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<mods:title>A systematic revision of the South African Gymnobisiidae (Pseudoscorpiones: Neobisioidea)</mods:title>
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<mods:affiliation>Terrestrial Invertebrates Department, National Museum, P. O. Box 266, Bloemfontein 9300, South Africa. ja. neethling @ nasmus. co. za; https: // orcid. org / 0000 - 0003 - 1702 - 9566</mods:affiliation>
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<mods:affiliation>Research Quality and Support, Clinglobal Limited, P. O. Box 11186, Bloemfontein 9321, South Africa. carmen. neethling @ clinglobal. com; https: // orcid. org / 0009 - 0003 - 6492 - 4798</mods:affiliation>
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Family
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Gymnobisiidae
<bibRefCitation author="Beier, M." box="[443,581,366,392]" pageId="3" pageNumber="504" pagination="285 - 339" refId="ref27468" refString="Beier, M. (1947) Zur kenntnis der Pseudoscorpionidenfauna des Sudlichen Afrika, inbesondere der Sudwestund Sudafrikanischen trockengebiete. Eos, 23, 285 - 339." type="journal article" year="1947">Beier, 1947</bibRefCitation>
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Gymnobisiidae
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;
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<bibRefCitation author="Harvey, M. S. &amp; Huey, J. &amp; Hillyer, M. &amp; McIntyre, E. &amp; Giribet, G." pageId="3" pageNumber="504" pagination="75 - 85" refId="ref28141" refString="Harvey, M. S., Huey, J., Hillyer, M., McIntyre, E. &amp; Giribet, G. (2016) The first troglobitic species of Gymnobisiidae (Pseudoscorpiones, Neobisioidea), from Table Mountain (Western Cape Province, South Africa) and its phylogenetic position. Invertebrate Systematics, 30, 75 - 85. https: // doi. org / 10.1071 / IS 15044" type="journal article" year="2016">
Harvey
<emphasis box="[1383,1437,435,459]" italics="true" pageId="3" pageNumber="504">et al.</emphasis>
2016: 7576
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, 79.
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<emphasis bold="true" box="[151,288,496,521]" pageId="3" pageNumber="504">
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Genus:
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<taxonomicName authority="Beier, 1931" authorityName="Beier" authorityYear="1931" box="[294,562,496,521]" class="Arachnida" family="Gymnobisiidae" genus="Gymnobisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="3" pageNumber="504" phylum="Arthropoda" rank="genus">
<emphasis box="[294,435,496,520]" italics="true" pageId="3" pageNumber="504">Gymnobisium</emphasis>
<bibRefCitation author="Beier, M." box="[441,562,496,521]" pageId="3" pageNumber="504" pagination="1 - 258" refId="ref27404" refString="Beier, M. (1931) Neue Pseudoscorpione der U. O. Neobisiinea. Mitteilungen aus dem Zoologischen Museum in Berlin, 17, 299 - 318. Beier, M. (1932) Pseudoscorpionidea I. Subord. Chthoniinea et Neobisiinea. Tierreich, 57, 1 - 258. https: // doi. org / 10.1515 / 9783111435107.1" type="journal article" year="1931">Beier, 1931</bibRefCitation>
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.
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<emphasis bold="true" box="[151,267,567,593]" pageId="3" pageNumber="504">Remarks.</emphasis>
According to
<bibRefCitation author="Harvey, M. S." box="[432,597,567,593]" pageId="3" pageNumber="504" pagination="1373 - 1435" refId="ref28020" refString="Harvey, M. S. (1992) The phylogeny and classification of the Pseudoscorpionida (Chelicerata: Arachnida). Invertebrate Taxonomy, 6, 1373 - 1435. https: // doi. org / 10.1071 / IT 9921373" type="journal article" year="1992">Harvey (1992)</bibRefCitation>
the
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have the following apomorphies: thickened spine-like seta present distally on movable chelal finger and prolaterally from the teeth, male genitalia that are greatly enlarged, and absence of a venom apparatus in the fixed chelal finger.
</paragraph>
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Further diagnostic characters include a movable chelal finger with a noticeably expanded nodus ramosus; both chelal fingers possessing lightly sclerotized, juxtadentate teeth on thin lamellae; the fixed chelal finger with a short row of highly sclerotized teeth, located terminally, and often accompanied by a single or multiple accessory teeth; the presence of a groove, proximal to the sclerotized teeth row, that receives the venedens of the movable finger; trichobothria
<emphasis box="[305,331,819,845]" italics="true" pageId="3" pageNumber="504">eb</emphasis>
,
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,
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,
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and
<emphasis box="[530,556,819,844]" italics="true" pageId="3" pageNumber="504">ist</emphasis>
clustered in a group at the base of the fixed chelal finger, with
<emphasis box="[1294,1325,819,844]" italics="true" pageId="3" pageNumber="504">est</emphasis>
,
<emphasis box="[1342,1358,819,844]" italics="true" pageId="3" pageNumber="504">it</emphasis>
and
<emphasis box="[1416,1436,819,844]" italics="true" pageId="3" pageNumber="504">et</emphasis>
grouping distally; the presence of a condyle on the retrolateral margin of each chelal hand; galea branched into multiple rami; suture line between leg IV femur and patella perpendicular (
<bibRefCitation author="Muchmore, W. B." box="[991,1183,891,917]" pageId="3" pageNumber="504" pagination="261 - 276" refId="ref28609" refString="Muchmore, W. B. (1972) New Diplosphyronid pseudoscorpions, mainly cavernicolous, from Mexico (Arachnida, Pseudoscorpionida). Transactions of the American Microscopical Society, 91, 261 - 276. https: // doi. org / 10.2307 / 3224874" type="journal article" year="1972">Muchmore 1972</bibRefCitation>
).
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