509 lines
52 KiB
XML
509 lines
52 KiB
XML
<document ID-DOI="10.11646/zootaxa.5256.6.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="7762456" ID-ZooBank="057285E1-92C9-4D76-921A-CE3C85D0D3B1" checkinTime="1679558471249" checkinUser="plazi" docAuthor="Neethling, J. A. & Neethling, C." docDate="2023" docId="EB305F51B517F664FF1EFB9BFC24FB94" docLanguage="en" docName="zootaxa.5256.6.1.pdf" docOrigin="Zootaxa 5256 (6)" docSource="http://dx.doi.org/10.11646/zootaxa.5256.6.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Gymnobisium hogsbackense Neethling & Neethling 2023, sp. nov." docType="treatment" docVersion="3" lastPageNumber="520" masterDocId="17092729B518F677FF89FF81FFD1FF85" masterDocTitle="A systematic revision of the South African Gymnobisiidae (Pseudoscorpiones: Neobisioidea)" masterLastPageNumber="543" masterPageNumber="501" pageNumber="516" updateTime="1679591149964" updateUser="juliana" zenodo-license-document="CLOSED">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A systematic revision of the South African Gymnobisiidae (Pseudoscorpiones: Neobisioidea)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Neethling, J. A.</mods:namePart>
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<mods:nameIdentifier type="ORCID">0000-0003-1702-9566</mods:nameIdentifier>
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<mods:affiliation>Terrestrial Invertebrates Department, National Museum, P. O. Box 266, Bloemfontein 9300, South Africa. ja. neethling @ nasmus. co. za; https: // orcid. org / 0000 - 0003 - 1702 - 9566</mods:affiliation>
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<mods:nameIdentifier type="email">ja.neethling@nasmus.co.za</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Neethling, C.</mods:namePart>
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<mods:nameIdentifier type="ORCID">0009-0003-6492-4798</mods:nameIdentifier>
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<mods:affiliation>Research Quality and Support, Clinglobal Limited, P. O. Box 11186, Bloemfontein 9321, South Africa. carmen. neethling @ clinglobal. com; https: // orcid. org / 0009 - 0003 - 6492 - 4798</mods:affiliation>
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<mods:nameIdentifier type="email">carmen.neethling@clinglobal.com</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-03-23</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>5256</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>6</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>501</mods:start>
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<mods:end>543</mods:end>
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</mods:extent>
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</mods:part>
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||
</mods:relatedItem>
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||
<mods:location>
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<mods:url>http://dx.doi.org/10.11646/zootaxa.5256.6.1</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.11646/zootaxa.5256.6.1</mods:identifier>
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<mods:identifier type="ISSN">1175-5326</mods:identifier>
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||
<mods:identifier type="Zenodo-Dep">7762456</mods:identifier>
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<mods:identifier type="ZooBank">057285E1-92C9-4D76-921A-CE3C85D0D3B1</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7762498" ID-Zenodo-Dep="7762498" LSID="urn:lsid:plazi:treatment:EB305F51B517F664FF1EFB9BFC24FB94" httpUri="http://treatment.plazi.org/id/EB305F51B517F664FF1EFB9BFC24FB94" lastPageId="19" lastPageNumber="520" pageId="15" pageNumber="516">
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<subSubSection box="[151,597,1050,1077]" pageId="15" pageNumber="516" type="nomenclature">
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||
<paragraph blockId="15.[151,597,1050,1113]" box="[151,597,1050,1077]" pageId="15" pageNumber="516">
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||
<heading bold="true" box="[151,597,1050,1077]" fontSize="11" level="1" pageId="15" pageNumber="516" reason="1">
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<emphasis bold="true" box="[151,597,1050,1077]" pageId="15" pageNumber="516">
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<taxonomicName authority="Neethling & Neethling, 2023" authorityName="Neethling & Neethling" authorityYear="2023" box="[151,496,1050,1077]" class="Arachnida" family="Gymnobisiidae" genus="Gymnobisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="15" pageNumber="516" phylum="Arthropoda" rank="species" species="hogsbackense" status="sp. nov.">
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<emphasis bold="true" box="[151,496,1050,1077]" italics="true" pageId="15" pageNumber="516">Gymnobisium hogsbackense</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[503,597,1051,1077]" pageId="15" pageNumber="516" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection box="[151,275,1087,1113]" pageId="15" pageNumber="516" type="description">
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||
<paragraph blockId="15.[151,597,1050,1113]" box="[151,275,1087,1113]" pageId="15" pageNumber="516">
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||
<figureCitation box="[151,275,1087,1113]" captionStart-0="FIGURE 11" captionStart-1="FIGURE 12" captionStart-2="FIGURE 13" captionStartId-0="16.[151,250,1877,1901]" captionStartId-1="17.[151,250,1721,1745]" captionStartId-2="18.[151,250,1939,1963]" captionTargetBox-0="[151,1436,181,1854]" captionTargetBox-1="[151,1434,1070,1698]" captionTargetBox-2="[170,1419,579,1897]" captionTargetId-0="figure-22@16.[151,1436,181,1854]" captionTargetId-1="figure-401@17.[151,1435,1065,1698]" captionTargetId-2="figure-192@18.[151,1436,561,1915]" captionTargetPageId-0="16" captionTargetPageId-1="17" captionTargetPageId-2="18" captionText-0="FIGURE 11. Digital microscope photographs of Gymnobisium hogsbackense sp. nov., Female (A, B) and Male (C, D). A, C. Dorsal view; B, D. Ventral view. Scale bar: 1.00 mm." captionText-1="FIGURE 12. Digital microscope photographs of Gymnobisium hogsbackense sp. nov. right chela: Female (A, B) and Male (C, D). A, C. Retrolateral view; B, D. Dorsal view. Scale bar: 1.00 mm." captionText-2="FIGURE 13. Gymnobisium hogsbackense sp. nov. A. Female right chela, retrolateral view; B. Galea; C. Female chelal fingers, anterior retrolateral view; D. Female right pedipalp, dorsal view; E. Same, male; F. Female left chelicera, dorsal view; G. Rallum. Scale bars: A, D, E: 0.50 mm; C, F: 0.20 mm; B, G: 0.10 mm." figureDoi-0="http://doi.org/10.5281/zenodo.7762492" figureDoi-1="http://doi.org/10.5281/zenodo.7762494" figureDoi-2="http://doi.org/10.5281/zenodo.7762496" httpUri-0="https://zenodo.org/record/7762492/files/figure.png" httpUri-1="https://zenodo.org/record/7762494/files/figure.png" httpUri-2="https://zenodo.org/record/7762496/files/figure.png" pageId="15" pageNumber="516">Figs 11–13</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="15" pageNumber="516" type="materials_examined">
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<paragraph blockId="15.[151,1437,1159,2013]" pageId="15" pageNumber="516">
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<materialsCitation collectingDate="2013-03-20" collectionCode="NMBAP" collectorName="Leaf & J. A. Neethling" country="South Africa" county="Hogsback" elevation="1174" latitude="-32.6" location="Afromontane forest" longLatPrecision="1209" longitude="26.916666" municipality="Hogsback State Forest" pageId="15" pageNumber="516" specimenCode="NMBAP 00341" specimenCount="1" specimenCount-female="1" stateProvince="Eastern Cape" typeStatus="holotype">
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<emphasis bold="true" box="[151,269,1159,1185]" pageId="15" pageNumber="516">
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<typeStatus box="[151,264,1159,1185]" pageId="15" pageNumber="516">Holotype</typeStatus>
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:
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</emphasis>
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<specimenCount box="[277,298,1159,1185]" pageId="15" pageNumber="516" type="female">♀</specimenCount>
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,
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<collectingCountry box="[312,517,1159,1185]" name="South Africa" pageId="15" pageNumber="516">SOUTH AFRICA</collectingCountry>
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,
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<collectingRegion box="[528,680,1159,1185]" country="South Africa" name="Eastern Cape" pageId="15" pageNumber="516">Eastern Cape</collectingRegion>
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,
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<collectingCounty box="[692,807,1159,1185]" pageId="15" pageNumber="516">Hogsback</collectingCounty>
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,
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<collectingMunicipality box="[818,1074,1159,1185]" pageId="15" pageNumber="516">Hogsback State Forest</collectingMunicipality>
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,
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<geoCoordinate box="[1085,1177,1160,1187]" degrees="32" direction="south" minutes="36" orientation="latitude" pageId="15" pageNumber="516" precision="925" value="-32.6">32°36′S</geoCoordinate>
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,
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<geoCoordinate box="[1188,1282,1160,1187]" degrees="26" direction="east" minutes="55" orientation="longitude" pageId="15" pageNumber="516" precision="925" value="26.916666">26°55′E</geoCoordinate>
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,
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<elevation box="[1293,1431,1159,1185]" metricMagnitude="3" metricUnit="m" metricValue="1.174" pageId="15" pageNumber="516" unit="m" value="1174.0">
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<quantity box="[1293,1370,1159,1185]" metricMagnitude="3" metricUnit="m" metricValue="1.174" pageId="15" pageNumber="516" unit="m" value="1174.0">1174m</quantity>
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a.s.l.
