364 lines
41 KiB
XML
364 lines
41 KiB
XML
<document id="CC60D8EC1765469F5BA8BC798F6FEA94" ID-DOI="10.11646/zootaxa.3947.4.4" ID-GBIF-Dataset="cf8bfab8-6721-4878-b6b4-28f4317ab703" ID-ISSN="1175-5326" ID-Zenodo-Dep="233730" ID-ZooBank="8FC05D43-724D-4A19-945F-6B5CE5DE2A0C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461042461294" checkinUser="plazi" docAuthor="Zaragoza-Caballero, Santiago & Zurita-García, Martín Leonel" docDate="2015" docId="A34F87832A570E6FFF4BFAAAFE35FB17" docLanguage="en" docName="zt03947p542.pdf" docOrigin="Zootaxa 3947 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Telegeusidae" docType="treatment" docVersion="7" lastPageNumber="539" masterDocId="5F76FFFB2A5C0E63FFDCFFF7FF86FFF0" masterDocTitle="A preliminary study on the phylogeny of the family Phengodidae (Insecta: Coleoptera)" masterLastPageNumber="542" masterPageNumber="527" pageNumber="538" updateTime="1698575927302" updateUser="plazi">
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<mods:mods id="1D9AECED9E20BE350955F95269C92FF5" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="4547E84972925803BA5A2F5E8F350B1D">
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<mods:title id="E183F7EDDCB9D896980CCEDB8522FCF4">A preliminary study on the phylogeny of the family Phengodidae (Insecta: Coleoptera)</mods:title>
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</mods:titleInfo>
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<mods:name id="6CC2DBDBDEBA82AD4EC6EC993F45871A" type="personal">
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<mods:namePart id="C21563C117F6F3FB7F729CB3510F57EA">Zaragoza-Caballero, Santiago</mods:namePart>
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<mods:name id="C27A16A8409FAA237AAC95B441FE2C6D" type="personal">
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<mods:roleTerm id="5CFB7662DF7B107AC171A5786CD53BDE">Author</mods:roleTerm>
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<mods:namePart id="580BF056F270DA8238F9E84B690E1197">Zurita-García, Martín Leonel</mods:namePart>
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<mods:typeOfResource id="020FF8906F776888DCC432A3D9169B2C">text</mods:typeOfResource>
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<mods:titleInfo id="148F1E9047DC31F2610E24D458B00847">
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<mods:title id="A19A48A3A9D513A0C779CAA2350244C0">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="2710D789ED9EC187589045603F0A3488">
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<mods:date id="7978BACC2B13F70BB7EE00F609752A2C">2015</mods:date>
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<mods:detail id="FDF6AFAF999918C7933D89707F0A6C78" type="volume">
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<mods:number id="6C93C3D83CE7900ADE728FEEC5A9C234">3947</mods:number>
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</mods:detail>
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<mods:detail id="78788D6400DDD7DFD8BD6445986FFE58" type="issue">
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<mods:number id="6378A28BA0CD379F8C3BADA3830415DF">4</mods:number>
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<mods:classification id="A4497E3EBB991B44BDADBFDAD218AA13">journal article</mods:classification>
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<mods:identifier id="DC56FD1E30AAD6BEC72EA8DF34CC82D7" type="DOI">10.11646/zootaxa.3947.4.4</mods:identifier>
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<treatment id="A34F87832A570E6FFF4BFAAAFE35FB17" ID-DOI="http://doi.org/10.5281/zenodo.6098551" ID-GBIF-Taxon="119610005" ID-Zenodo-Dep="6098551" LSID="urn:lsid:plazi:treatment:A34F87832A570E6FFF4BFAAAFE35FB17" httpUri="http://treatment.plazi.org/id/A34F87832A570E6FFF4BFAAAFE35FB17" lastPageId="12" lastPageNumber="539" pageId="11" pageNumber="538">
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<subSubSection id="63FC651E2A570E68FF4BFAAAFE59FA87" box="[151,479,1373,1399]" pageId="11" pageNumber="538" type="nomenclature">
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<paragraph id="2B5936952A570E68FF4BFAAAFE59FA87" blockId="11.[151,479,1373,1399]" box="[151,479,1373,1399]" pageId="11" pageNumber="538">
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<heading id="701181F92A570E68FF4BFAAAFE59FA87" bold="true" box="[151,479,1373,1399]" fontSize="11" level="1" pageId="11" pageNumber="538" reason="1">
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<emphasis id="1992EA872A570E68FF4BFAAAFE59FA87" bold="true" box="[151,479,1373,1399]" pageId="11" pageNumber="538">
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Taxonomy of
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<taxonomicName id="ECE64D162A570E68FE9DFAAAFE59FA87" ID-CoL="GYN" ID-ENA="195318" box="[321,479,1373,1399]" class="Insecta" family="Telegeusidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="538" phylum="Arthropoda" rank="family">Telegeusidae</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="63FC651E2A570E68FF4BFA53FC18F8D5" pageId="11" pageNumber="538" type="description">
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<paragraph id="2B5936952A570E68FF4BFA53FC18F8D5" blockId="11.[151,1436,1444,2009]" pageId="11" pageNumber="538">
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Our study corroborates a close relationship between the
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<taxonomicName id="ECE64D162A570E68FC80FA53FC68FA4D" box="[860,1006,1444,1469]" class="Insecta" family="Telegeusidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="538" phylum="Arthropoda" rank="family">Telegeusidae</taxonomicName>
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and Penicillophorinae.
