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<document ID-DOI="10.11646/zootaxa.5256.6.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="7762456" ID-ZooBank="057285E1-92C9-4D76-921A-CE3C85D0D3B1" checkinTime="1679558471249" checkinUser="plazi" docAuthor="Neethling, J. A. &amp; Neethling, C." docDate="2023" docId="EB305F51B51FF67CFF1EFAA0FA93FA60" docLanguage="en" docName="zootaxa.5256.6.1.pdf" docOrigin="Zootaxa 5256 (6)" docSource="http://dx.doi.org/10.11646/zootaxa.5256.6.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Gymnobisium capense Neethling &amp; Neethling 2023, sp. nov." docType="treatment" docVersion="3" lastPageNumber="512" masterDocId="17092729B518F677FF89FF81FFD1FF85" masterDocTitle="A systematic revision of the South African Gymnobisiidae (Pseudoscorpiones: Neobisioidea)" masterLastPageNumber="543" masterPageNumber="501" pageNumber="508" updateTime="1679591149964" updateUser="juliana" zenodo-license-document="CLOSED">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A systematic revision of the South African Gymnobisiidae (Pseudoscorpiones: Neobisioidea)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Neethling, J. A.</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0003-1702-9566</mods:nameIdentifier>
<mods:affiliation>Terrestrial Invertebrates Department, National Museum, P. O. Box 266, Bloemfontein 9300, South Africa. ja. neethling @ nasmus. co. za; https: // orcid. org / 0000 - 0003 - 1702 - 9566</mods:affiliation>
<mods:nameIdentifier type="email">ja.neethling@nasmus.co.za</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Neethling, C.</mods:namePart>
<mods:nameIdentifier type="ORCID">0009-0003-6492-4798</mods:nameIdentifier>
<mods:affiliation>Research Quality and Support, Clinglobal Limited, P. O. Box 11186, Bloemfontein 9321, South Africa. carmen. neethling @ clinglobal. com; https: // orcid. org / 0009 - 0003 - 6492 - 4798</mods:affiliation>
<mods:nameIdentifier type="email">carmen.neethling@clinglobal.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-03-23</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>5256</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>6</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>501</mods:start>
<mods:end>543</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.11646/zootaxa.5256.6.1</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/zootaxa.5256.6.1</mods:identifier>
<mods:identifier type="ISSN">1175-5326</mods:identifier>
<mods:identifier type="Zenodo-Dep">7762456</mods:identifier>
<mods:identifier type="ZooBank">057285E1-92C9-4D76-921A-CE3C85D0D3B1</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7762490" ID-Zenodo-Dep="7762490" LSID="urn:lsid:plazi:treatment:EB305F51B51FF67CFF1EFAA0FA93FA60" httpUri="http://treatment.plazi.org/id/EB305F51B51FF67CFF1EFAA0FA93FA60" lastPageId="11" lastPageNumber="512" pageId="7" pageNumber="508">
<subSubSection box="[151,525,1313,1340]" pageId="7" pageNumber="508" type="nomenclature">
<paragraph blockId="7.[151,525,1313,1376]" box="[151,525,1313,1340]" pageId="7" pageNumber="508">
<heading bold="true" box="[151,525,1313,1340]" fontSize="11" level="1" pageId="7" pageNumber="508" reason="1">
<emphasis bold="true" box="[151,525,1313,1340]" pageId="7" pageNumber="508">
<taxonomicName authority="Neethling &amp; Neethling, 2023" authorityName="Neethling &amp; Neethling" authorityYear="2023" box="[151,424,1313,1340]" class="Arachnida" family="Gymnobisiidae" genus="Gymnobisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="7" pageNumber="508" phylum="Arthropoda" rank="species" species="capense" status="sp. nov.">
<emphasis bold="true" box="[151,424,1313,1340]" italics="true" pageId="7" pageNumber="508">Gymnobisium capense</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[431,525,1313,1339]" pageId="7" pageNumber="508" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[151,249,1349,1376]" pageId="7" pageNumber="508" type="description">
<paragraph blockId="7.[151,525,1313,1376]" box="[151,249,1349,1376]" pageId="7" pageNumber="508">
<figureCitation box="[151,249,1349,1376]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStart-2="FIGURE 7" captionStartId-0="8.[151,250,1736,1760]" captionStartId-1="9.[151,250,1552,1576]" captionStartId-2="10.[151,250,1907,1931]" captionTargetBox-0="[151,1436,181,1712]" captionTargetBox-1="[151,1435,963,1529]" captionTargetBox-2="[169,1415,542,1865]" captionTargetId-0="figure-22@8.[151,1436,181,1712]" captionTargetId-1="figure-366@9.[151,1435,957,1529]" captionTargetId-2="figure-177@10.[151,1436,525,1884]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionTargetPageId-2="10" captionText-0="FIGURE 5. Digital microscope photographs of Gymnobisium capense sp. nov., Female (A, B) and Male (C, D). A, C. Dorsal view; B, D. Ventral view. Scale bar: 1.00 mm." captionText-1="FIGURE 6. Digital microscope photographs of Gymnobisium capense sp. nov. right chela: Female (A, B) and Male (C, D). A, C. Retrolateral view; B, D. Dorsal view. Scale bar: 1.00 mm." captionText-2="FIGURE 7. Gymnobisium capense sp. nov. A. Female right chela, retrolateral view; B. Female right pedipalp, dorsal view; C. Female left chelicera, dorsal view; D. Galea; E. Female chelal fingers, anterior retrolateral view; F. Male fixed chelal finger, anterior retrolateral view; G. Female rallum; H. Same; I. Male rallum. Scale bars: A, B: 0.50 mm; C, E, F: 0.20 mm; D, GI: 0.10 mm." figureDoi-0="http://doi.org/10.5281/zenodo.7762472" figureDoi-1="http://doi.org/10.5281/zenodo.7762476" figureDoi-2="http://doi.org/10.5281/zenodo.7762480" httpUri-0="https://zenodo.org/record/7762472/files/figure.png" httpUri-1="https://zenodo.org/record/7762476/files/figure.png" httpUri-2="https://zenodo.org/record/7762480/files/figure.png" pageId="7" pageNumber="508">Figs 57</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="508" type="materials_examined">
<paragraph blockId="7.[151,1437,1421,1988]" pageId="7" pageNumber="508">
<materialsCitation collectingDate="2015-12-07" collectionCode="NMBAP" collectorName="J. A. Neethling &amp; C. Neethling" country="South Africa" county="Rheenendal" elevation="308" latitude="-33.883335" location="Afromontane forest" longLatPrecision="1202" longitude="22.966667" municipality="Jubilee Creek Nature Reserve" pageId="7" pageNumber="508" specimenCode="NMBAP 00336" specimenCount="1" specimenCount-female="1" stateProvince="Western Cape" typeStatus="holotype">
<emphasis bold="true" box="[151,269,1421,1447]" pageId="7" pageNumber="508">
<typeStatus box="[151,264,1421,1447]" pageId="7" pageNumber="508">Holotype</typeStatus>
:
</emphasis>
<specimenCount box="[275,296,1422,1448]" pageId="7" pageNumber="508" type="female"></specimenCount>
,
<collectingCountry box="[308,511,1421,1447]" name="South Africa" pageId="7" pageNumber="508">SOUTH AFRICA</collectingCountry>
,
<collectingRegion box="[519,675,1421,1448]" country="South Africa" name="Western Cape" pageId="7" pageNumber="508">Western Cape</collectingRegion>
,
<collectingCounty box="[685,820,1421,1447]" pageId="7" pageNumber="508">Rheenendal</collectingCounty>
,
<collectingMunicipality box="[829,1160,1421,1448]" pageId="7" pageNumber="508">Jubilee Creek Nature Reserve</collectingMunicipality>
,
<geoCoordinate box="[1169,1260,1423,1450]" degrees="33" direction="south" minutes="53" orientation="latitude" pageId="7" pageNumber="508" precision="925" value="-33.883335">33°53S</geoCoordinate>
,
<geoCoordinate box="[1270,1363,1423,1450]" degrees="22" direction="east" minutes="58" orientation="longitude" pageId="7" pageNumber="508" precision="925" value="22.966667">22°58E</geoCoordinate>
,
<elevation metricMagnitude="2" metricUnit="m" metricValue="3.08" pageId="7" pageNumber="508" unit="m" value="308.0">
<quantity box="[1372,1436,1421,1447]" metricMagnitude="2" metricUnit="m" metricValue="3.08" pageId="7" pageNumber="508" unit="m" value="308.0">308m</quantity>
a.s.l.
</elevation>
,
<location LSID="urn:lsid:plazi:treatment:EB305F51B51FF67CFF1EFAA0FA93FA60:6646B89CB51FF670FF51FA30FE66FA4E" box="[216,439,1457,1483]" country="South Africa" county="Rheenendal" latitude="-33.883335" longLatPrecision="1202" longitude="22.966667" municipality="Jubilee Creek Nature Reserve" name="Afromontane forest" pageId="7" pageNumber="508" stateProvince="Western Cape">Afromontane forest</location>
,
<collectorName box="[450,501,1457,1483]" pageId="7" pageNumber="508">Leaf</collectorName>
litter sifting, leg.
<collectorName box="[702,864,1457,1484]" pageId="7" pageNumber="508">J.A. Neethling</collectorName>
&amp;
<collectorName box="[900,1046,1457,1483]" pageId="7" pageNumber="508">C. Neethling</collectorName>
,
<date box="[1057,1180,1457,1484]" pageId="7" pageNumber="508" value="2015-12-07">
<collectingDate box="[1057,1180,1457,1484]" pageId="7" pageNumber="508" value="2015-12-07">7.XII.2015</collectingDate>
</date>
(
<specimenCode box="[1194,1377,1457,1483]" collectionCode="NMBAP" pageId="7" pageNumber="508">NMBAP 00336</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph blockId="7.[151,1437,1421,1988]" box="[199,893,1493,1522]" pageId="7" pageNumber="508">
<materialsCitation box="[199,889,1493,1522]" collectingDate="2015-12-07" collectionCode="NMBAP" collectorName="J. A. Neethling &amp; C. Neethling" country="South Africa" county="Rheenendal" elevation="308" latitude="-33.883335" location="Afromontane forest" longLatPrecision="1202" longitude="22.966667" municipality="Jubilee Creek Nature Reserve" pageId="7" pageNumber="508" specimenCode="NMBAP 00337" specimenCount="4" specimenCount-female="4" stateProvince="Western Cape" typeStatus="paratype">
<emphasis bold="true" box="[199,328,1493,1519]" pageId="7" pageNumber="508">
<typeStatus box="[199,323,1493,1519]" pageId="7" pageNumber="508">Paratypes</typeStatus>
:
</emphasis>
<specimenCount box="[335,372,1493,1519]" pageId="7" pageNumber="508" type="female">4♀</specimenCount>
, 4♂, Same data as holotype (
<specimenCode box="[699,881,1493,1519]" collectionCode="NMBAP" pageId="7" pageNumber="508">NMBAP 00337</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="508" type="etymology">
<paragraph blockId="7.[151,1437,1421,1988]" pageId="7" pageNumber="508">
<emphasis bold="true" box="[199,335,1529,1556]" pageId="7" pageNumber="508">Etymology.</emphasis>
The Latin adjective
<taxonomicName authorityName="Neethling &amp; Neethling" authorityYear="2023" box="[572,662,1530,1555]" class="Arachnida" family="Gymnobisiidae" genus="Gymnobisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="7" pageNumber="508" phylum="Arthropoda" rank="species" species="capense">
<emphasis box="[572,662,1530,1555]" italics="true" pageId="7" pageNumber="508">capēnse</emphasis>
</taxonomicName>
means to belong to, or be from, the
<collectingRegion box="[1082,1247,1529,1555]" country="South Africa" name="Western Cape" pageId="7" pageNumber="508">Cape Province</collectingRegion>
in
<collectingCountry box="[1286,1433,1529,1555]" name="South Africa" pageId="7" pageNumber="508">South Africa</collectingCountry>
. The name refers to the main geographic distribution of this species.
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="508" type="diagnosis">
<paragraph blockId="7.[151,1437,1421,1988]" pageId="7" pageNumber="508">
<emphasis bold="true" box="[199,322,1601,1627]" pageId="7" pageNumber="508">Diagnosis.</emphasis>
Large species (chela length
<specimenCount box="[654,675,1602,1628]" pageId="7" pageNumber="508" type="female"></specimenCount>
<quantity box="[684,852,1601,1628]" metricMagnitude="-3" metricUnit="m" metricValue="1.865" metricValueMax="1.9" metricValueMin="1.83" pageId="7" pageNumber="508" unit="mm" value="1.865" valueMax="1.9" valueMin="1.83">1.831.90 mm</quantity>
,
<geoCoordinate box="[866,945,1601,1630]" degrees="1.59" direction="west" orientation="longitude" pageId="7" pageNumber="508" precision="555" value="-1.59">
<quantity box="[897,945,1601,1628]" metricMagnitude="-3" metricUnit="m" metricValue="1.665" metricValueMax="1.74" metricValueMin="1.59" pageId="7" pageNumber="508" unit="mm" value="1.665" valueMax="1.74" valueMin="1.59">1.59</quantity>
</geoCoordinate>
1.74 mm), with chelae significantly more bulbous in females than in males (chela ratio
<specimenCount box="[687,708,1638,1664]" pageId="7" pageNumber="508" type="female"></specimenCount>
2.963.16 versus
<geoCoordinate box="[922,1003,1637,1666]" degrees="3.70" direction="west" orientation="longitude" pageId="7" pageNumber="508" precision="555" value="-3.7">♂ 3.70</geoCoordinate>
3.88 times longer than broad); fixed finger multi-toothed structure with seven strongly sclerotized teeth; cheliceral hand with six setae; galea with six rami; rallum with seven to eight blades, distal blades split apically. Differs from all congeners, except
<taxonomicName authorityName="Harvey &amp; Giribet" authorityYear="2016" class="Arachnida" family="Gymnobisiidae" genus="Gymnobisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="7" pageNumber="508" phylum="Arthropoda" rank="species" species="inukshuk">
<emphasis italics="true" pageId="7" pageNumber="508">Gymnobisium inukshuk</emphasis>
</taxonomicName>
, by the presence of an extra basal seta on each chelicera. Differs from
<taxonomicName authorityName="Harvey &amp; Giribet" authorityYear="2016" box="[1042,1177,1745,1772]" class="Arachnida" family="Gymnobisiidae" genus="Gymnobisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="7" pageNumber="508" phylum="Arthropoda" rank="species" species="inukshuk">
<emphasis box="[1042,1177,1745,1772]" italics="true" pageId="7" pageNumber="508">G. inukshuk</emphasis>
</taxonomicName>
by the presence of two pairs of well-developed eyes.
</paragraph>
</subSubSection>
<subSubSection lastPageId="11" lastPageNumber="512" pageId="7" pageNumber="508" type="description">
<paragraph blockId="7.[151,1437,1421,1988]" box="[199,337,1817,1843]" pageId="7" pageNumber="508">
<emphasis bold="true" box="[199,337,1817,1843]" pageId="7" pageNumber="508">Description</emphasis>
</paragraph>
<paragraph blockId="7.[151,1437,1421,1988]" pageId="7" pageNumber="508">
<emphasis box="[199,319,1854,1880]" italics="true" pageId="7" pageNumber="508">Carapace:</emphasis>
Somewhat broader than long, subquadrate, overall smooth and without furrow. Dark brown in females, lighter in males. Two pairs of eyes located dorso-laterally at the front (
<figureCitation box="[1076,1204,1889,1916]" captionStart="FIGURE 5" captionStartId="8.[151,250,1736,1760]" captionTargetBox="[151,1436,181,1712]" captionTargetId="figure-22@8.[151,1436,181,1712]" captionTargetPageId="8" captionText="FIGURE 5. Digital microscope photographs of Gymnobisium capense sp. nov., Female (A, B) and Male (C, D). A, C. Dorsal view; B, D. Ventral view. Scale bar: 1.00 mm." figureDoi="http://doi.org/10.5281/zenodo.7762472" httpUri="https://zenodo.org/record/7762472/files/figure.png" pageId="7" pageNumber="508">Fig. 5A, C</figureCitation>
). Small seta located on the dorso-anterior margin of each of the posterior eyes. Female carapace with 27, male carapace with 28 setae, arranged 4: 6: 2: 6: 9(10).
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7762472" ID-Zenodo-Dep="7762472" httpUri="https://zenodo.org/record/7762472/files/figure.png" pageId="8" pageNumber="509" startId="8.[151,250,1736,1760]" targetBox="[151,1436,181,1712]" targetPageId="8">
<paragraph blockId="8.[151,1437,1736,1796]" pageId="8" pageNumber="509">
<emphasis bold="true" box="[151,275,1736,1760]" pageId="8" pageNumber="509">FIGURE 5.</emphasis>
Digital microscope photographs of
<taxonomicName authority="Neethling &amp; Neethling, 2023" authorityName="Neethling &amp; Neethling" authorityYear="2023" box="[642,871,1736,1760]" class="Arachnida" family="Gymnobisiidae" genus="Gymnobisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="8" pageNumber="509" phylum="Arthropoda" rank="species" species="capense" status="sp. nov.">
<emphasis box="[642,871,1736,1760]" italics="true" pageId="8" pageNumber="509">Gymnobisium capense</emphasis>
</taxonomicName>
<emphasis bold="true" box="[878,958,1736,1760]" pageId="8" pageNumber="509">
<taxonomicNameLabel box="[878,958,1736,1760]" pageId="8" pageNumber="509" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, Female (
<emphasis bold="true" box="[1062,1080,1736,1760]" pageId="8" pageNumber="509">A</emphasis>
,
<emphasis bold="true" box="[1093,1110,1736,1760]" pageId="8" pageNumber="509">B</emphasis>
) and Male (
<emphasis bold="true" box="[1235,1253,1736,1760]" pageId="8" pageNumber="509">C</emphasis>
,
<emphasis bold="true" box="[1266,1284,1736,1760]" pageId="8" pageNumber="509">D</emphasis>
).
<emphasis bold="true" box="[1306,1324,1736,1760]" pageId="8" pageNumber="509">A</emphasis>
,
<emphasis bold="true" box="[1337,1362,1736,1760]" pageId="8" pageNumber="509">C.</emphasis>
Dorsal view;
<emphasis bold="true" box="[214,231,1772,1796]" pageId="8" pageNumber="509">B</emphasis>
,
<emphasis bold="true" box="[243,268,1772,1796]" pageId="8" pageNumber="509">D.</emphasis>
Ventral view. Scale bar: 1.00 mm.
</paragraph>
</caption>
<paragraph blockId="8.[151,1437,1842,2013]" lastBlockId="9.[151,1437,150,321]" lastPageId="9" lastPageNumber="510" pageId="8" pageNumber="509">
<emphasis box="[199,314,1842,1868]" italics="true" pageId="8" pageNumber="509">Abdomen:</emphasis>
Tergites and sternites smooth, undivided and uniseriate. Female abdominal tergites uniformly dark brown. Male tergites same colour as carapace anteriorly and gradually darken posteriorly. Male tergites VIIIX often have a significantly darker central band (
<figureCitation box="[677,801,1914,1941]" captionStart="FIGURE 5" captionStartId="8.[151,250,1736,1760]" captionTargetBox="[151,1436,181,1712]" captionTargetId="figure-22@8.[151,1436,181,1712]" captionTargetPageId="8" captionText="FIGURE 5. Digital microscope photographs of Gymnobisium capense sp. nov., Female (A, B) and Male (C, D). A, C. Dorsal view; B, D. Ventral view. Scale bar: 1.00 mm." figureDoi="http://doi.org/10.5281/zenodo.7762472" httpUri="https://zenodo.org/record/7762472/files/figure.png" pageId="8" pageNumber="509">Fig. 5A, C</figureCitation>
). In both sexes the lateral and posterior edges of tergites IXI are distinctly lighter in colour, tergite XII uniformly lighter in colour. The tergal setae are located on the posterior edge of each tergite, within the lighter coloured band. In both sexes sternites lighten in colour moving from posterior to anterior of abdomen (
<figureCitation box="[585,706,150,177]" captionStart="FIGURE 5" captionStartId="8.[151,250,1736,1760]" captionTargetBox="[151,1436,181,1712]" captionTargetId="figure-22@8.[151,1436,181,1712]" captionTargetPageId="8" captionText="FIGURE 5. Digital microscope photographs of Gymnobisium capense sp. nov., Female (A, B) and Male (C, D). A, C. Dorsal view; B, D. Ventral view. Scale bar: 1.00 mm." figureDoi="http://doi.org/10.5281/zenodo.7762472" httpUri="https://zenodo.org/record/7762472/files/figure.png" pageId="9" pageNumber="510">Fig. 5B, D</figureCitation>
). In females the sternites lighten in colour from dark brown (same colour as tergites) to light or transparent brown, and in males they lighten from dark brown to tan. Sternites I and II are fused and hardly distinguishable. Female operculum with 14 setae, eight along the posterior margin and six additional setae clustered loosely alongside, near the centre. Males with 27 setae at the operculum cluster. Pleural membrane cream to light yellow in colour, faintly papillate.
</paragraph>
<paragraph blockId="9.[151,1436,367,429]" pageId="9" pageNumber="510">Tergal chaetotaxy: 7(8): 9(10): 10(12): 11(11): 12(12): 12(12): 11(12): 12(12): 11(11): 9(9): 11(12): 2(2). Sternal chaetotaxy: 14(27):?(?): 21(24): 21(16): 21(19): 19(16): 19(18): 17(15): 14(14): 13(13): 11(12): 2(2).</paragraph>
<paragraph blockId="9.[151,1437,475,933]" pageId="9" pageNumber="510">
<emphasis box="[199,300,475,501]" italics="true" pageId="9" pageNumber="510">Pedipalp</emphasis>
: All segments smooth and coloured orange-brown in both sexes. Trochanter cone-shaped and rounded anteriorly, curved slightly posteriorly. Base narrow, widening distally. Setae present on the anterior surface. Femur slender, 3.703.79 (
<specimenCount box="[375,393,547,573]" pageId="9" pageNumber="510" type="female"></specimenCount>
) to 3.863.95 (♂) times longer than wide, somewhat constricted basally, widening just distal of base. Shallow disto-prolateral excavation present. Setae absent from base, though distributed evenly over rest of surface. Patella constricted and slightly angled at base, widening markedly distal at around a quarter segment length, 2.572.75 (
<specimenCount box="[280,298,655,681]" pageId="9" pageNumber="510" type="female"></specimenCount>
) to 2.852.92 (♂) times longer than wide. Several small lyriform fissures present on the dorsal surface, just distal of base. Two larger lyriform fissures present latero-dorsally from these, where the segment starts to widen. Deep disto-prolateral excavation present. Setae sparse at base, more numerous on wide part of patella (
<figureCitation box="[1303,1392,727,753]" captionStart="FIGURE 7" captionStartId="10.[151,250,1907,1931]" captionTargetBox="[169,1415,542,1865]" captionTargetId="figure-177@10.[151,1436,525,1884]" captionTargetPageId="10" captionText="FIGURE 7. Gymnobisium capense sp. nov. A. Female right chela, retrolateral view; B. Female right pedipalp, dorsal view; C. Female left chelicera, dorsal view; D. Galea; E. Female chelal fingers, anterior retrolateral view; F. Male fixed chelal finger, anterior retrolateral view; G. Female rallum; H. Same; I. Male rallum. Scale bars: A, B: 0.50 mm; C, E, F: 0.20 mm; D, GI: 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.7762480" httpUri="https://zenodo.org/record/7762480/files/figure.png" pageId="9" pageNumber="510">Fig. 7B</figureCitation>
).
</paragraph>
<paragraph blockId="9.[151,1437,475,933]" pageId="9" pageNumber="510">
<emphasis box="[199,266,763,789]" italics="true" pageId="9" pageNumber="510">Chela</emphasis>
: In both sexes smooth and uniformly orange-brown, fingers may be slightly darker. Female hand bulbous, strongly convex on the prolateral edge, slightly less so on the retrolateral edge (
<figureCitation box="[1069,1195,799,825]" captionStart="FIGURE 6" captionStartId="9.[151,250,1552,1576]" captionTargetBox="[151,1435,963,1529]" captionTargetId="figure-366@9.[151,1435,957,1529]" captionTargetPageId="9" captionText="FIGURE 6. Digital microscope photographs of Gymnobisium capense sp. nov. right chela: Female (A, B) and Male (C, D). A, C. Retrolateral view; B, D. Dorsal view. Scale bar: 1.00 mm." figureDoi="http://doi.org/10.5281/zenodo.7762476" httpUri="https://zenodo.org/record/7762476/files/figure.png" pageId="9" pageNumber="510">Fig. 6A, B</figureCitation>
). Male hand smaller, markedly less convex prolaterally (
<figureCitation box="[550,675,835,861]" captionStart="FIGURE 6" captionStartId="9.[151,250,1552,1576]" captionTargetBox="[151,1435,963,1529]" captionTargetId="figure-366@9.[151,1435,957,1529]" captionTargetPageId="9" captionText="FIGURE 6. Digital microscope photographs of Gymnobisium capense sp. nov. right chela: Female (A, B) and Male (C, D). A, C. Retrolateral view; B, D. Dorsal view. Scale bar: 1.00 mm." figureDoi="http://doi.org/10.5281/zenodo.7762476" httpUri="https://zenodo.org/record/7762476/files/figure.png" pageId="9" pageNumber="510">Fig. 6C, D</figureCitation>
). Pedicel short in both sexes, retrolateral condyle small but distinct and rounded. Fingers narrow, same length as hand (with pedicel) and curved slightly prolaterally. Fixed and movable chelal fingers with eight and four trichobothria respectively (
<figureCitation box="[833,923,907,933]" captionStart="FIGURE 7" captionStartId="10.[151,250,1907,1931]" captionTargetBox="[169,1415,542,1865]" captionTargetId="figure-177@10.[151,1436,525,1884]" captionTargetPageId="10" captionText="FIGURE 7. Gymnobisium capense sp. nov. A. Female right chela, retrolateral view; B. Female right pedipalp, dorsal view; C. Female left chelicera, dorsal view; D. Galea; E. Female chelal fingers, anterior retrolateral view; F. Male fixed chelal finger, anterior retrolateral view; G. Female rallum; H. Same; I. Male rallum. Scale bars: A, B: 0.50 mm; C, E, F: 0.20 mm; D, GI: 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.7762480" httpUri="https://zenodo.org/record/7762480/files/figure.png" pageId="9" pageNumber="510">Fig. 7A</figureCitation>
).
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7762476" ID-Zenodo-Dep="7762476" httpUri="https://zenodo.org/record/7762476/files/figure.png" pageId="9" pageNumber="510" startId="9.[151,250,1552,1576]" targetBox="[151,1435,963,1529]" targetPageId="9">
<paragraph blockId="9.[151,1436,1552,1612]" pageId="9" pageNumber="510">
<emphasis bold="true" box="[151,275,1552,1576]" pageId="9" pageNumber="510">FIGURE 6.</emphasis>
Digital microscope photographs of
<taxonomicName authority="Neethling &amp; Neethling, 2023" authorityName="Neethling &amp; Neethling" authorityYear="2023" box="[639,868,1552,1576]" class="Arachnida" family="Gymnobisiidae" genus="Gymnobisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="9" pageNumber="510" phylum="Arthropoda" rank="species" species="capense" status="sp. nov.">
<emphasis box="[639,868,1552,1576]" italics="true" pageId="9" pageNumber="510">Gymnobisium capense</emphasis>
</taxonomicName>
<emphasis bold="true" box="[874,954,1552,1576]" pageId="9" pageNumber="510">
<taxonomicNameLabel box="[874,954,1552,1576]" pageId="9" pageNumber="510" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
right chela: Female (
<emphasis bold="true" box="[1170,1188,1552,1576]" pageId="9" pageNumber="510">A</emphasis>
,
<emphasis bold="true" box="[1201,1218,1552,1576]" pageId="9" pageNumber="510">B</emphasis>
) and Male (
<emphasis bold="true" box="[1342,1360,1552,1576]" pageId="9" pageNumber="510">C</emphasis>
,
<emphasis bold="true" box="[1372,1390,1552,1576]" pageId="9" pageNumber="510">D</emphasis>
).
<emphasis bold="true" box="[1412,1430,1552,1576]" pageId="9" pageNumber="510">A</emphasis>
,
<emphasis bold="true" box="[151,176,1588,1612]" pageId="9" pageNumber="510">C.</emphasis>
Retrolateral view;
<emphasis bold="true" box="[371,388,1588,1612]" pageId="9" pageNumber="510">B</emphasis>
,
<emphasis bold="true" box="[400,425,1588,1612]" pageId="9" pageNumber="510">D.</emphasis>
Dorsal view. Scale bar: 1.00 mm.
</paragraph>
</caption>
<paragraph blockId="9.[151,1436,1658,2009]" lastBlockId="10.[151,1436,151,501]" lastPageId="10" lastPageNumber="511" pageId="9" pageNumber="510">
Terminal multi-toothed structure of fixed finger extends beyond venom tooth of movable finger in both sexes, structure of female thicker, bent acutely downward (
<figureCitation box="[756,846,1694,1720]" captionStart="FIGURE 7" captionStartId="10.[151,250,1907,1931]" captionTargetBox="[169,1415,542,1865]" captionTargetId="figure-177@10.[151,1436,525,1884]" captionTargetPageId="10" captionText="FIGURE 7. Gymnobisium capense sp. nov. A. Female right chela, retrolateral view; B. Female right pedipalp, dorsal view; C. Female left chelicera, dorsal view; D. Galea; E. Female chelal fingers, anterior retrolateral view; F. Male fixed chelal finger, anterior retrolateral view; G. Female rallum; H. Same; I. Male rallum. Scale bars: A, B: 0.50 mm; C, E, F: 0.20 mm; D, GI: 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.7762480" httpUri="https://zenodo.org/record/7762480/files/figure.png" pageId="9" pageNumber="510">Fig. 7E</figureCitation>
), more slender and less bent in males (
<figureCitation box="[1302,1390,1694,1720]" captionStart="FIGURE 7" captionStartId="10.[151,250,1907,1931]" captionTargetBox="[169,1415,542,1865]" captionTargetId="figure-177@10.[151,1436,525,1884]" captionTargetPageId="10" captionText="FIGURE 7. Gymnobisium capense sp. nov. A. Female right chela, retrolateral view; B. Female right pedipalp, dorsal view; C. Female left chelicera, dorsal view; D. Galea; E. Female chelal fingers, anterior retrolateral view; F. Male fixed chelal finger, anterior retrolateral view; G. Female rallum; H. Same; I. Male rallum. Scale bars: A, B: 0.50 mm; C, E, F: 0.20 mm; D, GI: 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.7762480" httpUri="https://zenodo.org/record/7762480/files/figure.png" pageId="9" pageNumber="510">Fig. 7F</figureCitation>
). In both sexes, structure contains seven strongly sclerotized teeth; front tooth not significantly larger than tooth just behind it; five teeth in line behind front tooth, raised and slightly prolateral, most distal tooth expanded, reaching beyond front tooth; single large tooth on retrolateral surface, behind raised teeth. Both fingers with rounded, juxtadentate teeth situated on flexible lamellae. Lamella on fixed finger originating roughly one quarter finger length from base, on movable finger roughly one third finger length from base. Female fixed finger with 6365 lightly sclerotized, juxtadentate teeth, 1516 proximal to lamella base, together with seven strongly sclerotized teeth on distal structure. Male fixed finger with 6366 lightly sclerotized, juxtadentate teeth, 1920 proximal to lamella base, together with seven strongly sclerotized teeth on distal structure. Female movable finger with 5457 lightly sclerotized, juxtadentate teeth, 1920 proximal to lamella base. Male movable finger with 6164 lightly sclerotized, juxtadentate teeth, 2729 proximal to lamella base. Two thickened, spine-like setae present on distal end of movable finger in both sexes.
</paragraph>
<paragraph blockId="10.[151,1436,151,501]" pageId="10" pageNumber="511">
<emphasis box="[199,309,259,285]" italics="true" pageId="10" pageNumber="511">Chelicera</emphasis>
: Hand with six long and acuminate setae,
<emphasis box="[789,812,259,284]" italics="true" pageId="10" pageNumber="511">es</emphasis>
shorter, two basal setae,
<emphasis box="[1096,1127,259,285]" italics="true" pageId="10" pageNumber="511">bs</emphasis>
and
<emphasis box="[1183,1223,259,285]" italics="true" pageId="10" pageNumber="511">bs”</emphasis>
, present (
<figureCitation box="[1334,1423,259,285]" captionStart="FIGURE 7" captionStartId="10.[151,250,1907,1931]" captionTargetBox="[169,1415,542,1865]" captionTargetId="figure-177@10.[151,1436,525,1884]" captionTargetPageId="10" captionText="FIGURE 7. Gymnobisium capense sp. nov. A. Female right chela, retrolateral view; B. Female right pedipalp, dorsal view; C. Female left chelicera, dorsal view; D. Galea; E. Female chelal fingers, anterior retrolateral view; F. Male fixed chelal finger, anterior retrolateral view; G. Female rallum; H. Same; I. Male rallum. Scale bars: A, B: 0.50 mm; C, E, F: 0.20 mm; D, GI: 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.7762480" httpUri="https://zenodo.org/record/7762480/files/figure.png" pageId="10" pageNumber="511">Fig. 7C</figureCitation>
). Female fixed finger with 13 teeth, male with ten. Female moveable finger with ten teeth, male with nine. Galea of both sexes divided into two major rami, each terminating in three smaller rami (
<figureCitation box="[1100,1192,331,357]" captionStart="FIGURE 7" captionStartId="10.[151,250,1907,1931]" captionTargetBox="[169,1415,542,1865]" captionTargetId="figure-177@10.[151,1436,525,1884]" captionTargetPageId="10" captionText="FIGURE 7. Gymnobisium capense sp. nov. A. Female right chela, retrolateral view; B. Female right pedipalp, dorsal view; C. Female left chelicera, dorsal view; D. Galea; E. Female chelal fingers, anterior retrolateral view; F. Male fixed chelal finger, anterior retrolateral view; G. Female rallum; H. Same; I. Male rallum. Scale bars: A, B: 0.50 mm; C, E, F: 0.20 mm; D, GI: 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.7762480" httpUri="https://zenodo.org/record/7762480/files/figure.png" pageId="10" pageNumber="511">Fig. 7D</figureCitation>
). Female rallum with seven (
<figureCitation box="[231,320,367,393]" captionStart="FIGURE 7" captionStartId="10.[151,250,1907,1931]" captionTargetBox="[169,1415,542,1865]" captionTargetId="figure-177@10.[151,1436,525,1884]" captionTargetPageId="10" captionText="FIGURE 7. Gymnobisium capense sp. nov. A. Female right chela, retrolateral view; B. Female right pedipalp, dorsal view; C. Female left chelicera, dorsal view; D. Galea; E. Female chelal fingers, anterior retrolateral view; F. Male fixed chelal finger, anterior retrolateral view; G. Female rallum; H. Same; I. Male rallum. Scale bars: A, B: 0.50 mm; C, E, F: 0.20 mm; D, GI: 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.7762480" httpUri="https://zenodo.org/record/7762480/files/figure.png" pageId="10" pageNumber="511">Fig. 7G</figureCitation>
) to eight (
<figureCitation box="[437,525,367,393]" captionStart="FIGURE 7" captionStartId="10.[151,250,1907,1931]" captionTargetBox="[169,1415,542,1865]" captionTargetId="figure-177@10.[151,1436,525,1884]" captionTargetPageId="10" captionText="FIGURE 7. Gymnobisium capense sp. nov. A. Female right chela, retrolateral view; B. Female right pedipalp, dorsal view; C. Female left chelicera, dorsal view; D. Galea; E. Female chelal fingers, anterior retrolateral view; F. Male fixed chelal finger, anterior retrolateral view; G. Female rallum; H. Same; I. Male rallum. Scale bars: A, B: 0.50 mm; C, E, F: 0.20 mm; D, GI: 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.7762480" httpUri="https://zenodo.org/record/7762480/files/figure.png" pageId="10" pageNumber="511">Fig. 7H</figureCitation>
) long, acuminate blades; most distal blade with small, finger-like, protuberances clustered at distal end; when present on subdistal blade, only single protuberance present. Male rallum with seven long, acuminate blades; distal and subdistal blades with two to three, finger-like, protuberances at distal end, spaced apart (
<figureCitation box="[222,301,475,501]" captionStart="FIGURE 7" captionStartId="10.[151,250,1907,1931]" captionTargetBox="[169,1415,542,1865]" captionTargetId="figure-177@10.[151,1436,525,1884]" captionTargetPageId="10" captionText="FIGURE 7. Gymnobisium capense sp. nov. A. Female right chela, retrolateral view; B. Female right pedipalp, dorsal view; C. Female left chelicera, dorsal view; D. Galea; E. Female chelal fingers, anterior retrolateral view; F. Male fixed chelal finger, anterior retrolateral view; G. Female rallum; H. Same; I. Male rallum. Scale bars: A, B: 0.50 mm; C, E, F: 0.20 mm; D, GI: 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.7762480" httpUri="https://zenodo.org/record/7762480/files/figure.png" pageId="10" pageNumber="511">Fig. 7I</figureCitation>
). Lamina exterior absent in both sexes.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7762480" ID-Zenodo-Dep="7762480" httpUri="https://zenodo.org/record/7762480/files/figure.png" pageId="10" pageNumber="511" startId="10.[151,250,1907,1931]" targetBox="[169,1415,542,1865]" targetPageId="10">
<paragraph blockId="10.[151,1437,1907,2039]" pageId="10" pageNumber="511">
<emphasis bold="true" box="[151,269,1907,1931]" pageId="10" pageNumber="511">FIGURE 7</emphasis>
.
<taxonomicName authority="Neethling &amp; Neethling, 2023" authorityName="Neethling &amp; Neethling" authorityYear="2023" box="[281,510,1907,1931]" class="Arachnida" family="Gymnobisiidae" genus="Gymnobisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="10" pageNumber="511" phylum="Arthropoda" rank="species" species="capense" status="sp. nov.">
<emphasis box="[281,510,1907,1931]" italics="true" pageId="10" pageNumber="511">Gymnobisium capense</emphasis>
</taxonomicName>
<emphasis bold="true" box="[516,629,1907,1931]" pageId="10" pageNumber="511">
<taxonomicNameLabel box="[516,597,1907,1931]" pageId="10" pageNumber="511" rank="species">sp. nov.</taxonomicNameLabel>
A.
</emphasis>
Female right chela, retrolateral view;
<emphasis bold="true" box="[1016,1039,1907,1931]" pageId="10" pageNumber="511">B.</emphasis>
Female right pedipalp, dorsal view;
<emphasis bold="true" box="[1412,1436,1907,1931]" pageId="10" pageNumber="511">C.</emphasis>
Female left chelicera, dorsal view;
<emphasis bold="true" box="[514,539,1943,1967]" pageId="10" pageNumber="511">D.</emphasis>
Galea;
<emphasis bold="true" box="[620,644,1943,1967]" pageId="10" pageNumber="511">E.</emphasis>
Female chelal fingers, anterior retrolateral view;
<emphasis bold="true" box="[1155,1174,1943,1967]" pageId="10" pageNumber="511">F.</emphasis>
Male fixed chelal finger, anterior retrolateral view;
<emphasis bold="true" box="[419,445,1979,2003]" pageId="10" pageNumber="511">G.</emphasis>
Female rallum;
<emphasis bold="true" box="[614,640,1979,2003]" pageId="10" pageNumber="511">H.</emphasis>
Same;
<emphasis bold="true" box="[718,734,1979,2003]" pageId="10" pageNumber="511">I.</emphasis>
Male rallum. Scale bars: A, B: 0.50 mm; C, E, F: 0.20 mm; D, GI: 0.10 mm.
</paragraph>
</caption>
<paragraph blockId="11.[151,1437,151,1511]" pageId="11" pageNumber="512">
<emphasis box="[199,373,151,177]" italics="true" pageId="11" pageNumber="512">Coxae and legs</emphasis>
: Pedipalpal coxae same colour, or slightly lighter than rest of pedipalps. Coxae IIV tan. Legs IIV light brown to tan in colour. All legs diplotarsate with subterminal tarsal setae dentate distally; claws simple; arolium shorter than claws.
</paragraph>
<paragraph blockId="11.[151,1437,151,1511]" pageId="11" pageNumber="512">
<emphasis bold="true" box="[199,455,258,285]" pageId="11" pageNumber="512">Measurements (mm):</emphasis>
Body length
<specimenCount box="[607,628,259,285]" pageId="11" pageNumber="512" type="female"></specimenCount>
2.94
<geoCoordinate box="[698,775,259,287]" degrees="3.14" direction="west" orientation="longitude" pageId="11" pageNumber="512" precision="555" value="-3.14">3.14 ♂</geoCoordinate>
2.262.44; Carapace
<specimenCount box="[1019,1040,259,285]" pageId="11" pageNumber="512" type="female"></specimenCount>
0.820.85 x 0.820.89 (0.961.00)
<geoCoordinate box="[151,231,295,323]" degrees="0.62" direction="west" orientation="longitude" pageId="11" pageNumber="512" precision="555" value="-0.62">♂ 0.62</geoCoordinate>
0.68 x 0.700.73 (0.880.93); Chelicera
<specimenCount box="[712,733,295,321]" pageId="11" pageNumber="512" type="female"></specimenCount>
0.640.66 x 0.330.35 (1.891.94)
<geoCoordinate box="[1149,1228,294,323]" degrees="0.50" direction="west" orientation="longitude" pageId="11" pageNumber="512" precision="555" value="-0.5">♂ 0.50</geoCoordinate>
0.53 x 0.260.28 (1.891.92), movable finger length
<specimenCount box="[558,579,331,357]" pageId="11" pageNumber="512" type="female"></specimenCount>
0.38
<geoCoordinate box="[651,730,331,359]" degrees="0.40" direction="west" orientation="longitude" pageId="11" pageNumber="512" precision="555" value="-0.4">0.40 ♂</geoCoordinate>
0.310.33; Pedipalps: femur
<specimenCount box="[1070,1091,331,357]" pageId="11" pageNumber="512" type="female"></specimenCount>
1.061.11 x 0.280.30 (3.70 3.79)
<geoCoordinate box="[217,294,366,395]" degrees="0.85" direction="west" orientation="longitude" pageId="11" pageNumber="512" precision="555" value="-0.85">♂ 0.85</geoCoordinate>
0.91 x 0.220.23 (3.863.95), patella
<specimenCount box="[731,752,367,393]" pageId="11" pageNumber="512" type="female"></specimenCount>
0.880.90 x 0.320.35 (2.572.75)
<geoCoordinate box="[1156,1233,367,395]" degrees="0.74" direction="west" orientation="longitude" pageId="11" pageNumber="512" precision="555" value="-0.74">♂ 0.74</geoCoordinate>
0.79 x 0.260.27 (2.852.92), chela
<specimenCount box="[361,382,403,429]" pageId="11" pageNumber="512" type="female"></specimenCount>
1.831.90 x 0.580.64 (2.963.16)
<geoCoordinate box="[784,860,402,431]" degrees="1.59" direction="west" orientation="longitude" pageId="11" pageNumber="512" precision="555" value="-1.59">♂ 1.59</geoCoordinate>
1.74 x 0.410.47 (3.703.88), hand
<specimenCount box="[1276,1297,403,429]" pageId="11" pageNumber="512" type="female"></specimenCount>
0.940.97 x 0.580.64 (1.521.62)
<geoCoordinate box="[407,484,439,467]" degrees="0.79" direction="west" orientation="longitude" pageId="11" pageNumber="512" precision="555" value="-0.79">♂ 0.79</geoCoordinate>
0.89 x 0.410.47 (1.891.93), movable finger length
<specimenCount box="[1092,1113,439,465]" pageId="11" pageNumber="512" type="female"></specimenCount>
0.95
<geoCoordinate box="[1182,1259,438,467]" degrees="0.99" direction="west" orientation="longitude" pageId="11" pageNumber="512" precision="555" value="-0.99">0.99 ♂</geoCoordinate>
0.810.88; Leg I: femur
<specimenCount box="[253,274,475,501]" pageId="11" pageNumber="512" type="female"></specimenCount>
0.580.61 x 0.110.12 (5.085.27)
<geoCoordinate box="[687,765,474,503]" degrees="0.48" direction="west" orientation="longitude" pageId="11" pageNumber="512" precision="555" value="-0.48">♂ 0.48</geoCoordinate>
0.52 x 0.10 (4.805.20), patella
<specimenCount box="[1150,1171,475,501]" pageId="11" pageNumber="512" type="female"></specimenCount>
0.400.42 x 0.120.13 (3.233.33)
<geoCoordinate box="[289,366,511,539]" degrees="0.32" direction="west" orientation="longitude" pageId="11" pageNumber="512" precision="555" value="-0.32">♂ 0.32</geoCoordinate>
0.36 x 0.11 (2.913.27), tibia
<specimenCount box="[713,734,511,537]" pageId="11" pageNumber="512" type="female"></specimenCount>
0.450.47 x 0.10 (4.504.70)
<geoCoordinate box="[1075,1152,511,539]" degrees="0.36" direction="west" orientation="longitude" pageId="11" pageNumber="512" precision="555" value="-0.36">♂ 0.36</geoCoordinate>
0.40 x 0.08 (4.505.00), metatarsus
<specimenCount box="[278,299,547,573]" pageId="11" pageNumber="512" type="female"></specimenCount>
0.270.28 x 0.09 (3.003.11)
<geoCoordinate box="[639,716,547,575]" degrees="0.23" direction="west" orientation="longitude" pageId="11" pageNumber="512" precision="555" value="-0.23">♂ 0.23</geoCoordinate>
0.24 x 0.07 (3.283.43), tarsus
<specimenCount box="[1081,1102,547,573]" pageId="11" pageNumber="512" type="female"></specimenCount>
0.300.31 x 0.07 (4.294.43)
<geoCoordinate box="[151,229,582,611]" degrees="0.25" direction="west" orientation="longitude" pageId="11" pageNumber="512" precision="555" value="-0.25">♂ 0.25</geoCoordinate>
0.26 x 0.06 (4.174.33); Leg IV femoropatella
<specimenCount box="[782,803,583,609]" pageId="11" pageNumber="512" type="female"></specimenCount>
1.071.12 x 0.230.24 (4.654.67)
<geoCoordinate box="[1215,1293,582,611]" degrees="0.88" direction="west" orientation="longitude" pageId="11" pageNumber="512" precision="555" value="-0.88">♂ 0.88</geoCoordinate>
0.95 x 0.20 (4.404.75), tibia
<specimenCount box="[354,375,619,645]" pageId="11" pageNumber="512" type="female"></specimenCount>
0.880.89 x 0.16 (5.505.56)
<geoCoordinate box="[718,796,619,647]" degrees="0.69" direction="west" orientation="longitude" pageId="11" pageNumber="512" precision="555" value="-0.69">♂ 0.69</geoCoordinate>
0.76 x 0.13 (5.315.85), metatarsus
<specimenCount box="[1217,1238,619,645]" pageId="11" pageNumber="512" type="female"></specimenCount>
0.380.41 x 0.10 (3.804.10)
<geoCoordinate box="[292,371,654,683]" degrees="0.31" direction="west" orientation="longitude" pageId="11" pageNumber="512" precision="555" value="-0.31">♂ 0.31</geoCoordinate>
0.35 x 0.08 (3.884.36), tarsus
<specimenCount box="[751,772,655,681]" pageId="11" pageNumber="512" type="female"></specimenCount>
0.400.43 x 0.09 (4.444.78)
<geoCoordinate box="[1128,1208,655,683]" degrees="0.36" direction="west" orientation="longitude" pageId="11" pageNumber="512" precision="555" value="-0.36">♂ 0.36</geoCoordinate>
0.37 x 0.07 (5.14 5.29).
</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="512" type="discussion">
<paragraph blockId="11.[151,1437,151,1511]" pageId="11" pageNumber="512">
<emphasis bold="true" box="[199,315,727,753]" pageId="11" pageNumber="512">Remarks.</emphasis>
The first specimen of this species was collected from the forests near Knysna in 1947 by R.F. Lawrence, but was misidentified as
<taxonomicName baseAuthorityName="Tullgren" baseAuthorityYear="1907" box="[554,888,763,789]" class="Arachnida" family="Gymnobisiidae" genus="Gymnobisum" kingdom="Animalia" order="Pseudoscorpiones" pageId="11" pageNumber="512" phylum="Arthropoda" rank="species" species="quadrispinosum">
<emphasis box="[554,888,763,789]" italics="true" pageId="11" pageNumber="512">Gymnobisum quadrispinosum</emphasis>
</taxonomicName>
by
<bibRefCitation author="Beier, M." box="[931,1076,763,789]" pageId="11" pageNumber="512" pagination="285 - 339" refId="ref27468" refString="Beier, M. (1947) Zur kenntnis der Pseudoscorpionidenfauna des Sudlichen Afrika, inbesondere der Sudwestund Sudafrikanischen trockengebiete. Eos, 23, 285 - 339." type="journal article" year="1947">Beier (1947)</bibRefCitation>
.
<taxonomicName authority="Neethling &amp; Neethling, 2023" authorityName="Neethling &amp; Neethling" authorityYear="2023" box="[1087,1340,763,789]" class="Arachnida" family="Gymnobisiidae" genus="Gymnobisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="11" pageNumber="512" phylum="Arthropoda" rank="species" species="capense" status="sp. nov.">
<emphasis box="[1087,1340,763,789]" italics="true" pageId="11" pageNumber="512">Gymnobisium capense</emphasis>
</taxonomicName>
<emphasis bold="true" box="[1347,1436,763,789]" pageId="11" pageNumber="512">
<taxonomicNameLabel box="[1347,1436,763,789]" pageId="11" pageNumber="512" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is the only gymnobisiid other than
<taxonomicName authorityName="Harvey &amp; Giribet" authorityYear="2016" box="[542,676,799,825]" class="Arachnida" family="Gymnobisiidae" genus="Gymnobisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="11" pageNumber="512" phylum="Arthropoda" rank="species" species="inukshuk">
<emphasis box="[542,676,799,825]" italics="true" pageId="11" pageNumber="512">G. inukshuk</emphasis>
</taxonomicName>
with six setae on the cheliceral hand.
</paragraph>
<paragraph blockId="11.[151,1437,151,1511]" pageId="11" pageNumber="512">
<emphasis bold="true" box="[199,563,834,861]" pageId="11" pageNumber="512">Additional material examined.</emphasis>
<materialsCitation collectingDate="2012-12-05" collectionCode="NMBAP" collectorName="Leaf &amp; J. A. Neethling" country="South Africa" county="Tritonymphs" elevation="185" latitude="-33.983334" location="Stormsrivier" longLatPrecision="1202" longitude="23.9" municipality="Deutonymphs" pageId="11" pageNumber="512" specimenCode="NMBAP 00331" specimenCount="1" specimenCount-female="1" stateProvince="Eastern Cape">
<emphasis bold="true" box="[570,782,834,860]" pageId="11" pageNumber="512">
<collectingCountry box="[570,782,834,860]" name="South Africa" pageId="11" pageNumber="512">SOUTH AFRICA</collectingCountry>
</emphasis>
:
<emphasis box="[796,949,835,861]" italics="true" pageId="11" pageNumber="512">
<collectingRegion box="[796,949,835,861]" country="South Africa" name="Eastern Cape" pageId="11" pageNumber="512">Eastern Cape</collectingRegion>
</emphasis>
:
<specimenCount box="[963,998,835,861]" pageId="11" pageNumber="512" type="female">1♀</specimenCount>
, 2♂, 5
<collectingCounty box="[1080,1227,835,861]" pageId="11" pageNumber="512">Tritonymphs</collectingCounty>
, 10
<collectingMunicipality box="[1272,1432,835,861]" pageId="11" pageNumber="512">Deutonymphs</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:EB305F51B51FF67CFF1EFAA0FA93FA60:6646B89CB513F67CFF1EFCE6FEF4FC04" box="[151,293,871,897]" country="South Africa" county="Tritonymphs" latitude="-33.983334" longLatPrecision="1202" longitude="23.9" municipality="Deutonymphs" name="Stormsrivier" pageId="11" pageNumber="512" stateProvince="Eastern Cape">Stormsrivier</location>
,
<location LSID="urn:lsid:plazi:treatment:EB305F51B51FF67CFF1EFAA0FA93FA60:6646B89CB513F67CFEBCFCE6FD02FC04" box="[309,723,871,897]" country="South Africa" county="Tritonymphs" latitude="-33.983334" longLatPrecision="1202" longitude="23.9" municipality="Deutonymphs" name="Stormsrivier Forest Nature Reserve" pageId="11" pageNumber="512" stateProvince="Eastern Cape">Stormsrivier Forest Nature Reserve</location>
,
<geoCoordinate box="[739,830,872,899]" degrees="33" direction="south" minutes="59" orientation="latitude" pageId="11" pageNumber="512" precision="925" value="-33.983334">33°59S</geoCoordinate>
,
<geoCoordinate box="[846,940,872,899]" degrees="23" direction="east" minutes="54" orientation="longitude" pageId="11" pageNumber="512" precision="925" value="23.9">23°54E</geoCoordinate>
,
<elevation box="[955,1085,870,897]" metricMagnitude="2" metricUnit="m" metricValue="1.85" pageId="11" pageNumber="512" unit="m" value="185.0">
<quantity box="[955,1019,870,897]" metricMagnitude="2" metricUnit="m" metricValue="1.85" pageId="11" pageNumber="512" unit="m" value="185.0">185m</quantity>
a.s.l.
</elevation>
,
<location LSID="urn:lsid:plazi:treatment:EB305F51B51FF67CFF1EFAA0FA93FA60:6646B89CB513F67CFBC7FCE6FAC8FC04" box="[1102,1305,871,897]" country="South Africa" county="Tritonymphs" latitude="-33.983334" longLatPrecision="1202" longitude="23.9" municipality="Deutonymphs" name="Indigenous forest" pageId="11" pageNumber="512" stateProvince="Eastern Cape">Indigenous forest</location>
,
<collectorName box="[1320,1371,871,897]" pageId="11" pageNumber="512">Leaf</collectorName>
litter sifting, leg.
<collectorName box="[292,460,907,933]" pageId="11" pageNumber="512">J.A. Neethling</collectorName>
,
<date box="[475,598,906,933]" pageId="11" pageNumber="512" value="2012-12-05">
<collectingDate box="[475,598,906,933]" pageId="11" pageNumber="512" value="2012-12-05">5.XII.2012</collectingDate>
</date>
(
<specimenCode box="[615,801,907,933]" collectionCode="NMBAP" pageId="11" pageNumber="512">NMBAP 00331</specimenCode>
)
</materialsCitation>
;
<materialsCitation collectingDate="2012-12-05" collectionCode="NMBAP" collectorName="Leaf &amp; J. A. Neethling" country="South Africa" county="Tritonymph" elevation="47" latitude="-34.016666" location="Stormsrivier" longLatPrecision="1202" longitude="23.883333" municipality="Deutonymph" pageId="11" pageNumber="512" specimenCode="NMBAP 00330" specimenCount="1" specimenCount-female="1" stateProvince="Eastern Cape">
<specimenCount box="[824,859,907,933]" pageId="11" pageNumber="512" type="female">1♀</specimenCount>
, 1♂, 1
<collectingCounty box="[952,1088,907,933]" pageId="11" pageNumber="512">Tritonymph</collectingCounty>
, 1
<collectingMunicipality box="[1127,1276,907,933]" pageId="11" pageNumber="512">Deutonymph</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:EB305F51B51FF67CFF1EFAA0FA93FA60:6646B89CB513F67CFA83FC0AFA49FC20" box="[1290,1432,907,933]" country="South Africa" county="Tritonymph" latitude="-34.016666" longLatPrecision="1202" longitude="23.883333" municipality="Deutonymph" name="Stormsrivier" pageId="11" pageNumber="512" stateProvince="Eastern Cape">Stormsrivier</location>
,
<location LSID="urn:lsid:plazi:treatment:EB305F51B51FF67CFF1EFAA0FA93FA60:6646B89CB513F67CFF1EFC2EFE32FC4C" box="[151,483,943,969]" country="South Africa" county="Tritonymph" latitude="-34.016666" longLatPrecision="1202" longitude="23.883333" municipality="Deutonymph" name="Tsitsikamma Nature Reserve" pageId="11" pageNumber="512" stateProvince="Eastern Cape">Tsitsikamma Nature Reserve</location>
,
<geoCoordinate box="[497,574,944,971]" degrees="34" direction="south" minutes="1" orientation="latitude" pageId="11" pageNumber="512" precision="925" value="-34.016666">34°1S</geoCoordinate>
,
<geoCoordinate box="[589,682,944,971]" degrees="23" direction="east" minutes="53" orientation="longitude" pageId="11" pageNumber="512" precision="925" value="23.883333">23°53E</geoCoordinate>
,
<elevation box="[696,810,943,969]" metricMagnitude="1" metricUnit="m" metricValue="4.7" pageId="11" pageNumber="512" unit="m" value="47.0">
<quantity box="[696,746,943,969]" metricMagnitude="1" metricUnit="m" metricValue="4.7" pageId="11" pageNumber="512" unit="m" value="47.0">47m</quantity>
a.s.l.
</elevation>
,
<location LSID="urn:lsid:plazi:treatment:EB305F51B51FF67CFF1EFAA0FA93FA60:6646B89CB513F67CFCB0FC2EFC0BFC4C" box="[825,986,943,969]" country="South Africa" county="Tritonymph" latitude="-34.016666" longLatPrecision="1202" longitude="23.883333" municipality="Deutonymph" name="Coastal forest" pageId="11" pageNumber="512" stateProvince="Eastern Cape">Coastal forest</location>
,
<collectorName box="[1000,1051,943,969]" pageId="11" pageNumber="512">Leaf</collectorName>
litter sifting, leg.
<collectorName box="[1264,1432,943,969]" pageId="11" pageNumber="512">J.A. Neethling</collectorName>
,
<date box="[151,274,978,1005]" pageId="11" pageNumber="512" value="2012-12-05">
<collectingDate box="[151,274,978,1005]" pageId="11" pageNumber="512" value="2012-12-05">5.XII.2012</collectingDate>
</date>
(
<specimenCode box="[292,478,979,1005]" collectionCode="NMBAP" pageId="11" pageNumber="512">NMBAP 00330</specimenCode>
)
</materialsCitation>
;
<materialsCitation collectingDate="2013-12-20" collectionCode="NMBAP" collectorName="J. A. Neethling &amp; C. Neethling" country="South Africa" elevation="47" latitude="-34.016666" location="Coastal forest" longLatPrecision="1202" longitude="23.883333" municipality="Same" pageId="11" pageNumber="512" specimenCode="NMBAP 00329" specimenCount="1" specimenCount-female="1" stateProvince="Eastern Cape">
<specimenCount box="[501,536,979,1005]" pageId="11" pageNumber="512" type="female">1♀</specimenCount>
,
<collectingMunicipality box="[553,615,979,1005]" pageId="11" pageNumber="512">Same</collectingMunicipality>
locality,
<geoCoordinate box="[726,803,980,1007]" degrees="34" direction="south" minutes="1" orientation="latitude" pageId="11" pageNumber="512" precision="925" value="-34.016666">34°1S</geoCoordinate>
,
<geoCoordinate box="[818,911,980,1007]" degrees="23" direction="east" minutes="53" orientation="longitude" pageId="11" pageNumber="512" precision="925" value="23.883333">23°53E</geoCoordinate>
,
<elevation box="[926,1040,979,1005]" metricMagnitude="1" metricUnit="m" metricValue="4.7" pageId="11" pageNumber="512" unit="m" value="47.0">
<quantity box="[926,976,979,1005]" metricMagnitude="1" metricUnit="m" metricValue="4.7" pageId="11" pageNumber="512" unit="m" value="47.0">47m</quantity>
a.s.l.
</elevation>
,
<location LSID="urn:lsid:plazi:treatment:EB305F51B51FF67CFF1EFAA0FA93FA60:6646B89CB513F67CFBA9FC52FB13FC68" box="[1056,1218,979,1005]" country="South Africa" latitude="-34.016666" longLatPrecision="1202" longitude="23.883333" municipality="Same" name="Coastal forest" pageId="11" pageNumber="512" stateProvince="Eastern Cape">Coastal forest</location>
,
<collectorName box="[1232,1283,979,1005]" pageId="11" pageNumber="512">Leaf</collectorName>
litter sifting, leg.
<collectorName box="[200,363,1015,1041]" pageId="11" pageNumber="512">J.A. Neethling</collectorName>
&amp;
<collectorName box="[401,547,1015,1041]" pageId="11" pageNumber="512">C. Neethling</collectorName>
,
<date box="[559,696,1015,1041]" pageId="11" pageNumber="512" value="2013-12-20">
<collectingDate box="[559,696,1015,1041]" pageId="11" pageNumber="512" value="2013-12-20">20.XII.2013</collectingDate>
</date>
(
<specimenCode box="[711,895,1015,1041]" collectionCode="NMBAP" pageId="11" pageNumber="512">NMBAP 00329</specimenCode>
)
</materialsCitation>
;
<materialsCitation collectingDate="1985-11-13" collectorName="J. T. Doyen &amp; C. E. Griswold" country="South Africa" county="Knysna" elevation="548" latitude="-33.95" location="Indigenous forest" longLatPrecision="1202" longitude="23.166666" municipality="Diepwalle Forest" pageId="11" pageNumber="512" specimenCount="1" stateProvince="Western Cape">
<emphasis box="[915,1071,1015,1041]" italics="true" pageId="11" pageNumber="512">
<collectingRegion box="[915,1071,1015,1041]" country="South Africa" name="Western Cape" pageId="11" pageNumber="512">Western Cape</collectingRegion>
</emphasis>
: 1♂,
<collectingCounty box="[1136,1224,1015,1041]" pageId="11" pageNumber="512">Knysna</collectingCounty>
,
<collectingMunicipality box="[1236,1432,1015,1041]" pageId="11" pageNumber="512">Diepwalle Forest</collectingMunicipality>
,
<geoCoordinate box="[151,243,1052,1079]" degrees="33" direction="south" minutes="57" orientation="latitude" pageId="11" pageNumber="512" precision="925" value="-33.95">33°57S</geoCoordinate>
,
<geoCoordinate box="[257,349,1052,1079]" degrees="23" direction="east" minutes="10" orientation="longitude" pageId="11" pageNumber="512" precision="925" value="23.166666">23°10E</geoCoordinate>
,
<elevation box="[363,490,1050,1077]" metricMagnitude="2" metricUnit="m" metricValue="5.4799999999999995" pageId="11" pageNumber="512" unit="m" value="548.0">
<quantity box="[363,427,1050,1077]" metricMagnitude="2" metricUnit="m" metricValue="5.4799999999999995" pageId="11" pageNumber="512" unit="m" value="548.0">548m</quantity>
a.s.l.
</elevation>
,
<location LSID="urn:lsid:plazi:treatment:EB305F51B51FF67CFF1EFAA0FA93FA60:6646B89CB513F67CFE70FB9AFD13FBB0" box="[505,706,1051,1077]" country="South Africa" county="Knysna" latitude="-33.95" longLatPrecision="1202" longitude="23.166666" municipality="Diepwalle Forest" name="Indigenous forest" pageId="11" pageNumber="512" stateProvince="Western Cape">Indigenous forest</location>
,
<collectorName box="[719,805,1051,1077]" pageId="11" pageNumber="512">Burlese</collectorName>
funnels, leg.
<collectorName box="[964,1088,1051,1077]" pageId="11" pageNumber="512">J.T. Doyen</collectorName>
&amp;
<collectorName box="[1129,1295,1051,1077]" pageId="11" pageNumber="512">C.E. Griswold</collectorName>
,
<date box="[1309,1436,1050,1077]" pageId="11" pageNumber="512" value="1985-11-13">
<collectingDate box="[1309,1436,1050,1077]" pageId="11" pageNumber="512" value="1985-11-13">13.XI.1985</collectingDate>
</date>
(NM 19969)
</materialsCitation>
;
<materialsCitation collectionCode="SAMC" collectorName="R. F. Lawrence" country="South Africa" latitude="-33.933334" location="Diepwalle Forest" longLatPrecision="1202" longitude="23.116667" municipality="Same" pageId="11" pageNumber="512" specimenCode="B6911" specimenCount="1" specimenCount-female="1" stateProvince="Western Cape">
<specimenCount box="[311,346,1087,1113]" pageId="11" pageNumber="512" type="female">1♀</specimenCount>
,
<collectingMunicipality box="[362,424,1087,1113]" pageId="11" pageNumber="512">Same</collectingMunicipality>
location,
<location LSID="urn:lsid:plazi:treatment:EB305F51B51FF67CFF1EFAA0FA93FA60:6646B89CB513F67CFD92FBBEFD0FFBDC" box="[539,734,1087,1113]" country="South Africa" latitude="-33.933334" longLatPrecision="1202" longitude="23.116667" municipality="Same" name="Diepwalle Forest" pageId="11" pageNumber="512" stateProvince="Western Cape">Diepwalle Forest</location>
[
<geoCoordinate box="[750,843,1088,1115]" degrees="33" direction="south" minutes="56" orientation="latitude" pageId="11" pageNumber="512" precision="925" value="-33.933334">33°56S</geoCoordinate>
,
<geoCoordinate box="[857,938,1088,1115]" degrees="23" direction="east" minutes="7" orientation="longitude" pageId="11" pageNumber="512" precision="925" value="23.116667">23°7E</geoCoordinate>
], leg.
<collectorName box="[1009,1177,1087,1113]" pageId="11" pageNumber="512">R.F. Lawrence</collectorName>
, 1947 (
<collectionCode box="[1263,1344,1087,1113]" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" pageId="11" pageNumber="512" type="Museum">SAMC</collectionCode>
<specimenCode box="[1353,1428,1087,1113]" collectionCode="SAMC" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" pageId="11" pageNumber="512" type="Museum">B6911</specimenCode>
) (misidentified as
<taxonomicName baseAuthorityName="Tullgren" baseAuthorityYear="1907" box="[346,689,1123,1149]" class="Arachnida" family="Gymnobisiidae" genus="Gymnobisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="11" pageNumber="512" phylum="Arthropoda" rank="species" species="quadrispinosum">
<emphasis box="[346,689,1123,1149]" italics="true" pageId="11" pageNumber="512">Gymnobisium quadrispinosum</emphasis>
</taxonomicName>
)
</materialsCitation>
;
<materialsCitation collectingDate="2017-01-31" collectionCode="NMBAP" collectorName="Leaf &amp; J. A. Neethling" country="South Africa" county="Tritonymph" elevation="680" latitude="-34.9" location="Valley of Ferns" longLatPrecision="1197" longitude="23.133333" municipality="Same" pageId="11" pageNumber="512" specimenCode="NMBAP 00321" specimenCount="4" specimenCount-female="4" stateProvince="Western Cape">
<specimenCount box="[714,751,1123,1149]" pageId="11" pageNumber="512" type="female">4♀</specimenCount>
, 7♂, 1
<collectingCounty box="[834,970,1123,1149]" pageId="11" pageNumber="512">Tritonymph</collectingCounty>
,
<collectingMunicipality box="[981,1043,1123,1149]" pageId="11" pageNumber="512">Same</collectingMunicipality>
location,
<location LSID="urn:lsid:plazi:treatment:EB305F51B51FF67CFF1EFAA0FA93FA60:6646B89CB513F67CFB0BFBE2FAE0FBF8" box="[1154,1329,1123,1149]" country="South Africa" county="Tritonymph" latitude="-34.9" longLatPrecision="1197" longitude="23.133333" municipality="Same" name="Valley of Ferns" pageId="11" pageNumber="512" stateProvince="Western Cape">Valley of Ferns</location>
,
<geoCoordinate box="[1340,1432,1124,1151]" degrees="34" direction="south" minutes="54" orientation="latitude" pageId="11" pageNumber="512" precision="925" value="-34.9">34°54S</geoCoordinate>
,
<geoCoordinate box="[151,230,1160,1187]" degrees="23" direction="east" minutes="8" orientation="longitude" pageId="11" pageNumber="512" precision="925" value="23.133333">23°8E</geoCoordinate>
,
<elevation box="[246,374,1159,1185]" metricMagnitude="2" metricUnit="m" metricValue="6.8" pageId="11" pageNumber="512" unit="m" value="680.0">
<quantity box="[246,309,1159,1185]" metricMagnitude="2" metricUnit="m" metricValue="6.8" pageId="11" pageNumber="512" unit="m" value="680.0">680m</quantity>
a.s.l.
</elevation>
,
<location LSID="urn:lsid:plazi:treatment:EB305F51B51FF67CFF1EFAA0FA93FA60:6646B89CB513F67CFE0FFB06FD80FB24" box="[390,593,1159,1185]" country="South Africa" county="Tritonymph" latitude="-34.9" longLatPrecision="1197" longitude="23.133333" municipality="Same" name="Indigenous forest" pageId="11" pageNumber="512" stateProvince="Western Cape">Indigenous forest</location>
,
<collectorName box="[607,658,1159,1185]" pageId="11" pageNumber="512">Leaf</collectorName>
litter sifting, leg.
<collectorName box="[875,1044,1159,1185]" pageId="11" pageNumber="512">J.A. Neethling</collectorName>
,
<date box="[1059,1167,1159,1185]" pageId="11" pageNumber="512" value="2017-01-31">
<collectingDate box="[1059,1167,1159,1185]" pageId="11" pageNumber="512" value="2017-01-31">31.I.2017</collectingDate>
</date>
(
<specimenCode box="[1185,1371,1159,1185]" collectionCode="NMBAP" pageId="11" pageNumber="512">NMBAP 00321</specimenCode>
)
</materialsCitation>
;
<materialsCitation collectingDate="2011-12-20" collectionCode="NMBAP" collectorName="J. A. Neethling &amp; C. Neethling" country="South Africa" county="Rheenendal" elevation="308" latitude="-33.883335" location="Afromontane forest" longLatPrecision="1202" longitude="22.966667" municipality="Jubilee Creek Nature Reserve" pageId="11" pageNumber="512" specimenCode="NMBAP 00317" specimenCount="1" stateProvince="Western Cape">
1♂,
<collectingCounty box="[151,286,1195,1221]" pageId="11" pageNumber="512">Rheenendal</collectingCounty>
,
<collectingMunicipality box="[299,639,1195,1221]" pageId="11" pageNumber="512">Jubilee Creek Nature Reserve</collectingMunicipality>
,
<geoCoordinate box="[652,743,1196,1223]" degrees="33" direction="south" minutes="53" orientation="latitude" pageId="11" pageNumber="512" precision="925" value="-33.883335">33°53S</geoCoordinate>
,
<geoCoordinate box="[756,849,1196,1223]" degrees="22" direction="east" minutes="58" orientation="longitude" pageId="11" pageNumber="512" precision="925" value="22.966667">22°58E</geoCoordinate>
,
<elevation box="[862,988,1195,1221]" metricMagnitude="2" metricUnit="m" metricValue="3.08" pageId="11" pageNumber="512" unit="m" value="308.0">
<quantity box="[862,925,1195,1221]" metricMagnitude="2" metricUnit="m" metricValue="3.08" pageId="11" pageNumber="512" unit="m" value="308.0">308m</quantity>
a.s.l.
</elevation>
,
<location LSID="urn:lsid:plazi:treatment:EB305F51B51FF67CFF1EFAA0FA93FA60:6646B89CB513F67CFC61FB2AFB18FB40" box="[1000,1225,1195,1221]" country="South Africa" county="Rheenendal" latitude="-33.883335" longLatPrecision="1202" longitude="22.966667" municipality="Jubilee Creek Nature Reserve" name="Afromontane forest" pageId="11" pageNumber="512" stateProvince="Western Cape">Afromontane forest</location>
,
<collectorName box="[1237,1288,1195,1221]" pageId="11" pageNumber="512">Leaf</collectorName>
litter sifting, leg.
<collectorName box="[204,370,1231,1257]" pageId="11" pageNumber="512">J.A. Neethling</collectorName>
&amp;
<collectorName box="[414,564,1231,1257]" pageId="11" pageNumber="512">C. Neethling</collectorName>
,
<date box="[580,716,1231,1257]" pageId="11" pageNumber="512" value="2011-12-20">
<collectingDate box="[580,716,1231,1257]" pageId="11" pageNumber="512" value="2011-12-20">20.XII.2011</collectingDate>
</date>
(
<specimenCode box="[734,921,1231,1257]" collectionCode="NMBAP" pageId="11" pageNumber="512">NMBAP 00317</specimenCode>
)
</materialsCitation>
;
<materialsCitation collectingDate="2012-12-08" collectionCode="NMBAP" collectorName="Leaf &amp; J. A. Neethling" country="South Africa" elevation="308" latitude="-33.883335" location="Afromontane forest" longLatPrecision="1202" longitude="22.966667" municipality="Same" pageId="11" pageNumber="512" specimenCode="NMBAP 00316" specimenCount="3" specimenCount-female="3" stateProvince="Western Cape">
<specimenCount box="[944,980,1231,1257]" pageId="11" pageNumber="512" type="female">3♀</specimenCount>
, 3♂,
<collectingMunicipality box="[1050,1112,1231,1257]" pageId="11" pageNumber="512">Same</collectingMunicipality>
location,
<geoCoordinate box="[1232,1323,1232,1259]" degrees="33" direction="south" minutes="53" orientation="latitude" pageId="11" pageNumber="512" precision="925" value="-33.883335">33°53S</geoCoordinate>
,
<geoCoordinate box="[1339,1432,1232,1259]" degrees="22" direction="east" minutes="58" orientation="longitude" pageId="11" pageNumber="512" precision="925" value="22.966667">22°58E</geoCoordinate>
,
<elevation box="[151,275,1267,1293]" metricMagnitude="2" metricUnit="m" metricValue="3.08" pageId="11" pageNumber="512" unit="m" value="308.0">
<quantity box="[151,215,1267,1293]" metricMagnitude="2" metricUnit="m" metricValue="3.08" pageId="11" pageNumber="512" unit="m" value="308.0">308m</quantity>
a.s.l.
</elevation>
,
<location LSID="urn:lsid:plazi:treatment:EB305F51B51FF67CFF1EFAA0FA93FA60:6646B89CB513F67CFE94FB72FE2AFA88" box="[285,507,1267,1293]" country="South Africa" latitude="-33.883335" longLatPrecision="1202" longitude="22.966667" municipality="Same" name="Afromontane forest" pageId="11" pageNumber="512" stateProvince="Western Cape">Afromontane forest</location>
,
<collectorName box="[517,568,1267,1293]" pageId="11" pageNumber="512">Leaf</collectorName>
litter sifting, leg.
<collectorName box="[766,930,1267,1293]" pageId="11" pageNumber="512">J.A. Neethling</collectorName>
,
<date box="[941,1064,1267,1293]" pageId="11" pageNumber="512" value="2012-12-08">
<collectingDate box="[941,1064,1267,1293]" pageId="11" pageNumber="512" value="2012-12-08">8.XII.2012</collectingDate>
</date>
(
<specimenCode box="[1078,1259,1267,1293]" collectionCode="NMBAP" pageId="11" pageNumber="512">NMBAP 00316</specimenCode>
)
</materialsCitation>
;
<materialsCitation collectingDate="2013-12-18" collectionCode="NMBAP" collectorName="J. A. Neethling &amp; C. Neethling" country="South Africa" elevation="308" latitude="-33.883335" location="Afromontane forest" longLatPrecision="1202" longitude="22.966667" municipality="Same" pageId="11" pageNumber="512" specimenCode="NMBAP 00315" specimenCount="1" specimenCount-female="1" stateProvince="Western Cape">
<specimenCount box="[1278,1312,1267,1293]" pageId="11" pageNumber="512" type="female">1♀</specimenCount>
, 2♂,
<collectingMunicipality box="[1374,1436,1267,1293]" pageId="11" pageNumber="512">Same</collectingMunicipality>
location,
<geoCoordinate box="[256,347,1304,1331]" degrees="33" direction="south" minutes="53" orientation="latitude" pageId="11" pageNumber="512" precision="925" value="-33.883335">33°53S</geoCoordinate>
,
<geoCoordinate box="[359,452,1304,1331]" degrees="22" direction="east" minutes="58" orientation="longitude" pageId="11" pageNumber="512" precision="925" value="22.966667">22°58E</geoCoordinate>
,
<elevation box="[463,588,1303,1329]" metricMagnitude="2" metricUnit="m" metricValue="3.08" pageId="11" pageNumber="512" unit="m" value="308.0">
<quantity box="[463,527,1303,1329]" metricMagnitude="2" metricUnit="m" metricValue="3.08" pageId="11" pageNumber="512" unit="m" value="308.0">308m</quantity>
a.s.l.
</elevation>
,
<location LSID="urn:lsid:plazi:treatment:EB305F51B51FF67CFF1EFAA0FA93FA60:6646B89CB513F67CFDDEFA96FCE6FAB4" box="[599,823,1303,1329]" country="South Africa" latitude="-33.883335" longLatPrecision="1202" longitude="22.966667" municipality="Same" name="Afromontane forest" pageId="11" pageNumber="512" stateProvince="Western Cape">Afromontane forest</location>
,
<collectorName box="[834,885,1303,1329]" pageId="11" pageNumber="512">Leaf</collectorName>
litter sifting, leg.
<collectorName box="[1087,1249,1303,1329]" pageId="11" pageNumber="512">J.A. Neethling</collectorName>
&amp;
<collectorName box="[1286,1432,1303,1329]" pageId="11" pageNumber="512">C. Neethling</collectorName>
,
<date box="[151,289,1339,1365]" pageId="11" pageNumber="512" value="2013-12-18">
<collectingDate box="[151,289,1339,1365]" pageId="11" pageNumber="512" value="2013-12-18">18.XII.2013</collectingDate>
</date>
(
<specimenCode box="[300,480,1338,1365]" collectionCode="NMBAP" pageId="11" pageNumber="512">NMBAP 00315</specimenCode>
)
</materialsCitation>
;
<materialsCitation collectingDate="2015-12-07" collectionCode="NMBAP" collectorName="J. A. Neethling &amp; C. Neethling" country="South Africa" county="Tritonymphs" elevation="308" latitude="-33.883335" location="Protonymph" longLatPrecision="1202" longitude="22.966667" municipality="Deutonymphs" pageId="11" pageNumber="512" specimenCode="NMBAP 00318" specimenCount="1" stateProvince="Western Cape">
3♂, 2
<collectingCounty box="[560,707,1339,1365]" pageId="11" pageNumber="512">Tritonymphs</collectingCounty>
, 2
<collectingMunicipality box="[734,894,1339,1365]" pageId="11" pageNumber="512">Deutonymphs</collectingMunicipality>
, 1
<location LSID="urn:lsid:plazi:treatment:EB305F51B51FF67CFF1EFAA0FA93FA60:6646B89CB513F67CFC10FABAFBF7FAD0" box="[921,1062,1339,1365]" country="South Africa" county="Tritonymphs" latitude="-33.883335" longLatPrecision="1202" longitude="22.966667" municipality="Deutonymphs" name="Protonymph" pageId="11" pageNumber="512" stateProvince="Western Cape">Protonymph</location>
,
<location LSID="urn:lsid:plazi:treatment:EB305F51B51FF67CFF1EFAA0FA93FA60:6646B89CB513F67CFBA7FABAFBBDFAD0" box="[1070,1132,1339,1365]" country="South Africa" county="Tritonymphs" latitude="-33.883335" longLatPrecision="1202" longitude="22.966667" municipality="Deutonymphs" name="Same" pageId="11" pageNumber="512" stateProvince="Western Cape">Same</location>
location,
<geoCoordinate box="[1239,1330,1340,1367]" degrees="33" direction="south" minutes="53" orientation="latitude" pageId="11" pageNumber="512" precision="925" value="-33.883335">33°53S</geoCoordinate>
,
<geoCoordinate box="[1339,1432,1340,1367]" degrees="22" direction="east" minutes="58" orientation="longitude" pageId="11" pageNumber="512" precision="925" value="22.966667">22°58E</geoCoordinate>
,
<elevation box="[151,274,1375,1401]" metricMagnitude="2" metricUnit="m" metricValue="3.08" pageId="11" pageNumber="512" unit="m" value="308.0">
<quantity box="[151,215,1375,1401]" metricMagnitude="2" metricUnit="m" metricValue="3.08" pageId="11" pageNumber="512" unit="m" value="308.0">308m</quantity>
a.s.l.
</elevation>
,
<location LSID="urn:lsid:plazi:treatment:EB305F51B51FF67CFF1EFAA0FA93FA60:6646B89CB513F67CFE92FADEFE28FAFC" box="[283,505,1375,1401]" country="South Africa" county="Tritonymphs" latitude="-33.883335" longLatPrecision="1202" longitude="22.966667" municipality="Deutonymphs" name="Afromontane forest" pageId="11" pageNumber="512" stateProvince="Western Cape">Afromontane forest</location>
,
<collectorName box="[514,565,1375,1401]" pageId="11" pageNumber="512">Leaf</collectorName>
litter sifting, leg.
<collectorName box="[760,921,1375,1401]" pageId="11" pageNumber="512">J.A. Neethling</collectorName>
&amp;
<collectorName box="[954,1098,1375,1401]" pageId="11" pageNumber="512">C. Neethling</collectorName>
,
<date box="[1108,1230,1374,1401]" pageId="11" pageNumber="512" value="2015-12-07">
<collectingDate box="[1108,1230,1374,1401]" pageId="11" pageNumber="512" value="2015-12-07">7.XII.2015</collectingDate>
</date>
(
<specimenCode box="[1243,1424,1375,1401]" collectionCode="NMBAP" pageId="11" pageNumber="512">NMBAP 00318</specimenCode>
)
</materialsCitation>
;
<materialsCitation collectingDate="2012-12-08" collectionCode="NMBAP" collectorName="Leaf &amp; J. A. Neethling" country="South Africa" county="Wildernis" elevation="254" latitude="-33.933334" location="Woodville Forest" longLatPrecision="1202" longitude="22.633333" municipality="Hoekville" pageId="11" pageNumber="512" specimenCode="NMBAP 00320" specimenCount="1" specimenCount-female="1" stateProvince="Western Cape">
<specimenCount box="[151,186,1411,1437]" pageId="11" pageNumber="512" type="female">1♀</specimenCount>
, 1♂,
<collectingCounty box="[246,358,1411,1437]" pageId="11" pageNumber="512">Wildernis</collectingCounty>
,
<collectingMunicipality box="[368,481,1411,1437]" pageId="11" pageNumber="512">Hoekville</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:EB305F51B51FF67CFF1EFAA0FA93FA60:6646B89CB513F67CFE62FA02FD7FFA18" box="[491,686,1411,1437]" country="South Africa" county="Wildernis" latitude="-33.933334" longLatPrecision="1202" longitude="22.633333" municipality="Hoekville" name="Woodville Forest" pageId="11" pageNumber="512" stateProvince="Western Cape">Woodville Forest</location>
,
<geoCoordinate box="[696,787,1412,1439]" degrees="33" direction="south" minutes="56" orientation="latitude" pageId="11" pageNumber="512" precision="925" value="-33.933334">33°56S</geoCoordinate>
,
<geoCoordinate box="[797,891,1412,1439]" degrees="22" direction="east" minutes="38" orientation="longitude" pageId="11" pageNumber="512" precision="925" value="22.633333">22°38E</geoCoordinate>
,
<elevation box="[901,1024,1410,1437]" metricMagnitude="2" metricUnit="m" metricValue="2.54" pageId="11" pageNumber="512" unit="m" value="254.0">
<quantity box="[901,964,1410,1437]" metricMagnitude="2" metricUnit="m" metricValue="2.54" pageId="11" pageNumber="512" unit="m" value="254.0">254m</quantity>
a.s.l.
</elevation>
,
<location LSID="urn:lsid:plazi:treatment:EB305F51B51FF67CFF1EFAA0FA93FA60:6646B89CB513F67CFB82FA02FB01FA18" box="[1035,1232,1411,1437]" country="South Africa" county="Wildernis" latitude="-33.933334" longLatPrecision="1202" longitude="22.633333" municipality="Hoekville" name="Indigenous forest" pageId="11" pageNumber="512" stateProvince="Western Cape">Indigenous forest</location>
,
<collectorName box="[1242,1293,1411,1437]" pageId="11" pageNumber="512">Leaf</collectorName>
litter sifting, leg.
<collectorName box="[203,371,1447,1473]" pageId="11" pageNumber="512">J.A. Neethling</collectorName>
,
<date box="[386,509,1447,1473]" pageId="11" pageNumber="512" value="2012-12-08">
<collectingDate box="[386,509,1447,1473]" pageId="11" pageNumber="512" value="2012-12-08">8.XII.2012</collectingDate>
</date>
(
<specimenCode box="[527,712,1447,1473]" collectionCode="NMBAP" pageId="11" pageNumber="512">NMBAP 00320</specimenCode>
)
</materialsCitation>
;
<materialsCitation collectingDate="2017-01-29" collectionCode="NMBAP" collectorName="Leaf &amp; J. A. Neethling" country="South Africa" county="Tritonymphs" elevation="254" latitude="-33.933334" location="Same" longLatPrecision="1202" longitude="22.633333" municipality="Deutonymph" pageId="11" pageNumber="512" specimenCode="NMBAP 00319" specimenCount="4" specimenCount-female="4" stateProvince="Western Cape">
<specimenCount box="[735,772,1447,1473]" pageId="11" pageNumber="512" type="female">4♀</specimenCount>
, 2
<collectingCounty box="[811,958,1447,1473]" pageId="11" pageNumber="512">Tritonymphs</collectingCounty>
, 1
<collectingMunicipality box="[997,1146,1447,1473]" pageId="11" pageNumber="512">Deutonymph</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:EB305F51B51FF67CFF1EFAA0FA93FA60:6646B89CB513F67CFB01FA26FB17FA44" box="[1160,1222,1447,1473]" country="South Africa" county="Tritonymphs" latitude="-33.933334" longLatPrecision="1202" longitude="22.633333" municipality="Deutonymph" name="Same" pageId="11" pageNumber="512" stateProvince="Western Cape">Same</location>
location,
<geoCoordinate box="[1340,1432,1448,1475]" degrees="33" direction="south" minutes="56" orientation="latitude" pageId="11" pageNumber="512" precision="925" value="-33.933334">33°56S</geoCoordinate>
,
<geoCoordinate box="[151,245,1484,1511]" degrees="22" direction="east" minutes="38" orientation="longitude" pageId="11" pageNumber="512" precision="925" value="22.633333">22°38E</geoCoordinate>
,
<elevation box="[256,380,1482,1509]" metricMagnitude="2" metricUnit="m" metricValue="2.54" pageId="11" pageNumber="512" unit="m" value="254.0">
<quantity box="[256,319,1482,1509]" metricMagnitude="2" metricUnit="m" metricValue="2.54" pageId="11" pageNumber="512" unit="m" value="254.0">254m</quantity>
a.s.l.
</elevation>
,
<location LSID="urn:lsid:plazi:treatment:EB305F51B51FF67CFF1EFAA0FA93FA60:6646B89CB513F67CFE00FA4AFD9EFA60" box="[393,591,1483,1509]" country="South Africa" county="Tritonymphs" latitude="-33.933334" longLatPrecision="1202" longitude="22.633333" municipality="Deutonymph" name="Indigenous forest" pageId="11" pageNumber="512" stateProvince="Western Cape">Indigenous forest</location>
,
<collectorName box="[602,653,1483,1509]" pageId="11" pageNumber="512">Leaf</collectorName>
litter sifting, leg.
<collectorName box="[854,1019,1483,1509]" pageId="11" pageNumber="512">J.A. Neethling</collectorName>
,
<date box="[1030,1137,1483,1509]" pageId="11" pageNumber="512" value="2017-01-29">
<collectingDate box="[1030,1137,1483,1509]" pageId="11" pageNumber="512" value="2017-01-29">29.I.2017</collectingDate>
</date>
(
<specimenCode box="[1152,1334,1483,1509]" collectionCode="NMBAP" pageId="11" pageNumber="512">NMBAP 00319</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>