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</elevation>
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,
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<location LSID="urn:lsid:plazi:treatment:EB305F51B517F664FF1EFB9BFC24FB94:6646B89CB517F678FF1EFB2AFEA7FB40" box="[151,374,1195,1221]" country="South Africa" county="Hogsback" latitude="-32.6" longLatPrecision="1209" longitude="26.916666" municipality="Hogsback State Forest" name="Afromontane forest" pageId="15" pageNumber="516" stateProvince="Eastern Cape">Afromontane forest</location>
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,
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<collectorName box="[385,436,1195,1221]" pageId="15" pageNumber="516">Leaf</collectorName>
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litter sifting, leg.
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<collectorName box="[637,802,1195,1221]" pageId="15" pageNumber="516">J.A. Neethling</collectorName>
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,
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<date box="[814,940,1195,1221]" pageId="15" pageNumber="516" value="2013-03-20">
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<collectingDate box="[814,940,1195,1221]" pageId="15" pageNumber="516" value="2013-03-20">20.III.2013</collectingDate>
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</date>
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(
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<specimenCode box="[954,1137,1195,1221]" collectionCode="NMBAP" pageId="15" pageNumber="516">NMBAP 00341</specimenCode>
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).
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</materialsCitation>
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</paragraph>
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<paragraph blockId="15.[151,1437,1159,2013]" box="[199,819,1231,1259]" pageId="15" pageNumber="516">
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<materialsCitation box="[199,815,1231,1259]" collectingDate="2013-03-20" collectionCode="NMBAP" collectorName="Leaf & J. A. Neethling" country="South Africa" county="Hogsback" elevation="1174" latitude="-32.6" location="Afromontane forest" longLatPrecision="1209" longitude="26.916666" municipality="Hogsback State Forest" pageId="15" pageNumber="516" specimenCode="NMBAP 00340" specimenCount="1" stateProvince="Eastern Cape" typeStatus="paratype">
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<emphasis bold="true" box="[199,317,1231,1257]" pageId="15" pageNumber="516">
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<typeStatus box="[199,312,1231,1257]" pageId="15" pageNumber="516">Paratype</typeStatus>
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:
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</emphasis>
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♂, Same data as holotype (
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<specimenCode box="[625,807,1231,1257]" collectionCode="NMBAP" pageId="15" pageNumber="516">NMBAP 00340</specimenCode>
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)
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</materialsCitation>
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.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection pageId="15" pageNumber="516" type="etymology">
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<paragraph blockId="15.[151,1437,1159,2013]" pageId="15" pageNumber="516">
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<emphasis bold="true" box="[199,335,1267,1293]" pageId="15" pageNumber="516">Etymology.</emphasis>
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<emphasis box="[340,502,1267,1293]" italics="true" pageId="15" pageNumber="516">Hogsbackense</emphasis>
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refers to the
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<typeStatus box="[639,687,1267,1293]" pageId="15" pageNumber="516">type</typeStatus>
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locality of this species, Hogsback, and is in honour of the indigenous Afromontane forest in the area that has high levels of biodiversity.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="15" pageNumber="516" type="diagnosis">
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<paragraph blockId="15.[151,1437,1159,2013]" pageId="15" pageNumber="516">
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<emphasis bold="true" box="[199,322,1338,1364]" pageId="15" pageNumber="516">Diagnosis.</emphasis>
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Moderately sized species (chela length
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<specimenCount box="[773,794,1339,1365]" pageId="15" pageNumber="516" type="female">♀</specimenCount>
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<quantity box="[802,967,1339,1365]" metricMagnitude="-3" metricUnit="m" metricValue="1.475" metricValueMax="1.49" metricValueMin="1.46" pageId="15" pageNumber="516" unit="mm" value="1.475" valueMax="1.49" valueMin="1.46">1.46–1.49 mm</quantity>
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,
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<geoCoordinate box="[980,1057,1339,1367]" degrees="1.38" direction="west" orientation="longitude" pageId="15" pageNumber="516" precision="555" value="-1.38">
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♂
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<quantity box="[1008,1057,1339,1365]" metricMagnitude="-3" metricUnit="m" metricValue="1.395" metricValueMax="1.41" metricValueMin="1.38" pageId="15" pageNumber="516" unit="mm" value="1.395" valueMax="1.41" valueMin="1.38">1.38</quantity>
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</geoCoordinate>
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–1.41 mm), with chelae distinctly and significantly more bulbous in females than in males (chela ratio
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<specimenCount box="[922,943,1375,1401]" pageId="15" pageNumber="516" type="female">♀</specimenCount>
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2.98–3.04 versus
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<geoCoordinate box="[1149,1224,1375,1403]" degrees="3.71" direction="west" orientation="longitude" pageId="15" pageNumber="516" precision="555" value="-3.71">♂ 3.71</geoCoordinate>
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–3.94 times longer than broad); fixed finger multi-toothed structure with seven strongly sclerotized teeth; cheliceral hand with five setae; galea with six rami; rallum with eight blades. Differs from all congeners, except
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<taxonomicName authority="Neethling & Neethling, 2023" authorityName="Neethling & Neethling" authorityYear="2023" box="[1141,1394,1447,1473]" class="Arachnida" family="Gymnobisiidae" genus="Gymnobisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="15" pageNumber="516" phylum="Arthropoda" rank="species" species="capense" status="sp. nov.">
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<emphasis box="[1141,1394,1447,1473]" italics="true" pageId="15" pageNumber="516">Gymnobisium capense</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="15" pageNumber="516">
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<taxonomicNameLabel pageId="15" pageNumber="516" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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, by having seven teeth on the fixed finger multi-toothed structure; differs from
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<taxonomicName authority="Neethling & Neethling, 2023" authorityName="Neethling & Neethling" authorityYear="2023" box="[1091,1215,1483,1509]" class="Arachnida" family="Gymnobisiidae" genus="Gymnobisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="15" pageNumber="516" phylum="Arthropoda" rank="species" species="capense" status="sp. nov.">
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<emphasis box="[1091,1215,1483,1509]" italics="true" pageId="15" pageNumber="516">G. capense</emphasis>
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</taxonomicName>
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<emphasis bold="true" box="[1223,1312,1483,1509]" pageId="15" pageNumber="516">
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<taxonomicNameLabel box="[1223,1312,1483,1509]" pageId="15" pageNumber="516" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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by lacking an extra basal seta on the cheliceral hand.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="19" lastPageNumber="520" pageId="15" pageNumber="516" type="description">
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<paragraph blockId="15.[151,1437,1159,2013]" box="[199,337,1554,1580]" pageId="15" pageNumber="516">
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<emphasis bold="true" box="[199,337,1554,1580]" pageId="15" pageNumber="516">Description</emphasis>
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</paragraph>
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<paragraph blockId="15.[151,1437,1159,2013]" pageId="15" pageNumber="516">
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<emphasis box="[199,319,1591,1617]" italics="true" pageId="15" pageNumber="516">Carapace:</emphasis>
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Somewhat longer than broad, subquadrate, overall smooth and without furrow. Light or pale brown in both sexes, slightly darker in males. Two pairs of eyes located dorso-laterally at the front. Small seta located on the dorso-anterior margin of each of the posterior eyes. The carapace of both sexes with 20 setae, arranged 4: 4: 4: 2: 6.
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</paragraph>
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<paragraph blockId="15.[151,1437,1159,2013]" lastBlockId="17.[151,1436,151,285]" lastPageId="17" lastPageNumber="518" pageId="15" pageNumber="516">
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<emphasis box="[199,314,1735,1761]" italics="true" pageId="15" pageNumber="516">Abdomen:</emphasis>
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Elongate and mostly of uniform width. Tergites and sternites smooth, undivided and uniseriate. Abdominal tergites uniformly light brown in both sexes, slightly darker in males (
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<figureCitation box="[1084,1222,1771,1797]" captionStart="FIGURE 11" captionStartId="16.[151,250,1877,1901]" captionTargetBox="[151,1436,181,1854]" captionTargetId="figure-22@16.[151,1436,181,1854]" captionTargetPageId="16" captionText="FIGURE 11. Digital microscope photographs of Gymnobisium hogsbackense sp. nov., Female (A, B) and Male (C, D). A, C. Dorsal view; B, D. Ventral view. Scale bar: 1.00 mm." figureDoi="http://doi.org/10.5281/zenodo.7762492" httpUri="https://zenodo.org/record/7762492/files/figure.png" pageId="15" pageNumber="516">Fig. 11A, C</figureCitation>
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). In both sexes the posterior margin of tergite I–XI presents a thin, slightly darker band. Tergal setae located on the posterior edge of each tergite within the darkened band. In both sternites remain mostly uniform in colour across the abdomen (
|
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<figureCitation captionStart="FIGURE 11" captionStartId="16.[151,250,1877,1901]" captionTargetBox="[151,1436,181,1854]" captionTargetId="figure-22@16.[151,1436,181,1854]" captionTargetPageId="16" captionText="FIGURE 11. Digital microscope photographs of Gymnobisium hogsbackense sp. nov., Female (A, B) and Male (C, D). A, C. Dorsal view; B, D. Ventral view. Scale bar: 1.00 mm." figureDoi="http://doi.org/10.5281/zenodo.7762492" httpUri="https://zenodo.org/record/7762492/files/figure.png" pageId="15" pageNumber="516">Fig. 11B, D</figureCitation>
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||
). In females the sternites are light brown and in some specimens almost translucent. Male sternites also light brown in colour, lighter than male tergites and somewhat darker than the female sternites, and remain uniformly coloured across the abdomen. Some male specimens present slightly darker posterior sternites. Sternites I and II are fused and hardly distinguishable. Female operculum with 15 setae, six along the posterior margin of sternites II, and seven additional setae clustered loosely alongside, near the centre. Males with 21 setae at the operculum cluster, nine centrally along the poster margin of sternites II, 12 clustered centrally alongside. Male sternites III with 19 small setae situated, in two loose rows, centrally along the anterior margin at the genital opening. Pleural membrane of both sexes cream in colour, faintly papillate.
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</paragraph>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.7762492" ID-Zenodo-Dep="7762492" httpUri="https://zenodo.org/record/7762492/files/figure.png" pageId="16" pageNumber="517" startId="16.[151,250,1877,1901]" targetBox="[151,1436,181,1854]" targetPageId="16">
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||
<paragraph blockId="16.[151,1436,1877,1938]" pageId="16" pageNumber="517">
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||
<emphasis bold="true" box="[151,287,1877,1901]" pageId="16" pageNumber="517">FIGURE 11.</emphasis>
|
||
Digital microscope photographs of
|
||
<taxonomicName authority="Neethling & Neethling, 2023" authorityName="Neethling & Neethling" authorityYear="2023" box="[655,943,1878,1902]" class="Arachnida" family="Gymnobisiidae" genus="Gymnobisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="16" pageNumber="517" phylum="Arthropoda" rank="species" species="hogsbackense" status="sp. nov.">
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<emphasis box="[655,943,1878,1902]" italics="true" pageId="16" pageNumber="517">Gymnobisium hogsbackense</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[950,1031,1878,1902]" pageId="16" pageNumber="517">
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||
<taxonomicNameLabel box="[950,1031,1878,1902]" pageId="16" pageNumber="517" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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, Female (
|
||
<emphasis bold="true" box="[1134,1152,1877,1901]" pageId="16" pageNumber="517">A</emphasis>
|
||
,
|
||
<emphasis bold="true" box="[1166,1183,1878,1902]" pageId="16" pageNumber="517">B</emphasis>
|
||
) and Male (
|
||
<emphasis bold="true" box="[1309,1327,1877,1901]" pageId="16" pageNumber="517">C</emphasis>
|
||
,
|
||
<emphasis bold="true" box="[1340,1358,1878,1902]" pageId="16" pageNumber="517">D</emphasis>
|
||
).
|
||
<emphasis bold="true" box="[1380,1398,1877,1901]" pageId="16" pageNumber="517">A</emphasis>
|
||
,
|
||
<emphasis bold="true" box="[1412,1436,1877,1901]" pageId="16" pageNumber="517">C.</emphasis>
|
||
Dorsal view;
|
||
<emphasis bold="true" box="[288,305,1914,1938]" pageId="16" pageNumber="517">B</emphasis>
|
||
,
|
||
<emphasis bold="true" box="[317,342,1914,1938]" pageId="16" pageNumber="517">D.</emphasis>
|
||
Ventral view. Scale bar: 1.00 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="17.[151,1436,331,393]" pageId="17" pageNumber="518">Tergal chaetotaxy: 6(8): 9(12): 10(12): 12(12): 12(11): 12(12): 12(12): 12(11): 11(12): 8(9): 10(10): 2(2). Sternal chaetotaxy: 15(21):?(?): 20(20): 20(17): 18(16): 15(15): 15(14): 13(14): 13(13): 12(13): 10(10): 2(2).</paragraph>
|
||
<paragraph blockId="17.[151,1437,439,1041]" pageId="17" pageNumber="518">
|
||
<emphasis box="[199,300,439,465]" italics="true" pageId="17" pageNumber="518">Pedipalp</emphasis>
|
||
: All segments smooth and coloured orange-brown in both sexes. Trochanter cone-shaped and rounded anteriorly, curved slightly posteriorly. Base narrow, widening distally. Setae present on the anterior surface. Femur slender, 3.11–3.25 (
|
||
<specimenCount box="[374,392,511,537]" pageId="17" pageNumber="518" type="female">♀</specimenCount>
|
||
) to 3.36–3.54 (♂) times longer than wide, somewhat constricted basally, widening just distal of base. Deep disto-prolateral excavation present. Setae absent from base, though distributed evenly over rest of surface. Patella constricted and slightly angled at base, widening markedly in females, less in males, at around quarter segment length from base, 2.44–2.61 (
|
||
<specimenCount box="[660,678,619,645]" pageId="17" pageNumber="518" type="female">♀</specimenCount>
|
||
) to 3.08–3.17 (♂) times longer than wide. This gives female patellae a much thicker appearance, while male patellae appear more as elongate tapered cones. Several small lyriform fissures present on the dorsal surface, just distal of base. One to two larger lyriform fissures present latero-dorsally from these, where the segment starts to widen. Disto-prolateral excavation present and deeper on female patellae. Setae sparse at base, more numerous on wide part of patella.