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<bibRefCitation id="4F774B642A570E68FAC9FA53FA1AFA4C" author="Ivie" box="[1301,1436,1444,1469]" pageId="11" pageNumber="538" refString="Ivie, M. A. (2002) The transfer of Pseudokarumia Pic from Dascillidae (Karumiinae) to Telegeusidae, with a key to world genera of Telegusidae. The Coleopterists Bulletin, 56, 582 - 584. http: // dx. doi. org / 10.1649 / 0010 - 065 X (2002) 056 [0582: TTOPPF] 2.0. CO; 2" type="journal article" year="2002">Ivie (2002)</bibRefCitation>
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transferred
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<emphasis id="1992EA872A570E68FEC1FA30FE4BFA10" box="[285,461,1479,1504]" italics="true" pageId="11" pageNumber="538">Pseudokarumia</emphasis>
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Pic from
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<taxonomicName id="ECE64D162A570E68FD92FA30FD4AFA10" box="[590,716,1479,1504]" class="Insecta" family="Dascillidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="538" phylum="Arthropoda" rank="family">Dascillidae</taxonomicName>
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to
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<taxonomicName id="ECE64D162A570E68FD20FA30FC17FA10" box="[764,913,1479,1504]" class="Insecta" family="Telegeusidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="538" phylum="Arthropoda" rank="family">Telegeusidae</taxonomicName>
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, and provided a key to separate
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<taxonomicName id="ECE64D162A570E68FAFDFA30FA14FA10" box="[1313,1426,1479,1504]" class="Insecta" family="Telegeusidae" genus="Telegeusis" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="538" phylum="Arthropoda" rank="genus">
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<emphasis id="1992EA872A570E68FAFDFA30FA14FA10" box="[1313,1426,1479,1504]" italics="true" pageId="11" pageNumber="538">Telegeusis</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="ECE64D162A570E68FF4BFA1BFEDEF9F5" box="[151,344,1516,1541]" class="Insecta" family="Telegeusidae" genus="Pseudotelegeusis" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="538" phylum="Arthropoda" rank="genus">
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<emphasis id="1992EA872A570E68FF4BFA1BFEDEF9F5" box="[151,344,1516,1541]" italics="true" pageId="11" pageNumber="538">Pseudotelegeusis</emphasis>
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</taxonomicName>
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and
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<emphasis id="1992EA872A570E68FE4BFA1BFC8FF9F5" box="[407,777,1516,1541]" italics="true" pageId="11" pageNumber="538">Pseudokarumia. Pseudokarumia</emphasis>
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species share the following characters diagnostic of the family (
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<bibRefCitation id="4F774B642A570E68FF33F9E7FE62F9D8" author="Lawrence" box="[239,484,1551,1576]" pageId="11" pageNumber="538" refString="Lawrence, J. F., Hastings, A. M., Dallwitz, M. J., Paine, T. A. & Zurcher, E. J. (1999) Beetles of the world: a key and information system for families and subfamilies. CSIRO, Melbourne. CD-ROM version 1. 0 for MS-Windows." type="book chapter" year="1999">
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Lawrence
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<emphasis id="1992EA872A570E68FEBBF9E6FE1CF9D8" box="[359,410,1551,1576]" italics="true" pageId="11" pageNumber="538">et al</emphasis>
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. 1999
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</bibRefCitation>
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;
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<bibRefCitation id="4F774B642A570E68FE2CF9F8FDFDF9D8" author="Miller" box="[496,635,1551,1576]" pageId="11" pageNumber="538" refString="Miller, R. S. (2002) Family 68. Telegeusidae Leng 1920. In: Arnett, R. H., Thomas, M. C., Frank, J. H. & Skelley, P. E. (Eds.), American beetles. Vol. 2. CRC, Boca Raton, Florida, pp. 179 - 180." type="book chapter" year="2002">Miller 2002</bibRefCitation>
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): body narrow and large; labrum tri-lobed; last maxillary palpomere as long as antennae; pronotal disk with lateral impressions; elytra short; mesotibial spur serrate; abdomen with 8 ventrites.
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<bibRefCitation id="4F774B642A570E68FEDBF9A0FE00F980" author="Ivie" box="[263,390,1623,1648]" pageId="11" pageNumber="538" refString="Ivie, M. A. (2002) The transfer of Pseudokarumia Pic from Dascillidae (Karumiinae) to Telegeusidae, with a key to world genera of Telegusidae. The Coleopterists Bulletin, 56, 582 - 584. http: // dx. doi. org / 10.1649 / 0010 - 065 X (2002) 056 [0582: TTOPPF] 2.0. CO; 2" type="journal article" year="2002">Ivie (2002)</bibRefCitation>
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described
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<emphasis id="1992EA872A570E68FDDFF9A0FD35F980" box="[515,691,1623,1648]" italics="true" pageId="11" pageNumber="538">Pseudokarumia</emphasis>
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having eyes small; antennal insertions separated, mesotibial spur serrated; postocular space (1.5 to 3 times the length of eye), maxillary palps long; pronotal disk with lateral impressions; elytra short: abdomen with eight ventrites. He also segregated
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<taxonomicName id="ECE64D162A570E68FB9AF968FAF3F948" authority="Wittmer" authorityName="Wittmer" box="[1094,1397,1695,1720]" class="Insecta" family="Telegeusidae" genus="Pseudotelegeusis" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="538" phylum="Arthropoda" rank="genus">
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<emphasis id="1992EA872A570E68FB9AF968FA81F948" box="[1094,1287,1695,1720]" italics="true" pageId="11" pageNumber="538">Pseudotelegeusis</emphasis>
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Wittmer
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</taxonomicName>
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of
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<emphasis id="1992EA872A570E68FF4BF933FEC1F92D" box="[151,327,1732,1757]" italics="true" pageId="11" pageNumber="538">Pseudokarumia</emphasis>
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Pic by the shape of antennae (serrate or filiform), distance between eyes and posterior margin of the head (reduced or wide), number of labial palpomeres (1 or 1–2). Our results support the transfer of
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<emphasis id="1992EA872A570E68FF4BF8FBFEC1F8D5" box="[151,327,1804,1829]" italics="true" pageId="11" pageNumber="538">Pseudokarumia</emphasis>
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to
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<taxonomicName id="ECE64D162A570E68FEB6F8FBFE7AF8D5" box="[362,508,1804,1829]" class="Insecta" family="Telegeusidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="538" phylum="Arthropoda" rank="family">Telegeusidae</taxonomicName>
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and so corroborate the Ivie proposal.
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</paragraph>
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</subSubSection>
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<subSubSection id="63FC651E2A570E6FFF1BF8D8FE35FB17" lastPageId="12" lastPageNumber="539" pageId="11" pageNumber="538" type="reference_group">
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<paragraph id="2B5936952A570E6FFF1BF8D8FAA0FD3D" blockId="11.[151,1436,1444,2009]" lastBlockId="12.[151,1437,151,1256]" lastPageId="12" lastPageNumber="539" pageId="11" pageNumber="538">
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The
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<taxonomicName id="ECE64D162A570E68FEDCF8D8FE14F8B8" ID-CoL="GYN" box="[256,402,1839,1864]" class="Insecta" family="Telegeusidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="538" phylum="Arthropoda" rank="family">Telegeusidae</taxonomicName>
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as designated by
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<bibRefCitation id="4F774B642A570E68FDA8F8C7FC80F8B8" author="Leng" box="[628,774,1840,1864]" pageId="11" pageNumber="538" refString="Leng, C. W. (1920) Catalogue of Coleoptera from America North of Mexico. John D. Sherman, Jr. Mount Vernon, New York, 470 pp." type="book" year="1920">Leng (1920)</bibRefCitation>
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is comprised of
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<taxonomicName id="ECE64D162A570E68FC00F8D8FB11F8B7" ID-CoL="647BH" authority="Horn" authorityName="Horn" box="[988,1175,1839,1864]" class="Insecta" family="Telegeusidae" genus="Telegeusis" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="538" phylum="Arthropoda" rank="genus">
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<emphasis id="1992EA872A570E68FC00F8D8FBD6F8B8" box="[988,1104,1839,1864]" italics="true" pageId="11" pageNumber="538">Telegeusis</emphasis>
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Horn
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</taxonomicName>
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and
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<taxonomicName id="ECE64D162A570E68FB07F8D8FE0DF89D" ID-CoL="73RY" authority="Wittmer. Horn (1895)" authorityName="Wittmer. Horn" authorityYear="1895" class="Insecta" family="Telegeusidae" genus="Pseudotelegeusis" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="538" phylum="Arthropoda" rank="genus">
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<emphasis id="1992EA872A570E68FB07F8D8FA1AF8B8" box="[1243,1436,1839,1864]" italics="true" pageId="11" pageNumber="538">Pseudotelegeusis</emphasis>
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Wittmer.
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<bibRefCitation id="4F774B642A570E68FEDCF8A2FE0DF89D" author="Horn" box="[256,395,1876,1901]" pageId="11" pageNumber="538" refString="Horn, G. H. (1895) Coleoptera of Baja California (Supplement 1). Proceedings of the California Academy of Science, Series 2, 5, 242 - 243." type="journal article" year="1895">Horn (1895)</bibRefCitation>
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</taxonomicName>
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originally described and included
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<taxonomicName id="ECE64D162A570E68FCCCF8A3FC02F89D" ID-CoL="647BH" box="[784,900,1876,1901]" class="Insecta" family="Telegeusidae" genus="Telegeusis" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="538" phylum="Arthropoda" rank="genus">
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<emphasis id="1992EA872A570E68FCCCF8A3FC02F89D" box="[784,900,1876,1901]" italics="true" pageId="11" pageNumber="538">Telegeusis</emphasis>
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</taxonomicName>
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in the
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<taxonomicName id="ECE64D162A570E68FC0EF8A3FBDAF89D" ID-CoL="BRN" box="[978,1116,1876,1901]" class="Insecta" family="Lampyridae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="538" phylum="Arthropoda" rank="family">Lampyridae</taxonomicName>
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, and in the same publication this genus was associated with
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<taxonomicName id="ECE64D162A570E68FE23F880FDCBF860" box="[511,589,1911,1936]" class="Insecta" family="Elateridae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="538" phylum="Arthropoda" rank="tribe" tribe="Drilini">Drilini</taxonomicName>
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.
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<bibRefCitation id="4F774B642A570E68FD86F880FD7CF860" author="Barber" box="[602,762,1911,1936]" pageId="11" pageNumber="538" refString="Barber, H. S. (1913) Observations on the life history of Micromalthus debilis LeC. (Coleoptera). Proceedings of the Entomological Society of Washington, 15, 31 - 38." type="journal article" year="1913">Barber (1913)</bibRefCitation>
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related
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<taxonomicName id="ECE64D162A570E68FC85F880FC4DF860" ID-CoL="647BH" box="[857,971,1911,1936]" class="Insecta" family="Telegeusidae" genus="Telegeusis" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="538" phylum="Arthropoda" rank="genus">
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<emphasis id="1992EA872A570E68FC85F880FC4DF860" box="[857,971,1911,1936]" italics="true" pageId="11" pageNumber="538">Telegeusis</emphasis>
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</taxonomicName>
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with Lymexyloidea and as a monobasic genus. This proposal was rejected by
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<bibRefCitation id="4F774B642A570E68FDBFF86BFC81F844" author="Martin" box="[611,775,1948,1973]" pageId="11" pageNumber="538" refString="Martin, J. O. (1931) A new Telegeusis from Arizona (Coleoptera). The Pan-Pacific Entomologist, 8, 91 - 92." type="journal article" year="1931">Martin (1931)</bibRefCitation>
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who transferred it to
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<taxonomicName id="ECE64D162A570E68FBC5F86BFB22F845" ID-CoL="7P3" box="[1049,1188,1948,1973]" class="Insecta" family="Cantharidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="538" phylum="Arthropoda" rank="family">Cantharidae</taxonomicName>
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||
.
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<bibRefCitation id="4F774B642A570E68FB69F86BFAC3F845" author="King" box="[1205,1349,1948,1973]" pageId="11" pageNumber="538" refString="King, E. W. (1955) The phylogenetic position of Atractocerus Palis. The Coleopterists Bulletin, 9, 65 - 74." type="journal article" year="1955">King (1955)</bibRefCitation>
|
||
placed
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<taxonomicName id="ECE64D162A570E68FF4BF836FEFBF829" ID-CoL="359N" authority="Palisot" authorityName="Palisot" box="[151,381,1984,2009]" class="Insecta" family="Lymexylidae" genus="Atractocerus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="538" phylum="Arthropoda" rank="genus">
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<emphasis id="1992EA872A570E68FF4BF836FEAEF828" box="[151,296,1985,2008]" italics="true" pageId="11" pageNumber="538">Atractocerus</emphasis>
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Palisot
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</taxonomicName>
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in
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<taxonomicName id="ECE64D162A570E68FE7AF837FDBEF829" ID-CoL="C9T" box="[422,568,1984,2009]" class="Insecta" family="Lymexylidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="538" phylum="Arthropoda" rank="family">Lymexylidae</taxonomicName>
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and suggested an association with
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<taxonomicName id="ECE64D162A570E68FC0CF837FBC2F829" ID-CoL="647BH" box="[976,1092,1984,2009]" class="Insecta" family="Telegeusidae" genus="Telegeusis" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="538" phylum="Arthropoda" rank="genus">
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<emphasis id="1992EA872A570E68FC0CF837FBC2F829" box="[976,1092,1984,2009]" italics="true" pageId="11" pageNumber="538">Telegeusis</emphasis>
|
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</taxonomicName>
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.
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<bibRefCitation id="4F774B642A570E68FB89F837FA88F829" author="Crowson" box="[1109,1294,1984,2009]" pageId="11" pageNumber="538" refString="Crowson, R. A. (1955) The natural classification of the families of Coleoptera. Nathaniel Lloyd, London, 187 pp." type="book" year="1955">Crowson (1955)</bibRefCitation>
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related it to
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<taxonomicName id="ECE64D162A500E6FFF4BFF60FE94FF40" ID-CoL="6N6K" box="[151,274,151,176]" class="Insecta" family="Phengodidae" genus="Phengodes" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="539" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1992EA872A500E6FFF4BFF60FE94FF40" box="[151,274,151,176]" italics="true" pageId="12" pageNumber="539">Phengodes</emphasis>
|
||
</taxonomicName>
|
||
, based on the shared same configuration of male genitalia.
|
||
<bibRefCitation id="4F774B642A500E6FFC28FF6FFB34FF40" author="Crowson" box="[1012,1202,152,176]" pageId="12" pageNumber="539" refString="Crowson, R. A. (1972) A review of the classification of Cantharoidea (Coleoptera), with the definition of two new families, Cneoglossidae and Omethidae. Revista de la Universidad de Madrid, 21, 35 - 77." type="journal article" year="1972">Crowson (1972)</bibRefCitation>
|
||
defined the family
|
||
<taxonomicName id="ECE64D162A500E6FFF4BFF4BFEAFFF25" ID-CoL="GYN" box="[151,297,188,213]" class="Insecta" family="Telegeusidae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="539" phylum="Arthropoda" rank="family">Telegeusidae</taxonomicName>
|
||
based on the structure of the aedeagus and tentorium.
|
||
<bibRefCitation id="4F774B642A500E6FFC1AFF4AFB00FF24" author="Zaragoza" box="[966,1158,188,213]" pageId="12" pageNumber="539" refString="Zaragoza-Caballero, S. (1990) La familia Phengodidae en los " Tuxtlas ", Veracruz, Mexico. Anales Instituto de Biologia Nacional Autonoma de Mexico (Zoologia), 61 (2), 307 - 312." type="journal article" year="1990">Zaragoza (1990)</bibRefCitation>
|
||
noted that the reduced venation of the hind wing in the genera
|
||
<taxonomicName id="ECE64D162A500E6FFD54FF28FD7AFF08" ID-CoL="647BH" box="[648,764,223,248]" class="Insecta" family="Telegeusidae" genus="Telegeusis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="539" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1992EA872A500E6FFD54FF28FD7AFF08" box="[648,764,223,248]" italics="true" pageId="12" pageNumber="539">Telegeusis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="ECE64D162A500E6FFCCCFF28FB04FF08" ID-CoL="73RY" box="[784,1154,223,248]" class="Insecta" family="Telegeusidae" genus="Pseudotelegeusis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="539" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1992EA872A500E6FFCCCFF28FC57FF08" box="[784,977,223,248]" italics="true" pageId="12" pageNumber="539">Pseudotelegeusis</emphasis>
|
||
(Telegeusidae)
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="1992EA872A500E6FFB1AFF28FAA8FF08" box="[1222,1326,223,248]" italics="true" pageId="12" pageNumber="539">Titthonyx</emphasis>
|
||
LeConte (
|
||
<taxonomicName id="ECE64D162A500E6FFF42FEF3FEAFFEED" ID-CoL="7P3" box="[158,297,260,285]" class="Insecta" family="Cantharidae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="539" phylum="Arthropoda" rank="family">Cantharidae</taxonomicName>
|
||
) is a pattern shared with
|
||
<taxonomicName id="ECE64D162A500E6FFD8DFEF3FAA7FEED" authority="Zaragoza and Tarsakanthus Zaragoza" authorityName="Zaragoza and Tarsakanthus Zaragoza" box="[593,1313,260,285]" class="Insecta" family="Phengodidae" genus="Walterius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="539" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1992EA872A500E6FFD8DFEF3FD3DFEED" box="[593,699,260,285]" italics="true" pageId="12" pageNumber="539">Walterius</emphasis>
|
||
Zaragoza and
|
||
<emphasis id="1992EA872A500E6FFCB5FEF3FC78FEED" box="[873,1022,260,285]" italics="true" pageId="12" pageNumber="539">Tarsakanthus</emphasis>
|
||
Zaragoza (Phengodidae)
|
||
</taxonomicName>
|
||
. The same author noted that
|
||
<taxonomicName id="ECE64D162A500E6FFEBCFED0FE54FEB0" ID-CoL="647BH" box="[352,466,295,320]" class="Insecta" family="Telegeusidae" genus="Telegeusis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="539" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1992EA872A500E6FFEBCFED0FE54FEB0" box="[352,466,295,320]" italics="true" pageId="12" pageNumber="539">Telegeusis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="ECE64D162A500E6FFE3FFED0FD61FEB0" ID-CoL="3K5P" authority="LeConte" authorityName="LeConte" box="[483,743,295,320]" class="Insecta" family="Phengodidae" genus="Cenophengus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="539" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1992EA872A500E6FFE3FFED0FDFDFEB0" box="[483,635,295,320]" italics="true" pageId="12" pageNumber="539">Cenophengus</emphasis>
|
||
LeConte
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="ECE64D162A500E6FFD28FED0FBA5FEB0" ID-CoL="6335G" authority="Wittmer" authorityName="Wittmer" box="[756,1059,295,320]" class="Insecta" family="Phengodidae" genus="Distremocephalus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="539" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1992EA872A500E6FFD28FED0FC39FEB0" box="[756,959,295,320]" italics="true" pageId="12" pageNumber="539">Distremocephalus</emphasis>
|
||
Wittmer
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="ECE64D162A500E6FFB80FED0FB42FEB0" box="[1116,1220,295,320]" class="Insecta" family="Phengodidae" genus="Walterius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="539" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1992EA872A500E6FFB80FED0FB42FEB0" box="[1116,1220,295,320]" italics="true" pageId="12" pageNumber="539">Walterius</emphasis>
|
||
</taxonomicName>
|
||
have two tentorial pits whereas the genera
|
||
<taxonomicName id="ECE64D162A500E6FFE78FEBBFD99FE95" ID-CoL="6N6K" box="[420,543,332,357]" class="Insecta" family="Phengodidae" genus="Phengodes" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="539" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1992EA872A500E6FFE78FEBBFD99FE95" box="[420,543,332,357]" italics="true" pageId="12" pageNumber="539">Phengodes</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="ECE64D162A500E6FFDF1FEBBFC11FE95" ID-CoL="63X7X" authority="LeConte, Tarsakanthus" authorityName="LeConte, Tarsakanthus" box="[557,919,332,357]" class="Insecta" family="Phengodidae" genus="Zarhipis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="539" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1992EA872A500E6FFDF1FEBBFD0AFE95" box="[557,652,332,357]" italics="true" pageId="12" pageNumber="539">Zarhipis</emphasis>
|
||
LeConte,
|
||
<emphasis id="1992EA872A500E6FFCDDFEBBFC11FE95" box="[769,919,332,357]" italics="true" pageId="12" pageNumber="539">Tarsakanthus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="ECE64D162A500E6FFC79FEBBFBE0FE95" ID-CoL="73RY" box="[933,1126,332,357]" class="Insecta" family="Telegeusidae" genus="Pseudotelegeusis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="539" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1992EA872A500E6FFC79FEBBFBE0FE95" box="[933,1126,332,357]" italics="true" pageId="12" pageNumber="539">Pseudotelegeusis</emphasis>
|
||
</taxonomicName>
|
||
and other taxa have one pit only.
|
||
<bibRefCitation id="4F774B642A500E6FFF04FE87FD9BFE78" author="Lawrence" box="[216,541,367,392]" pageId="12" pageNumber="539" refString="Lawrence, J. F. & Newton, A. F., Jr. (1995) Families and subfamilies of Coleoptera (with selected genera, notes, references and data on family-group names). In: Pakaluk, J. & Slipinski, S. A. (Eds.), Biology, Phylogeny, and Classification of Coleoptera: Papers Celebrating the 80 th Birthday of Roy A. Crowson. Museum i Instytut Zoologii PAN, Warszawa, pp. 779 - 1006." type="book chapter" year="1995">Lawrence & Newton (1995)</bibRefCitation>
|
||
defined the superfamily Elateroidea (incl.
|
||
<taxonomicName id="ECE64D162A500E6FFBD2FE98FB19FE78" ID-CoL="6268M" box="[1038,1183,367,392]" class="Insecta" family="Phengodidae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="539" phylum="Arthropoda" rank="family">Phengodidae</taxonomicName>
|
||
and
|
||
<taxonomicName id="ECE64D162A500E6FFB07FE98FAE9FE78" ID-CoL="GYN" box="[1243,1391,367,392]" class="Insecta" family="Telegeusidae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="539" phylum="Arthropoda" rank="family">Telegeusidae</taxonomicName>
|
||
) on wing configuration and they recognized two patterns, an elateroid
|
||
<typeStatus id="F45D88372A500E6FFCA4FE62FC2EFE5D" box="[888,936,405,429]" pageId="12" pageNumber="539">type</typeStatus>
|
||
and another with reduced venation. This last pattern, presence of tentorial pits and antennal configuration are characters that suggest a close relationship between
|
||
<taxonomicName id="ECE64D162A500E6FFEDDFE2BFEF3FE05" ID-CoL="647BH" box="[257,373,476,501]" class="Insecta" family="Telegeusidae" genus="Telegeusis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="539" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1992EA872A500E6FFEDDFE2BFEF3FE05" box="[257,373,476,501]" italics="true" pageId="12" pageNumber="539">Telegeusis</emphasis>
|
||
</taxonomicName>
|
||
and the subfamily Penicillophorinae. Recent work based on molecular evidence suggests different hypotheses about the relationships of
|
||
<taxonomicName id="ECE64D162A500E6FFD62FE08FCD5FDE8" ID-CoL="GYN" box="[702,851,511,536]" class="Insecta" family="Telegeusidae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="539" phylum="Arthropoda" rank="family">Telegeusidae</taxonomicName>
|
||
, with Bocakova
|
||
<emphasis id="1992EA872A500E6FFBC7FDF6FBD6FDE8" box="[1051,1104,511,536]" italics="true" pageId="12" pageNumber="539">et al</emphasis>
|
||
. (2007),
|
||
<bibRefCitation id="4F774B642A500E6FFB1CFE08FF66FDCC" author="Kundrata" pageId="12" pageNumber="539" refString="Kundrata, R. & Bocak, L. (2011) The phylogeny and limits of Elateridae (Insecta, Coleoptera): is there a common tendency of click beetles to soft-bodiedness and neoteny? Zoologica Scripta, 40, 364 - 378." type="journal article" year="2011">Kundrata & Bocak (2011)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="4F774B642A500E6FFECBFDD3FD97FDCC" box="[279,529,548,573]" pageId="12" pageNumber="539" refString="Kundrata, R., Bocakova, M. & Bocak, L. (2013) The phylogenetic position of Artematopodidae (Coleoptera: Elateroidea), with description of the first two Eurypogon species from China. Contribution to Zoology, 82 (4), 199 - 208." type="journal article">
|
||
Kundrata
|
||
<emphasis id="1992EA872A500E6FFE5AFDD2FE3EFDCD" box="[390,440,548,573]" italics="true" pageId="12" pageNumber="539">et al</emphasis>
|
||
. (2013)
|
||
</bibRefCitation>
|
||
relating
|
||
<taxonomicName id="ECE64D162A500E6FFDA8FDD3FD6EFDCD" ID-CoL="647BH" box="[628,744,548,573]" class="Insecta" family="Telegeusidae" genus="Telegeusis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="539" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1992EA872A500E6FFDA8FDD3FD6EFDCD" box="[628,744,548,573]" italics="true" pageId="12" pageNumber="539">Telegeusis</emphasis>
|
||
</taxonomicName>
|
||
with
|
||
<taxonomicName id="ECE64D162A500E6FFCF4FDD3FC21FDCD" ID-CoL="DKY" box="[808,935,548,573]" class="Insecta" family="Omethidae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="539" phylum="Arthropoda" rank="family">Omethidae</taxonomicName>
|
||
. In the study of Hunt
|
||
<emphasis id="1992EA872A500E6FFB47FDD2FB4DFDCD" box="[1179,1227,548,573]" italics="true" pageId="12" pageNumber="539">et al</emphasis>
|
||
. (2007), we found different hypotheses, with either under the parsimony criterion
|
||
<taxonomicName id="ECE64D162A500E6FFC84FDB0FC4FFD90" ID-CoL="647BH" box="[856,969,583,608]" class="Insecta" family="Telegeusidae" genus="Telegeusis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="539" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1992EA872A500E6FFC84FDB0FC4FFD90" box="[856,969,583,608]" italics="true" pageId="12" pageNumber="539">Telegeusis</emphasis>
|
||
</taxonomicName>
|
||
is related with
|
||
<taxonomicName id="ECE64D162A500E6FFBABFDB0FA8DFD90" ID-CoL="6268M" box="[1143,1291,583,608]" class="Insecta" family="Phengodidae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="539" phylum="Arthropoda" rank="family">Phengodidae</taxonomicName>
|
||
, or under the Bayesian approach it is associated with
|
||
<taxonomicName id="ECE64D162A500E6FFD5CFD9BFD79FD75" ID-CoL="DKY" box="[640,767,620,645]" class="Insecta" family="Omethidae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="539" phylum="Arthropoda" rank="family">Omethidae</taxonomicName>
|
||
.
|
||
<bibRefCitation id="4F774B642A500E6FFCCCFD9BFB98FD74" box="[784,1054,620,645]" pageId="12" pageNumber="539" refString="Kundrata, R., Bocakova, M. & Bocak, L. (2014) The comprehensive phylogeny of the superfamily Elateroidea (Coleoptera: Elateriformia). Molecular Phylogenetics and Evolution, 76, 162 - 171." type="journal article">
|
||
Kundrata
|
||
<emphasis id="1992EA872A500E6FFC5AFD9AFC38FD75" box="[902,958,620,645]" italics="true" pageId="12" pageNumber="539">et al</emphasis>
|
||
. (2014)
|
||
</bibRefCitation>
|
||
also associated
|
||
<taxonomicName id="ECE64D162A500E6FFB34FD9BFADAFD75" ID-CoL="647BH" box="[1256,1372,620,645]" class="Insecta" family="Telegeusidae" genus="Telegeusis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="539" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1992EA872A500E6FFB34FD9BFADAFD75" box="[1256,1372,620,645]" italics="true" pageId="12" pageNumber="539">Telegeusis</emphasis>
|
||
</taxonomicName>
|
||
with
|
||
<taxonomicName id="ECE64D162A500E6FFF4BFD78FE94FD58" ID-CoL="DKY" box="[151,274,655,680]" class="Insecta" family="Omethidae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="539" phylum="Arthropoda" rank="family">Omethidae</taxonomicName>
|
||
and mentioned that the similar shape of labrum is a character that supports this relationship. The present study based on morphological evidence does not corroborate the conclusions of these molecular studies.
|
||
</paragraph>
|
||
<paragraph id="2B5936952A500E6FFF1BFD2FFB43FC38" blockId="12.[151,1437,151,1256]" pageId="12" pageNumber="539">
|
||
<bibRefCitation id="4F774B642A500E6FFF1BFD2FFECAFD00" author="Jeng" box="[199,332,728,752]" pageId="12" pageNumber="539" refString="Jeng, M. - L. (2008) Comprehensive phylogenetics, systematics, and evolution of neoteny of Lampyridae (Insecta: Coleoptera). Phd Thesis, Graduate Faculty of the University of Kansas, 388 pp." type="book" year="2008">Jeng (2008)</bibRefCitation>
|
||
studied the evolution of neoteny in cantharoid-elateroid lineage with a full data set of 220 species and 410 characters ((incl.
|
||
<taxonomicName id="ECE64D162A500E6FFE00FD0BFDDDFCE5" ID-CoL="DKY" box="[476,603,764,789]" class="Insecta" family="Omethidae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="539" phylum="Arthropoda" rank="family">Omethidae</taxonomicName>
|
||
:
|
||
<taxonomicName id="ECE64D162A500E6FFDB2FD0BFC7EFCE5" authority="Kiensewetter" authorityName="Kiensewetter" box="[622,1016,764,789]" class="Insecta" family="Omethidae" genus="Drilonius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="539" phylum="Arthropoda" rank="species" species="striatulus">
|
||
<emphasis id="1992EA872A500E6FFDB2FD0BFCD4FCE5" box="[622,850,764,789]" italics="true" pageId="12" pageNumber="539">Drilonius striatulus</emphasis>
|
||
Kiensewetter
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="ECE64D162A500E6FFBD7FD0BFAEBFCE4" authority="LeConte" authorityName="LeConte" box="[1035,1389,764,789]" class="Insecta" family="Omethidae" genus="Omethes" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="539" phylum="Arthropoda" rank="species" species="marginatus">
|
||
<emphasis id="1992EA872A500E6FFBD7FD0BFB7DFCE4" box="[1035,1275,764,789]" italics="true" pageId="12" pageNumber="539">Omethes marginatus</emphasis>
|
||
LeConte
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="ECE64D162A500E6FFA5DFD0AFEE7FCC8" authority="Lewis" authorityName="Lewis" class="Insecta" family="Omethidae" genus="Omethes" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="539" phylum="Arthropoda" rank="species" species="rugiceps">
|
||
<emphasis id="1992EA872A500E6FFA5DFD0AFF71FCC8" italics="true" pageId="12" pageNumber="539">
|
||
<collectingCountry id="53F176052A500E6FFA5DFD0AFA1EFCE4" box="[1409,1432,765,788]" name="Iceland" pageId="12" pageNumber="539">O</collectingCountry>
|
||
. rugiceps
|
||
</emphasis>
|
||
(Lewis)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="ECE64D162A500E6FFEABFCE8FD45FCC8" authority="Fender" authorityName="Fender" box="[375,707,799,824]" class="Insecta" family="Omethidae" genus="Troglomethes" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="539" phylum="Arthropoda" rank="species" species="leechi">
|
||
<emphasis id="1992EA872A500E6FFEABFCE8FDE4FCC8" box="[375,610,799,824]" italics="true" pageId="12" pageNumber="539">Troglomethes leechi</emphasis>
|
||
Fender
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="ECE64D162A500E6FFD04FCE8FC68FCC8" box="[728,1006,799,824]" class="Insecta" family="Omethidae" genus="Malthomethes" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="539" phylum="Arthropoda" rank="species" species="oregonus">
|
||
<emphasis id="1992EA872A500E6FFD04FCE8FC68FCC8" box="[728,1006,799,824]" italics="true" pageId="12" pageNumber="539">Malthomethes oregonus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="ECE64D162A500E6FFBDAFCE8FAE4FCC8" authority="LeConte" authorityName="LeConte" box="[1030,1378,799,824]" class="Insecta" family="Omethidae" genus="Matheteus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="539" phylum="Arthropoda" rank="species" species="theveneti">
|
||
<emphasis id="1992EA872A500E6FFBDAFCE8FB77FCC8" box="[1030,1265,799,824]" italics="true" pageId="12" pageNumber="539">Matheteus theveneti</emphasis>
|
||
LeConte
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="ECE64D162A500E6FFF4BFCB3FDC3FCAD" authority="Van Dyke" authorityName="Van Dyke" box="[151,581,836,861]" class="Insecta" family="Omethidae" genus="Ginglymocladus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="539" phylum="Arthropoda" rank="species" species="luteicollis">
|
||
<emphasis id="1992EA872A500E6FFF4BFCB3FE40FCAD" box="[151,454,836,861]" italics="true" pageId="12" pageNumber="539">Ginglymocladus luteicollis</emphasis>
|
||
Van Dyke
|
||
</taxonomicName>
|
||
)), and
|
||
<bibRefCitation id="4F774B642A500E6FFD44FCB2FC18FCAC" box="[664,926,836,861]" pageId="12" pageNumber="539" refString="Lawrence, J. F., Slipinski, A., Seago, A. E., Thayer, M. K., Newton, A. F. & Marvaldi, A. E. (2011) Phylogeny of the coleoptera based on morphological characters of adults and larvae. Annales Zoologici, 61 (1), 1 - 217." type="journal article">
|
||
Lawrence
|
||
<emphasis id="1992EA872A500E6FFCCCFCB2FCC4FCAD" box="[784,834,836,861]" italics="true" pageId="12" pageNumber="539">et al</emphasis>
|
||
. (2011)
|
||
</bibRefCitation>
|
||
focused in the phylogeny of the
|
||
<taxonomicName id="ECE64D162A500E6FFAFDFCB3FA1AFCAD" ID-CoL="C2L" box="[1313,1436,836,861]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="539" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
|
||
based on morphological characters of adults and larvae (incl.
|
||
<taxonomicName id="ECE64D162A500E6FFCACFC90FC69FC70" ID-CoL="DKY" box="[880,1007,871,896]" class="Insecta" family="Omethidae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="539" phylum="Arthropoda" rank="family">Omethidae</taxonomicName>
|
||
:
|
||
<taxonomicName id="ECE64D162A500E6FFC21FC90FAD5FC70" authority="LeConte" authorityName="LeConte" box="[1021,1363,871,896]" class="Insecta" family="Omethidae" genus="Matheteus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="539" phylum="Arthropoda" rank="species" species="theveneti">
|
||
<emphasis id="1992EA872A500E6FFC21FC90FB64FC70" box="[1021,1250,871,896]" italics="true" pageId="12" pageNumber="539">Matheteus theveneti</emphasis>
|
||
LeConte
|
||
</taxonomicName>
|
||
), both concluded that
|
||
<taxonomicName id="ECE64D162A500E6FFE9FFC7BFE32FC55" ID-CoL="647BH" box="[323,436,908,933]" class="Insecta" family="Telegeusidae" genus="Telegeusis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="539" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1992EA872A500E6FFE9FFC7BFE32FC55" box="[323,436,908,933]" italics="true" pageId="12" pageNumber="539">Telegeusis</emphasis>
|
||
</taxonomicName>
|
||
is nested into
|
||
<taxonomicName id="ECE64D162A500E6FFD85FC7BFD6CFC55" ID-CoL="6268M" box="[601,746,908,933]" class="Insecta" family="Phengodidae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="539" phylum="Arthropoda" rank="family">Phengodidae</taxonomicName>
|
||
(and
|
||
<taxonomicName id="ECE64D162A500E6FFCF7FC7BFC20FC55" ID-CoL="DKY" box="[811,934,908,933]" class="Insecta" family="Omethidae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="539" phylum="Arthropoda" rank="family">Omethidae</taxonomicName>
|
||
with
|
||
<taxonomicName id="ECE64D162A500E6FFC3AFC7BFBB9FC55" ID-CoL="624HT" box="[998,1087,908,933]" class="Insecta" family="Lycidae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="539" phylum="Arthropoda" rank="family">Lycidae</taxonomicName>
|
||
and
|
||
<taxonomicName id="ECE64D162A500E6FFBAAFC7BFB7BFC55" ID-CoL="7P3" box="[1142,1277,908,933]" class="Insecta" family="Cantharidae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="539" phylum="Arthropoda" rank="family">Cantharidae</taxonomicName>
|
||
respectively). All these works (molecular and morphological) lacked representatives of the Penicillophorinae.
|
||
</paragraph>
|
||
<paragraph id="2B5936952A500E6FFF1BFC22FE35FB17" blockId="12.[151,1437,151,1256]" pageId="12" pageNumber="539">
|
||
Here, we suggest a close relationship between
|
||
<taxonomicName id="ECE64D162A500E6FFD08FC23FCE3FC1D" ID-CoL="6268M" box="[724,869,980,1005]" class="Insecta" family="Phengodidae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="539" phylum="Arthropoda" rank="family">Phengodidae</taxonomicName>
|
||
and
|
||
<taxonomicName id="ECE64D162A500E6FFC40FC23FBA8FC1D" ID-CoL="GYN" box="[924,1070,980,1005]" class="Insecta" family="Telegeusidae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="539" phylum="Arthropoda" rank="family">Telegeusidae</taxonomicName>
|
||
(both strictly from New World), specifically with the genera of the subfamily Penicillophorinae, demonstrated as monophyletic. The fact that
|
||
<taxonomicName id="ECE64D162A500E6FFF4BFBEBFEAFFBC5" ID-CoL="GYN" box="[151,297,1052,1077]" class="Insecta" family="Telegeusidae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="539" phylum="Arthropoda" rank="family">Telegeusidae</taxonomicName>
|
||
+Penicillophorinae are nested into the clade that contains the terminals of external group could be an artifact of our sampling of outgroups.
|
||
<taxonomicName id="ECE64D162A500E6FFD4FFBC8FCA3FBA8" ID-CoL="GYN" box="[659,805,1087,1112]" class="Insecta" family="Telegeusidae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="539" phylum="Arthropoda" rank="family">Telegeusidae</taxonomicName>
|
||
was recovered as non-monophyletic due to that
|
||
<taxonomicName id="ECE64D162A500E6FFF4BFB93FEDEFB8D" ID-CoL="73RY" box="[151,344,1124,1149]" class="Insecta" family="Telegeusidae" genus="Pseudotelegeusis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="539" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1992EA872A500E6FFF4BFB93FEDEFB8D" box="[151,344,1124,1149]" italics="true" pageId="12" pageNumber="539">Pseudotelegeusis</emphasis>
|
||
</taxonomicName>
|
||
is sister to Penicillophorinae.
|
||
<taxonomicName id="ECE64D162A500E6FFD6CFB93FCF7FB8D" ID-CoL="73RY" box="[688,881,1124,1149]" class="Insecta" family="Telegeusidae" genus="Pseudotelegeusis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="539" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1992EA872A500E6FFD6CFB93FCF7FB8D" box="[688,881,1124,1149]" italics="true" pageId="12" pageNumber="539">Pseudotelegeusis</emphasis>
|
||
</taxonomicName>
|
||
is characterized as having the maxillary palps 4- segmented, the last palpomere as long as the three previous, labial palps 1-segmented and one tentorial pit. The main difference between
|
||
<taxonomicName id="ECE64D162A500E6FFE6AFB5BFDF1FB35" ID-CoL="73RY" box="[438,631,1196,1221]" class="Insecta" family="Telegeusidae" genus="Pseudotelegeusis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="539" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1992EA872A500E6FFE6AFB5BFDF1FB35" box="[438,631,1196,1221]" italics="true" pageId="12" pageNumber="539">Pseudotelegeusis</emphasis>
|
||
</taxonomicName>
|
||
and the penicillophorines is the length of the last maxillary palpomere relative to the basal ones.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |