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<document id="52511312B6C793F599468D4D9501E66E" ID-CLB-Dataset="45456" ID-DOI="10.11646/zootaxa.3815.4.4" ID-GBIF-Dataset="b2e39573-c2df-4ad5-9dec-93eb67d58e6c" ID-ISSN="1175-5326" ID-Zenodo-Dep="228698" ID-ZooBank="172C594B-7321-4F1C-B8CC-158A195A7D73" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460438354307" checkinUser="plazi" docAuthor="Väisänen, Rauno" docDate="2014" docId="03998794FFF3F70BFF4063A1FE12FA8C" docLanguage="en" docName="zt03815p540.pdf" docOrigin="Zootaxa 3815 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Mycomya nakanishii Väisänen, 2014, sp. n." docType="treatment" docVersion="8" lastPageNumber="532" masterDocId="FFA0FFECFFF5F70DFFD76339FFB3FF99" masterDocTitle="New Mycomya species from South-East Asia (Diptera, Mycetophilidae)" masterLastPageNumber="540" masterPageNumber="526" pageNumber="532" updateTime="1698332543223" updateUser="ExternalLinkService">
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<mods:title id="10C3835393BD570B26CCC07B420A3482">New Mycomya species from South-East Asia (Diptera, Mycetophilidae)</mods:title>
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<mods:namePart id="E7F432397E53092F26544F48BDE734AF">Väisänen, Rauno</mods:namePart>
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<mods:title id="DF4BA07832BEC01FBB1557559C3159C1">Zootaxa</mods:title>
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<mods:date id="095B4434C67DC5ACD46FA4474D64061A">2014</mods:date>
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<mods:classification id="B7DC72A8EC826EBDE7ECBD57997B7DA3">journal article</mods:classification>
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<mods:identifier id="6AB2D3C9AF46F9E14003E38C2201CB30" type="DOI">10.11646/zootaxa.3815.4.4</mods:identifier>
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<treatment id="03998794FFF3F70BFF4063A1FE12FA8C" ID-DOI="http://doi.org/10.5281/zenodo.6134637" ID-GBIF-Taxon="119507184" ID-Zenodo-Dep="6134637" LSID="urn:lsid:plazi:treatment:03998794FFF3F70BFF4063A1FE12FA8C" httpUri="http://treatment.plazi.org/id/03998794FFF3F70BFF4063A1FE12FA8C" lastPageNumber="532" pageId="6" pageNumber="532">
<subSubSection id="C32A6509FFF3F70BFF4063A1FEAAFF4D" pageId="6" pageNumber="532" type="nomenclature">
<paragraph id="8B8F3682FFF3F70BFF4063A1FE7CFF29" blockId="6.[151,463,151,213]" box="[151,463,151,176]" pageId="6" pageNumber="532">
<heading id="D0C781EEFFF3F70BFF4063A1FE7CFF29" bold="true" box="[151,463,151,176]" fontSize="10" level="2" pageId="6" pageNumber="532" reason="2">
<emphasis id="B944EA90FFF3F70BFF4063A1FE7CFF29" bold="true" box="[151,463,151,176]" pageId="6" pageNumber="532">
<taxonomicName id="4C304D01FFF3F70BFF4063A1FE3AFF29" box="[151,393,151,176]" class="Insecta" family="Mycetophilidae" genus="Mycomya" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="532" phylum="Arthropoda" rank="species" species="nakanishii" status="sp. nov.">
<emphasis id="B944EA90FFF3F70BFF4063A1FE3AFF29" bold="true" box="[151,393,151,176]" italics="true" pageId="6" pageNumber="532">Mycomya nakanishii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757EBFFF3F70BFE4763AEFE7CFF29" box="[400,463,151,176]" pageId="6" pageNumber="532" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8F3682FFF3F70BFF406385FEAAFF4D" blockId="6.[151,463,151,213]" box="[151,281,188,213]" pageId="6" pageNumber="532">
<figureCitation id="130B2A07FFF3F70BFF406385FF51FF4D" box="[151,226,188,213]" captionStart="FIGURE 4" captionStartId="6.[151,250,1974,1996]" captionTargetBox="[180,1405,1359,1944]" captionTargetId="figure@6.[176,1411,1356,1952]" captionTargetPageId="6" captionText="FIGURE 4. Mycomya nakanishii sp. n., male hypopygium. A – C tergal, sternal and lateral views, D apex of aedeagus, lateral view, twice enlarged. Scale 0.2 mm." httpUri="https://zenodo.org/record/228702/files/figure.png" pageId="6" pageNumber="532">Figs. 4</figureCitation>
A–D
</paragraph>
</subSubSection>
<subSubSection id="C32A6509FFF3F70BFF40623DFAEAFED9" pageId="6" pageNumber="532" type="materials_examined">
<paragraph id="8B8F3682FFF3F70BFF40623DFAEAFED9" blockId="6.[151,1437,260,1301]" pageId="6" pageNumber="532">
<emphasis id="B944EA90FFF3F70BFF40623DFEDAFE84" bold="true" box="[151,361,260,285]" pageId="6" pageNumber="532">Material studied.</emphasis>
<emphasis id="B944EA90FFF3F70BFEA2623DFE50FE85" box="[373,483,260,284]" italics="true" pageId="6" pageNumber="532">
<typeStatus id="548B8820FFF3F70BFEA2623DFE6CFE85" box="[373,479,260,284]" pageId="6" pageNumber="532" type="holotype">Holotype</typeStatus>
.
</emphasis>
♂.
<collectingCountry id="F3277612FFF3F70BFDC1623DFD0DFE85" box="[534,702,260,284]" name="Philippines" pageId="6" pageNumber="532">PHILIPPINES</collectingCountry>
, Mindanao, Mt. Apo, Agko – Lake Binado
<quantity id="4CC89B67FFF3F70BFB1A623DFADEFE85" box="[1229,1389,260,285]" metricMagnitude="3" metricUnit="m" metricValue="1.825" metricValueMax="2.3" metricValueMin="1.35" pageId="6" pageNumber="532" unit="m" value="1825.0" valueMax="2300.0" valueMin="1350.0">1350–2300 m</quantity>
alt. (
<typeStatus id="548B8820FFF3F70BFF49621EFEB1FED9" box="[158,258,295,320]" pageId="6" pageNumber="532" type="holotype">holotype</typeStatus>
collected on the way between the two localities)
<date id="FF8E1042FFF3F70BFCFC621EFC14FED9" box="[811,935,295,320]" pageId="6" pageNumber="532" value="1978-08-03">3.viii.1978</date>
, Nakanishi, A. &amp; Yata,
<collectingCountry id="F3277612FFF3F70BFB666211FB7BFED9" box="[1201,1224,296,320]" name="Iceland" pageId="6" pageNumber="532">O</collectingCountry>
. leg. (KUC).
</paragraph>
</subSubSection>
<subSubSection id="C32A6509FFF3F70BFF106275FB18FCFC" pageId="6" pageNumber="532" type="description">
<paragraph id="8B8F3682FFF3F70BFF106275FB18FCFC" blockId="6.[151,1437,260,1301]" pageId="6" pageNumber="532">
<emphasis id="B944EA90FFF3F70BFF106275FEEAFEFC" bold="true" box="[199,345,332,357]" pageId="6" pageNumber="532">Description.</emphasis>
♂.
<emphasis id="B944EA90FFF3F70BFE5B6275FE60FEFC" bold="true" box="[396,467,332,357]" pageId="6" pageNumber="532">Head.</emphasis>
Palp and other mouthparts yellow, face brownish, posterior parts of head brown. Antenna brownish to brown, scape, pedicel and base of 1st flagellomere yellow. 1st flagellomere about 2.5x its width, 2nd 1.5x its width.
<emphasis id="B944EA90FFF3F70BFE1362ADFD90FE34" bold="true" box="[452,547,404,429]" pageId="6" pageNumber="532">Thorax.</emphasis>
Pronotum yellow, with 3 long setae. Scutum brown with or without indistinct yellowish anterior margin. Anepisternum yellow to yellowish. Preepisternum yellow to brownish. Scutellum brown, with 4 long setae. Laterotergite brownish to brown. Mediotergite brown, bare.
<emphasis id="B944EA90FFF3F70BFB8F62E5FB2CFE6C" bold="true" box="[1112,1183,476,501]" pageId="6" pageNumber="532">Wing.</emphasis>
Length 4.0 mm. Wing hyaline. Sc ending in R1 distinctly distad of middle of small cell, Sc1 missing or broken (spur of Sc1 not reaching C). Apical part of Sc bearing about 12 small setae. Small cell about
<date id="FF8E1042FFF3F70BFC58611EFC19FDA6" box="[911,938,551,575]" pageId="6" pageNumber="532">2x</date>
as long as wide. Cu fork slightly distal to M fork. M ratios: 0.74, 1.00. Cu ratios: 0.82, 1.28. Small setae: M petiole: 0; M1: 4–5; M2: 4–5; Cu petiole: 0; Cu1: 1; Cu2: 1. Halter yellowish, apex brownish.
<emphasis id="B944EA90FFF3F70BFDB66148FD13FD13" bold="true" box="[609,672,625,650]" pageId="6" pageNumber="532">Legs.</emphasis>
Coxae and femora yellow, tibiae and tarsi brownish. Coxa 2 without spur. Leg ratios: bt1:t1 = 0.82, bt2:t2 = 0.65, bt3:t3 = 0.58.
<emphasis id="B944EA90FFF3F70BFCB661A1FC66FD28" bold="true" box="[865,981,664,689]" pageId="6" pageNumber="532">Abdomen</emphasis>
. Tergites brownish to brown. Sternites brownish.
<emphasis id="B944EA90FFF3F70BFEDB6185FE1CFD4C" bold="true" box="[268,431,700,725]" pageId="6" pageNumber="532">Hypopygium.</emphasis>
<figureCitation id="130B2A07FFF3F70BFE626185FDB2FD4C" box="[437,513,700,725]" captionStart="FIGURE 4" captionStartId="6.[151,250,1974,1996]" captionTargetBox="[180,1405,1359,1944]" captionTargetId="figure@6.[176,1411,1356,1952]" captionTargetPageId="6" captionText="FIGURE 4. Mycomya nakanishii sp. n., male hypopygium. A – C tergal, sternal and lateral views, D apex of aedeagus, lateral view, twice enlarged. Scale 0.2 mm." httpUri="https://zenodo.org/record/228702/files/figure.png" pageId="6" pageNumber="532">Figs. 4</figureCitation>
A–D, yellow. Sternal lateral appendage setose, short and rounded (
<figureCitation id="130B2A07FFF3F70BFB316185FA99FD4C" box="[1254,1322,700,725]" captionStart="FIGURE 4" captionStartId="6.[151,250,1974,1996]" captionTargetBox="[180,1405,1359,1944]" captionTargetId="figure@6.[176,1411,1356,1952]" captionTargetPageId="6" captionText="FIGURE 4. Mycomya nakanishii sp. n., male hypopygium. A – C tergal, sternal and lateral views, D apex of aedeagus, lateral view, twice enlarged. Scale 0.2 mm." httpUri="https://zenodo.org/record/228702/files/figure.png" pageId="6" pageNumber="532">Fig. 4</figureCitation>
C). Apical 1/3 of sternal synsclerite (including sternal lateral appendage) covered by strong setae, with some short, thin setae extending along submedian margin mid-way towards base of sternal submedian filament. Sternal submedian filament (stemming from basal-middle part of deeply bilobed sternal synsclerite) straight, longer than sternal lateral appendage (
<figureCitation id="130B2A07FFF3F70BFEC96075FED1FCFC" box="[286,354,844,869]" captionStart="FIGURE 4" captionStartId="6.[151,250,1974,1996]" captionTargetBox="[180,1405,1359,1944]" captionTargetId="figure@6.[176,1411,1356,1952]" captionTargetPageId="6" captionText="FIGURE 4. Mycomya nakanishii sp. n., male hypopygium. A – C tergal, sternal and lateral views, D apex of aedeagus, lateral view, twice enlarged. Scale 0.2 mm." httpUri="https://zenodo.org/record/228702/files/figure.png" pageId="6" pageNumber="532">Fig. 4</figureCitation>
B). Gonostylus with two apical teeth. Apex of aedeagus rounded (
<figureCitation id="130B2A07FFF3F70BFB936075FB3BFCFC" box="[1092,1160,844,869]" captionStart="FIGURE 4" captionStartId="6.[151,250,1974,1996]" captionTargetBox="[180,1405,1359,1944]" captionTargetId="figure@6.[176,1411,1356,1952]" captionTargetPageId="6" captionText="FIGURE 4. Mycomya nakanishii sp. n., male hypopygium. A – C tergal, sternal and lateral views, D apex of aedeagus, lateral view, twice enlarged. Scale 0.2 mm." httpUri="https://zenodo.org/record/228702/files/figure.png" pageId="6" pageNumber="532">Fig. 4</figureCitation>
D).
</paragraph>
</subSubSection>
<subSubSection id="C32A6509FFF3F70BFF106049FBAEFB55" pageId="6" pageNumber="532" type="discussion">
<paragraph id="8B8F3682FFF3F70BFF106049FBAEFB55" blockId="6.[151,1437,260,1301]" pageId="6" pageNumber="532">
<emphasis id="B944EA90FFF3F70BFF106049FEFFFC10" bold="true" box="[199,332,880,905]" pageId="6" pageNumber="532">Discussion.</emphasis>
<taxonomicName id="4C304D01FFF3F70BFE846048FD8FFC11" box="[339,572,880,904]" class="Insecta" family="Mycetophilidae" genus="Mycomya" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="532" phylum="Arthropoda" rank="species" species="nakanishii">
<emphasis id="B944EA90FFF3F70BFE846048FD8FFC11" box="[339,572,880,904]" italics="true" pageId="6" pageNumber="532">Mycomya nakanishii</emphasis>
</taxonomicName>
belongs to the subgenus
<taxonomicName id="4C304D01FFF3F70BFC8D6048FC79FC11" box="[858,970,881,904]" class="Insecta" family="Mycetophilidae" genus="Cymomya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="532" phylum="Arthropoda" rank="genus">
<emphasis id="B944EA90FFF3F70BFC8D6048FC79FC11" box="[858,970,881,904]" italics="true" pageId="6" pageNumber="532">Cymomya</emphasis>
</taxonomicName>
(as discussed for
<taxonomicName id="4C304D01FFF3F70BFB436048FB49FC11" box="[1172,1274,880,904]" class="Insecta" family="Mycetophilidae" genus="Mycomya" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="532" phylum="Arthropoda" rank="species" species="klossi">
<emphasis id="B944EA90FFF3F70BFB436048FB49FC11" box="[1172,1274,880,904]" italics="true" pageId="6" pageNumber="532">M. klossi</emphasis>
</taxonomicName>
above). It is a dark species, without the typical contrasting colour pattern of most species of the subgenus. It is quite similar to the Nepalese
<taxonomicName id="4C304D01FFF3F70BFED06080FE48FC48" authority="Vaisanen" authorityName="Vaisanen" box="[263,507,952,977]" class="Insecta" family="Mycetophilidae" genus="Mycomya" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="532" phylum="Arthropoda" rank="species" species="paguma">
<emphasis id="B944EA90FFF3F70BFED06080FE3AFC49" box="[263,393,953,976]" italics="true" pageId="6" pageNumber="532">M. paguma</emphasis>
Väisänen
</taxonomicName>
(2013: figs. 6A–C) in having a short and rounded sternal submedian appendage (
<figureCitation id="130B2A07FFF3F70BFF4860E5FF55FC6C" box="[159,230,988,1013]" captionStart="FIGURE 4" captionStartId="6.[151,250,1974,1996]" captionTargetBox="[180,1405,1359,1944]" captionTargetId="figure@6.[176,1411,1356,1952]" captionTargetPageId="6" captionText="FIGURE 4. Mycomya nakanishii sp. n., male hypopygium. A – C tergal, sternal and lateral views, D apex of aedeagus, lateral view, twice enlarged. Scale 0.2 mm." httpUri="https://zenodo.org/record/228702/files/figure.png" pageId="6" pageNumber="532">Fig. 4</figureCitation>
C) and its sternal submedian filament being longer than the sternal lateral appendage (
<figureCitation id="130B2A07FFF3F70BFB0560E5FA97FC6C" box="[1234,1316,988,1013]" captionStart="FIGURE 4" captionStartId="6.[151,250,1974,1996]" captionTargetBox="[180,1405,1359,1944]" captionTargetId="figure@6.[176,1411,1356,1952]" captionTargetPageId="6" captionText="FIGURE 4. Mycomya nakanishii sp. n., male hypopygium. A – C tergal, sternal and lateral views, D apex of aedeagus, lateral view, twice enlarged. Scale 0.2 mm." httpUri="https://zenodo.org/record/228702/files/figure.png" pageId="6" pageNumber="532">Figs. 4</figureCitation>
B–C). The setosity of the sternal synsclerite extends widely from the apex to the middle part in
<taxonomicName id="4C304D01FFF3F70BFB986738FB7DFB81" box="[1103,1230,1025,1048]" class="Insecta" family="Mycetophilidae" genus="Mycomya" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="532" phylum="Arthropoda" rank="species" species="paguma">
<emphasis id="B944EA90FFF3F70BFB986738FB7DFB81" box="[1103,1230,1025,1048]" italics="true" pageId="6" pageNumber="532">M. paguma</emphasis>
</taxonomicName>
, whereas only the apical third of sternal synsclerite has setae in
<taxonomicName id="4C304D01FFF3F70BFD77671CFC8FFBA5" box="[672,828,1060,1084]" class="Insecta" family="Mycetophilidae" genus="Mycomya" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="532" phylum="Arthropoda" rank="species" species="nakanishii">
<emphasis id="B944EA90FFF3F70BFD77671CFC8FFBA5" box="[672,828,1060,1084]" italics="true" pageId="6" pageNumber="532">M. nakanishii</emphasis>
</taxonomicName>
(excluding the small, thin setae along the submedian margins). The sternal lateral appendage of
<taxonomicName id="4C304D01FFF3F70BFDA56770FD42FBF9" box="[626,753,1097,1120]" class="Insecta" family="Mycetophilidae" genus="Mycomya" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="532" phylum="Arthropoda" rank="species" species="paguma">
<emphasis id="B944EA90FFF3F70BFDA56770FD42FBF9" box="[626,753,1097,1120]" italics="true" pageId="6" pageNumber="532">M. paguma</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFA14B73FFF3F70BFD286771FBA3FBF8" author="Vaisanen" box="[767,1040,1096,1121]" pageId="6" pageNumber="532" refString="Vaisanen, R. (2013 a) New Mycomya species from the Himalayas (Diptera, Mycetophilidae): 2. Subgenera Calomycomya, Cymomya, Neomycomya and Pavomya subg. n. Zootaxa, 3666 (3), 301 - 318. http: // dx. doi. org / 10.11646 / zootaxa. 3666.3.1" type="journal article" year="2013" yearSuffix="a">Väisänen 2013a: fig. 6B</bibRefCitation>
) is relatively larger and longer than in
<taxonomicName id="4C304D01FFF3F70BFF606754FEE6FB1D" box="[183,341,1132,1156]" class="Insecta" family="Mycetophilidae" genus="Mycomya" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="532" phylum="Arthropoda" rank="species" species="nakanishii">
<emphasis id="B944EA90FFF3F70BFF606754FEE6FB1D" box="[183,341,1132,1156]" italics="true" pageId="6" pageNumber="532">M. nakanishii</emphasis>
</taxonomicName>
(
<figureCitation id="130B2A07FFF3F70BFEBE6755FE03FB1C" box="[361,432,1132,1157]" captionStart="FIGURE 4" captionStartId="6.[151,250,1974,1996]" captionTargetBox="[180,1405,1359,1944]" captionTargetId="figure@6.[176,1411,1356,1952]" captionTargetPageId="6" captionText="FIGURE 4. Mycomya nakanishii sp. n., male hypopygium. A – C tergal, sternal and lateral views, D apex of aedeagus, lateral view, twice enlarged. Scale 0.2 mm." httpUri="https://zenodo.org/record/228702/files/figure.png" pageId="6" pageNumber="532">Fig. 4</figureCitation>
C), when the appendage is compared to the apical submedian extension of the sternal synsclerite on the other (medial) side of the base of gonostylus. The sternal submedian filament of
<taxonomicName id="4C304D01FFF3F70BFAD767A8FA33FB31" box="[1280,1408,1169,1192]" class="Insecta" family="Mycetophilidae" genus="Mycomya" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="532" phylum="Arthropoda" rank="species" species="paguma">
<emphasis id="B944EA90FFF3F70BFAD767A8FA33FB31" box="[1280,1408,1169,1192]" italics="true" pageId="6" pageNumber="532">M. paguma</emphasis>
</taxonomicName>
is very long, extending beyond the apex of the gonostylus, unlike in
<taxonomicName id="4C304D01FFF3F70BFCAB678CFBA6FB55" box="[892,1045,1204,1228]" class="Insecta" family="Mycetophilidae" genus="Mycomya" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="532" phylum="Arthropoda" rank="species" species="nakanishii">
<emphasis id="B944EA90FFF3F70BFCAB678CFBA6FB55" box="[892,1045,1204,1228]" italics="true" pageId="6" pageNumber="532">M. nakanishii</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C32A6509FFF3F70BFF1067E1FE12FA8C" pageId="6" pageNumber="532" type="etymology">
<paragraph id="8B8F3682FFF3F70BFF1067E1FE12FA8C" blockId="6.[151,1437,260,1301]" pageId="6" pageNumber="532">
<emphasis id="B944EA90FFF3F70BFF1067E1FEFDFB68" bold="true" box="[199,334,1240,1265]" pageId="6" pageNumber="532">Etymology.</emphasis>
The species is named after the Japanese entomologist, A. Nakanishi, who collected the
<typeStatus id="548B8820FFF3F70BFAED67E1FA2FFB68" box="[1338,1436,1240,1265]" pageId="6" pageNumber="532" type="holotype">holotype</typeStatus>
(together with
<collectingCountry id="F3277612FFF3F70BFEEB67C4FEE0FA8C" box="[316,339,1277,1301]" name="Iceland" pageId="6" pageNumber="532">O</collectingCountry>
. Yata).
</paragraph>
</subSubSection>
</treatment>
</document>
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