|
||
</paragraph>
|
||
<paragraph blockId="17.[151,1437,439,1041]" pageId="17" pageNumber="518">
|
||
<emphasis box="[199,266,799,825]" italics="true" pageId="17" pageNumber="518">Chela</emphasis>
|
||
: In both sexes smooth and uniformly orange-brown, fingers may be slightly darker. Female hand significantly more bulbous, strongly convex on the prolateral edge, slightly less so on the retrolateral edge (
|
||
<figureCitation captionStart="FIGURE 13" captionStartId="18.[151,250,1939,1963]" captionTargetBox="[170,1419,579,1897]" captionTargetId="figure-192@18.[151,1436,561,1915]" captionTargetPageId="18" captionText="FIGURE 13. Gymnobisium hogsbackense sp. nov. A. Female right chela, retrolateral view; B. Galea; C. Female chelal fingers, anterior retrolateral view; D. Female right pedipalp, dorsal view; E. Same, male; F. Female left chelicera, dorsal view; G. Rallum. Scale bars: A, D, E: 0.50 mm; C, F: 0.20 mm; B, G: 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.7762496" httpUri="https://zenodo.org/record/7762496/files/figure.png" pageId="17" pageNumber="518">Fig. 13D</figureCitation>
|
||
). Male hand much thinner, markedly less convex prolaterally (
|
||
<figureCitation box="[925,1028,871,897]" captionStart="FIGURE 13" captionStartId="18.[151,250,1939,1963]" captionTargetBox="[170,1419,579,1897]" captionTargetId="figure-192@18.[151,1436,561,1915]" captionTargetPageId="18" captionText="FIGURE 13. Gymnobisium hogsbackense sp. nov. A. Female right chela, retrolateral view; B. Galea; C. Female chelal fingers, anterior retrolateral view; D. Female right pedipalp, dorsal view; E. Same, male; F. Female left chelicera, dorsal view; G. Rallum. Scale bars: A, D, E: 0.50 mm; C, F: 0.20 mm; B, G: 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.7762496" httpUri="https://zenodo.org/record/7762496/files/figure.png" pageId="17" pageNumber="518">Fig. 13E</figureCitation>
|
||
). Pedicel taller than wide and short in females (
|
||
<figureCitation box="[287,428,907,933]" captionStart="FIGURE 12" captionStartId="17.[151,250,1721,1745]" captionTargetBox="[151,1434,1070,1698]" captionTargetId="figure-401@17.[151,1435,1065,1698]" captionTargetPageId="17" captionText="FIGURE 12. Digital microscope photographs of Gymnobisium hogsbackense sp. nov. right chela: Female (A, B) and Male (C, D). A, C. Retrolateral view; B, D. Dorsal view. Scale bar: 1.00 mm." figureDoi="http://doi.org/10.5281/zenodo.7762494" httpUri="https://zenodo.org/record/7762494/files/figure.png" pageId="17" pageNumber="518">Fig. 12A, B</figureCitation>
|
||
), markedly longer and more rounded in males (
|
||
<figureCitation box="[981,1122,907,933]" captionStart="FIGURE 12" captionStartId="17.[151,250,1721,1745]" captionTargetBox="[151,1434,1070,1698]" captionTargetId="figure-401@17.[151,1435,1065,1698]" captionTargetPageId="17" captionText="FIGURE 12. Digital microscope photographs of Gymnobisium hogsbackense sp. nov. right chela: Female (A, B) and Male (C, D). A, C. Retrolateral view; B, D. Dorsal view. Scale bar: 1.00 mm." figureDoi="http://doi.org/10.5281/zenodo.7762494" httpUri="https://zenodo.org/record/7762494/files/figure.png" pageId="17" pageNumber="518">Fig. 12C, D</figureCitation>
|
||
), retrolateral condyle small but distinct and rounded. Female chelal fingers thicker, shorter than hand (without pedicel), while male fingers are narrow and longer than hand (without pedicel). Fingers curved slightly prolaterally in both sexes. Fixed and movable chelal fingers with eight and four trichobothria respectively (
|
||
<figureCitation box="[936,1041,1015,1041]" captionStart="FIGURE 13" captionStartId="18.[151,250,1939,1963]" captionTargetBox="[170,1419,579,1897]" captionTargetId="figure-192@18.[151,1436,561,1915]" captionTargetPageId="18" captionText="FIGURE 13. Gymnobisium hogsbackense sp. nov. A. Female right chela, retrolateral view; B. Galea; C. Female chelal fingers, anterior retrolateral view; D. Female right pedipalp, dorsal view; E. Same, male; F. Female left chelicera, dorsal view; G. Rallum. Scale bars: A, D, E: 0.50 mm; C, F: 0.20 mm; B, G: 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.7762496" httpUri="https://zenodo.org/record/7762496/files/figure.png" pageId="17" pageNumber="518">Fig. 13A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7762494" ID-Zenodo-Dep="7762494" httpUri="https://zenodo.org/record/7762494/files/figure.png" pageId="17" pageNumber="518" startId="17.[151,250,1721,1745]" targetBox="[151,1434,1070,1698]" targetPageId="17">
|
||
<paragraph blockId="17.[151,1436,1721,1782]" pageId="17" pageNumber="518">
|
||
<emphasis bold="true" box="[151,287,1721,1746]" pageId="17" pageNumber="518">FIGURE 12.</emphasis>
|
||
Digital microscope photographs of
|
||
<taxonomicName authority="Neethling & Neethling, 2023" authorityName="Neethling & Neethling" authorityYear="2023" box="[651,939,1722,1746]" class="Arachnida" family="Gymnobisiidae" genus="Gymnobisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="17" pageNumber="518" phylum="Arthropoda" rank="species" species="hogsbackense" status="sp. nov.">
|
||
<emphasis box="[651,939,1722,1746]" italics="true" pageId="17" pageNumber="518">Gymnobisium hogsbackense</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[944,1025,1722,1746]" pageId="17" pageNumber="518">
|
||
<taxonomicNameLabel box="[944,1025,1722,1746]" pageId="17" pageNumber="518" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
right chela: Female (
|
||
<emphasis bold="true" box="[1240,1258,1721,1745]" pageId="17" pageNumber="518">A</emphasis>
|
||
,
|
||
<emphasis bold="true" box="[1271,1288,1722,1746]" pageId="17" pageNumber="518">B</emphasis>
|
||
) and Male (
|
||
<emphasis bold="true" box="[1412,1430,1721,1745]" pageId="17" pageNumber="518">C</emphasis>
|
||
,
|
||
<emphasis bold="true" box="[151,169,1758,1782]" pageId="17" pageNumber="518">D</emphasis>
|
||
).
|
||
<emphasis bold="true" box="[191,209,1757,1781]" pageId="17" pageNumber="518">A</emphasis>
|
||
,
|
||
<emphasis bold="true" box="[222,246,1757,1781]" pageId="17" pageNumber="518">C.</emphasis>
|
||
Retrolateral view;
|
||
<emphasis bold="true" box="[441,458,1758,1782]" pageId="17" pageNumber="518">B</emphasis>
|
||
,
|
||
<emphasis bold="true" box="[471,495,1758,1782]" pageId="17" pageNumber="518">D.</emphasis>
|
||
Dorsal view. Scale bar: 1.00 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="17.[151,1437,1828,2034]" lastBlockId="18.[151,1437,151,537]" lastPageId="18" lastPageNumber="519" pageId="17" pageNumber="518">
|
||
Terminal multi-toothed structure of fixed finger extends beyond venom tooth of movable finger in both sexes, bent slightly downward, thicker in females, more slender in males. In both sexes, this structure contains seven strongly sclerotized teeth; front tooth with a minute protuberance on the retrolateral surface (
|
||
<figureCitation box="[1208,1312,1900,1926]" captionStart="FIGURE 13" captionStartId="18.[151,250,1939,1963]" captionTargetBox="[170,1419,579,1897]" captionTargetId="figure-192@18.[151,1436,561,1915]" captionTargetPageId="18" captionText="FIGURE 13. Gymnobisium hogsbackense sp. nov. A. Female right chela, retrolateral view; B. Galea; C. Female chelal fingers, anterior retrolateral view; D. Female right pedipalp, dorsal view; E. Same, male; F. Female left chelicera, dorsal view; G. Rallum. Scale bars: A, D, E: 0.50 mm; C, F: 0.20 mm; B, G: 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.7762496" httpUri="https://zenodo.org/record/7762496/files/figure.png" pageId="17" pageNumber="518">Fig. 13C</figureCitation>
|
||
). Proximal and just prolateral of the larger front tooth, five smaller, raised teeth located in a straight line, followed by a larger tooth situated retrolaterally on the structures surface. Both fingers with rounded, juxtadentate teeth situated on flexible lamellae. Lamella on fixed finger of female chela originating roughly one fifth finger length from base, originating roughly midway on movable finger. On male chela, lamellae originate midway on both fingers. Female fixed finger with 49–50 lightly sclerotized, juxtadentate teeth, eight to nine proximal to lamella base, together with seven strongly sclerotized teeth on distal structure. Male fixed finger with 50–51 lightly sclerotized, juxtadentate teeth, 20–22 proximal to lamella base, together with seven strongly sclerotized teeth on distal structure. Female movable finger with 44–46 lightly sclerotized, juxtadentate teeth, 16–17 proximal to lamella base. Male movable finger with 44–46 lightly sclerotized, juxtadentate teeth, 20–22 proximal to lamella base. Two thickened, spine-like setae present on distal end of movable finger in both sexes.
|
||
</paragraph>
|
||
<paragraph blockId="18.[151,1437,151,537]" pageId="18" pageNumber="519">
|
||
<emphasis box="[199,309,403,429]" italics="true" pageId="18" pageNumber="519">Chelicera</emphasis>
|
||
: Hand with five long and acuminate setae,
|
||
<emphasis box="[804,827,403,428]" italics="true" pageId="18" pageNumber="519">es</emphasis>
|
||
shorter (
|
||
<figureCitation box="[929,1030,403,429]" captionStart="FIGURE 13" captionStartId="18.[151,250,1939,1963]" captionTargetBox="[170,1419,579,1897]" captionTargetId="figure-192@18.[151,1436,561,1915]" captionTargetPageId="18" captionText="FIGURE 13. Gymnobisium hogsbackense sp. nov. A. Female right chela, retrolateral view; B. Galea; C. Female chelal fingers, anterior retrolateral view; D. Female right pedipalp, dorsal view; E. Same, male; F. Female left chelicera, dorsal view; G. Rallum. Scale bars: A, D, E: 0.50 mm; C, F: 0.20 mm; B, G: 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.7762496" httpUri="https://zenodo.org/record/7762496/files/figure.png" pageId="18" pageNumber="519">Fig. 13F</figureCitation>
|
||
). Female fixed finger with 13 teeth, male with 11. Female moveable finger with ten teeth, male with eight. Galea of both sexes divided into two major rami, each terminating in three smaller rami (
|
||
<figureCitation box="[673,777,475,501]" captionStart="FIGURE 13" captionStartId="18.[151,250,1939,1963]" captionTargetBox="[170,1419,579,1897]" captionTargetId="figure-192@18.[151,1436,561,1915]" captionTargetPageId="18" captionText="FIGURE 13. Gymnobisium hogsbackense sp. nov. A. Female right chela, retrolateral view; B. Galea; C. Female chelal fingers, anterior retrolateral view; D. Female right pedipalp, dorsal view; E. Same, male; F. Female left chelicera, dorsal view; G. Rallum. Scale bars: A, D, E: 0.50 mm; C, F: 0.20 mm; B, G: 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.7762496" httpUri="https://zenodo.org/record/7762496/files/figure.png" pageId="18" pageNumber="519">Fig. 13B</figureCitation>
|
||
). Rallum of both sexes with eight long, acuminate blades (
|
||
<figureCitation box="[159,263,511,537]" captionStart="FIGURE 13" captionStartId="18.[151,250,1939,1963]" captionTargetBox="[170,1419,579,1897]" captionTargetId="figure-192@18.[151,1436,561,1915]" captionTargetPageId="18" captionText="FIGURE 13. Gymnobisium hogsbackense sp. nov. A. Female right chela, retrolateral view; B. Galea; C. Female chelal fingers, anterior retrolateral view; D. Female right pedipalp, dorsal view; E. Same, male; F. Female left chelicera, dorsal view; G. Rallum. Scale bars: A, D, E: 0.50 mm; C, F: 0.20 mm; B, G: 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.7762496" httpUri="https://zenodo.org/record/7762496/files/figure.png" pageId="18" pageNumber="519">Fig. 13G</figureCitation>
|
||
). Lamina exterior absent in both sexes.
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7762496" ID-Zenodo-Dep="7762496" httpUri="https://zenodo.org/record/7762496/files/figure.png" pageId="18" pageNumber="519" startId="18.[151,250,1939,1963]" targetBox="[170,1419,579,1897]" targetPageId="18">
|
||
<paragraph blockId="18.[151,1437,1939,2035]" pageId="18" pageNumber="519">
|
||
<emphasis bold="true" box="[151,281,1939,1963]" pageId="18" pageNumber="519">FIGURE 13</emphasis>
|
||
.
|
||
<taxonomicName authority="Neethling & Neethling, 2023" authorityName="Neethling & Neethling" authorityYear="2023" box="[293,581,1939,1963]" class="Arachnida" family="Gymnobisiidae" genus="Gymnobisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="18" pageNumber="519" phylum="Arthropoda" rank="species" species="hogsbackense" status="sp. nov.">
|
||
<emphasis box="[293,581,1939,1963]" italics="true" pageId="18" pageNumber="519">Gymnobisium hogsbackense</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[586,697,1939,1963]" pageId="18" pageNumber="519">
|
||
<taxonomicNameLabel box="[586,666,1939,1963]" pageId="18" pageNumber="519" rank="species">sp. nov.</taxonomicNameLabel>
|
||
A.
|
||
</emphasis>
|
||
Female right chela, retrolateral view;
|
||
<emphasis bold="true" box="[1081,1105,1939,1963]" pageId="18" pageNumber="519">B.</emphasis>
|
||
Galea;
|
||
<emphasis bold="true" box="[1182,1207,1939,1963]" pageId="18" pageNumber="519">C.</emphasis>
|
||
Female chelal fingers, anterior retrolateral view;
|
||
<emphasis bold="true" box="[425,450,1975,1999]" pageId="18" pageNumber="519">D.</emphasis>
|
||
Female right pedipalp, dorsal view;
|
||
<emphasis bold="true" box="[840,864,1975,1999]" pageId="18" pageNumber="519">E.</emphasis>
|
||
Same, male;
|
||
<emphasis bold="true" box="[1010,1029,1975,1999]" pageId="18" pageNumber="519">F.</emphasis>
|
||
Female left chelicera, dorsal view;
|
||
<emphasis bold="true" box="[1410,1436,1975,1999]" pageId="18" pageNumber="519">G.</emphasis>
|
||
Rallum. Scale bars: A, D, E: 0.50 mm; C, F: 0.20 mm; B, G: 0.10 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="19.[151,1437,151,1041]" pageId="19" pageNumber="520">
|
||
<emphasis box="[199,373,151,177]" italics="true" pageId="19" pageNumber="520">Coxae and legs</emphasis>
|
||
: Pedipalpal coxae same colour, or slightly lighter than rest of pedipalps. Coxae I–IV tan. Legs I–IV light brown to tan in colour. All legs diplotarsate with subterminal tarsal setae dentate distally; claws simple; arolium shorter than claws.
|
||
</paragraph>
|
||
<paragraph blockId="19.[151,1437,151,1041]" pageId="19" pageNumber="520">
|
||
<emphasis bold="true" box="[199,455,258,285]" pageId="19" pageNumber="520">Measurements (mm):</emphasis>
|
||
Body length
|
||
<specimenCount box="[607,628,259,285]" pageId="19" pageNumber="520" type="female">♀</specimenCount>
|
||
2.19–
|
||
<geoCoordinate box="[699,775,259,287]" degrees="2.29" direction="west" orientation="longitude" pageId="19" pageNumber="520" precision="555" value="-2.29">2.29 ♂</geoCoordinate>
|
||
2.16–2.31; Carapace
|
||
<specimenCount box="[1020,1041,259,285]" pageId="19" pageNumber="520" type="female">♀</specimenCount>
|
||
0.74–0.79 x 0.67–0.71 (1.10–1.11)
|
||
<geoCoordinate box="[151,229,295,323]" degrees="0.64" direction="west" orientation="longitude" pageId="19" pageNumber="520" precision="555" value="-0.64">♂ 0.64</geoCoordinate>
|
||
–0.67 x 0.57–0.61 (1.10–1.12); Chelicera
|
||
<specimenCount box="[705,726,295,321]" pageId="19" pageNumber="520" type="female">♀</specimenCount>
|
||
0.50–0.52 x 0.24–0.26 (2.00–2.08)
|
||
<geoCoordinate box="[1135,1213,295,323]" degrees="0.40" direction="west" orientation="longitude" pageId="19" pageNumber="520" precision="555" value="-0.4">♂ 0.40</geoCoordinate>
|
||
–0.43 x 0.22 (1.82– 1.95), movable finger length
|
||
<specimenCount box="[482,503,331,357]" pageId="19" pageNumber="520" type="female">♀</specimenCount>
|
||
0.30–
|
||
<geoCoordinate box="[575,652,331,359]" degrees="0.31" direction="west" orientation="longitude" pageId="19" pageNumber="520" precision="555" value="-0.31">0.31 ♂</geoCoordinate>
|
||
0.25–0.26; Pedipalps: femur
|
||
<specimenCount box="[987,1008,331,357]" pageId="19" pageNumber="520" type="female">♀</specimenCount>
|
||
0.78–0.81 x 0.24–0.26 (3.11–3.25)
|
||
<geoCoordinate degrees="0.74" direction="west" orientation="longitude" pageId="19" pageNumber="520" precision="555" value="-0.74">♂ 0.74</geoCoordinate>
|
||
–0.78 x 0.22 (3.36–3.54), patella
|
||
<specimenCount box="[578,599,367,393]" pageId="19" pageNumber="520" type="female">♀</specimenCount>
|
||
0.73–0.78 x 0.28–0.32 (2.44–2.61)
|
||
<geoCoordinate box="[1008,1086,367,395]" degrees="0.76" direction="west" orientation="longitude" pageId="19" pageNumber="520" precision="555" value="-0.76">♂ 0.76</geoCoordinate>
|
||
–0.80 x 0.24–0.26 (3.08–3.17), chela
|
||
<specimenCount box="[217,238,403,429]" pageId="19" pageNumber="520" type="female">♀</specimenCount>
|
||
1.46–1.49 x 0.49 (2.98–3.04)
|
||
<geoCoordinate box="[575,651,403,431]" degrees="1.38" direction="west" orientation="longitude" pageId="19" pageNumber="520" precision="555" value="-1.38">♂ 1.38</geoCoordinate>
|
||
–1.41 x 0.35–0.38 (3.71–3.94), hand
|
||
<specimenCount box="[1064,1085,403,429]" pageId="19" pageNumber="520" type="female">♀</specimenCount>
|
||
0.87 x 0.49 (1.77)
|
||
<geoCoordinate box="[1297,1373,403,431]" degrees="0.74" direction="west" orientation="longitude" pageId="19" pageNumber="520" precision="555" value="-0.74">♂ 0.74</geoCoordinate>
|
||
–0.77 x 0.35–0.38 (2.03–2.11), movable finger length
|
||
<specimenCount box="[695,716,439,465]" pageId="19" pageNumber="520" type="female">♀</specimenCount>
|
||
0.65–
|
||
<geoCoordinate box="[787,866,438,467]" degrees="0.66" direction="west" orientation="longitude" pageId="19" pageNumber="520" precision="555" value="-0.66">0.66 ♂</geoCoordinate>
|
||
0.61–0.62; Leg I: femur
|
||
<specimenCount box="[1153,1174,439,465]" pageId="19" pageNumber="520" type="female">♀</specimenCount>
|
||
0.38–0.40 x 0.09–0.10 (4.00–4.22)
|
||
<geoCoordinate box="[288,365,475,503]" degrees="0.36" direction="west" orientation="longitude" pageId="19" pageNumber="520" precision="555" value="-0.36">♂ 0.36</geoCoordinate>
|
||
–0.37 x 0.09–0.10 (3.70–4.00), patella
|
||
<specimenCount box="[799,820,475,501]" pageId="19" pageNumber="520" type="female">♀</specimenCount>
|
||
0.27–0.30 x 0.09–0.10 (2.70–3.00)
|
||
<geoCoordinate box="[1221,1297,475,503]" degrees="0.26" direction="west" orientation="longitude" pageId="19" pageNumber="520" precision="555" value="-0.26">♂ 0.26</geoCoordinate>
|
||
–0.29 x 0.09 (2.89–3.22), tibia
|
||
<specimenCount box="[356,377,511,537]" pageId="19" pageNumber="520" type="female">♀</specimenCount>
|
||
0.35–0.37 x 0.08–0.09 (4.11–4.37)
|
||
<geoCoordinate box="[789,867,511,539]" degrees="0.34" direction="west" orientation="longitude" pageId="19" pageNumber="520" precision="555" value="-0.34">♂ 0.34</geoCoordinate>
|
||
–0.36 x 0.08 (4.25–4.50), metatarsus
|
||
<specimenCount box="[1295,1316,511,537]" pageId="19" pageNumber="520" type="female">♀</specimenCount>
|
||
0.19–0.20 x 0.06 (3.17–3.33)
|
||
<geoCoordinate box="[375,454,547,575]" degrees="0.18" direction="west" orientation="longitude" pageId="19" pageNumber="520" precision="555" value="-0.18">♂ 0.18</geoCoordinate>
|
||
–0.20 x 0.06 (3.00–3.33), tarsus
|
||
<specimenCount box="[834,855,547,573]" pageId="19" pageNumber="520" type="female">♀</specimenCount>
|
||
0.25–0.26 x 0.05 (5.00–5.20)
|
||
<geoCoordinate box="[1210,1289,547,575]" degrees="0.21" direction="west" orientation="longitude" pageId="19" pageNumber="520" precision="555" value="-0.21">♂ 0.21</geoCoordinate>
|
||
–0.24 x 0.05 (4.20–4.80); Leg IV femoropatella
|
||
<specimenCount box="[558,579,583,609]" pageId="19" pageNumber="520" type="female">♀</specimenCount>
|
||
0.67–0.73 x 0.16–0.17 (4.19–4.29)
|
||
<geoCoordinate box="[999,1078,583,611]" degrees="0.67" direction="west" orientation="longitude" pageId="19" pageNumber="520" precision="555" value="-0.67">♂ 0.67</geoCoordinate>
|
||
–0.70 x 0.15–0.17 (4.12–4.47), tibia
|
||
<specimenCount box="[210,231,619,645]" pageId="19" pageNumber="520" type="female">♀</specimenCount>
|
||
0.55–0.64 x 0.10–0.12 (5.33–5.50)
|
||
<geoCoordinate box="[644,723,618,647]" degrees="0.59" direction="west" orientation="longitude" pageId="19" pageNumber="520" precision="555" value="-0.59">♂ 0.59</geoCoordinate>
|
||
–0.63 x 0.10–0.11 (5.73–5.90), metatarsus
|
||
<specimenCount box="[1213,1234,619,645]" pageId="19" pageNumber="520" type="female">♀</specimenCount>
|
||
0.21–0.25 x 0.08 (2.62–3.12)
|
||
<geoCoordinate box="[289,366,655,683]" degrees="0.24" direction="west" orientation="longitude" pageId="19" pageNumber="520" precision="555" value="-0.24">♂ 0.24</geoCoordinate>
|
||
x 0.07 (3.43), tarsus
|
||
<specimenCount box="[604,625,655,681]" pageId="19" pageNumber="520" type="female">♀</specimenCount>
|
||
0.32–0.35 x 0.06 (5.33–5.83)
|
||
<geoCoordinate box="[965,1041,655,683]" degrees="0.31" direction="west" orientation="longitude" pageId="19" pageNumber="520" precision="555" value="-0.31">♂ 0.31</geoCoordinate>
|
||
–0.33 x 0.06 (5.17–5.50).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="19" pageNumber="520" type="discussion">
|
||
<paragraph blockId="19.[151,1437,151,1041]" pageId="19" pageNumber="520">
|
||
<emphasis bold="true" box="[199,315,691,717]" pageId="19" pageNumber="520">Remarks.</emphasis>
|
||
Both
|
||
<taxonomicName authority="Neethling & Neethling, 2023" authorityName="Neethling & Neethling" authorityYear="2023" box="[381,651,691,717]" class="Arachnida" family="Gymnobisiidae" genus="Gymnobisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="19" pageNumber="520" phylum="Arthropoda" rank="species" species="cuneatum" status="sp. nov.">
|
||
<emphasis box="[381,651,691,717]" italics="true" pageId="19" pageNumber="520">Gymnobisium cuneatum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[656,744,691,717]" pageId="19" pageNumber="520">
|
||
<taxonomicNameLabel box="[656,744,691,717]" pageId="19" pageNumber="520" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName authority="Neethling & Neethling, 2023" authorityName="Neethling & Neethling" authorityYear="2023" box="[796,984,691,717]" class="Arachnida" family="Gymnobisiidae" genus="Gymnobisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="19" pageNumber="520" phylum="Arthropoda" rank="species" species="hogsbackense" status="sp. nov.">
|
||
<emphasis box="[796,816,691,717]" italics="true" pageId="19" pageNumber="520">G</emphasis>
|
||
.
|
||
<emphasis box="[828,984,691,717]" italics="true" pageId="19" pageNumber="520">hogsbackense</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[989,1078,691,717]" pageId="19" pageNumber="520">
|
||
<taxonomicNameLabel box="[989,1078,691,717]" pageId="19" pageNumber="520" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
have, so far, only been collected at the
|
||
<typeStatus box="[219,267,727,753]" pageId="19" pageNumber="520">type</typeStatus>
|
||
locality, lending further support to the indigenous Afromontane forests of Hogsback being an endemism hotspot.
|
||
<bibRefCitation author="Harvey, M. S. & Huey, J. & Hillyer, M. & McIntyre, E. & Giribet, G." box="[246,470,763,789]" pageId="19" pageNumber="520" pagination="75 - 85" refId="ref28141" refString="Harvey, M. S., Huey, J., Hillyer, M., McIntyre, E. & Giribet, G. (2016) The first troglobitic species of Gymnobisiidae (Pseudoscorpiones, Neobisioidea), from Table Mountain (Western Cape Province, South Africa) and its phylogenetic position. Invertebrate Systematics, 30, 75 - 85. https: // doi. org / 10.1071 / IS 15044" type="journal article" year="2016">
|
||
Harvey
|
||
<emphasis box="[335,390,763,789]" italics="true" pageId="19" pageNumber="520">et al.</emphasis>
|
||
(2016)
|
||
</bibRefCitation>
|
||
provided sequence data for specimens from Hogsback that can likely be referred to this species. The reason is that
|
||
<taxonomicName authority="Neethling & Neethling, 2023" authorityName="Neethling & Neethling" authorityYear="2023" box="[454,598,799,825]" class="Arachnida" family="Gymnobisiidae" genus="Gymnobisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="19" pageNumber="520" phylum="Arthropoda" rank="species" species="cuneatum" status="sp. nov.">
|
||
<emphasis box="[454,598,799,825]" italics="true" pageId="19" pageNumber="520">G. cuneatum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[606,696,799,825]" pageId="19" pageNumber="520">
|
||
<taxonomicNameLabel box="[606,696,799,825]" pageId="19" pageNumber="520" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
was sampled only from the indigenous state forest, located along the mountain side, while
|
||
<taxonomicName authorityName="Neethling & Neethling" authorityYear="2023" box="[434,624,835,861]" class="Arachnida" family="Gymnobisiidae" genus="Gymnobisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="19" pageNumber="520" phylum="Arthropoda" rank="species" species="hogsbackense">
|
||
<emphasis box="[434,624,835,861]" italics="true" pageId="19" pageNumber="520">G. hogsbackense</emphasis>
|
||
</taxonomicName>
|
||
specimens were also sampled in the pine plantation, located close to the sampling site of the specimens referred to in
|
||
<bibRefCitation author="Harvey, M. S. & Huey, J. & Hillyer, M. & McIntyre, E. & Giribet, G." box="[655,885,871,897]" pageId="19" pageNumber="520" pagination="75 - 85" refId="ref28141" refString="Harvey, M. S., Huey, J., Hillyer, M., McIntyre, E. & Giribet, G. (2016) The first troglobitic species of Gymnobisiidae (Pseudoscorpiones, Neobisioidea), from Table Mountain (Western Cape Province, South Africa) and its phylogenetic position. Invertebrate Systematics, 30, 75 - 85. https: // doi. org / 10.1071 / IS 15044" type="journal article" year="2016">
|
||
Harvey
|
||
<emphasis box="[744,801,871,897]" italics="true" pageId="19" pageNumber="520">et al.</emphasis>
|
||
(2016)
|
||
</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="19.[151,1437,151,1041]" pageId="19" pageNumber="520">
|
||
<emphasis bold="true" box="[199,558,906,933]" pageId="19" pageNumber="520">Additional material examined.</emphasis>
|
||
<materialsCitation collectingDate="2013-03-20" collectionCode="NMBAP" collectorName="Leaf & J. A. Neethling" country="South Africa" county="Tritonymph" elevation="1080" latitude="-32.6" location="Hogsback State Forest" longLatPrecision="1209" longitude="26.933332" municipality="Hogsback" pageId="19" pageNumber="520" specimenCode="NMBAP 00327" specimenCount="2" specimenCount-female="2" stateProvince="Eastern Cape">
|
||
<emphasis bold="true" box="[563,772,906,932]" pageId="19" pageNumber="520">
|
||
<collectingCountry box="[563,772,906,932]" name="South Africa" pageId="19" pageNumber="520">SOUTH AFRICA</collectingCountry>
|
||
</emphasis>
|
||
:
|
||
<emphasis box="[784,935,907,933]" italics="true" pageId="19" pageNumber="520">
|
||
<collectingRegion box="[784,935,907,933]" country="South Africa" name="Eastern Cape" pageId="19" pageNumber="520">Eastern Cape</collectingRegion>
|
||
</emphasis>
|
||
:
|
||
<specimenCount box="[947,983,907,933]" pageId="19" pageNumber="520" type="female">2♀</specimenCount>
|
||
, 1♂, 1
|
||
<collectingCounty box="[1057,1193,907,933]" pageId="19" pageNumber="520">Tritonymph</collectingCounty>
|
||
,
|
||
<collectingMunicipality box="[1201,1316,907,933]" pageId="19" pageNumber="520">Hogsback</collectingMunicipality>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:EB305F51B517F664FF1EFB9BFC24FB94:6646B89CB50BF664FAA5FC0AFECEFC4C" country="South Africa" county="Tritonymph" latitude="-32.6" longLatPrecision="1209" longitude="26.933332" municipality="Hogsback" name="Hogsback State Forest" pageId="19" pageNumber="520" stateProvince="Eastern Cape">Hogsback State Forest</location>
|
||
,
|
||
<geoCoordinate box="[297,389,944,971]" degrees="32" direction="south" minutes="36" orientation="latitude" pageId="19" pageNumber="520" precision="925" value="-32.6">32°36′S</geoCoordinate>
|
||
,
|
||
<geoCoordinate box="[400,492,944,971]" degrees="26" direction="east" minutes="56" orientation="longitude" pageId="19" pageNumber="520" precision="925" value="26.933332">26°56′E</geoCoordinate>
|
||
,
|
||
<elevation box="[503,641,943,969]" metricMagnitude="3" metricUnit="m" metricValue="1.08" pageId="19" pageNumber="520" unit="m" value="1080.0">
|
||
<quantity box="[503,581,943,969]" metricMagnitude="3" metricUnit="m" metricValue="1.08" pageId="19" pageNumber="520" unit="m" value="1080.0">1080m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:EB305F51B517F664FF1EFB9BFC24FB94:6646B89CB50BF664FD05FC2EFCBAFC4C" box="[652,875,943,969]" country="South Africa" county="Tritonymph" latitude="-32.6" longLatPrecision="1209" longitude="26.933332" municipality="Hogsback" name="Afromontane forest" pageId="19" pageNumber="520" stateProvince="Eastern Cape">Afromontane forest</location>
|
||
,
|
||
<collectorName box="[885,936,943,969]" pageId="19" pageNumber="520">Leaf</collectorName>
|
||
litter sifting, leg.
|
||
<collectorName box="[1135,1299,943,969]" pageId="19" pageNumber="520">J.A. Neethling</collectorName>
|
||
,
|
||
<date box="[1310,1437,943,969]" pageId="19" pageNumber="520" value="2013-03-20">
|
||
<collectingDate box="[1310,1437,943,969]" pageId="19" pageNumber="520" value="2013-03-20">20.III.2013</collectingDate>
|
||
</date>
|
||
(
|
||
<specimenCode box="[159,343,979,1005]" collectionCode="NMBAP" pageId="19" pageNumber="520">NMBAP 00327</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation collectingDate="2013-03-21" collectionCode="NMBAP" collectorName="U. F. S. Entomology Students" country="South Africa" elevation="1201" latitude="-32.583332" location="Pine Plantation" longLatPrecision="1209" longitude="26.933332" municipality="Same" pageId="19" pageNumber="520" specimenCode="NMBAP 00232" specimenCount="1" specimenCount-female="1" stateProvince="Eastern Cape">
|
||
<specimenCount box="[364,399,979,1005]" pageId="19" pageNumber="520" type="female">1♀</specimenCount>
|
||
, 4♂,
|
||
<collectingMunicipality box="[466,528,979,1005]" pageId="19" pageNumber="520">Same</collectingMunicipality>
|
||
location,
|
||
<location LSID="urn:lsid:plazi:treatment:EB305F51B517F664FF1EFB9BFC24FB94:6646B89CB50BF664FD0DFC52FCE2FC68" box="[644,819,979,1005]" country="South Africa" latitude="-32.583332" longLatPrecision="1209" longitude="26.933332" municipality="Same" name="Pine Plantation" pageId="19" pageNumber="520" stateProvince="Eastern Cape">Pine Plantation</location>
|
||
,
|
||
<geoCoordinate box="[832,923,980,1007]" degrees="32" direction="south" minutes="35" orientation="latitude" pageId="19" pageNumber="520" precision="925" value="-32.583332">32°35′S</geoCoordinate>
|
||
,
|
||
<geoCoordinate box="[937,1029,980,1007]" degrees="26" direction="east" minutes="56" orientation="longitude" pageId="19" pageNumber="520" precision="925" value="26.933332">26°56′E</geoCoordinate>
|
||
,
|
||
<elevation box="[1043,1184,979,1005]" metricMagnitude="3" metricUnit="m" metricValue="1.201" pageId="19" pageNumber="520" unit="m" value="1201.0">
|
||
<quantity box="[1043,1121,979,1005]" metricMagnitude="3" metricUnit="m" metricValue="1.201" pageId="19" pageNumber="520" unit="m" value="1201.0">1201m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
,
|
||
<collectorName box="[1198,1248,979,1005]" pageId="19" pageNumber="520">Pine</collectorName>
|
||
plantation,
|
||
<collectorName box="[1385,1436,979,1005]" pageId="19" pageNumber="520">Leaf</collectorName>
|
||
litter sifting, leg.
|
||
<collectorName box="[345,666,1015,1041]" pageId="19" pageNumber="520">U.F.S. Entomology Students</collectorName>
|
||
,
|
||
<date box="[678,804,1015,1041]" pageId="19" pageNumber="520" value="2013-03-21">
|
||
<collectingDate box="[678,804,1015,1041]" pageId="19" pageNumber="520" value="2013-03-21">21.III.2013</collectingDate>
|
||
</date>
|
||
(
|
||
<specimenCode box="[818,1001,1015,1041]" collectionCode="NMBAP" pageId="19" pageNumber="520">NMBAP 00232</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |