasis id="B944EA9EFF96FFD6FEA8FAC2FE120404" box="[312,424,1332,1357]" italics="true" pageId="8" pageNumber="9">Remarks:</emphasis>
There is some discrepancy between the published data and those on the the three ‘type’ specimens of these two species. First, Lewis does not report numbers of specimens for either species. The BMNH presently has two specimens labelled, apparently by Lewis himself, as
<taxonomicName id="4C304D0FFF96FFD6FE72FA5AFD02048C" box="[482,696,1452,1477]" class="Insecta" family="Histeridae" genus="Hister" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="laevimargo">
<emphasis id="B944EA9EFF96FFD6FE72FA5AFD02048C" box="[482,696,1452,1477]" italics="true" pageId="8" pageNumber="9">Hister laevimargo</emphasis>
</taxonomicName>
, one collected by Donckier, labelled the ‘Type’, and one by P. Biolley, from Barba Esmeralda, as the ‘Co­type’. Only the latter data is cited in the original description for that species. The one specimen of
<taxonomicName id="4C304D0FFF96FFD6FB84FA0AFB7C075C" box="[1044,1222,1532,1557]" class="Insecta" family="Histeridae" genus="Hister" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="planimargo">
<emphasis id="B944EA9EFF96FFD6FB84FA0AFB7C075C" box="[1044,1222,1532,1557]" italics="true" pageId="8" pageNumber="9">H. planimargo</emphasis>
</taxonomicName>
bears a “
<collectingCountry id="F327761CFF96FFD6FE82F9D2FE230777" box="[274,409,1572,1598]" name="Costa Rica" pageId="8" pageNumber="9">Costa Rica</collectingCountry>
, (Donckier)” label, as cited. It seems that Lewis confused these specimens in the labelling, and that the specimen of
<taxonomicName id="4C304D0FFF96FFD6FD70F9BAFC30072C" box="[736,906,1612,1637]" class="Insecta" family="Histeridae" genus="Hister" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="laevimargo">
<emphasis id="B944EA9EFF96FFD6FD70F9BAFC30072C" box="[736,906,1612,1637]" italics="true" pageId="8" pageNumber="9">H. laevimargo</emphasis>
</taxonomicName>
labelled Co­type (with the correct label data) should be considered the primary
<typeStatus id="548B882EFF96FFD6FCBBF982FCE407C7" box="[811,862,1652,1678]" pageId="8" pageNumber="9">type</typeStatus>
of the species, and it is designated as such. The other two specimens, both collected by Donckier, are considered
<typeStatus id="548B882EFF96FFD6FB10F96AFB5D07FF" box="[1152,1255,1692,1718]" pageId="8" pageNumber="9" type="syntype">syntypes</typeStatus>
of
<taxonomicName id="4C304D0FFF96FFD6FA9EF96AFE2A0794" class="Insecta" family="Histeridae" genus="Hister" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="planimargo">
<emphasis id="B944EA9EFF96FFD6FA9EF96AFE2A0794" italics="true" pageId="8" pageNumber="9">H. planimargo</emphasis>
</taxonomicName>
, despite the fact that only one bears Lewis’
<taxonomicName id="4C304D0FFF96FFD6FC3EF932FBE10794" box="[942,1115,1732,1757]" class="Insecta" family="Histeridae" genus="Hister" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="planimargo">
<emphasis id="B944EA9EFF96FFD6FC3EF932FBE10794" box="[942,1115,1732,1757]" italics="true" pageId="8" pageNumber="9">H. planimargo</emphasis>
</taxonomicName>
label. This latter specimen is designated the
<typeStatus id="548B882EFF96FFD6FDD8F91AFD0D064F" box="[584,695,1772,1798]" pageId="8" pageNumber="9" type="lectotype">lectotype</typeStatus>
, and the other is not specifically designated.
</paragraph>
<paragraph id="8B8F368CFF96FFD7FEA8F8E2FB54009F" blockId="8.[264,1323,1332,1878]" lastBlockId="9.[264,1323,284,470]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
Describing
<taxonomicName id="4C304D0FFF96FFD6FE2FF8E2FD7B0667" authority="Lewis" authorityName="Lewis" box="[447,705,1812,1838]" class="Insecta" family="Histeridae" genus="Hister" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="planimargo">
<emphasis id="B944EA9EFF96FFD6FE2FF8E2FDD00664" box="[447,618,1812,1837]" italics="true" pageId="8" pageNumber="9">H. planimargo</emphasis>
, Lewis
</taxonomicName>
notes only that it is related to
<taxonomicName id="4C304D0FFF96FFD6FBB3F8E2FB760664" box="[1059,1228,1812,1837]" class="Insecta" family="Histeridae" genus="Hister" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="laevimargo">
<emphasis id="B944EA9EFF96FFD6FBB3F8E2FB760664" box="[1059,1228,1812,1837]" italics="true" pageId="8" pageNumber="9">H. laevimargo</emphasis>
</taxonomicName>
(and
<taxonomicName id="4C304D0FFF96FFD6FA9EF8E2FEC0061C" class="Insecta" family="Histeridae" genus="Hister" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="latimargo">
<emphasis id="B944EA9EFF96FFD6FA9EF8E2FEC0061C" italics="true" pageId="8" pageNumber="9">H. latimargo</emphasis>
</taxonomicName>
), but does not specify characters by which they differ. The most obvious difference in their descriptions is how Lewis describes the frontal stria, &quot;sinuous behind the mandibles, angulate above the eyes&quot; in
<taxonomicName id="4C304D0FFF97FFD7FD71FEB2FC310014" box="[737,907,324,349]" class="Insecta" family="Histeridae" genus="Hister" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="laevimargo">
<emphasis id="B944EA9EFF97FFD7FD71FEB2FC310014" box="[737,907,324,349]" italics="true" pageId="9" pageNumber="10">H. laevimargo</emphasis>
</taxonomicName>
, and &quot;nearly straight anteriorly&quot; in
<taxonomicName id="4C304D0FFF97FFD7FE98FE9AFE0300CC" box="[264,441,364,389]" class="Insecta" family="Histeridae" genus="Hister" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="planimargo">
<emphasis id="B944EA9EFF97FFD7FE98FE9AFE0300CC" box="[264,441,364,389]" italics="true" pageId="9" pageNumber="10">H. planimargo</emphasis>
</taxonomicName>
. However, elsewhere in their descriptions different language is used to describe apparently similar conditions, making it difficult to determine what he considered significant. In any case, in none of these characters can I detect any clear differences.
</paragraph>
</subSubSection>
</treatment>
</document>
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