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<document id="2AF61140EE8E1D72A3E7B24A8853BBAC" ID-CLB-Dataset="99303" ID-DOI="10.5852/ejt.2022.830.1865" ID-GBIF-Dataset="deb635f8-c685-4637-b2a0-d028b1cba0ab" ID-ISSN="2118-9773" ID-Zenodo-Dep="6839847" ID-ZooBank="C1F47AD9-ECF6-4F0F-9928-55A45332FF4B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1657891195425" checkinUser="felipe" docAuthor="Kolesnichenko, Kirill A. &amp; Kotlobay, Anatoly A." docDate="2022" docId="EB42F330FFC0B33C913AFAF8FADBFD92" docLanguage="en" docName="ejt-830_1-60.pdf" docOrigin="European Journal of Taxonomy 830 (1)" docSource="http://zoobank.org/c1f47ad9-ecf6-4f0f-9928-55a45332ff4b" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Melitaea timandra subsp. binaludica Kolesnichenko &amp; Kotlobay 2022, subsp. nov." docType="treatment" docUuid="62F42B37-68FD-4A67-8E4F-FBAA2ED93033" docUuidSource="ZooBank" docVersion="9" lastPageNumber="32" masterDocId="177B8B48FFD7B323931AFF82FFD2FF6B" masterDocTitle="Review of the fritillary species systematically close to Melitaea lutko Evans, 1932 (Lepidoptera: Nymphalidae) with analysis of their geographic distribution and interrelations with host plants" masterLastPageNumber="60" masterPageNumber="1" pageNumber="24" updateTime="1699360064085" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="FA762353012F66E3EAF0182954C4373A">Review of the fritillary species systematically close to Melitaea lutko Evans, 1932 (Lepidoptera: Nymphalidae) with analysis of their geographic distribution and interrelations with host plants</mods:title>
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<mods:name id="30B627986A20A91B996CE5E88CC1A1B2" type="personal">
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<mods:namePart id="0CC87CBFA6317541B4DBA351FDCD9C6B">Kolesnichenko, Kirill A.</mods:namePart>
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<mods:affiliation id="524BAED8527DBA98AB07B936C4E6FFEC">Department of Entomology, Faculty of Biology, Lomonosov Moscow State University, Leninskie Gory I / 12, Moscow 119991, Russia.</mods:affiliation>
<mods:nameIdentifier id="B20CAFB97F72E0A90EEED702718C9D33" type="email">kkolesnichenko@gmail.com</mods:nameIdentifier>
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<mods:namePart id="416C2677CE9E67794C80E832B9AD946A">Kotlobay, Anatoly A.</mods:namePart>
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<mods:affiliation id="09D13E550BCA9D1172C9576F45462AB2">Department of Entomology, Faculty of Biology, Lomonosov Moscow State University, Leninskie Gory I / 12, Moscow 119991, Russia.</mods:affiliation>
<mods:nameIdentifier id="3BF8ADAB1FB7784A0A9EB38A56A44495" type="email">an_kotlobay@mail.ru</mods:nameIdentifier>
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<mods:title id="CDB30F8B5AB7E5B6538D64DF6B3948A9">European Journal of Taxonomy</mods:title>
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<mods:date id="802ADDE2905596199B2336DADBE98ADF">2022</mods:date>
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<mods:number id="0C9608BE5E6874FC995FBE8285BC453F">2022-07-15</mods:number>
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<treatment id="EB42F330FFC0B33C913AFAF8FADBFD92" ID-DOI="http://doi.org/10.5281/zenodo.6855860" ID-GBIF-Taxon="197817567" ID-Zenodo-Dep="6855860" LSID="urn:lsid:zoobank.org:act:62F42B37-68FD-4A67-8E4F-FBAA2ED93033" httpUri="http://treatment.plazi.org/id/EB42F330FFC0B33C913AFAF8FADBFD92" lastPageId="31" lastPageNumber="32" pageId="23" pageNumber="24">
<subSubSection id="2BF111ADFFC0B334913AFAF8FBC1FAFE" box="[544,1043,1402,1429]" pageId="23" pageNumber="24" type="nomenclature">
<paragraph id="63544226FFC0B334913AFAF8FBC1FAFE" blockId="23.[224,1364,1402,1499]" box="[544,1043,1402,1429]" pageId="23" pageNumber="24">
<heading id="381CF54AFFC0B334913AFAF8FBC1FAFE" box="[544,1043,1402,1429]" centered="true" fontSize="11" level="2" pageId="23" pageNumber="24" reason="2">
<taxonomicName id="A4EB39A5FFC0B334913AFAF8FC5BFAFE" authority="Kolesnichenko &amp; Kotlobay, 2022" authorityName="Kolesnichenko &amp; Kotlobay" authorityYear="2022" box="[544,905,1402,1429]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="subSpecies" species="timandra" status="subsp. nov." subSpecies="binaludica">
<emphasis id="519F9E34FFC0B334913AFAF8FC5BFAFE" bold="true" box="[544,905,1402,1429]" italics="true" pageId="23" pageNumber="24">Melitaea timandra binaludica</emphasis>
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<taxonomicNameLabel id="4AAC234FFFC0B334908AFAF8FBC1FAFE" box="[912,1043,1402,1429]" pageId="23" pageNumber="24" rank="subSpecies">subsp. nov.</taxonomicNameLabel>
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<subSubSection id="2BF111ADFFC0B33493FAFA42FA86FAB1" box="[224,1364,1472,1499]" pageId="23" pageNumber="24" type="description">
<paragraph id="63544226FFC0B33493FAFA42FA86FAB1" blockId="23.[224,1364,1402,1499]" box="[224,1364,1472,1499]" pageId="23" pageNumber="24">
<figureCitation id="FBD05EA3FFC0B33493FAFA42FDF2FAB0" box="[224,544,1472,1499]" captionStart="Fig" captionStartId="6.[190,233,1671,1697]" captionTargetBox="[192,1394,744,1633]" captionTargetId="figure-129@6.[189,1398,741,1634]" captionTargetPageId="6" captionText="Fig. 2. Adults of Melitaea lutko Evans, 1932 and M. timandra binaluduca subsp. nov. A–H. UPS. I–P. UNS. A, I. M. lutko, ♂, Chitral Gol National Park, Chitral, Pakistan, alt. 2700 m. B, J. M. timandra binaluduca subsp. nov., ♂, Iran, S Khorasan Prov., 35 km N Birjant t. C, K. M. timandra binaluduca subsp. nov., ♂, Central Afghanistan, Bamian Prov., Band-e-Amir, Dzhudoi-Kvak Gorge, alt. 3200 m. D, L. M. timandra binaluduca subsp. nov., ♂, Afghanistan, Bamian Prov., Band-e-Amir, alt. 3200 m. E, M. M. lutko, ♀, Chitral Gol National Park, Chitral, Pakistan, alt. 2700 m. F, N. M. timandra binaluduca subsp. nov., ♂, Iran, Mazandaran Prov., S macroslopes of Albors Mts, 80 km SE of Sari, 5 km NE Foulad Mahhaleh v., E slopes of Sultan Kuh Mt., alt. 2000 m. G, O. M. timandra binaluduca subsp. nov., ♀, Afghanistan, Bamian Prov., Band-e-Amir, alt. 3200 m. H, P. M. timandra binaluduca subsp. nov., ♀, Central Afghanistan, Bamian Prov., Band-e-Amir, Dzhudoi-Kvak Gorge, alt. 3200 m." figureDoi="http://doi.org/10.5281/zenodo.6839851" httpUri="https://zenodo.org/record/6839851/files/figure.png" pageId="23" pageNumber="24">
Figs 2B
<emphasis id="519F9E34FFC0B3349226FA43FE99FAB1" box="[316,331,1473,1498]" italics="true" pageId="23" pageNumber="24">–</emphasis>
D, F–H, J–L, N–P
</figureCitation>
,
<figureCitation id="FBD05EA3FFC0B3349136FA43FD57FAB0" box="[556,645,1473,1499]" captionStart="Fig" captionStartId="18.[189,232,1819,1845]" captionTargetBox="[216,1371,268,1778]" captionTargetId="figure-16@18.[214,1372,265,1781]" captionTargetPageId="18" captionText="Fig. 10. Male genitalia (a = valva; b = aedeagus). A–F. Melitaea timandra timandra Coutsis &amp; van Oorschot, 2014. G–O. M. timandra binaludica subsp. nov. A–C. S Turkmenistan, Sary-Yazy, alt. 300 m. D. Paratype, Turkmenistan, Badhyz Reserve, Kepeli. E–F. S Turkmenistan, Sary-Yazy, alt. 300 m. G–L. Iran, Rezavi Khorassan Prov., Kuh-e-Binalud Mts, Dorrud v. vicinity, alt. 2430 m. M. Central Afghanistan, Bamian Prov., Band-e-Amir, Dzhudoi-Kvak Gorge, alt. 3200 m. N. Bande-Amir, Hazarajat, Afghanistan. O. Afghanistan, Bamian Prov., Band-e-Amir, alt. 3200 m." figureDoi="http://doi.org/10.5281/zenodo.6839869" httpUri="https://zenodo.org/record/6839869/files/figure.png" pageId="23" pageNumber="24">
10G
<emphasis id="519F9E34FFC0B3349144FA43FDBFFAB1" box="[606,621,1473,1498]" italics="true" pageId="23" pageNumber="24">–</emphasis>
O
</figureCitation>
,
<figureCitation id="FBD05EA3FFC0B334918BFA43FD0FFAB0" box="[657,733,1473,1499]" captionStart="Fig" captionStartId="19.[189,232,1732,1758]" captionTargetBox="[192,1393,270,1668]" captionTargetId="figure-11@19.[189,1398,265,1675]" captionTargetPageId="19" captionText="Fig. 11. Female genitalia (without the anal papillae, copulatory bursa, and anterior and posterior apophyses). A–C. Melitaea timandra timandra Coutsis &amp; van Oorschot, 2014, S Turkmenistan, Sary- Yazy, alt. 300 m. D–I. M. timandra binaludica subsp. nov. D–F. Iran, Rezavi Khorassan Prov., Kuh-e- Binalud Mts, Dorrud v. vicinity, alt. 2430 m. G–H. Iran, Mazandaran Prov., S macroslopes of Albors Mts, 80 km SE of Sari, 5 km NE of Foulad Mahhaleh v., E slopes of Sultan Kuh Mt., alt. 2000 m. I. Afghanistan, Bamian Prov., Band-e-Amir, alt. 3200 m." figureDoi="http://doi.org/10.5281/zenodo.6839871" httpUri="https://zenodo.org/record/6839871/files/figure.png" pageId="23" pageNumber="24">
11D
<emphasis id="519F9E34FFC0B33491D8FA43FD03FAB1" box="[706,721,1473,1498]" italics="true" pageId="23" pageNumber="24">–</emphasis>
I
</figureCitation>
,
<figureCitation id="FBD05EA3FFC0B33491F3FA42FCE7FAB0" box="[745,821,1472,1499]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="23.[189,232,910,936]" captionStartId-1="24.[189,232,1891,1917]" captionTargetBox-0="[189,1398,267,872]" captionTargetBox-1="[194,1395,1026,1847]" captionTargetId-0="figure-261@23.[189,1398,267,872]" captionTargetId-1="figure-252@24.[189,1398,1022,1851]" captionTargetPageId-0="23" captionTargetPageId-1="24" captionText-0="Fig. 14. Distribution of Melitaea timandra timandra Coutsis &amp; van Oorschot, 2014, M. timandra binaludica subsp. nov. and M. shahvarica sp. nov. For a description of the points by letters, see Fig. 30. = Melitaea timandra timandra; = Melitaea timandra binaludica subsp. nov.; = Melitaea shahvarica sp. nov.;? = unconfirmed finds of Melitaea timandra; = Melitaea timandra with uncertain subspecies status." captionText-1="Fig. 15. Adults of Melitaea timandra binaludica subsp. nov. A–H. UPS. I–P. UNS. A, I. Holotype, ♂ (SDM). B–H, J–P. Paratypes (EDMSU). A–D, I–L. ♂, Iran, Rezavi Khorassan Prov., Kuh-e Binalud Mts, Dorrud v. vicinity, alt. 2430 m. E–H, M–P. ♀, same data as for preceding." figureDoi-0="http://doi.org/10.5281/zenodo.6839881" figureDoi-1="http://doi.org/10.5281/zenodo.6839883" httpUri-0="https://zenodo.org/record/6839881/files/figure.png" httpUri-1="https://zenodo.org/record/6839883/files/figure.png" pageId="23" pageNumber="24">14–15</figureCitation>
,
<figureCitation id="FBD05EA3FFC0B334905BFA43FC5CFAB0" box="[833,910,1473,1499]" captionStart="Fig" captionStartId="26.[189,232,1847,1873]" captionTargetBox="[276,1315,271,1797]" captionTargetId="figure-17@26.[261,1325,263,1804]" captionTargetPageId="26" captionText="Fig. 17. Male genitalia (without aedeagus) in dorsal projection. A–C. Melitaea timandra timandra Coutsis &amp; van Oorschot, 2014. D–I. M. timandra binaludica subsp. nov. A–C. S Turkmenistan, Sary- Yazy, alt. 300 m. D–F. Iran, Rezavi Khorassan Prov., Kuh-e-Binalud Mts, Dorrud v. vicinity, alt. 2430 m. G. Iran, Horossan Prov., 35 km N of Birjant t. H. Afghanistan, Bamian Prov., Band-e-Amir, alt. 3200 m. I. Central Afghanistan, Bamian Prov., Band-e-Amir, Dzhudoi-Kvak Gorge, alt. 3200 m." figureDoi="http://doi.org/10.5281/zenodo.6839887" httpUri="https://zenodo.org/record/6839887/files/figure.png" pageId="23" pageNumber="24">
17D
<emphasis id="519F9E34FFC0B334906EFA43FC51FAB1" box="[884,899,1473,1498]" italics="true" pageId="23" pageNumber="24">–</emphasis>
I
</figureCitation>
,
<figureCitation id="FBD05EA3FFC0B3349081FA43FC3AFAB0" box="[923,1000,1473,1499]" captionStart="Fig" captionStartId="28.[189,232,1853,1879]" captionTargetBox="[192,1398,692,1812]" captionTargetId="figure-178@28.[189,1398,689,1813]" captionTargetPageId="28" captionText="Fig. 19. Male genitalia and harpe. A–C. Melitaea timandra timandra Coutsis &amp; van Oorschot, 2014. D–I. M. timandra binaludica subsp. nov. A–C. Turkmenistan, Sary-Yazy, alt. 300 m. D–F. Iran, Rezavi Khorassan Prov., Kuh-e-Binalud Mts, Dorrud v. vicinity, alt. 2430 m. G. Afghanistan, Bamian Prov., Band-e-Amir, alt. 3200 m. H. Afghanistan, Bamian Prov., Band-e-Amir, Dzhudoi-Kvak Gorge, alt. 3200 m. I. Afghanistan, Band-e-Amir, Hazarajat." figureDoi="http://doi.org/10.5281/zenodo.6839893" httpUri="https://zenodo.org/record/6839893/files/figure.png" pageId="23" pageNumber="24">
19D
<emphasis id="519F9E34FFC0B33490D7FA43FC0EFAB1" box="[973,988,1473,1498]" italics="true" pageId="23" pageNumber="24">–</emphasis>
I
</figureCitation>
,
<figureCitation id="FBD05EA3FFC0B33490EEFA43FBC6FAB0" box="[1012,1044,1473,1499]" captionStart="Fig" captionStartId="30.[189,232,1954,1980]" captionTargetBox="[189,1398,554,1911]" captionTargetId="figure-99@30.[189,1398,554,1911]" captionTargetPageId="30" captionText="Fig. 20. Eggs of Melitaea timandra binaludica subsp. nov., Iran, Kuh-e-Binalud Mts. A–C. Lateral view. D–F. View from above. G–I. Micropile area." figureDoi="http://doi.org/10.5281/zenodo.6839895" httpUri="https://zenodo.org/record/6839895/files/figure.png" pageId="23" pageNumber="24">20</figureCitation>
,
<figureCitation id="FBD05EA3FFC0B334973AFA43FB4EFAB0" box="[1056,1180,1473,1499]" captionStart="Fig" captionStartId="47.[189,232,1750,1776]" captionTargetBox="[273,1315,270,1661]" captionTargetId="figure-11@47.[270,1317,265,1713]" captionTargetPageId="47" captionText="Fig.29.Differencesinthestructureofvalvaandaedeagusofthelutko speciesgroup.A.Melitaeashahvarica sp. nov. B, D, H. M. timandra binaludica subsp. nov. C–E. M. mimetica Higgins, 1940. F. M. lutko Evans, 1932. G. M. timandra timandra Coutsis &amp;van Oorschot, 2014. A. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mts, alt. 2200–2400 m. B. Iran, Rezavi Khorassan Prov., Kuh-e-Binalud Mts, Dorrud v. vicinity, alt. 2430 m. C. Afghanistan, Bamian Prov., Punjub Distr., 10 km NE of Varas v., alt. 2400 m. D. Afghanistan, Band-i-Amir, Hazarajat. E. Pakistan, Balochistan, Quetta, Urak, alt. 2400– 2700 m. F. Pakistan, Chitral, Gol National Park, alt. 2700 m. G. Turkmenistan, Sary-Yazy, alt. 700 m. H. Iran, Rezavi Khorassan Prov., Kuh-e-Binalud Mts, Dorrud v. vicinity, alt. 2430 m." figureDoi="http://doi.org/10.5281/zenodo.6839915" httpUri="https://zenodo.org/record/6839915/files/figure.png" pageId="23" pageNumber="24">29B, D, H</figureCitation>
,
<figureCitation id="FBD05EA3FFC0B33497B2FA43FB1BFAB0" box="[1192,1225,1473,1499]" captionStart="Fig" captionStartId="51.[189,232,1039,1065]" captionTargetBox="[189,1398,269,992]" captionTargetId="figure-11@51.[189,1398,269,992]" captionTargetPageId="51" captionText="Fig. 30 (continued on next page). Distribution of the lutko species group fritillaries in combination with the distribution of known and suspected species of host plants from the genus Phlomoides Blume. = Melitaea lutko Evans, 1932; = M. timandra timandra Coutsis &amp; Oorschot, 2014; = M. timandra binaludica subsp. nov.; = M. shahvarica sp. nov.; = M. mimetica mimetica Higgins, 1940; = M. mimetica delerei Heidemann, 1954;? = unconfirmed findings of M. timandra; = M. timandra with an unclear subspecies status; = Phlomoides regeliana (Aitch. &amp; Hemsl.) Adylov, Kamelin &amp; Makhm.; = Phlomoides boissieriana (Regel) Adylov, Kamelin &amp; Makhm.; = Phlomoides laciniata (L.) Kamelin &amp; Makhm.; = Phlomoides labiosiformis (Popov) Adylov, Kamelin &amp; Makhm.; = Phlomoides loasifolia (Benth.) Kamelin &amp; Makhm.; = Phlomoides molucelloides (Bunge) Salmaki; = Phlomoides acaulis(Beck ex Rech.f.) Salmaki; = Phlomoides labiosa(Bunge) Adylov, Kamelin &amp; Makhm. A. Pakistan, Chitral, Chaghbini CGNP, alt. 2700–3000 m. B. Pakistan, Khyber Pakhtunkhwa, Drosh. C. Pakistan, Khyber Pakhtunkhwa, Keon Nullah. D. Pakistan, Khyber Pakhtunkhwa, Malakand. E. Pakistan, Khyber Pakhtunkhwa, Birmoglasht. F. Turkmenistan, Badkhyz, Kepeli, alt. 700 m. G. Turkmenistan, Badkhyz, Kyzyl-Jar, alt. 700 m. H. Turkmenistan, Kushka, alt. 700 m. I. Turkmenistan, Murgab river, Sary-Yazy, alt. 300 m. J. Turkmenistan, 30 km E of BairamAli, Zahmet, alt. 240 m. K. Turkmenistan, Bairam-Ali, alt. 230 m. L. Turkmenistan, Kara-Kum desert, 30 km W of Mary, alt. 200 m.M. Turkmenistan, Dushak, alt. 250 m.N. Turkmenistan, Chaacha, alt. 400 m. O. Turkmenistan, Bakharden, alt. 200 m. P. Iran, Khorossan Razavi, Kuh-e-Binalud Mts, Qadamgah area, Gerina, alt. 2000 m. Q. Iran, Khorasan Razavi, Kuh-e-Binalud Mts, 15 km SW of Zoshk, alt. 2300– 2500 m. R. Iran, S Khorosan, 75 km N of Birjant, Sedeh, alt. 1500 m. S. Iran, S Khorosan, 35 km N of Birjant, alt. 1500 m. T. Afghanistan, Bamian, Band-e-Amir, Dzhudoi-Kvak Gorge, alt. 3200 m. U. Afghanistan, Bamian, Band-e-Amir, Hazarajat, alt. 3000–3200 m. V. Afghanistan, Bamian, Koh-iBaba Mts, Joshanak, alt. 2800 m. W. Afghanistan, Heart, Qala-i-Naw, Kashka pass. X. Iran, Semnan, Shahvar Mt., alt. 2200–2500 m. Y. Turkmenistan, Kara-Kala, Monjukly Ridge, 300–700 m. Z. Iran, Golestan, E Maraveh Tappeh, N Ghazan Ghayeh, Palizan Mts. A”. Pakistan, Balochistan, Quetta, Urak, alt. 2500 m. B”. Pakistan, Balochistan, Ziarat, alt. 2500 m. C”. Pakistan, Balochistan, Khojak, alt. 1700 m. D”. Pakistan, Balochistan, Zaghum, alt. 1600 m; E”. Pakistan, Punjab, Gawar, alt. 500 m. F”. Pakistan, Balochistan, Sheik Wazil, alt. 1600 m. G”. Afghanistan, Bamian, Hushkak, alt. 2700– 2800 m. H”. Afghanistan, Bamian, Punjub Distr., 10 km NE of Varas, alt. 2400 m. I”. Afghanistan, Ghor, 17 km E of Changcharan, 15 km S of Bandi-Ali, Gazak Mts, alt. 2400 m. J”. Afghanistan, Ghor, Bayan Range, 15 km S of Changcharan, Kindival valley, alt. 2700 m. K”. Afghanistan, Bamiyan, Kohi-Baba Mts, Panjao, alt. 3000 m. Afghanistan, Bamiyan, Koh-i-Baba Mts, Shah-tu-Kotal, alt. 4000 m. L”. Afghanistan, Kapisa, Pandshir valley, alt. 2200–2800 m. M”. Afghanistan, Kabul. N”. Iran, Tehran, Elburz Ridge, Demavend Mt., Ask, alt. 1800 m. O”. Iran, Semnan, Foulad Mohaleh, alt. 2200 m. P”. Pakistan, Punjab, Murree." figureDoi="http://doi.org/10.5281/zenodo.6841901" httpUri="https://zenodo.org/record/6841901/files/figure.png" pageId="23" pageNumber="24">30</figureCitation>
;
<tableCitation id="2E69779DFFC0B33497CEFA42FAE5FAB0" box="[1236,1335,1472,1499]" captionStart="Table 3" captionStartId="29.[189,259,265,291]" captionTargetPageId="29" captionText="Table 3. Measurements of the egg of M. timandra binaludica subsp. nov." httpUri="http://table.plazi.org/id/379412AEFFCAB33E93A7FE8BFBCBFE4F" pageId="23" pageNumber="24" tableUuid="379412AEFFCAB33E93A7FE8BFBCBFE4F">Tables 3</tableCitation>
,
<tableCitation id="2E69779DFFC0B334965FFA42FA86FAB1" box="[1349,1364,1472,1498]" captionStart="Table 6" captionStartId="40.[189,259,265,291]" captionTargetPageId="40" captionText="Table 6. Distance between the bristles on the head capsule of M. timandra binaludica subsp. nov." pageId="23" pageNumber="24">6</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="2BF111ADFFC0B33493A7F980FF2CF82A" pageId="23" pageNumber="24" type="diagnosis">
<paragraph id="63544226FFC0B33493A7F980FE1BF977" blockId="23.[189,457,1538,1564]" box="[189,457,1538,1564]" pageId="23" pageNumber="24">
<heading id="381CF54AFFC0B33493A7F980FE1BF977" bold="true" box="[189,457,1538,1564]" fontSize="11" level="3" pageId="23" pageNumber="24" reason="3">
<emphasis id="519F9E34FFC0B33493A7F980FE1BF977" bold="true" box="[189,457,1538,1564]" pageId="23" pageNumber="24">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="63544226FFC0B33493A7F9B2FF2CF82A" blockId="23.[187,1400,1584,1857]" pageId="23" pageNumber="24">
<taxonomicName id="A4EB39A5FFC0B33493A7F9B2FDCDF921" authority="Kolesnichenko &amp; Kotlobay, 2022" authorityName="Kolesnichenko &amp; Kotlobay" authorityYear="2022" box="[189,543,1584,1610]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="subSpecies" species="timandra" status="subsp. nov." subSpecies="binaludica">
<emphasis id="519F9E34FFC0B33493A7F9B2FDCDF921" box="[189,543,1584,1610]" italics="true" pageId="23" pageNumber="24">Melitaea timandra binaludica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4AAC234FFFC0B3349132F9B2FD7EF920" box="[552,684,1584,1611]" pageId="23" pageNumber="24" rank="subSpecies">subsp. nov.</taxonomicNameLabel>
differs from the nominative subspecies in appearance by its smaller size, the absence or weakly expressed UPF postdiscal pale area in males, the presence of UPH discal row, the developed UPH basal suffusion and the presence of UNH dark scales both in males and females. In the structure of the male genitalia, the main distinguishing features should be considered a noticeably thinner caudal process of the valva (on average thinner than in the nominative subspecies) and a noticeably less wide central part of the harpe (1.5–2 times narrower than in the nominative subspecies). In addition, unlike the nominative subspecies, the harpe on its inner surface carries a smaller number of teeth.
</paragraph>
</subSubSection>
<subSubSection id="2BF111ADFFC0B33493A7F8EAFEFDF8BF" pageId="23" pageNumber="24" type="etymology">
<paragraph id="63544226FFC0B33493A7F8EAFE96F8E9" blockId="23.[189,324,1896,1922]" box="[189,324,1896,1922]" pageId="23" pageNumber="24">
<heading id="381CF54AFFC0B33493A7F8EAFE96F8E9" bold="true" box="[189,324,1896,1922]" fontSize="11" level="3" pageId="23" pageNumber="24" reason="3">
<emphasis id="519F9E34FFC0B33493A7F8EAFE96F8E9" bold="true" box="[189,324,1896,1922]" pageId="23" pageNumber="24">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="63544226FFC0B33493A7F815FEFDF8BF" blockId="23.[189,1399,1943,2004]" pageId="23" pageNumber="24">
The name is toponymic and denotes the name of the mountain Ridge on which the
<typeStatus id="BC50FC84FFC0B33497A4F815FB23F8DA" box="[1214,1265,1943,1969]" pageId="23" pageNumber="24">type</typeStatus>
series was collected.
</paragraph>
</subSubSection>
<subSubSection id="2BF111ADFFCFB33893A7FE8BFC6FFEE6" lastPageId="27" lastPageNumber="28" pageId="24" pageNumber="25" type="materials_examined">
<paragraph id="63544226FFCFB33B93A7FE8BFEBEFE49" blockId="24.[189,364,264,291]" box="[189,364,264,291]" pageId="24" pageNumber="25">
<heading id="381CF54AFFCFB33B93A7FE8BFEBEFE49" bold="true" box="[189,364,264,291]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
<emphasis id="519F9E34FFCFB33B93A7FE8BFEBEFE49" bold="true" box="[189,364,264,291]" pageId="24" pageNumber="25">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="63544226FFCFB33B93F0FEB5FDB6FEF3" blockId="24.[189,1399,311,408]" pageId="24" pageNumber="25">
<materialsCitation id="D383487BFFCFB33B93F0FEB5FDB2FEF3" ID-GBIF-Occurrence="3863902324" collectingDate="2017-05-12" collectingDateMax="2017-05-13" collectingDateMin="2017-05-12" collectionCode="SDM" collectorName="K. Kolesnichenko" country="Iran" county="Rezavi" elevation="2430" location="Dorrud" municipality="Kuh-e Binalud Mts" pageId="24" pageNumber="25" specimenCount="1" specimenCount-male="1" stateProvince="Khorassan Prov." typeStatus="holotype">
<emphasis id="519F9E34FFCFB33B93F0FEB5FE8EFE3A" bold="true" box="[234,348,311,337]" pageId="24" pageNumber="25">
<typeStatus id="BC50FC84FFCFB33B93F0FEB5FE8EFE3A" box="[234,348,311,337]" pageId="24" pageNumber="25" type="holotype">Holotype</typeStatus>
</emphasis>
(
<figureCitation id="FBD05EA3FFCFB33B9276FEB5FE3CFE3A" box="[364,494,311,338]" captionStart="Fig" captionStartId="24.[189,232,1891,1917]" captionTargetBox="[194,1395,1026,1847]" captionTargetId="figure-252@24.[189,1398,1022,1851]" captionTargetPageId="24" captionText="Fig. 15. Adults of Melitaea timandra binaludica subsp. nov. A–H. UPS. I–P. UNS. A, I. Holotype, ♂ (SDM). B–H, J–P. Paratypes (EDMSU). A–D, I–L. ♂, Iran, Rezavi Khorassan Prov., Kuh-e Binalud Mts, Dorrud v. vicinity, alt. 2430 m. E–H, M–P. ♀, same data as for preceding." figureDoi="http://doi.org/10.5281/zenodo.6839883" httpUri="https://zenodo.org/record/6839883/files/figure.png" pageId="24" pageNumber="25">Fig. 15A, I</figureCitation>
)
<collectingCountry id="1BFC02B6FFCFB33B93A7FED8FED7FE1F" box="[189,261,346,372]" name="Iran" pageId="24" pageNumber="25">IRAN</collectingCountry>
•
<specimenCount id="75ED89AFFFCFB33B9238FED9FEE9FE1E" box="[290,315,347,373]" pageId="24" pageNumber="25" type="male">♂</specimenCount>
;
<collectingCounty id="8A353AAAFFCFB33B9253FED8FE48FE1F" box="[329,410,346,372]" pageId="24" pageNumber="25">Rezavi</collectingCounty>
<collectingRegion id="A12F8CC4FFCFB33B92BEFED8FDBBFE1E" box="[420,617,346,373]" country="Iran" name="Khorasan-e Shomali" pageId="24" pageNumber="25">Khorassan Prov.</collectingRegion>
,
<collectingMunicipality id="8330D85CFFCFB33B916CFED8FC8DFE1E" box="[630,863,346,373]" pageId="24" pageNumber="25">Kuh-e Binalud Mts</collectingMunicipality>
,
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<taxonomicName id="A4EB39A5FFCFB33B90D1FED9FB94FE1F" box="[971,1094,346,373]" pageId="24" pageNumber="25" rank="variety" variety="vicinity">v. vicinity</taxonomicName>
; alt.
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;
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<collectingDate id="07119D0EFFCFB33B97F4FED8FF29FEF3" pageId="24" pageNumber="25" value="2017-05-12" valueMax="2017-05-13" valueMin="2017-05-12">12–13 May 2017</collectingDate>
</date>
;
<collectorName id="CE1E27F0FFCFB33B921DFEFCFE09FEFC" box="[263,475,381,408]" pageId="24" pageNumber="25">K. Kolesnichenko</collectorName>
leg.;
<collectionCode id="05FADAE3FFCFB33B9107FEFCFDB2FEF3" box="[541,608,382,408]" country="United Kingdom" httpUri="http://grbio.org/cool/gw1v-bpc3" name="Stroud and District Museum" pageId="24" pageNumber="25">SDM</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="63544226FFCFB33B93F0FE3DFD9FFEB2" blockId="24.[189,1399,447,896]" box="[234,589,447,474]" pageId="24" pageNumber="25">
<emphasis id="519F9E34FFCFB33B93F0FE3DFEB5FEB2" bold="true" box="[234,359,447,473]" pageId="24" pageNumber="25">
<typeStatus id="BC50FC84FFCFB33B93F0FE3DFEB5FEB2" box="[234,359,447,473]" pageId="24" pageNumber="25" type="paratype">Paratypes</typeStatus>
</emphasis>
(
<figureCitation id="FBD05EA3FFCFB33B9262FE3DFD94FEB2" box="[376,582,447,474]" captionStart="Fig" captionStartId="24.[189,232,1891,1917]" captionTargetBox="[194,1395,1026,1847]" captionTargetId="figure-252@24.[189,1398,1022,1851]" captionTargetPageId="24" captionText="Fig. 15. Adults of Melitaea timandra binaludica subsp. nov. A–H. UPS. I–P. UNS. A, I. Holotype, ♂ (SDM). B–H, J–P. Paratypes (EDMSU). A–D, I–L. ♂, Iran, Rezavi Khorassan Prov., Kuh-e Binalud Mts, Dorrud v. vicinity, alt. 2430 m. E–H, M–P. ♀, same data as for preceding." figureDoi="http://doi.org/10.5281/zenodo.6839883" httpUri="https://zenodo.org/record/6839883/files/figure.png" pageId="24" pageNumber="25">Fig. 15B– H, J–P</figureCitation>
)
</paragraph>
<paragraph id="63544226FFCFB33B93A7FE60FE03FC14" blockId="24.[189,1399,447,896]" pageId="24" pageNumber="25">
<materialsCitation id="D383487BFFCFB33B93A7FE60FB1BFD74" ID-GBIF-Occurrence="3863902301" collectingDate="2017-05-12" collectingDateMax="2017-05-13" collectingDateMin="2017-05-12" collectionCode="EDMSU" collectorName="K. Kolesnichenko" country="Iran" elevation="2430" location="Dorrud" municipality="Kuh-e Binalud Mts" pageId="24" pageNumber="25" specimenCount="100" specimenCount-female="18" specimenCount-male="82" stateProvince="Khorasan-e Shomali" typeStatus="paratype">
<collectingCountry id="1BFC02B6FFCFB33B93A7FE60FED7FE97" box="[189,261,482,508]" name="Iran" pageId="24" pageNumber="25">IRAN</collectingCountry>
–
<emphasis id="519F9E34FFCFB33B9232FE60FD8BFE97" bold="true" box="[296,601,482,508]" pageId="24" pageNumber="25">
Rezavi
<collectingRegion id="A12F8CC4FFCFB33B929DFE60FD8BFE97" box="[391,601,482,508]" country="Iran" name="Khorasan-e Shomali" pageId="24" pageNumber="25">Khorassan Prov.</collectingRegion>
</emphasis>
•
<specimenCount id="75ED89AFFFCFB33B9162FE60FD1EFE96" box="[632,716,482,509]" pageId="24" pageNumber="25" type="male">82 ♂♂</specimenCount>
(7 dissected),
<specimenCount id="75ED89AFFFCFB33B9065FE60FC01FE96" box="[895,979,482,509]" pageId="24" pageNumber="25" type="female">18 ♀♀</specimenCount>
(6 dissected);
<collectingMunicipality id="8330D85CFFCFB33B979DFE60FAA3FE96" box="[1159,1393,482,509]" pageId="24" pageNumber="25">Kuh-e Binalud Mts</collectingMunicipality>
,
<location id="663414FDFFCFB33B93A7FD87FEC0FD74" LSID="urn:lsid:plazi:treatment:EB42F330FFC0B33C913AFAF8FADBFD92:663414FDFFCFB33B93A7FD87FEC0FD74" box="[189,274,517,543]" country="Iran" municipality="Kuh-e Binalud Mts" name="Dorrud" pageId="24" pageNumber="25" stateProvince="Khorasan-e Shomali">Dorrud</location>
<taxonomicName id="A4EB39A5FFCFB33B9205FD84FE4EFD74" box="[287,412,517,544]" pageId="24" pageNumber="25" rank="variety" variety="vicinity">v. vicinity</taxonomicName>
; alt.
<quantity id="A413EFC3FFCFB33B92FAFD87FD93FD4B" box="[480,577,517,544]" metricMagnitude="3" metricUnit="m" metricValue="2.43" pageId="24" pageNumber="25" unit="m" value="2430.0">
<elevation id="E8C6A515FFCFB33B92FAFD87FD93FD4B" box="[480,577,517,544]" metricMagnitude="3" metricUnit="m" metricValue="2.43" pageId="24" pageNumber="25" unit="m" value="2430.0">2430 m</elevation>
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;
<date id="175564E6FFCFB33B9149FD87FCF8FD74" box="[595,810,517,544]" pageId="24" pageNumber="25" value="2017-05-12" valueMax="2017-05-13" valueMin="2017-05-12">
<collectingDate id="07119D0EFFCFB33B9149FD87FCF8FD74" box="[595,810,517,544]" pageId="24" pageNumber="25" value="2017-05-12" valueMax="2017-05-13" valueMin="2017-05-12">12–13 May 2017</collectingDate>
</date>
;
<collectorName id="CE1E27F0FFCFB33B9021FD84FBC7FD74" box="[827,1045,517,544]" pageId="24" pageNumber="25">K. Kolesnichenko</collectorName>
leg.;
<collectionCode id="05FADAE3FFCFB33B9778FD87FB1BFD74" box="[1122,1225,517,543]" pageId="24" pageNumber="25">EDMSU</collectionCode>
</materialsCitation>
•
<materialsCitation id="D383487BFFCFB33B97F6FD87FDEAFD0D" ID-GBIF-Occurrence="3863902345" collectingDate="2009-06-04" collectionCode="EDMSU" collectorName="K. Kolesnichenko" country="Iran" elevation="2000" location="Dorrud" municipality="Qadamgan area" pageId="24" pageNumber="25" specimenCount="38" specimenCount-female="10" specimenCount-male="28" stateProvince="Rezavi" typeStatus="paratype">
<specimenCount id="75ED89AFFFCFB33B97F6FD87FA90FD4B" box="[1260,1346,517,544]" pageId="24" pageNumber="25" type="male">28 ♂♂</specimenCount>
(20 dissected),
<specimenCount id="75ED89AFFFCFB33B925AFDABFE42FD28" box="[320,400,553,579]" pageId="24" pageNumber="25" type="female">10 ♀♀</specimenCount>
(7 dissected);
<collectingMunicipality id="8330D85CFFCFB33B9121FDAAFD26FD28" box="[571,756,552,579]" pageId="24" pageNumber="25">Qadamgan area</collectingMunicipality>
,
<location id="663414FDFFCFB33B91E5FDAAFC86FD29" LSID="urn:lsid:plazi:treatment:EB42F330FFC0B33C913AFAF8FADBFD92:663414FDFFCFB33B91E5FDAAFC86FD29" box="[767,852,552,578]" country="Iran" municipality="Qadamgan area" name="Dorrud" pageId="24" pageNumber="25" stateProvince="Rezavi">Dorrud</location>
<taxonomicName id="A4EB39A5FFCFB33B9041FDABFBF9FD29" authority=", Gerina" authorityName="Gerina" box="[859,1067,552,579]" pageId="24" pageNumber="25" rank="variety" variety="vicinity">v. vicinity, Gerina</taxonomicName>
v.; alt.
<quantity id="A413EFC3FFCFB33B979AFDABFB0EFD28" box="[1152,1244,553,579]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="24" pageNumber="25" unit="m" value="2000.0">
<elevation id="E8C6A515FFCFB33B979AFDABFB0EFD28" box="[1152,1244,553,579]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="24" pageNumber="25" unit="m" value="2000.0">2000 m</elevation>
</quantity>
;
<date id="175564E6FFCFB33B97FDFDABFAA0FD28" box="[1255,1394,553,579]" pageId="24" pageNumber="25" value="2009-06-04">
<collectingDate id="07119D0EFFCFB33B97FDFDABFAA0FD28" box="[1255,1394,553,579]" pageId="24" pageNumber="25" value="2009-06-04">4 Jun. 2009</collectingDate>
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;
<collectorName id="CE1E27F0FFCFB33B93A7FDCEFE42FD0D" box="[189,400,588,614]" pageId="24" pageNumber="25">K. Kolesnichenko</collectorName>
leg.;
<collectionCode id="05FADAE3FFCFB33B92CBFDCEFDEAFD0D" box="[465,568,588,614]" pageId="24" pageNumber="25">EDMSU</collectionCode>
</materialsCitation>
•
<materialsCitation id="D383487BFFCFB33B9155FDCEFD25FDE2" ID-GBIF-Occurrence="3863902343" collectingDate="1999-06-07" collectionCode="EDMSU" collectorName="Eckweiler" country="Iran" elevation="2400" location="Kuh-e-Binalud" pageId="24" pageNumber="25" specimenCount="2" specimenCount-male="2" stateProvince="Rezavi" typeStatus="paratype">
<specimenCount id="75ED89AFFFCFB33B9155FDCEFD43FD0D" box="[591,657,588,614]" pageId="24" pageNumber="25" type="male">2 ♂♂</specimenCount>
(all dissected);
<location id="663414FDFFCFB33B9050FDCEFC2FFD0D" LSID="urn:lsid:plazi:treatment:EB42F330FFC0B33C913AFAF8FADBFD92:663414FDFFCFB33B9050FDCEFC2FFD0D" box="[842,1021,588,614]" country="Iran" name="Kuh-e-Binalud" pageId="24" pageNumber="25" stateProvince="Rezavi">Kuh-e-Binalud</location>
,
<locationDeviation id="72E65B04FFCFB33B9712FDC9FB24FD0D" box="[1032,1270,587,614]" location="Zoshk" pageId="24" pageNumber="25">
<quantity id="A413EFC3FFCFB33B9712FDC9FB83FD0D" box="[1032,1105,587,614]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="24" pageNumber="25" unit="km" value="15.0">15 km</quantity>
SW of Zoshk
</locationDeviation>
; alt.
<quantity id="A413EFC3FFCFB33B9637FDCEFECFFDE2" metricMagnitude="3" metricUnit="m" metricValue="2.4" metricValueMax="2.5" metricValueMin="2.3" pageId="24" pageNumber="25" unit="m" value="2400.0" valueMax="2500.0" valueMin="2300.0">
<elevation id="E8C6A515FFCFB33B9637FDCEFECFFDE2" metricMagnitude="3" metricUnit="m" metricValue="2.4" metricValueMax="2.5" metricValueMin="2.3" pageId="24" pageNumber="25" unit="m" value="2400.0" valueMax="2500.0" valueMin="2300.0">2300– 2500 m</elevation>
</quantity>
;
<date id="175564E6FFCFB33B9237FDEDFE12FDE2" box="[301,448,623,649]" pageId="24" pageNumber="25" value="1999-06-07">
<collectingDate id="07119D0EFFCFB33B9237FDEDFE12FDE2" box="[301,448,623,649]" pageId="24" pageNumber="25" value="1999-06-07">7 Jun. 1999</collectingDate>
</date>
;
<collectorName id="CE1E27F0FFCFB33B92D5FDEDFD94FDE2" box="[463,582,623,649]" pageId="24" pageNumber="25">Eckweiler</collectorName>
leg.;
<collectionCode id="05FADAE3FFCFB33B918AFDEDFD25FDE2" box="[656,759,623,649]" pageId="24" pageNumber="25">EDMSU</collectionCode>
</materialsCitation>
•
<materialsCitation id="D383487BFFCFB33B900DFDEDFE12FDA4" ID-GBIF-Occurrence="3863902325" collectingDate="2019-05-09" collectorName="A. A. Kotlobay &amp; A. A. Kotlobay." country="Iran" elevation="2015" latitude="36.154324" location="Kuh-e-Binalud Ridge" longLatPrecision="1" longitude="59.18167" pageId="24" pageNumber="25" specimenCount="11" specimenCount-male="11" stateProvince="Mazandaran" typeStatus="paratype">
<specimenCount id="75ED89AFFFCFB33B900DFDEDFCBFFDE2" box="[791,877,623,649]" pageId="24" pageNumber="25" type="male">11 ♂♂</specimenCount>
;
<locationDeviation id="72E65B04FFCFB33B9067FDEDFB8CFDE2" box="[893,1118,623,649]" location="Gerine" pageId="24" pageNumber="25">
<quantity id="A413EFC3FFCFB33B9067FDEDFC6EFDE2" box="[893,956,623,649]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="24" pageNumber="25" unit="km" value="6.0">6 km</quantity>
N of Gerine
</locationDeviation>
,
<location id="663414FDFFCFB33B9777FDEDFAA0FDE2" LSID="urn:lsid:plazi:treatment:EB42F330FFC0B33C913AFAF8FADBFD92:663414FDFFCFB33B9777FDEDFAA0FDE2" box="[1133,1394,623,649]" country="Iran" latitude="36.154324" longLatPrecision="1" longitude="59.18167" name="Kuh-e-Binalud Ridge" pageId="24" pageNumber="25" stateProvince="Mazandaran">Kuh-e-Binalud Ridge</location>
;
<geoCoordinate id="06DF24E1FFCFB33B93A7FD12FEA8FDC6" box="[189,378,656,685]" degrees="36" direction="north" minutes="09" orientation="latitude" pageId="24" pageNumber="25" precision="1" seconds="15.56" value="36.154324">36°09′15.56″ N</geoCoordinate>
,
<geoCoordinate id="06DF24E1FFCFB33B9292FD12FD90FDC6" box="[392,578,656,685]" degrees="59" direction="east" minutes="10" orientation="longitude" pageId="24" pageNumber="25" precision="1" seconds="54.01" value="59.18167">59°10′54.01″ E</geoCoordinate>
; alt.
<quantity id="A413EFC3FFCFB33B9165FD10FD0FFDC6" box="[639,733,658,685]" metricMagnitude="3" metricUnit="m" metricValue="2.0149999999999997" pageId="24" pageNumber="25" unit="m" value="2015.0">
<elevation id="E8C6A515FFCFB33B9165FD10FD0FFDC6" box="[639,733,658,685]" metricMagnitude="3" metricUnit="m" metricValue="2.0149999999999997" pageId="24" pageNumber="25" unit="m" value="2015.0">2015 m</elevation>
</quantity>
;
<date id="175564E6FFCFB33B91F6FD10FC50FDC7" box="[748,898,658,685]" pageId="24" pageNumber="25" value="2019-05-09">
<collectingDate id="07119D0EFFCFB33B91F6FD10FC50FDC7" box="[748,898,658,685]" pageId="24" pageNumber="25" value="2019-05-09">9 May 2019</collectingDate>
</date>
;
<collectorName id="CE1E27F0FFCFB33B9095FD10FBEDFDC7" box="[911,1087,658,684]" pageId="24" pageNumber="25">A.A. Kotlobay</collectorName>
leg.; coll.
<collectorName id="CE1E27F0FFCFB33B97DBFD10FAA5FDC7" box="[1217,1399,658,684]" pageId="24" pageNumber="25">A.A. Kotlobay.</collectorName>
–
<emphasis id="519F9E34FFCFB33B93CEFD37FE12FDA4" bold="true" box="[212,448,693,719]" pageId="24" pageNumber="25">
<collectingRegion id="A12F8CC4FFCFB33B93CEFD37FE12FDA4" box="[212,448,693,719]" country="Iran" name="Mazandaran" pageId="24" pageNumber="25">Mazandaran Prov.</collectingRegion>
</emphasis>
</materialsCitation>
•
<materialsCitation id="D383487BFFCFB33B92C1FD37FEF6FC7D" ID-GBIF-Occurrence="3863902320" collectingDate="2018-05-02" collectionCode="EDMSU" collectorName="K. Kolesnichenko" country="Iran" county="Albors Mts" elevation="2000" location="Sultan Kuh Mt." municipality="Foulad Mahhaleh" pageId="24" pageNumber="25" specimenCount="1" specimenCount-female="1" stateProvince="Rezavi" typeStatus="paratype">
<specimenCount id="75ED89AFFFCFB33B92C1FD37FDDAFDBB" box="[475,520,693,720]" pageId="24" pageNumber="25" type="female">1 ♀</specimenCount>
(dissected); S macroslopes of
<collectingCounty id="8A353AAAFFCFB33B906EFD37FC2FFDBB" box="[884,1021,693,720]" pageId="24" pageNumber="25">Albors Mts</collectingCounty>
,
<locationDeviation id="72E65B04FFCFB33B9710FD37FB0EFDA4" box="[1034,1244,693,719]" location="Sari" pageId="24" pageNumber="25">
<quantity id="A413EFC3FFCFB33B9710FD37FB87FDA4" box="[1034,1109,693,719]" metricMagnitude="4" metricUnit="m" metricValue="8.0" pageId="24" pageNumber="25" unit="km" value="80.0">80 km</quantity>
SE of Sari
</locationDeviation>
,
<quantity id="A413EFC3FFCFB33B97F3FD37FAF7FDA4" box="[1257,1317,693,720]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="24" pageNumber="25" unit="km" value="5.0">5 km</quantity>
NE of
<collectingMunicipality id="8330D85CFFCFB33B93A7FD5AFE5CFD99" box="[189,398,728,754]" pageId="24" pageNumber="25">Foulad Mahhaleh</collectingMunicipality>
v., E slopes of
<location id="663414FDFFCFB33B914BFD5AFCC2FD98" LSID="urn:lsid:plazi:treatment:EB42F330FFC0B33C913AFAF8FADBFD92:663414FDFFCFB33B914BFD5AFCC2FD98" box="[593,784,728,755]" country="Iran" county="Albors Mts" municipality="Foulad Mahhaleh" name="Sultan Kuh Mt." pageId="24" pageNumber="25" stateProvince="Rezavi">Sultan Kuh Mt.</location>
; alt.
<quantity id="A413EFC3FFCFB33B9054FD5BFC7FFD98" box="[846,941,729,755]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="24" pageNumber="25" unit="m" value="2000.0">
<elevation id="E8C6A515FFCFB33B9054FD5BFC7FFD98" box="[846,941,729,755]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="24" pageNumber="25" unit="m" value="2000.0">2000 m</elevation>
</quantity>
;
<date id="175564E6FFCFB33B90A6FD5BFB81FD98" box="[956,1107,729,755]" pageId="24" pageNumber="25" value="2018-05-02">
<collectingDate id="07119D0EFFCFB33B90A6FD5BFB81FD98" box="[956,1107,729,755]" pageId="24" pageNumber="25" value="2018-05-02">2 May 2018</collectingDate>
</date>
;
<collectorName id="CE1E27F0FFCFB33B9778FD5BFAEBFD99" box="[1122,1337,728,755]" pageId="24" pageNumber="25">K. Kolesnichenko</collectorName>
leg.;
<collectionCode id="05FADAE3FFCFB33B93A7FD7EFEF6FC7D" box="[189,292,764,790]" pageId="24" pageNumber="25">EDMSU</collectionCode>
</materialsCitation>
•
<materialsCitation id="D383487BFFCFB33B9226FD7EFAA1FC7D" ID-GBIF-Occurrence="3863902348" box="[316,1395,764,790]" collectingDate="2019-05-12" collectionCode="EDMSU" collectorName="A. Kotlobay" country="Iran" county="Albors Mts" elevation="2000" location="Sultan Kuh Mt." municipality="Foulad Mahhaleh" pageId="24" pageNumber="25" specimenCount="1" specimenCount-female="1" stateProvince="Rezavi" typeStatus="paratype">
<specimenCount id="75ED89AFFFCFB33B9226FD7EFEB5FC7D" box="[316,359,764,790]" pageId="24" pageNumber="25" type="female">1 ♀</specimenCount>
(dissected); same locality as for preceding but
<date id="175564E6FFCFB33B9094FD7EFBFFFC7D" box="[910,1069,764,790]" pageId="24" pageNumber="25" value="2019-05-12">
<collectingDate id="07119D0EFFCFB33B9094FD7EFBFFFC7D" box="[910,1069,764,790]" pageId="24" pageNumber="25" value="2019-05-12">12 May 2019</collectingDate>
</date>
;
<collectorName id="CE1E27F0FFCFB33B972DFD7EFB15FC7D" box="[1079,1223,764,790]" pageId="24" pageNumber="25">A. Kotlobay</collectorName>
leg.;
<collectionCode id="05FADAE3FFCFB33B9612FD7EFAA1FC7D" box="[1288,1395,764,790]" pageId="24" pageNumber="25">EDMSU</collectionCode>
</materialsCitation>
. –
<materialsCitation id="D383487BFFCFB33B93CDFC9DFEF6FC37" ID-GBIF-Occurrence="3863902349" collectingDate="2006-04-28" collectionCode="EDMSU" collectorName="K. Kolesnichenko" country="Iran" location="35 km N of Birjant" pageId="24" pageNumber="25" specimenCount="1" specimenCount-male="1" stateProvince="Horossan Prov." typeStatus="paratype">
<emphasis id="519F9E34FFCFB33B93CDFC9DFE49FC52" bold="true" box="[215,411,799,825]" pageId="24" pageNumber="25">
<collectingRegion id="A12F8CC4FFCFB33B93CDFC9DFE49FC52" box="[215,411,799,825]" pageId="24" pageNumber="25">Horossan Prov.</collectingRegion>
</emphasis>
•
<specimenCount id="75ED89AFFFCFB33B92A6FC9DFE3FFC52" box="[444,493,799,825]" pageId="24" pageNumber="25" type="male">1 ♂</specimenCount>
(dissected);
<location id="663414FDFFCFB33B9191FC9DFCA9FC52" LSID="urn:lsid:plazi:treatment:EB42F330FFC0B33C913AFAF8FADBFD92:663414FDFFCFB33B9191FC9DFCA9FC52" box="[651,891,799,826]" country="Iran" name="35 km N of Birjant" pageId="24" pageNumber="25" stateProvince="Horossan Prov.">
<quantity id="A413EFC3FFCFB33B9191FC9DFD0BFC52" box="[651,729,799,826]" metricMagnitude="4" metricUnit="m" metricValue="3.5" pageId="24" pageNumber="25" unit="km" value="35.0">35 km</quantity>
N of Birjant
</location>
t.;
<date id="175564E6FFCFB33B90B1FC9DFB9DFC52" box="[939,1103,799,825]" pageId="24" pageNumber="25" value="2006-04-28">
<collectingDate id="07119D0EFFCFB33B90B1FC9DFB9DFC52" box="[939,1103,799,825]" pageId="24" pageNumber="25" value="2006-04-28">28 Apr. 2006</collectingDate>
</date>
;
<collectorName id="CE1E27F0FFCFB33B9745FC9DFAEAFC52" box="[1119,1336,799,825]" pageId="24" pageNumber="25">K. Kolesnichenko</collectorName>
leg.;
<collectionCode id="05FADAE3FFCFB33B93A7FCC0FEF6FC37" box="[189,292,834,860]" pageId="24" pageNumber="25">EDMSU</collectionCode>
</materialsCitation>
•
<materialsCitation id="D383487BFFCFB33B925EFCC0FE1FFC14" ID-GBIF-Occurrence="3863902309" collectingDate="2005-04-27" collectionCode="EDMSU" collectorName="R. Nazarov" country="Iran" elevation="1500" location="Sedeh" pageId="24" pageNumber="25" specimenCount="1" specimenCount-male="1" stateProvince="Horossan Prov." typeStatus="paratype">
<specimenCount id="75ED89AFFFCFB33B925EFCC0FEA1FC36" box="[324,371,834,861]" pageId="24" pageNumber="25" type="male">1 ♂</specimenCount>
(dissected);
<locationDeviation id="72E65B04FFCFB33B910BFCC0FD2CFC37" box="[529,766,834,861]" location="Birjant" pageId="24" pageNumber="25">
<quantity id="A413EFC3FFCFB33B910BFCC0FD8CFC37" box="[529,606,834,861]" metricMagnitude="4" metricUnit="m" metricValue="7.5" pageId="24" pageNumber="25" unit="km" value="75.0">75 km</quantity>
N of Birjant
</locationDeviation>
t.,
<location id="663414FDFFCFB33B9031FCC0FCA1FC37" LSID="urn:lsid:plazi:treatment:EB42F330FFC0B33C913AFAF8FADBFD92:663414FDFFCFB33B9031FCC0FCA1FC37" box="[811,883,834,860]" country="Iran" name="Sedeh" pageId="24" pageNumber="25" stateProvince="Horossan Prov.">Sedeh</location>
<taxonomicName id="A4EB39A5FFCFB33B9064FCC1FC28FC37" box="[894,1018,834,861]" pageId="24" pageNumber="25" rank="variety" variety="vicinity">v. vicinity</taxonomicName>
; alt.
<quantity id="A413EFC3FFCFB33B9723FCC0FB4BFC36" box="[1081,1177,834,861]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="24" pageNumber="25" unit="m" value="1500.0">
<elevation id="E8C6A515FFCFB33B9723FCC0FB4BFC36" box="[1081,1177,834,861]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="24" pageNumber="25" unit="m" value="1500.0">1500 m</elevation>
</quantity>
;
<date id="175564E6FFCFB33B97B3FCC0FA9EFC36" box="[1193,1356,834,861]" pageId="24" pageNumber="25" value="2005-04-27">
<collectingDate id="07119D0EFFCFB33B97B3FCC0FA9EFC36" box="[1193,1356,834,861]" pageId="24" pageNumber="25" value="2005-04-27">27 Apr. 2005</collectingDate>
</date>
;
<collectorName id="CE1E27F0FFCFB33B9646FCC1FEF2FCEB" pageId="24" pageNumber="25">R. Nazarov</collectorName>
leg.;
<collectionCode id="05FADAE3FFCFB33B9278FCE7FE1FFC14" box="[354,461,869,895]" pageId="24" pageNumber="25">EDMSU</collectionCode>
</materialsCitation>
.
</paragraph>
<caption id="379412AEFFCFB33B93A7F8E1FBB1F8AF" ID-DOI="http://doi.org/10.5281/zenodo.6839883" ID-Zenodo-Dep="6839883" httpUri="https://zenodo.org/record/6839883/files/figure.png" pageId="24" pageNumber="25" startId="24.[189,232,1891,1917]" targetBox="[194,1395,1026,1847]" targetPageId="24">
<paragraph id="63544226FFCFB33B93A7F8E1FBB1F8AF" blockId="24.[189,1399,1891,1989]" pageId="24" pageNumber="25">
<emphasis id="519F9E34FFCFB33B93A7F8E1FEC9F816" bold="true" box="[189,283,1891,1917]" pageId="24" pageNumber="25">Fig. 15.</emphasis>
Adults of
<taxonomicName id="A4EB39A5FFCFB33B9281F8E6FD2EF815" authority="Kolesnichenko &amp; Kotlobay, 2022" authorityName="Kolesnichenko &amp; Kotlobay" authorityYear="2022" box="[411,764,1892,1918]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="subSpecies" species="timandra" status="subsp. nov." subSpecies="binaludica">
<emphasis id="519F9E34FFCFB33B9281F8E6FD2EF815" box="[411,764,1892,1918]" italics="true" pageId="24" pageNumber="25">Melitaea timandra binaludica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4AAC234FFFCFB33B901EF8E6FC5AF815" box="[772,904,1892,1918]" pageId="24" pageNumber="25" rank="subSpecies">subsp. nov.</taxonomicNameLabel>
<emphasis id="519F9E34FFCFB33B908AF8E1FC18F815" bold="true" box="[912,970,1891,1918]" pageId="24" pageNumber="25">
A
<emphasis id="519F9E34FFCFB33B90BFF8E6FC66F815" bold="true" box="[933,948,1892,1918]" italics="true" pageId="24" pageNumber="25">–</emphasis>
H
</emphasis>
. UPS.
<emphasis id="519F9E34FFCFB33B973AF8E6FB99F815" bold="true" box="[1056,1099,1892,1918]" pageId="24" pageNumber="25">
I
<emphasis id="519F9E34FFCFB33B9730F8E6FBEBF815" bold="true" box="[1066,1081,1892,1918]" italics="true" pageId="24" pageNumber="25">–</emphasis>
P
</emphasis>
. UNS.
<emphasis id="519F9E34FFCFB33B97BEF8E1FB06F815" bold="true" box="[1188,1236,1891,1918]" pageId="24" pageNumber="25">A, I</emphasis>
. Holotype, ♂ (SDM).
<emphasis id="519F9E34FFCFB33B923AF805FE8BF8CA" bold="true" box="[288,345,1927,1953]" pageId="24" pageNumber="25">B–H</emphasis>
,
<emphasis id="519F9E34FFCFB33B9270F805FE4BF8CA" bold="true" box="[362,409,1927,1953]" pageId="24" pageNumber="25">J–P</emphasis>
. Paratypes (EDMSU).
<emphasis id="519F9E34FFCFB33B91AAF805FCF4F8CA" bold="true" box="[688,806,1927,1953]" pageId="24" pageNumber="25">A–D, I–L</emphasis>
. ♂, Iran, Rezavi Khorassan Prov., Kuh-e Binalud Mts, Dorrud
<taxonomicName id="A4EB39A5FFCFB33B924FF829FE1EF8AF" box="[341,460,1962,1989]" pageId="24" pageNumber="25" rank="variety" variety="vicinity">v. vicinity</taxonomicName>
, alt. 2430 m.
<emphasis id="519F9E34FFCFB33B9171F828FD3DF8AF" bold="true" box="[619,751,1962,1988]" pageId="24" pageNumber="25">E–H, M–P</emphasis>
. ♀, same data as for preceding.
</paragraph>
</caption>
<caption id="379412AEFFCEB33A93A7F956FC02F88E" ID-DOI="http://doi.org/10.5281/zenodo.6839885" ID-Zenodo-Dep="6839885" httpUri="https://zenodo.org/record/6839885/files/figure.png" pageId="25" pageNumber="26" startId="25.[189,232,1748,1774]" targetBox="[311,1284,274,1694]" targetPageId="25">
<paragraph id="63544226FFCEB33A93A7F956FC02F88E" blockId="25.[189,1399,1748,2021]" pageId="25" pageNumber="26">
<emphasis id="519F9E34FFCEB33A93A7F956FECCF985" bold="true" box="[189,286,1748,1774]" pageId="25" pageNumber="26">Fig. 16.</emphasis>
Male genitalia (without aedeagus) in dorsal projection.
<emphasis id="519F9E34FFCEB33A90D7F956FBD4F985" bold="true" box="[973,1030,1748,1774]" pageId="25" pageNumber="26">A–C</emphasis>
.
<taxonomicName id="A4EB39A5FFCEB33A9702F956FADFF985" authority="Kolesnichenko &amp; Kotlobay, 2022" authorityName="Kolesnichenko &amp; Kotlobay" authorityYear="2022" box="[1048,1293,1748,1774]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="shahvarica" status="sp. nov.">
<emphasis id="519F9E34FFCEB33A9702F956FADFF985" box="[1048,1293,1748,1774]" italics="true" pageId="25" pageNumber="26">Melitaea shahvarica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4AAC234FFFCEB33A9603F957FAA5F984" box="[1305,1399,1749,1775]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="519F9E34FFCEB33A93A7F975FF2AF87A" bold="true" box="[189,248,1783,1809]" pageId="25" pageNumber="26">D–H</emphasis>
.
<taxonomicName id="A4EB39A5FFCEB33A9213F97AFD86F87A" authority="Higgins, 1940" authorityName="Higgins" authorityYear="1940" baseAuthorityName="Higgins" baseAuthorityYear="1940" box="[265,596,1783,1809]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="mimetica">
<emphasis id="519F9E34FFCEB33A9213F97AFE4DF87A" box="[265,415,1784,1809]" italics="true" pageId="25" pageNumber="26">M. mimetica</emphasis>
<bibRefCitation id="077A3FD7FFCEB33A92B2F975FD86F87A" author="Higgins L. G." box="[424,596,1783,1809]" pageId="25" pageNumber="26" pagination="51 - 53" refId="ref29676" refString="Higgins L. G. 1940. A new species and two new subspecies of Melitaea (Lep. Nymphalidae). The Entomologist 73: 51 - 53." type="journal article" year="1940">Higgins, 1940</bibRefCitation>
</taxonomicName>
.
<emphasis id="519F9E34FFCEB33A9178F975FDBFF87A" bold="true" box="[610,621,1783,1809]" pageId="25" pageNumber="26">I</emphasis>
.
<taxonomicName id="A4EB39A5FFCEB33A9165F97AFC56F87A" authority="Evans, 1932" authorityName="Evans" authorityYear="1932" box="[639,900,1783,1810]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="lutko">
<emphasis id="519F9E34FFCEB33A9165F97AFD31F87A" box="[639,739,1783,1809]" italics="true" pageId="25" pageNumber="26">M. lutko</emphasis>
<bibRefCitation id="077A3FD7FFCEB33A91F7F97AFC56F87A" author="Evans W. H." box="[749,900,1783,1810]" pageId="25" pageNumber="26" refId="ref29423" refString="Evans W. H. 1932. The Identification of Indian Butterflies. The Bombay Natural History Society, Diocesan Press, Madras." type="book" year="1932">Evans, 1932</bibRefCitation>
</taxonomicName>
.
<emphasis id="519F9E34FFCEB33A908BF975FC19F87A" bold="true" box="[913,971,1783,1809]" pageId="25" pageNumber="26">A–C</emphasis>
. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mts, alt. 2200–2400 m.
<emphasis id="519F9E34FFCEB33A9001F899FCE2F85E" bold="true" box="[795,816,1819,1845]" pageId="25" pageNumber="26">D</emphasis>
. Pakistan, Balochistan, Quetta, Urak, alt. 2400– 2700 m.
<emphasis id="519F9E34FFCEB33A9229F8BCFE95F833" bold="true" box="[307,327,1854,1880]" pageId="25" pageNumber="26">E</emphasis>
. Afghanistan, Bamian Prov., 10 km S of Bamian t., Hushkak
<taxonomicName id="A4EB39A5FFCEB33A9770F8BCFB35F833" box="[1130,1255,1854,1880]" pageId="25" pageNumber="26" rank="variety" variety="vicinity">v. vicinity</taxonomicName>
, alt, 2700– 2800 m.
<emphasis id="519F9E34FFCEB33A9233F8E3FEEBF810" bold="true" box="[297,313,1889,1915]" pageId="25" pageNumber="26">F</emphasis>
. Central Afghanistan, Ghor Prov., 17 km E of Changcharan, 15 km S of Bandi-Ali, Gazak Mts, alt. 2400 m.
<emphasis id="519F9E34FFCEB33A9283F806FE62F8F5" bold="true" box="[409,432,1924,1950]" pageId="25" pageNumber="26">G</emphasis>
. Afghanistan, Bamian Prov., Punjub Distr., 10 km NE of Varas v., alt. 2400 m.
<emphasis id="519F9E34FFCEB33A93A7F825FF06F8AA" bold="true" box="[189,212,1959,1985]" pageId="25" pageNumber="26">H</emphasis>
. Afghanistan, Ghor Prov., 16 km E of Changcharan, Bandi-Ali
<taxonomicName id="A4EB39A5FFCEB33A90C8F82AFB98F8AA" box="[978,1098,1959,1986]" pageId="25" pageNumber="26" rank="variety" variety="vicinity">v. vicinity</taxonomicName>
, alt. 2400 m.
<emphasis id="519F9E34FFCEB33A97E8F825FB2EF8AA" bold="true" box="[1266,1276,1959,1985]" pageId="25" pageNumber="26">I</emphasis>
. Pakistan, Chitral, Chaghbini CGNP [Chitral Gol National Park], alt. 2700 m.
</paragraph>
</caption>
<caption id="379412AEFFCDB33993A7F8B5FB11F8B4" ID-DOI="http://doi.org/10.5281/zenodo.6839887" ID-Zenodo-Dep="6839887" httpUri="https://zenodo.org/record/6839887/files/figure.png" pageId="26" pageNumber="27" startId="26.[189,232,1847,1873]" targetBox="[276,1315,271,1797]" targetPageId="26">
<paragraph id="63544226FFCDB33993A7F8B5FB11F8B4" blockId="26.[187,1399,1847,2015]" pageId="26" pageNumber="27">
<emphasis id="519F9E34FFCDB33993A7F8B5FECDF83A" bold="true" box="[189,287,1847,1873]" pageId="26" pageNumber="27">Fig. 17.</emphasis>
Male genitalia (without aedeagus) in dorsal projection.
<emphasis id="519F9E34FFCDB33990C9F8B5FBDEF83A" bold="true" box="[979,1036,1847,1873]" pageId="26" pageNumber="27">A–C</emphasis>
.
<taxonomicName id="A4EB39A5FFCDB339973AF8B5FDFBF81E" authority="Coutsis &amp; van Oorschot, 2014" authorityName="Coutsis &amp; van Oorschot" authorityYear="2014" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="subSpecies" species="timandra" subSpecies="timandra">
<emphasis id="519F9E34FFCDB339973AF8B5FAA5F83A" box="[1056,1399,1847,1873]" italics="true" pageId="26" pageNumber="27">Melitaea timandra timandra</emphasis>
Coutsis &amp;
<bibRefCitation id="077A3FD7FFCDB3399227F8D9FDFBF81E" author="Van Oorschot H. &amp; Coutsis J. G." box="[317,553,1882,1909]" pageId="26" pageNumber="27" refId="ref31081" refString="Van Oorschot H. &amp; Coutsis J. G. 2014. The Genus Melitaea Fabricius, 1807 (Lepidoptera, Nymphalidae, Nymphalinae). Taxonomy and Systematics with Special Reference to the Male Genitalia. Tshikolovets publication, Pardubice." type="book" year="2014">van Oorschot, 2014</bibRefCitation>
</taxonomicName>
.
<emphasis id="519F9E34FFCDB339912DF8D9FDB7F81E" bold="true" box="[567,613,1883,1909]" pageId="26" pageNumber="27">D–I</emphasis>
.
<taxonomicName id="A4EB39A5FFCDB339916FF8D9FC5CF81F" authority="Kolesnichenko &amp; Kotlobay, 2022" authorityName="Kolesnichenko &amp; Kotlobay" authorityYear="2022" box="[629,910,1882,1908]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="subSpecies" species="timandra" status="subsp. nov." subSpecies="binaludica">
<emphasis id="519F9E34FFCDB339916FF8D9FC5CF81F" box="[629,910,1882,1908]" italics="true" pageId="26" pageNumber="27">M. timandra binaludica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4AAC234FFFCDB3399082F8D8FBCFF81E" box="[920,1053,1882,1909]" pageId="26" pageNumber="27" rank="subSpecies">subsp. nov.</taxonomicNameLabel>
<emphasis id="519F9E34FFCDB339973FF8D8FB8FF81F" bold="true" box="[1061,1117,1882,1908]" pageId="26" pageNumber="27">A–C</emphasis>
. S Turkmenistan, SaryYazy, alt. 300 m.
<emphasis id="519F9E34FFCDB339929BF8FCFE6BF8F3" bold="true" box="[385,441,1918,1944]" pageId="26" pageNumber="27">
D
<emphasis id="519F9E34FFCDB3399282F8FCFE75F8F3" bold="true" box="[408,423,1918,1944]" italics="true" pageId="26" pageNumber="27">–</emphasis>
F
</emphasis>
. Iran, Rezavi Khorassan Prov., Kuh-e-Binalud Mts, Dorrud
<taxonomicName id="A4EB39A5FFCDB3399775F8FCFB36F8F3" box="[1135,1252,1918,1944]" pageId="26" pageNumber="27" rank="variety" variety="vicinity">v. vicinity</taxonomicName>
, alt. 2430 m.
<emphasis id="519F9E34FFCDB33993A1F823FF00F8D0" bold="true" box="[187,210,1953,1979]" pageId="26" pageNumber="27">G</emphasis>
. Iran, Horossan Prov., 35 km N of Birjant t.
<emphasis id="519F9E34FFCDB33991CCF823FD3FF8D0" bold="true" box="[726,749,1953,1979]" pageId="26" pageNumber="27">H</emphasis>
. Afghanistan, Bamian Prov., Band-e-Amir, alt. 3200 m.
<emphasis id="519F9E34FFCDB33993A6F846FF15F8B5" bold="true" box="[188,199,1988,2014]" pageId="26" pageNumber="27">I</emphasis>
. Central Afghanistan, Bamian Prov., Band-e-Amir, Dzhudoi-Kvak Gorge, alt. 3200 m.
</paragraph>
</caption>
<paragraph id="63544226FFCCB33893A7FE8BFC6FFEE6" blockId="27.[189,1398,265,397]" pageId="27" pageNumber="28">
<materialsCitation id="D383487BFFCCB33893A7FE8BFCBFFE2D" ID-GBIF-Occurrence="3863902351" collectingDate="2009-07-01" collectingDateMax="2009-07-02" collectingDateMin="2009-07-01" collectionCode="EDMSU" collectorName="O. Pak" country="Afghanistan" elevation="3200" location="Dzhudoi-Kvak Gorge" municipality="Band-e-Amir" pageId="27" pageNumber="28" specimenCount="5" specimenCount-female="1" specimenCount-male="4" stateProvince="Bamyan">
<collectingCountry id="1BFC02B6FFCCB33893A7FE8BFE59FE48" box="[189,395,265,291]" name="Afghanistan" pageId="27" pageNumber="28">AFGHANISTAN</collectingCountry>
–
<emphasis id="519F9E34FFCCB33892BCFE8BFD82FE48" bold="true" box="[422,592,265,291]" pageId="27" pageNumber="28">
<collectingRegion id="A12F8CC4FFCCB33892BCFE8BFD82FE48" box="[422,592,265,291]" country="Afghanistan" name="Bamyan" pageId="27" pageNumber="28">Bamian Prov.</collectingRegion>
</emphasis>
•
<specimenCount id="75ED89AFFFCCB3389172FE8BFD7BFE48" box="[616,681,265,291]" pageId="27" pageNumber="28" type="male">4 ♂♂</specimenCount>
(all dissected),
<specimenCount id="75ED89AFFFCCB338907AFE8BFC59FE48" box="[864,907,265,291]" pageId="27" pageNumber="28" type="female">1 ♀</specimenCount>
(dissected);
<collectingMunicipality id="8330D85CFFCCB3389705FE8BFB6FFE48" box="[1055,1213,265,291]" pageId="27" pageNumber="28">Band-e-Amir</collectingMunicipality>
,
<location id="663414FDFFCCB33897D2FE8BFED8FE2D" LSID="urn:lsid:plazi:treatment:EB42F330FFC0B33C913AFAF8FADBFD92:663414FDFFCCB33897D2FE8BFED8FE2D" country="Afghanistan" municipality="Band-e-Amir" name="Dzhudoi-Kvak Gorge" pageId="27" pageNumber="28" stateProvince="Bamyan">Dzhudoi-Kvak Gorge</location>
; alt.
<quantity id="A413EFC3FFCCB338925DFEAEFE74FE2C" box="[327,422,300,327]" metricMagnitude="3" metricUnit="m" metricValue="3.2" pageId="27" pageNumber="28" unit="m" value="3200.0">
<elevation id="E8C6A515FFCCB338925DFEAEFE74FE2C" box="[327,422,300,327]" metricMagnitude="3" metricUnit="m" metricValue="3.2" pageId="27" pageNumber="28" unit="m" value="3200.0">3200 m</elevation>
</quantity>
;
<date id="175564E6FFCCB33892AFFEAEFD8EFE2D" box="[437,604,300,326]" pageId="27" pageNumber="28" value="2009-07-01" valueMax="2009-07-02" valueMin="2009-07-01">
<collectingDate id="07119D0EFFCCB33892AFFEAEFD8EFE2D" box="[437,604,300,326]" pageId="27" pageNumber="28" value="2009-07-01" valueMax="2009-07-02" valueMin="2009-07-01">1–2 Jul. 2009</collectingDate>
</date>
;
<collectorName id="CE1E27F0FFCCB3389171FEAEFD6CFE2D" box="[619,702,300,326]" pageId="27" pageNumber="28">O. Pak</collectorName>
leg.;
<collectionCode id="05FADAE3FFCCB338901CFEAEFCBFFE2D" box="[774,877,300,326]" pageId="27" pageNumber="28">EDMSU</collectionCode>
</materialsCitation>
•
<materialsCitation id="D383487BFFCCB3389091FEAEFC71FE02" ID-GBIF-Occurrence="3863902355" collectingDate="1960-06-09" collectingDateMax="1960-06-10" collectingDateMin="1960-06-09" collectionCode="EDMSU" collectorName="Colin Wyatt" country="Afghanistan" elevation="3109" location="Band-i-Amir" municipality="Hazarajat" pageId="27" pageNumber="28" specimenCount="1" specimenCount-male="1" stateProvince="Bamyan">
<specimenCount id="75ED89AFFFCCB3389091FEAEFC68FE2C" box="[907,954,300,327]" pageId="27" pageNumber="28" type="male">1 ♂</specimenCount>
(dissected);
<collectingMunicipality id="8330D85CFFCCB338974FFEAEFB1BFE2D" box="[1109,1225,300,326]" pageId="27" pageNumber="28">Hazarajat</collectingMunicipality>
,
<location id="663414FDFFCCB33897CDFEAEFAA0FE2D" LSID="urn:lsid:plazi:treatment:EB42F330FFC0B33C913AFAF8FADBFD92:663414FDFFCCB33897CDFEAEFAA0FE2D" box="[1239,1394,300,326]" country="Afghanistan" municipality="Hazarajat" name="Band-i-Amir" pageId="27" pageNumber="28" stateProvince="Bamyan">Band-i-Amir</location>
; alt.
<quantity id="A413EFC3FFCCB33893F0FECDFE4AFE02" box="[234,408,335,361]" metricMagnitude="3" metricUnit="m" metricValue="3.10896" metricValueMax="3.3528" metricValueMin="2.86512" pageId="27" pageNumber="28" unit="ft" value="10200.0" valueMax="11000.0" valueMin="9400.0">
<elevation id="E8C6A515FFCCB33893F0FECDFE4AFE02" box="[234,408,335,361]" metricMagnitude="3" metricUnit="m" metricValue="3.10896" metricValueMax="3.3528" metricValueMin="2.86512" pageId="27" pageNumber="28" unit="ft" value="10200.0" valueMax="11000.0" valueMin="9400.0">9400–11000 ft</elevation>
</quantity>
;
<date id="175564E6FFCCB33892BFFECDFD8CFE02" box="[421,606,335,362]" pageId="27" pageNumber="28" value="1960-06-09" valueMax="1960-06-10" valueMin="1960-06-09">
<collectingDate id="07119D0EFFCCB33892BFFECDFD8CFE02" box="[421,606,335,362]" pageId="27" pageNumber="28" value="1960-06-09" valueMax="1960-06-10" valueMin="1960-06-09">9–10 Jun. 1960</collectingDate>
</date>
;
<collectorName id="CE1E27F0FFCCB3389170FECDFD2BFE01" box="[618,761,335,362]" pageId="27" pageNumber="28">Colin Wyatt</collectorName>
leg.;
<collectionCode id="05FADAE3FFCCB3389026FECDFC71FE02" box="[828,931,335,361]" pageId="27" pageNumber="28">EDMSU</collectionCode>
</materialsCitation>
•
<materialsCitation id="D383487BFFCCB33890A7FECDFC68FEE6" ID-GBIF-Occurrence="3863902321" collectingDate="2009-07-05" collectionCode="EDMSU" collectorName="I. Pljushtch" country="Afghanistan" elevation="3200" location="Band-e-Amir" pageId="27" pageNumber="28" specimenCount="6" specimenCount-female="1" specimenCount-male="5" stateProvince="Bamyan">
<specimenCount id="75ED89AFFFCCB33890A7FECDFC2DFE01" box="[957,1023,335,362]" pageId="27" pageNumber="28" type="male">5 ♂♂</specimenCount>
(all dissected),
<specimenCount id="75ED89AFFFCCB33897A1FECDFB35FE01" box="[1211,1255,335,362]" pageId="27" pageNumber="28" type="female">1 ♀</specimenCount>
(dissected);
<location id="663414FDFFCCB33893A7FEF0FE8FFEE7" LSID="urn:lsid:plazi:treatment:EB42F330FFC0B33C913AFAF8FADBFD92:663414FDFFCCB33893A7FEF0FE8FFEE7" box="[189,349,370,396]" country="Afghanistan" name="Band-e-Amir" pageId="27" pageNumber="28" stateProvince="Bamyan">Band-e-Amir</location>
; alt.
<quantity id="A413EFC3FFCCB338928FFEF1FE23FEE6" box="[405,497,371,397]" metricMagnitude="3" metricUnit="m" metricValue="3.2" pageId="27" pageNumber="28" unit="m" value="3200.0">
<elevation id="E8C6A515FFCCB338928FFEF1FE23FEE6" box="[405,497,371,397]" metricMagnitude="3" metricUnit="m" metricValue="3.2" pageId="27" pageNumber="28" unit="m" value="3200.0">3200 m</elevation>
</quantity>
;
<date id="175564E6FFCCB33892E7FEF0FD50FEE6" box="[509,642,370,397]" pageId="27" pageNumber="28" value="2009-07-05">
<collectingDate id="07119D0EFFCCB33892E7FEF0FD50FEE6" box="[509,642,370,397]" pageId="27" pageNumber="28" value="2009-07-05">5 Jul. 2009</collectingDate>
</date>
;
<collectorName id="CE1E27F0FFCCB3389194FEF1FCC6FEE7" box="[654,788,370,397]" pageId="27" pageNumber="28">I. Pljushtch</collectorName>
leg;
<collectionCode id="05FADAE3FFCCB3389055FEF1FC68FEE6" box="[847,954,371,397]" pageId="27" pageNumber="28">EDMSU</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="2BF111ADFFCCB33D93A7FE36FB3DFEDB" lastPageId="30" lastPageNumber="31" pageId="27" pageNumber="28" type="description">
<paragraph id="63544226FFCCB33893A7FE36FE9CFEA5" blockId="27.[189,334,436,462]" box="[189,334,436,462]" pageId="27" pageNumber="28">
<heading id="381CF54AFFCCB33893A7FE36FE9CFEA5" bold="true" box="[189,334,436,462]" fontSize="11" level="3" pageId="27" pageNumber="28" reason="3">
<emphasis id="519F9E34FFCCB33893A7FE36FE9CFEA5" bold="true" box="[189,334,436,462]" pageId="27" pageNumber="28">Description</emphasis>
</heading>
</paragraph>
<paragraph id="63544226FFCCB33893F0FE61FD4FFE96" blockId="27.[189,1399,482,615]" box="[234,669,482,509]" pageId="27" pageNumber="28">
<emphasis id="519F9E34FFCCB33893F0FE61FEF8FE96" bold="true" box="[234,298,483,509]" pageId="27" pageNumber="28">Male</emphasis>
(
<figureCitation id="FBD05EA3FFCCB3389223FE60FE2AFE96" box="[313,504,482,509]" captionStart="Fig" captionStartId="6.[190,233,1671,1697]" captionTargetBox="[192,1394,744,1633]" captionTargetId="figure-129@6.[189,1398,741,1634]" captionTargetPageId="6" captionText="Fig. 2. Adults of Melitaea lutko Evans, 1932 and M. timandra binaluduca subsp. nov. A–H. UPS. I–P. UNS. A, I. M. lutko, ♂, Chitral Gol National Park, Chitral, Pakistan, alt. 2700 m. B, J. M. timandra binaluduca subsp. nov., ♂, Iran, S Khorasan Prov., 35 km N Birjant t. C, K. M. timandra binaluduca subsp. nov., ♂, Central Afghanistan, Bamian Prov., Band-e-Amir, Dzhudoi-Kvak Gorge, alt. 3200 m. D, L. M. timandra binaluduca subsp. nov., ♂, Afghanistan, Bamian Prov., Band-e-Amir, alt. 3200 m. E, M. M. lutko, ♀, Chitral Gol National Park, Chitral, Pakistan, alt. 2700 m. F, N. M. timandra binaluduca subsp. nov., ♂, Iran, Mazandaran Prov., S macroslopes of Albors Mts, 80 km SE of Sari, 5 km NE Foulad Mahhaleh v., E slopes of Sultan Kuh Mt., alt. 2000 m. G, O. M. timandra binaluduca subsp. nov., ♀, Afghanistan, Bamian Prov., Band-e-Amir, alt. 3200 m. H, P. M. timandra binaluduca subsp. nov., ♀, Central Afghanistan, Bamian Prov., Band-e-Amir, Dzhudoi-Kvak Gorge, alt. 3200 m." figureDoi="http://doi.org/10.5281/zenodo.6839851" httpUri="https://zenodo.org/record/6839851/files/figure.png" pageId="27" pageNumber="28">Figs 2B–D, J–L</figureCitation>
,
<figureCitation id="FBD05EA3FFCCB338911EFE60FD44FE96" box="[516,662,482,509]" captionStart="Fig" captionStartId="24.[189,232,1891,1917]" captionTargetBox="[194,1395,1026,1847]" captionTargetId="figure-252@24.[189,1398,1022,1851]" captionTargetPageId="24" captionText="Fig. 15. Adults of Melitaea timandra binaludica subsp. nov. A–H. UPS. I–P. UNS. A, I. Holotype, ♂ (SDM). B–H, J–P. Paratypes (EDMSU). A–D, I–L. ♂, Iran, Rezavi Khorassan Prov., Kuh-e Binalud Mts, Dorrud v. vicinity, alt. 2430 m. E–H, M–P. ♀, same data as for preceding." figureDoi="http://doi.org/10.5281/zenodo.6839883" httpUri="https://zenodo.org/record/6839883/files/figure.png" pageId="27" pageNumber="28">15A–D, I–L</figureCitation>
)
</paragraph>
<paragraph id="63544226FFCCB33F93A7FD84FD16FD0B" blockId="27.[189,1399,482,615]" lastBlockId="28.[189,1401,265,608]" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28">
WINGS. FW length is
<quantity id="A413EFC3FFCCB33892D8FD84FD8EFD4B" box="[450,604,518,545]" metricMagnitude="-2" metricUnit="m" metricValue="1.725" metricValueMax="1.8" metricValueMin="1.65" pageId="27" pageNumber="28" unit="mm" value="17.25" valueMax="18.0" valueMin="16.5">16.5–18 mm</quantity>
, the
<typeStatus id="BC50FC84FFCCB3389183FD84FCD2FD4B" box="[665,768,518,544]" pageId="27" pageNumber="28" type="holotype">holotype</typeStatus>
is
<quantity id="A413EFC3FFCCB338903DFD84FC52FD4B" box="[807,896,518,544]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="27" pageNumber="28" unit="mm" value="18.0">18 mm</quantity>
. UPS ground color is bright orange. UPS black pattern well developed; UPS thin black marginal border with well-defined proximal marginal spots along the outer edge of the wings; UPF submarginal row formed by black strokes, UPH – by thin lunules, usually not connected to the black marginal border; UPF discal row represented by well-defined black spots fused closer to the costa, UPH discal row is more or less developed and formed by small black spots reduced closer to the costa; UPF postdiscal pale-yellow area, located behind discal row, is weakly expressed or absent, usually there are 2–3 pale macules near the UPF costal edge; UPF paleyellowish macule in the distal part of the discoidal cell is usually absent; UPH discal row is formed by fine black spots; UPH basal suffusion covers up to ⅓ of the wing surface; UNF is bright orange with well-developed pale area along the outer edge of the wing between the veins Sc and M2, UNF pale macules in the postdiscal area is usually absent; UNH ground color is white, with an admixture of dark scales. UNH lunules forming the proximal edge of submarginal orange fascia outwardly concave and sharply pointed between veins M3 and Cu2.
</paragraph>
<caption id="379412AEFFCCB33893A7F8BFFC76F88E" ID-DOI="http://doi.org/10.5281/zenodo.6839891" ID-Zenodo-Dep="6839891" httpUri="https://zenodo.org/record/6839891/files/figure.png" pageId="27" pageNumber="28" startId="27.[189,232,1853,1879]" targetBox="[192,1398,690,1814]" targetPageId="27">
<paragraph id="63544226FFCCB33893A7F8BFFC76F88E" blockId="27.[188,1399,1853,2021]" pageId="27" pageNumber="28">
<emphasis id="519F9E34FFCCB33893A7F8BFFECCF83C" bold="true" box="[189,286,1853,1879]" pageId="27" pageNumber="28">Fig. 18.</emphasis>
Male genitalia and harpe.
<emphasis id="519F9E34FFCCB3389170F8BFFD71F83C" bold="true" box="[618,675,1853,1879]" pageId="27" pageNumber="28">A–C</emphasis>
.
<taxonomicName id="A4EB39A5FFCCB33891AFF8BCFC79F833" authority="Kolesnichenko &amp; Kotlobay, 2022" authorityName="Kolesnichenko &amp; Kotlobay" authorityYear="2022" box="[693,939,1854,1880]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="shahvarica" status="sp. nov.">
<emphasis id="519F9E34FFCCB33891AFF8BCFC79F833" box="[693,939,1854,1880]" italics="true" pageId="27" pageNumber="28">Melitaea shahvarica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4AAC234FFFCCB33890ACF8BCFBC7F833" box="[950,1045,1854,1880]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="519F9E34FFCCB338973AF8BCFB85F833" bold="true" box="[1056,1111,1854,1880]" pageId="27" pageNumber="28">D–E</emphasis>
.
<taxonomicName id="A4EB39A5FFCCB3389773F8BCFAA0F833" authority="Evans, 1932" authorityName="Evans" authorityYear="1932" box="[1129,1394,1854,1880]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="lutko">
<emphasis id="519F9E34FFCCB3389773F8BCFB1DF833" box="[1129,1231,1854,1880]" italics="true" pageId="27" pageNumber="28">M. lutko</emphasis>
<bibRefCitation id="077A3FD7FFCCB33897C0F8BCFAA0F833" author="Evans W. H." box="[1242,1394,1854,1880]" pageId="27" pageNumber="28" refId="ref29423" refString="Evans W. H. 1932. The Identification of Indian Butterflies. The Bombay Natural History Society, Diocesan Press, Madras." type="book" year="1932">Evans, 1932</bibRefCitation>
</taxonomicName>
.
<emphasis id="519F9E34FFCCB33893A7F8E3FF3BF810" bold="true" box="[189,233,1889,1915]" pageId="27" pageNumber="28">F–I</emphasis>
.
<taxonomicName id="A4EB39A5FFCCB33893ECF8E0FDE7F810" authority="Higgins, 1940" authorityName="Higgins" authorityYear="1940" baseAuthorityName="Higgins" baseAuthorityYear="1940" box="[246,565,1889,1915]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="mimetica">
<emphasis id="519F9E34FFCCB33893ECF8E0FE5AF810" box="[246,392,1890,1915]" italics="true" pageId="27" pageNumber="28">M. mimetica</emphasis>
<bibRefCitation id="077A3FD7FFCCB3389297F8E3FDE7F810" author="Higgins L. G." box="[397,565,1889,1915]" pageId="27" pageNumber="28" pagination="51 - 53" refId="ref29676" refString="Higgins L. G. 1940. A new species and two new subspecies of Melitaea (Lep. Nymphalidae). The Entomologist 73: 51 - 53." type="journal article" year="1940">Higgins, 1940</bibRefCitation>
</taxonomicName>
.
<emphasis id="519F9E34FFCCB3389125F8E3FDAAF810" bold="true" box="[575,632,1889,1915]" pageId="27" pageNumber="28">A–C</emphasis>
. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mts, alt. 2200–2400 m.
<emphasis id="519F9E34FFCCB3389287F806FE01F8F5" bold="true" box="[413,467,1924,1950]" pageId="27" pageNumber="28">D–E</emphasis>
. Pakistan, Chitral, Chaghbini, CGNP [Chitral Gol National Park], alt. 2700 m.
<emphasis id="519F9E34FFCCB33893A4F825FF26F8AA" bold="true" box="[190,244,1959,1985]" pageId="27" pageNumber="28">F–G</emphasis>
. Afghanistan, Bamian Prov., Punjub Distr., 10 km. NE Varas v., alt. 2400 m.
<emphasis id="519F9E34FFCCB3389785F825FB02F8AA" bold="true" box="[1183,1232,1959,1985]" pageId="27" pageNumber="28">H–I</emphasis>
. Afghanistan, Bamian Prov., Panjub Distr., 10 km. NE Varas vil., alt. 2400 m.
</paragraph>
</caption>
<caption id="379412AEFFCBB33F93A7F8BFFCFBF88F" ID-DOI="http://doi.org/10.5281/zenodo.6839893" ID-Zenodo-Dep="6839893" httpUri="https://zenodo.org/record/6839893/files/figure.png" pageId="28" pageNumber="29" startId="28.[189,232,1853,1879]" targetBox="[192,1398,692,1812]" targetPageId="28">
<paragraph id="63544226FFCBB33F93A7F8BFFCFBF88F" blockId="28.[189,1401,1853,2021]" pageId="28" pageNumber="29">
<emphasis id="519F9E34FFCBB33F93A7F8BFFECEF83C" bold="true" box="[189,284,1853,1879]" pageId="28" pageNumber="29">Fig. 19.</emphasis>
Male genitalia and harpe.
<emphasis id="519F9E34FFCBB33F917AF8BFFD45F83C" bold="true" box="[608,663,1853,1879]" pageId="28" pageNumber="29">A–C</emphasis>
.
<taxonomicName id="A4EB39A5FFCBB33F91B2F8BCFAA6F833" authority="Coutsis &amp; van Oorschot, 2014" authorityName="Coutsis &amp; van Oorschot" authorityYear="2014" box="[680,1396,1854,1880]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="subSpecies" species="timandra" subSpecies="timandra">
<emphasis id="519F9E34FFCBB33F91B2F8BCFC29F833" box="[680,1019,1854,1880]" italics="true" pageId="28" pageNumber="29">Melitaea timandra timandra</emphasis>
Coutsis &amp;
<bibRefCitation id="077A3FD7FFCBB33F979DF8BCFAA6F833" author="Van Oorschot H. &amp; Coutsis J. G." box="[1159,1396,1854,1880]" pageId="28" pageNumber="29" refId="ref31081" refString="Van Oorschot H. &amp; Coutsis J. G. 2014. The Genus Melitaea Fabricius, 1807 (Lepidoptera, Nymphalidae, Nymphalinae). Taxonomy and Systematics with Special Reference to the Male Genitalia. Tshikolovets publication, Pardubice." type="book" year="2014">van Oorschot, 2014</bibRefCitation>
</taxonomicName>
.
<emphasis id="519F9E34FFCBB33F93A5F8E3FF3EF810" bold="true" box="[191,236,1889,1915]" pageId="28" pageNumber="29">D–I</emphasis>
.
<taxonomicName id="A4EB39A5FFCBB33F921BF8E0FDF7F810" authority="Kolesnichenko &amp; Kotlobay, 2022" authorityName="Kolesnichenko &amp; Kotlobay" authorityYear="2022" box="[257,549,1889,1915]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="subSpecies" species="timandra" status="subsp. nov." subSpecies="binaludica">
<emphasis id="519F9E34FFCBB33F921BF8E0FDF7F810" box="[257,549,1889,1915]" italics="true" pageId="28" pageNumber="29">M. timandra binaludica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4AAC234FFFCBB33F9129F8E3FD6FF810" box="[563,701,1889,1915]" pageId="28" pageNumber="29" rank="subSpecies">subsp. nov.</taxonomicNameLabel>
<emphasis id="519F9E34FFCBB33F91D0F8E3FCD1F810" bold="true" box="[714,771,1889,1915]" pageId="28" pageNumber="29">A–C</emphasis>
. Turkmenistan, Sary-Yazy, alt. 300 m.
<emphasis id="519F9E34FFCBB33F97EFF8E3FAF9F810" bold="true" box="[1269,1323,1889,1915]" pageId="28" pageNumber="29">
D
<emphasis id="519F9E34FFCBB33F9610F8E3FACBF810" bold="true" box="[1290,1305,1889,1915]" italics="true" pageId="28" pageNumber="29">–</emphasis>
F
</emphasis>
. Iran, Rezavi Khorassan Prov., Kuh-e-Binalud Mts, Dorrud
<taxonomicName id="A4EB39A5FFCBB33F9021F807FC61F8F5" box="[827,947,1924,1951]" pageId="28" pageNumber="29" rank="variety" variety="vicinity">v. vicinity</taxonomicName>
, alt. 2430 m.
<emphasis id="519F9E34FFCBB33F9742F806FBBCF8F5" bold="true" box="[1112,1134,1924,1950]" pageId="28" pageNumber="29">G</emphasis>
. Afghanistan, Bamian Prov., Band-e-Amir, alt. 3200 m.
<emphasis id="519F9E34FFCBB33F9155F825FDB6F8AA" bold="true" box="[591,612,1959,1985]" pageId="28" pageNumber="29">H</emphasis>
. Afghanistan, Bamian Prov., Band-e-Amir, Dzhudoi-Kvak Gorge, alt. 3200 m.
<emphasis id="519F9E34FFCBB33F924AF849FE89F88E" bold="true" box="[336,347,1995,2021]" pageId="28" pageNumber="29">I</emphasis>
. Afghanistan, Band-e-Amir, Hazarajat.
</paragraph>
</caption>
<caption id="379412AEFFCAB33E93A7FE8BFBCBFE4F" ID-Table-UUID="379412AEFFCAB33E93A7FE8BFBCBFE4F" box="[189,1049,265,292]" httpUri="http://table.plazi.org/id/379412AEFFCAB33E93A7FE8BFBCBFE4F" pageId="29" pageNumber="30" startId="29.[189,259,265,291]" targetBox="[200,1385,358,568]" targetIsTable="true" targetPageId="29">
<paragraph id="63544226FFCAB33E93A7FE8BFBCBFE4F" blockId="29.[189,1049,265,292]" box="[189,1049,265,292]" pageId="29" pageNumber="30">
<emphasis id="519F9E34FFCAB33E93A7FE8BFEF2FE48" bold="true" box="[189,288,265,291]" pageId="29" pageNumber="30">Table 3.</emphasis>
Measurements of the egg of
<taxonomicName id="A4EB39A5FFCAB33E916DFE88FC5CFE48" authority="Kolesnichenko &amp; Kotlobay, 2022" authorityName="Kolesnichenko &amp; Kotlobay" authorityYear="2022" box="[631,910,265,291]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="subSpecies" species="timandra" status="subsp. nov." subSpecies="binaludica">
<emphasis id="519F9E34FFCAB33E916DFE88FC5CFE48" box="[631,910,265,291]" italics="true" pageId="29" pageNumber="30">M. timandra binaludica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4AAC234FFFCAB33E908FFE8BFBCBFE4F" box="[917,1049,265,292]" pageId="29" pageNumber="30" rank="subSpecies">subsp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="63544226FFCAB33E9263FEE4FAE0FD7E" pageId="29" pageNumber="30">
<table id="11EBB086FFCA4CDC93D2FEE4FABBFD53" box="[200,1385,358,568]" gridcols="7" gridrows="3" pageId="29" pageNumber="30">
<tr id="DDDB4064FFCA4CDC93D2FEE4FABBFECA" box="[200,1385,358,417]" gridrow="0" pageId="29" pageNumber="30" rowspan-0="1">
<th id="9E0A2918FFCA4CDC9263FEE4FE21FECA" box="[377,499,358,417]" gridcol="1" gridrow="0" pageId="29" pageNumber="30">
<emphasis id="519F9E34FFCAB33E9263FEE4FE00FECB" bold="true" pageId="29" pageNumber="30">Egg height (μm)</emphasis>
</th>
<th id="9E0A2918FFCA4CDC9116FEE4FD52FECA" box="[524,640,358,417]" gridcol="2" gridrow="0" pageId="29" pageNumber="30">
<emphasis id="519F9E34FFCAB33E9116FEE4FDB0FECB" bold="true" pageId="29" pageNumber="30">Egg width (μm)</emphasis>
</th>
<th id="9E0A2918FFCA4CDC9187FEE4FC8CFECA" box="[669,862,358,417]" gridcol="3" gridrow="0" pageId="29" pageNumber="30">
<emphasis id="519F9E34FFCAB33E9187FEE4FC82FECB" bold="true" pageId="29" pageNumber="30">Micropile rosette diameter (μm)</emphasis>
</th>
<th id="9E0A2918FFCA4CDC906CFEE4FBCBFECA" box="[886,1049,358,417]" gridcol="4" gridrow="0" pageId="29" pageNumber="30">
<emphasis id="519F9E34FFCAB33E908AFEE4FBCBFECB" bold="true" pageId="29" pageNumber="30">Micropile diameter (μm)</emphasis>
</th>
<th id="9E0A2918FFCA4CDC972BFEE4FB6DFECA" box="[1073,1215,358,417]" gridcol="5" gridrow="0" pageId="29" pageNumber="30">
<emphasis id="519F9E34FFCAB33E972BFEE4FB6AFECB" bold="true" pageId="29" pageNumber="30">Primary cell width (μm)</emphasis>
</th>
<th id="9E0A2918FFCA4CDC97C1FEE4FABBFECA" box="[1243,1385,358,417]" gridcol="6" gridrow="0" pageId="29" pageNumber="30">
<emphasis id="519F9E34FFCAB33E97C1FEE4FAB7FECB" bold="true" pageId="29" pageNumber="30">Primary cell length (μm)</emphasis>
</th>
</tr>
<tr id="DDDB4064FFCA4CDC93D2FE3BFABBFEB9" box="[200,1385,441,466]" gridrow="1" pageId="29" pageNumber="30">
<th id="9E0A2918FFCA4CDC93D2FE3BFEB3FEB9" box="[200,353,441,466]" gridcol="0" gridrow="1" pageId="29" pageNumber="30">Average value</th>
<td id="9E0A2918FFCA4CDC9263FE3BFE21FEB9" box="[377,499,441,466]" gridcol="1" gridrow="1" pageId="29" pageNumber="30">792.6</td>
<td id="9E0A2918FFCA4CDC9116FE3BFD52FEB9" box="[524,640,441,466]" gridcol="2" gridrow="1" pageId="29" pageNumber="30">666.0</td>
<td id="9E0A2918FFCA4CDC9187FE3BFC8CFEB9" box="[669,862,441,466]" gridcol="3" gridrow="1" pageId="29" pageNumber="30">78.2</td>
<td id="9E0A2918FFCA4CDC906CFE3BFBCBFEB9" box="[886,1049,441,466]" gridcol="4" gridrow="1" pageId="29" pageNumber="30">7.8</td>
<td id="9E0A2918FFCA4CDC972BFE3BFB6DFEB9" box="[1073,1215,441,466]" gridcol="5" gridrow="1" pageId="29" pageNumber="30">10.5</td>
<td id="9E0A2918FFCA4CDC97C1FE3BFABBFEB9" box="[1243,1385,441,466]" gridcol="6" gridrow="1" pageId="29" pageNumber="30">19.4</td>
</tr>
<tr id="DDDB4064FFCA4CDC93D2FE7FFABBFD53" box="[200,1385,509,568]" gridrow="2" pageId="29" pageNumber="30">
<th id="9E0A2918FFCA4CDC93D2FE7FFEB3FD53" box="[200,353,509,568]" gridcol="0" gridrow="2" pageId="29" pageNumber="30">Standard deviation</th>
<td id="9E0A2918FFCA4CDC9263FE7FFE21FD53" box="[377,499,509,568]" gridcol="1" gridrow="2" pageId="29" pageNumber="30">33.8</td>
<td id="9E0A2918FFCA4CDC9116FE7FFD52FD53" box="[524,640,509,568]" gridcol="2" gridrow="2" pageId="29" pageNumber="30">18.7</td>
<td id="9E0A2918FFCA4CDC9187FE7FFC8CFD53" box="[669,862,509,568]" gridcol="3" gridrow="2" pageId="29" pageNumber="30">6.3</td>
<td id="9E0A2918FFCA4CDC906CFE7FFBCBFD53" box="[886,1049,509,568]" gridcol="4" gridrow="2" pageId="29" pageNumber="30">1.2</td>
<td id="9E0A2918FFCA4CDC972BFE7FFB6DFD53" box="[1073,1215,509,568]" gridcol="5" gridrow="2" pageId="29" pageNumber="30">2.6</td>
<td id="9E0A2918FFCA4CDC97C1FE7FFABBFD53" box="[1243,1385,509,568]" gridcol="6" gridrow="2" pageId="29" pageNumber="30">5.5</td>
</tr>
</table>
</paragraph>
<paragraph id="63544226FFCAB33E93A7FD07FE3FFCD3" blockId="29.[188,1400,645,952]" pageId="29" pageNumber="30">
MALE GENITALIA (
<figureCitation id="FBD05EA3FFCAB33E929CFD07FDC8FDF4" box="[390,538,645,671]" captionStart="Fig" captionStartId="18.[189,232,1819,1845]" captionTargetBox="[216,1371,268,1778]" captionTargetId="figure-16@18.[214,1372,265,1781]" captionTargetPageId="18" captionText="Fig. 10. Male genitalia (a = valva; b = aedeagus). A–F. Melitaea timandra timandra Coutsis &amp; van Oorschot, 2014. G–O. M. timandra binaludica subsp. nov. A–C. S Turkmenistan, Sary-Yazy, alt. 300 m. D. Paratype, Turkmenistan, Badhyz Reserve, Kepeli. E–F. S Turkmenistan, Sary-Yazy, alt. 300 m. G–L. Iran, Rezavi Khorassan Prov., Kuh-e-Binalud Mts, Dorrud v. vicinity, alt. 2430 m. M. Central Afghanistan, Bamian Prov., Band-e-Amir, Dzhudoi-Kvak Gorge, alt. 3200 m. N. Bande-Amir, Hazarajat, Afghanistan. O. Afghanistan, Bamian Prov., Band-e-Amir, alt. 3200 m." figureDoi="http://doi.org/10.5281/zenodo.6839869" httpUri="https://zenodo.org/record/6839869/files/figure.png" pageId="29" pageNumber="30">
FigS 10G
<emphasis id="519F9E34FFCAB33E92E9FD0AFDD0FDF5" box="[499,514,648,670]" italics="true" pageId="29" pageNumber="30">–</emphasis>
O
</figureCitation>
,
<figureCitation id="FBD05EA3FFCAB33E913CFD07FDA0FDF4" box="[550,626,645,671]" captionStart="Fig" captionStartId="26.[189,232,1847,1873]" captionTargetBox="[276,1315,271,1797]" captionTargetId="figure-17@26.[261,1325,263,1804]" captionTargetPageId="26" captionText="Fig. 17. Male genitalia (without aedeagus) in dorsal projection. A–C. Melitaea timandra timandra Coutsis &amp; van Oorschot, 2014. D–I. M. timandra binaludica subsp. nov. A–C. S Turkmenistan, Sary- Yazy, alt. 300 m. D–F. Iran, Rezavi Khorassan Prov., Kuh-e-Binalud Mts, Dorrud v. vicinity, alt. 2430 m. G. Iran, Horossan Prov., 35 km N of Birjant t. H. Afghanistan, Bamian Prov., Band-e-Amir, alt. 3200 m. I. Central Afghanistan, Bamian Prov., Band-e-Amir, Dzhudoi-Kvak Gorge, alt. 3200 m." figureDoi="http://doi.org/10.5281/zenodo.6839887" httpUri="https://zenodo.org/record/6839887/files/figure.png" pageId="29" pageNumber="30">
17D
<emphasis id="519F9E34FFCAB33E9143FD0AFDBAFDF5" box="[601,616,648,670]" italics="true" pageId="29" pageNumber="30">–</emphasis>
I
</figureCitation>
,
<figureCitation id="FBD05EA3FFCAB33E919AFD07FD1EFDF4" box="[640,716,645,671]" captionStart="Fig" captionStartId="28.[189,232,1853,1879]" captionTargetBox="[192,1398,692,1812]" captionTargetId="figure-178@28.[189,1398,689,1813]" captionTargetPageId="28" captionText="Fig. 19. Male genitalia and harpe. A–C. Melitaea timandra timandra Coutsis &amp; van Oorschot, 2014. D–I. M. timandra binaludica subsp. nov. A–C. Turkmenistan, Sary-Yazy, alt. 300 m. D–F. Iran, Rezavi Khorassan Prov., Kuh-e-Binalud Mts, Dorrud v. vicinity, alt. 2430 m. G. Afghanistan, Bamian Prov., Band-e-Amir, alt. 3200 m. H. Afghanistan, Bamian Prov., Band-e-Amir, Dzhudoi-Kvak Gorge, alt. 3200 m. I. Afghanistan, Band-e-Amir, Hazarajat." figureDoi="http://doi.org/10.5281/zenodo.6839893" httpUri="https://zenodo.org/record/6839893/files/figure.png" pageId="29" pageNumber="30">
19D
<emphasis id="519F9E34FFCAB33E91A9FD0AFD10FDF5" box="[691,706,648,670]" italics="true" pageId="29" pageNumber="30">–</emphasis>
I
</figureCitation>
,
<figureCitation id="FBD05EA3FFCAB33E91C0FD07FC85FDF4" box="[730,855,645,671]" captionStart="Fig" captionStartId="47.[189,232,1750,1776]" captionTargetBox="[273,1315,270,1661]" captionTargetId="figure-11@47.[270,1317,265,1713]" captionTargetPageId="47" captionText="Fig.29.Differencesinthestructureofvalvaandaedeagusofthelutko speciesgroup.A.Melitaeashahvarica sp. nov. B, D, H. M. timandra binaludica subsp. nov. C–E. M. mimetica Higgins, 1940. F. M. lutko Evans, 1932. G. M. timandra timandra Coutsis &amp;van Oorschot, 2014. A. Iran, Semnan Prov., Shahrud area, S macroslope of Shahvar Mts, alt. 2200–2400 m. B. Iran, Rezavi Khorassan Prov., Kuh-e-Binalud Mts, Dorrud v. vicinity, alt. 2430 m. C. Afghanistan, Bamian Prov., Punjub Distr., 10 km NE of Varas v., alt. 2400 m. D. Afghanistan, Band-i-Amir, Hazarajat. E. Pakistan, Balochistan, Quetta, Urak, alt. 2400– 2700 m. F. Pakistan, Chitral, Gol National Park, alt. 2700 m. G. Turkmenistan, Sary-Yazy, alt. 700 m. H. Iran, Rezavi Khorassan Prov., Kuh-e-Binalud Mts, Dorrud v. vicinity, alt. 2430 m." figureDoi="http://doi.org/10.5281/zenodo.6839915" httpUri="https://zenodo.org/record/6839915/files/figure.png" pageId="29" pageNumber="30">29B, D, H</figureCitation>
). In general, the size of the genitalia is smaller than that of the nominative subspecies. Valva with a relatively narrow (on average narrower than that of the nominative subspecies) caudal process, usually with several small spines on the dorsal surface. The relatively short harpe is somewhat expanded in the central part and bears a small number of noticeable teeth on the inner surface. The expanded part of the harpe is 1.5–2 times narrower than that of the nominative subspecies. Aedeagus with a rounded convex dorsal edge, its posterior part is mostly located at an angle to the anterior part and is directed downward. When both parts of the aedeagus are joined, there is a well-defined protrusion on the ventral side. Narrow saccus is distally pointed, its width is 2–3 times less than the length.
</paragraph>
<paragraph id="63544226FFCAB33E93F0FC62FCD2FC91" blockId="29.[188,1399,992,1159]" box="[234,768,992,1019]" pageId="29" pageNumber="30">
<emphasis id="519F9E34FFCAB33E93F0FC62FE97FC91" bold="true" box="[234,325,992,1018]" pageId="29" pageNumber="30">Female</emphasis>
(
<figureCitation id="FBD05EA3FFCAB33E924FFC62FDC8FC91" box="[341,538,992,1018]" captionStart="Fig" captionStartId="6.[190,233,1671,1697]" captionTargetBox="[192,1394,744,1633]" captionTargetId="figure-129@6.[189,1398,741,1634]" captionTargetPageId="6" captionText="Fig. 2. Adults of Melitaea lutko Evans, 1932 and M. timandra binaluduca subsp. nov. A–H. UPS. I–P. UNS. A, I. M. lutko, ♂, Chitral Gol National Park, Chitral, Pakistan, alt. 2700 m. B, J. M. timandra binaluduca subsp. nov., ♂, Iran, S Khorasan Prov., 35 km N Birjant t. C, K. M. timandra binaluduca subsp. nov., ♂, Central Afghanistan, Bamian Prov., Band-e-Amir, Dzhudoi-Kvak Gorge, alt. 3200 m. D, L. M. timandra binaluduca subsp. nov., ♂, Afghanistan, Bamian Prov., Band-e-Amir, alt. 3200 m. E, M. M. lutko, ♀, Chitral Gol National Park, Chitral, Pakistan, alt. 2700 m. F, N. M. timandra binaluduca subsp. nov., ♂, Iran, Mazandaran Prov., S macroslopes of Albors Mts, 80 km SE of Sari, 5 km NE Foulad Mahhaleh v., E slopes of Sultan Kuh Mt., alt. 2000 m. G, O. M. timandra binaluduca subsp. nov., ♀, Afghanistan, Bamian Prov., Band-e-Amir, alt. 3200 m. H, P. M. timandra binaluduca subsp. nov., ♀, Central Afghanistan, Bamian Prov., Band-e-Amir, Dzhudoi-Kvak Gorge, alt. 3200 m." figureDoi="http://doi.org/10.5281/zenodo.6839851" httpUri="https://zenodo.org/record/6839851/files/figure.png" pageId="29" pageNumber="30">Figs 2F–H, N–P</figureCitation>
;
<figureCitation id="FBD05EA3FFCAB33E913CFC62FD2AFC91" box="[550,760,992,1019]" captionStart="Fig" captionStartId="24.[189,232,1891,1917]" captionTargetBox="[194,1395,1026,1847]" captionTargetId="figure-252@24.[189,1398,1022,1851]" captionTargetPageId="24" captionText="Fig. 15. Adults of Melitaea timandra binaludica subsp. nov. A–H. UPS. I–P. UNS. A, I. Holotype, ♂ (SDM). B–H, J–P. Paratypes (EDMSU). A–D, I–L. ♂, Iran, Rezavi Khorassan Prov., Kuh-e Binalud Mts, Dorrud v. vicinity, alt. 2430 m. E–H, M–P. ♀, same data as for preceding." figureDoi="http://doi.org/10.5281/zenodo.6839883" httpUri="https://zenodo.org/record/6839883/files/figure.png" pageId="29" pageNumber="30">Fig. 15E–H, M–P</figureCitation>
)
</paragraph>
<paragraph id="63544226FFCAB33E93A7FB86FD26FBEC" blockId="29.[188,1399,992,1159]" pageId="29" pageNumber="30">
WINGS. FW length is
<quantity id="A413EFC3FFCAB33E92A3FB81FD80FB75" box="[441,594,1027,1054]" metricMagnitude="-2" metricUnit="m" metricValue="1.875" metricValueMax="2.0" metricValueMin="1.75" pageId="29" pageNumber="30" unit="mm" value="18.75" valueMax="20.0" valueMin="17.5">17.5–20 mm</quantity>
. Externally the female is similar to the male, but the pattern is more contrasting. UPF postdiscal pale area is more developed than in the male. There is a well-developed pale macule in the discoilal cell between the first and second discal spots. UPH submarginal row is usually reduced. UNS pattern is similar to that of males.
</paragraph>
<paragraph id="63544226FFCAB33E93A7FB2DFE90FA59" blockId="29.[187,1399,1198,1330]" pageId="29" pageNumber="30">
FEMALE GENITALIA (
<figureCitation id="FBD05EA3FFCAB33E9282FB2CFDC9FBA3" box="[408,539,1198,1225]" captionStart="Fig" captionStartId="19.[189,232,1732,1758]" captionTargetBox="[192,1393,270,1668]" captionTargetId="figure-11@19.[189,1398,265,1675]" captionTargetPageId="19" captionText="Fig. 11. Female genitalia (without the anal papillae, copulatory bursa, and anterior and posterior apophyses). A–C. Melitaea timandra timandra Coutsis &amp; van Oorschot, 2014, S Turkmenistan, Sary- Yazy, alt. 300 m. D–I. M. timandra binaludica subsp. nov. D–F. Iran, Rezavi Khorassan Prov., Kuh-e- Binalud Mts, Dorrud v. vicinity, alt. 2430 m. G–H. Iran, Mazandaran Prov., S macroslopes of Albors Mts, 80 km SE of Sari, 5 km NE of Foulad Mahhaleh v., E slopes of Sultan Kuh Mt., alt. 2000 m. I. Afghanistan, Bamian Prov., Band-e-Amir, alt. 3200 m." figureDoi="http://doi.org/10.5281/zenodo.6839871" httpUri="https://zenodo.org/record/6839871/files/figure.png" pageId="29" pageNumber="30">Fig. 11D–I</figureCitation>
). In general, the genitalia are similar to those of the nominative subspecies. The postvaginal plate is roundly triangular in shape. The antevaginal plate is narrow in the dorsoventral direction, its outer edge does not extend beyond the boundaries of the bend of the postvaginal plate (auricules).
</paragraph>
<paragraph id="63544226FFCAB33E93F0FADBFD36FA18" blockId="29.[189,1399,1369,1466]" box="[234,740,1369,1395]" pageId="29" pageNumber="30">
<emphasis id="519F9E34FFCAB33E93F0FADBFDC2FA18" bold="true" box="[234,528,1369,1395]" pageId="29" pageNumber="30">Preimaginal stages: egg</emphasis>
(
<figureCitation id="FBD05EA3FFCAB33E913BFADBFDAAFA18" box="[545,632,1369,1395]" captionStart="Fig" captionStartId="30.[189,232,1954,1980]" captionTargetBox="[189,1398,554,1911]" captionTargetId="figure-99@30.[189,1398,554,1911]" captionTargetPageId="30" captionText="Fig. 20. Eggs of Melitaea timandra binaludica subsp. nov., Iran, Kuh-e-Binalud Mts. A–C. Lateral view. D–F. View from above. G–I. Micropile area." figureDoi="http://doi.org/10.5281/zenodo.6839895" httpUri="https://zenodo.org/record/6839895/files/figure.png" pageId="29" pageNumber="30">Fig. 20</figureCitation>
,
<tableCitation id="2E69779DFFCAB33E9199FADBFD08FA18" box="[643,730,1369,1395]" captionStart="Table 3" captionStartId="29.[189,259,265,291]" captionTargetPageId="29" captionText="Table 3. Measurements of the egg of M. timandra binaludica subsp. nov." httpUri="http://table.plazi.org/id/379412AEFFCAB33E93A7FE8BFBCBFE4F" pageId="29" pageNumber="30" tableUuid="379412AEFFCAB33E93A7FE8BFBCBFE4F">Table 3</tableCitation>
)
</paragraph>
<paragraph id="63544226FFCAB33E93A7FAFEFDF8FAD1" blockId="29.[189,1399,1369,1466]" pageId="29" pageNumber="30">
Material examined:
<specimenCount id="75ED89AFFFCAB33E92AAFAFEFD94FAFD" box="[432,582,1404,1430]" pageId="29" pageNumber="30" type="generic">4 specimens</specimenCount>
,
<specimenCount id="75ED89AFFFCAB33E914FFAFEFD67FAFC" box="[597,693,1404,1431]" pageId="29" pageNumber="30" type="egg">27 eggs</specimenCount>
,
<collectingCountry id="1BFC02B6FFCAB33E91DEFAFFFD24FAFC" box="[708,758,1405,1431]" name="Iran" pageId="29" pageNumber="30">Iran</collectingCountry>
, Rezavi
<collectingRegion id="A12F8CC4FFCAB33E9078FAFEFBFAFAFC" box="[866,1064,1404,1431]" country="Iran" name="Khorasan-e Shomali" pageId="29" pageNumber="30">Khorassan Prov.</collectingRegion>
, Qadamgan area, Dorrud
<taxonomicName id="A4EB39A5FFCAB33E9678FAFFFEA1FAD2" authority=", Gerina" authorityName="Gerina" pageId="29" pageNumber="30" rank="variety" variety="vicinity">v. vicinity, Gerina</taxonomicName>
v., alt.
<quantity id="A413EFC3FFCAB33E92D3FA22FDF7FAD1" box="[457,549,1440,1466]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="29" pageNumber="30" unit="m" value="2000.0">2000 m</quantity>
.
</paragraph>
<paragraph id="63544226FFCAB33E93A7FA63FC0CF92A" blockId="29.[188,1399,1505,1602]" pageId="29" pageNumber="30">
The description of the egg of
<taxonomicName id="A4EB39A5FFCAB33E913DFA60FC91FA90" authority="Kolesnichenko &amp; Kotlobay, 2022" authorityName="Kolesnichenko &amp; Kotlobay" authorityYear="2022" box="[551,835,1505,1531]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="subSpecies" species="timandra" status="subsp. nov." subSpecies="binaludica">
<emphasis id="519F9E34FFCAB33E913DFA60FC91FA90" box="[551,835,1505,1531]" italics="true" pageId="29" pageNumber="30">M. timandra binaludica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4AAC234FFFCAB33E9054FA63FC06FA97" box="[846,980,1505,1532]" pageId="29" pageNumber="30" rank="subSpecies">subsp. nov.</taxonomicNameLabel>
was given earlier based on
<specimenCount id="75ED89AFFFCAB33E9632FA63FAA5FA97" box="[1320,1399,1505,1532]" pageId="29" pageNumber="30" type="egg">5 eggs</specimenCount>
examined using a scanning electron microscope (
<bibRefCitation id="077A3FD7FFCAB33E91E2F986FB50F975" author="Kolesnichenko K. A. &amp; Kotlobay A. A." box="[760,1154,1540,1566]" pageId="29" pageNumber="30" pagination="566 - 574" refId="ref30158" refString="Kolesnichenko K. A. &amp; Kotlobay A. A. 2020. Early stages and biology of Melitaea timandra Coutsis et Oorschot, 2014 (Lepidoptera, Nymphalidae) under the conditions of Kuh-e-Binalud Mountain (Iran). Zootaxa 4729 (4): 566 - 574. https: // doi. org / 10.11646 / zootaxa. 4729.4.7" type="journal article" year="2020">Kolesnichenko &amp; Kotlobay 2020</bibRefCitation>
). Additional material made it possible to more accurately identify real diagnostic features.
</paragraph>
<paragraph id="63544226FFCAB33E93A7F9EBFDA1F8AB" blockId="29.[188,1400,1641,1984]" pageId="29" pageNumber="30">
The egg is the largest among the representatives of the
<taxonomicName id="A4EB39A5FFCAB33E9050F9EBFC57F9E8" authorityName="Evans" authorityYear="1932" box="[842,901,1641,1667]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="lutko">
<emphasis id="519F9E34FFCAB33E9050F9EBFC57F9E8" box="[842,901,1641,1667]" italics="true" pageId="29" pageNumber="30">lutko</emphasis>
</taxonomicName>
group. It is oval and noticeably elongated in the dorso-ventral direction (
<bibRefCitation id="077A3FD7FFCAB33E9132F90EFC6BF9CD" author="Kolesnichenko K. A. &amp; Kotlobay A. A." box="[552,953,1676,1702]" pageId="29" pageNumber="30" pagination="566 - 574" refId="ref30158" refString="Kolesnichenko K. A. &amp; Kotlobay A. A. 2020. Early stages and biology of Melitaea timandra Coutsis et Oorschot, 2014 (Lepidoptera, Nymphalidae) under the conditions of Kuh-e-Binalud Mountain (Iran). Zootaxa 4729 (4): 566 - 574. https: // doi. org / 10.11646 / zootaxa. 4729.4.7" type="journal article" year="2020">Kolesnichenko &amp; Kotlobay 2020</bibRefCitation>
). The height of the egg varies greatly from 761.0 µm to 855.0 µm, the width is from 627.5 µm to 703.0 µm (
<tableCitation id="2E69779DFFCAB33E970EF92DFBA2F9A2" box="[1044,1136,1711,1737]" captionStart="Table 3" captionStartId="29.[189,259,265,291]" captionTargetPageId="29" captionText="Table 3. Measurements of the egg of M. timandra binaludica subsp. nov." httpUri="http://table.plazi.org/id/379412AEFFCAB33E93A7FE8BFBCBFE4F" pageId="29" pageNumber="30" tableUuid="379412AEFFCAB33E93A7FE8BFBCBFE4F">Table 3</tableCitation>
). The sculpture of the micropilar area is formed from four to five rows of penta-hexagonal cells of various lengths and widths. The diameter of the micropilar rosette in the widest part varies from 78.0 µm to 90.0 µm. The micropilar rosette is formed by 7–9 (in rare cases 10) primary quadro-pentahedral cells with a width from 6.0 µm to 15.0 µm and a length from 9.0 µm to 27.5 µm. The micropile is rounded in shape, with an average diameter of about 8.0 µm. Lateral longitudinal ribs from 26 to 30 limit the micropile area and fall to ⅓ of the egg surface. The transverse ribs are expressed in the first third of the egg surface. Below the lateral ribs, the chorion is relatively smooth.
</paragraph>
<paragraph id="63544226FFC9B33D93A7FE8BFB3DFEDB" blockId="30.[187,1399,265,433]" pageId="30" pageNumber="31">
In general, the eggs of
<taxonomicName id="A4EB39A5FFC9B33D92A6FE88FD1CFE48" authority="Kolesnichenko &amp; Kotlobay, 2022" authorityName="Kolesnichenko &amp; Kotlobay" authorityYear="2022" box="[444,718,265,291]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="subSpecies" species="timandra" status="subsp. nov." subSpecies="binaludica">
<emphasis id="519F9E34FFC9B33D92A6FE88FD1CFE48" box="[444,718,265,291]" italics="true" pageId="30" pageNumber="31">M. timandra binaludica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4AAC234FFFC9B33D91CCFE8BFC85FE48" box="[726,855,265,291]" pageId="30" pageNumber="31" rank="subSpecies">subsp. nov.</taxonomicNameLabel>
are similar to those of the nominate subspecies. A distinctive feature of the morphology of the eggs of the new subspecies is their well-pronounced elongated-oval shape. In the nominate subspecies, the egg shape is also oval, but not so elongated dorsoventrally. In addition, the eggs of
<taxonomicName id="A4EB39A5FFC9B33D9163FEF1FC5EFEE7" authority="Kolesnichenko &amp; Kotlobay, 2022" authorityName="Kolesnichenko &amp; Kotlobay" authorityYear="2022" box="[633,908,370,396]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="subSpecies" species="timandra" status="subsp. nov." subSpecies="binaludica">
<emphasis id="519F9E34FFC9B33D9163FEF1FC5EFEE7" box="[633,908,370,396]" italics="true" pageId="30" pageNumber="31">M. timandra binaludica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4AAC234FFFC9B33D908EFEF0FBC7FEE6" box="[916,1045,370,397]" pageId="30" pageNumber="31" rank="subSpecies">subsp. nov.</taxonomicNameLabel>
are the largest, their maximum height is 850.0 µm, while the height of those of the nominate subspecies is about 700.0 µm.
</paragraph>
</subSubSection>
<caption id="379412AEFFC9B33D93A7F820FCC7F88B" ID-DOI="http://doi.org/10.5281/zenodo.6839895" ID-Zenodo-Dep="6839895" httpUri="https://zenodo.org/record/6839895/files/figure.png" pageId="30" pageNumber="31" startId="30.[189,232,1954,1980]" targetBox="[189,1398,554,1911]" targetPageId="30">
<paragraph id="63544226FFC9B33D93A7F820FCC7F88B" blockId="30.[189,1398,1954,2016]" pageId="30" pageNumber="31">
<emphasis id="519F9E34FFC9B33D93A7F820FECCF8D7" bold="true" box="[189,286,1954,1980]" pageId="30" pageNumber="31">Fig. 20.</emphasis>
Eggs of
<taxonomicName id="A4EB39A5FFC9B33D928BF820FD2AF8D7" authority="Kolesnichenko &amp; Kotlobay, 2022" authorityName="Kolesnichenko &amp; Kotlobay" authorityYear="2022" box="[401,760,1954,1980]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="subSpecies" species="timandra" status="subsp. nov." subSpecies="binaludica">
<emphasis id="519F9E34FFC9B33D928BF820FD2AF8D7" box="[401,760,1954,1980]" italics="true" pageId="30" pageNumber="31">Melitaea timandra binaludica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4AAC234FFFC9B33D9019F820FC5FF8D6" box="[771,909,1954,1981]" pageId="30" pageNumber="31" rank="subSpecies">subsp. nov.</taxonomicNameLabel>
, Iran, Kuh-e-Binalud Mts.
<emphasis id="519F9E34FFC9B33D97C2F820FAC3F8D7" bold="true" box="[1240,1297,1954,1980]" pageId="30" pageNumber="31">A–C</emphasis>
. Lateral view.
<emphasis id="519F9E34FFC9B33D9219F844FEEAF88B" bold="true" box="[259,312,1990,2016]" pageId="30" pageNumber="31">D–F</emphasis>
. View from above.
<emphasis id="519F9E34FFC9B33D913BF847FD80F8B4" bold="true" box="[545,594,1989,2015]" pageId="30" pageNumber="31">G–I</emphasis>
. Micropile area.
</paragraph>
</caption>
<subSubSection id="2BF111ADFFC8B33C93A7FE8AFC11FE1F" pageId="31" pageNumber="32" type="biology_ecology">
<paragraph id="63544226FFC8B33C93A7FE8AFECEFE49" blockId="31.[189,284,264,290]" box="[189,284,264,290]" pageId="31" pageNumber="32">
<heading id="381CF54AFFC8B33C93A7FE8AFECEFE49" bold="true" box="[189,284,264,290]" fontSize="11" level="3" pageId="31" pageNumber="32" reason="3">
<emphasis id="519F9E34FFC8B33C93A7FE8AFECEFE49" bold="true" box="[189,284,264,290]" pageId="31" pageNumber="32">Biology</emphasis>
</heading>
</paragraph>
<paragraph id="63544226FFC8B33C93A7FEB5FC11FE1F" blockId="31.[189,1398,311,372]" pageId="31" pageNumber="32">
The biology of the new subspecies in the Kuh-e-Binalud mountains and the early stages were described in detail by the authors earlier (
<bibRefCitation id="077A3FD7FFC8B33C9131FED8FC65FE1F" author="Kolesnichenko K. A. &amp; Kotlobay A. A." box="[555,951,346,372]" pageId="31" pageNumber="32" pagination="566 - 574" refId="ref30158" refString="Kolesnichenko K. A. &amp; Kotlobay A. A. 2020. Early stages and biology of Melitaea timandra Coutsis et Oorschot, 2014 (Lepidoptera, Nymphalidae) under the conditions of Kuh-e-Binalud Mountain (Iran). Zootaxa 4729 (4): 566 - 574. https: // doi. org / 10.11646 / zootaxa. 4729.4.7" type="journal article" year="2020">Kolesnichenko &amp; Kotlobay 2020</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="2BF111ADFFC8B33C93A7FE19FADEFE8F" pageId="31" pageNumber="32" type="distribution">
<paragraph id="63544226FFC8B33C93A7FE19FE16FEDD" blockId="31.[189,452,411,438]" box="[189,452,411,438]" pageId="31" pageNumber="32">
<emphasis id="519F9E34FFC8B33C93A7FE19FE84FEDE" bold="true" box="[189,342,411,437]" pageId="31" pageNumber="32">Distribution</emphasis>
(
<figureCitation id="FBD05EA3FFC8B33C927CFE1EFE69FEDD" box="[358,443,412,438]" captionStart="Fig" captionStartId="23.[189,232,910,936]" captionTargetBox="[189,1398,267,872]" captionTargetId="figure-261@23.[189,1398,267,872]" captionTargetPageId="23" captionText="Fig. 14. Distribution of Melitaea timandra timandra Coutsis &amp; van Oorschot, 2014, M. timandra binaludica subsp. nov. and M. shahvarica sp. nov. For a description of the points by letters, see Fig. 30. = Melitaea timandra timandra; = Melitaea timandra binaludica subsp. nov.; = Melitaea shahvarica sp. nov.;? = unconfirmed finds of Melitaea timandra; = Melitaea timandra with uncertain subspecies status." figureDoi="http://doi.org/10.5281/zenodo.6839881" httpUri="https://zenodo.org/record/6839881/files/figure.png" pageId="31" pageNumber="32">Fig. 14</figureCitation>
)
</paragraph>
<paragraph id="63544226FFC8B33C93A7FE49FADEFE8F" blockId="31.[189,1292,458,485]" box="[189,1292,458,485]" pageId="31" pageNumber="32">
<collectingCountry id="1BFC02B6FFC8B33C93A7FE49FF23FE8E" box="[189,241,459,485]" name="Iran" pageId="31" pageNumber="32">Iran</collectingCountry>
: Turkmen-Khorassan Mts, Kuh-e-Binalud Ridge, Kayen Mts;
<collectingCountry id="1BFC02B6FFC8B33C90D0FE48FB8CFE8F" box="[970,1118,458,484]" name="Afghanistan" pageId="31" pageNumber="32">Afghanistan</collectingCountry>
: Band-e-Amir.
</paragraph>
</subSubSection>
<subSubSection id="2BF111ADFFC8B33C93A7FD8EFADBFD92" pageId="31" pageNumber="32" type="discussion">
<paragraph id="63544226FFC8B33C93A7FD8EFADBFD92" blockId="31.[186,1398,524,761]" pageId="31" pageNumber="32">
The described subspecies
<taxonomicName id="A4EB39A5FFC8B33C92FEFD8EFD24FD4D" authority="Kolesnichenko &amp; Kotlobay, 2022" authorityName="Kolesnichenko &amp; Kotlobay" authorityYear="2022" box="[484,758,524,550]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="subSpecies" species="timandra" status="subsp. nov." subSpecies="binaludica">
<emphasis id="519F9E34FFC8B33C92FEFD8EFD24FD4D" box="[484,758,524,550]" italics="true" pageId="31" pageNumber="32">M. timandra binaludica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4AAC234FFFC8B33C91E4FD8EFCACFD4D" box="[766,894,524,550]" pageId="31" pageNumber="32" rank="subSpecies">subsp. nov.</taxonomicNameLabel>
undoubtedly should also include specimens of
<taxonomicName id="A4EB39A5FFC8B33C93C7FDB2FEA2FD22" baseAuthorityName="van Oorschot &amp; Coutsis" baseAuthorityYear="2014" box="[221,368,559,585]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="timandra">
<emphasis id="519F9E34FFC8B33C93C7FDB2FEA2FD22" box="[221,368,559,585]" italics="true" pageId="31" pageNumber="32">M. timandra</emphasis>
</taxonomicName>
(
<taxonomicName id="A4EB39A5FFC8B33C929AFDB2FD87FD22" authorityName="Higgins" authorityYear="1940" box="[384,597,559,585]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="subSpecies" species="lutko" subSpecies="mimetica">
<emphasis id="519F9E34FFC8B33C929AFDB2FD87FD22" box="[384,597,559,585]" italics="true" pageId="31" pageNumber="32">M. lutko mimetica</emphasis>
</taxonomicName>
from the authors), caught by V. Eckweiler in
<collectingCountry id="1BFC02B6FFC8B33C9771FDADFB48FD22" box="[1131,1178,559,585]" name="Iran" pageId="31" pageNumber="32">Iran</collectingCountry>
on Kuh-e-Binalud Ridge in the vicinity of Zoshk and Firizi (
<bibRefCitation id="077A3FD7FFC8B33C91A0FDD0FC31FD07" author="Tshikolovets V. &amp; Naderi A. &amp; Eckweiler W." box="[698,995,594,620]" pageId="31" pageNumber="32" refId="ref30958" refString="Tshikolovets V., Naderi A. &amp; Eckweiler W. 2014. The Butterflies of Iran and Iraq. Tshikolovets Publications, Pardubice." type="book" year="2014">
Tshikolovets
<emphasis id="519F9E34FFC8B33C9046FDD1FC4BFD07" box="[860,921,594,620]" italics="true" pageId="31" pageNumber="32">et al.</emphasis>
2014
</bibRefCitation>
). The butterflies on the published photos (
<bibRefCitation id="077A3FD7FFC8B33C9204FDF7FD98FDE4" author="Tshikolovets V. &amp; Naderi A. &amp; Eckweiler W." box="[286,586,629,656]" pageId="31" pageNumber="32" refId="ref30958" refString="Tshikolovets V., Naderi A. &amp; Eckweiler W. 2014. The Butterflies of Iran and Iraq. Tshikolovets Publications, Pardubice." type="book" year="2014">
Tshikolovets
<emphasis id="519F9E34FFC8B33C92DBFDF4FE2AFDE4" box="[449,504,629,655]" italics="true" pageId="31" pageNumber="32">et al</emphasis>
. 2014
</bibRefCitation>
: tables lxi, 1–2) have an external similarity with the specimens of
<taxonomicName id="A4EB39A5FFC8B33C93A7FD1BFE0EFDD9" authorityName="Kolesnichenko &amp; Kotlobay" authorityYear="2022" box="[189,476,664,690]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="subSpecies" species="timandra" subSpecies="binaludica">
<emphasis id="519F9E34FFC8B33C93A7FD1BFE0EFDD9" box="[189,476,664,690]" italics="true" pageId="31" pageNumber="32">M. timandra binaludica</emphasis>
</taxonomicName>
available to us from the same place, caught at the same time by the same collector. The butterflies from the Bandi Amir Valley, identified by the authors (
<bibRefCitation id="077A3FD7FFC8B33C974EFD3EFABCFDBD" author="Tshikolovets V. &amp; Pliushch I. &amp; Pak O. &amp; Skrylnik Y." box="[1108,1390,700,726]" pageId="31" pageNumber="32" refId="ref30983" refString="Tshikolovets V., Pliushch I., Pak O. &amp; Skrylnik Y. 2018. The Butterflies of Afghanistan. Tshikolovets Publications, Pardubice." type="book" year="2018">
Tshikolovets
<emphasis id="519F9E34FFC8B33C97E8FD3EFAF7FDBD" box="[1266,1317,700,726]" italics="true" pageId="31" pageNumber="32">et al</emphasis>
. 2018
</bibRefCitation>
) as
<taxonomicName id="A4EB39A5FFC8B33C93C7FD62FEA2FD92" authorityName="Higgins" authorityYear="1940" baseAuthorityName="Higgins" baseAuthorityYear="1940" box="[221,368,736,761]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="mimetica">
<emphasis id="519F9E34FFC8B33C93C7FD62FEA2FD92" box="[221,368,736,761]" italics="true" pageId="31" pageNumber="32">M. mimetica</emphasis>
</taxonomicName>
, should also be attributed to the described subspecies
<taxonomicName id="A4EB39A5FFC8B33C90F1FD62FAD0FD92" authorityName="Kolesnichenko &amp; Kotlobay" authorityYear="2022" box="[1003,1282,735,761]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="subSpecies" species="timandra" subSpecies="binaludica">
<emphasis id="519F9E34FFC8B33C90F1FD62FAD0FD92" box="[1003,1282,735,761]" italics="true" pageId="31" pageNumber="32">M. timandra binaludica</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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