231 lines
34 KiB
XML
231 lines
34 KiB
XML
<document id="F9E5371D19FBDB0D55AB5F2E6D6EF35B" ID-CLB-Dataset="55535" ID-DOI="10.5281/zenodo.5193661" ID-GBIF-Dataset="c2346a92-3813-4b3d-b139-f3b149b50f4a" ID-ISSN="0253-116X" ID-Zenodo-Dep="5193661" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628802610787" checkinUser="felipe" docAuthor="Jaeger, Bernd" docDate="2015" docId="5C580E35A81DF11AFF5EFEC4FE699D28" docLanguage="en" docName="LinzerbiolBeitr.47.2.1361-1396.pdf" docOrigin="Linzer biologische Beiträge 47 (2)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Anthracus javaensis Jaeger 2015, nov.sp." docType="treatment" docVersion="11" lastPageNumber="1374" masterDocId="A061764DA811F117FFECFFFCFFF39965" masterDocTitle="Revision of the maculate species of the Anthracus annamensis group from the East Palaearctic and Oriental Regions. Part 2. A redescription of Anthracus nesophilus (ANDREWES, 1936) and six new species from Nepal, India and SE Asia (Coleoptera, Carabidae, Harpalini, Stenolophina)" masterLastPageNumber="1396" masterPageNumber="1361" pageNumber="1373" updateTime="1699052547504" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="C7BC51290E9CF9F8E36625EF1FC2DB22">
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<mods:title id="856B615FC3C9B26044F696304CBEF5C5">Revision of the maculate species of the Anthracus annamensis group from the East Palaearctic and Oriental Regions. Part 2. A redescription of Anthracus nesophilus (ANDREWES, 1936) and six new species from Nepal, India and SE Asia (Coleoptera, Carabidae, Harpalini, Stenolophina)</mods:title>
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<mods:namePart id="FB00D9B16035B21EF96D1664B89CBF03">Jaeger, Bernd</mods:namePart>
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<mods:title id="EB48E475968D8F2230DF4800F1133AF7">Linzer biologische Beiträge</mods:title>
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<mods:part id="FE8C89D56C16BF2DAAB9867413C9C7A4">
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<mods:date id="11A40EFD637549152061B671EC446C40">2015</mods:date>
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<mods:number id="A999FC91992572DF0D901F1D46C86575">2015-12-30</mods:number>
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<mods:number id="DB6D5FCE76FABB7B4D06776FD4DCFBFF">47</mods:number>
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<mods:start id="68F535792C31AD007342173B4C065C81">1361</mods:start>
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<mods:identifier id="B3B49060EC25821328795B81C5CBFB57" type="DOI">10.5281/zenodo.5193661</mods:identifier>
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<treatment id="5C580E35A81DF11AFF5EFEC4FE699D28" ID-DOI="http://doi.org/10.5281/zenodo.6546858" ID-GBIF-Taxon="195305590" ID-Zenodo-Dep="6546858" LSID="urn:lsid:plazi:treatment:5C580E35A81DF11AFF5EFEC4FE699D28" httpUri="http://treatment.plazi.org/id/5C580E35A81DF11AFF5EFEC4FE699D28" lastPageId="13" lastPageNumber="1374" pageId="12" pageNumber="1373">
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<subSubSection id="9CEBECA8A81DF11BFF5EFEC4FE2B982B" box="[178,472,312,334]" pageId="12" pageNumber="1373" type="nomenclature">
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<paragraph id="D44EBF23A81DF11BFF5EFEC4FE2B982B" blockId="12.[178,719,312,334]" box="[178,472,312,334]" pageId="12" pageNumber="1373">
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<heading id="8F06084FA81DF11BFF5EFEC4FE2B982B" box="[178,472,312,334]" fontSize="8" level="2" pageId="12" pageNumber="1373" reason="2">
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<emphasis id="E6856331A81DF11BFF5EFEC4FE2B982B" bold="true" box="[178,472,312,334]" pageId="12" pageNumber="1373">
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<taxonomicName id="13F1C4A0A81DF11BFF5EFEC4FE769828" ID-CoL="67H79" authority="Jaeger, 2015" authorityName="Jaeger" authorityYear="2015" box="[178,389,312,334]" class="Insecta" family="Carabidae" genus="Anthracus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="1373" phylum="Arthropoda" rank="species" species="javaensis" status="sp. nov.">
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<emphasis id="E6856331A81DF11BFF5EFEC4FE769828" bold="true" box="[178,389,312,334]" italics="true" pageId="12" pageNumber="1373">Anthracus javaensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="FDB6DE4AA81DF11BFE60FEC5FE2B982B" box="[396,472,313,334]" pageId="12" pageNumber="1373" rank="species">nov.sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="9CEBECA8A81DF11BFE32FEC4FD3C982B" box="[478,719,312,334]" pageId="12" pageNumber="1373" type="description">
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<paragraph id="D44EBF23A81DF11BFE32FEC4FD3C982B" blockId="12.[178,719,312,334]" box="[478,719,312,334]" pageId="12" pageNumber="1373">
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<heading id="8F06084FA81DF11BFE32FEC4FD3C982B" box="[478,719,312,334]" fontSize="8" level="2" pageId="12" pageNumber="1373" reason="2">
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<emphasis id="E6856331A81DF11BFE32FEC4FD3C982B" bold="true" box="[478,719,312,334]" pageId="12" pageNumber="1373">
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(
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<figureCitation id="4CCAA3A6A81DF11BFE09FEC4FDA59828" box="[485,598,312,334]" captionStart="Figs 19-20" captionStartId="23.[178,218,1493,1515]" captionTargetBox="[178,1085,212,1573]" captionTargetId="figure-0@23.[178,1085,212,1573]" captionTargetPageId="23" captionText="Figs 19-20: Anthracus javaensis nov.sp. Habitus, head and pronotum. (19-20) HT." figureDoi="http://doi.org/10.5281/zenodo.5193673" httpUri="https://zenodo.org/record/5193673/files/figure.png" pageId="12" pageNumber="1373">Figs 19-20</figureCitation>
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,
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<figureCitation id="4CCAA3A6A81DF11BFD8EFEC4FD8E9828" box="[610,637,312,333]" captionStart="Figs 62-67" captionStartId="31.[178,218,1520,1542]" captionTargetBox="[150,1112,174,1495]" captionTargetId="figure-0@31.[150,1112,174,1617]" captionTargetPageId="31" captionText="Figs 62-67: Median lobe of aedoeagus, lateral aspect. Anthracus javaensis nov.sp. (62) HT and A. siamensis nov.sp. (63, 64, 67) PT, Laos, Pakkading, (65) HT, (66) Thailand, Ban Khoun." figureDoi="http://doi.org/10.5281/zenodo.6546849" httpUri="https://zenodo.org/record/6546849/files/figure.png" pageId="12" pageNumber="1373">62</figureCitation>
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,
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<figureCitation id="4CCAA3A6A81DF11BFD64FEC4FD519828" box="[648,674,312,333]" captionStart="Figs 68-73" captionStartId="32.[178,218,1420,1442]" captionTargetBox="[150,1112,159,1602]" captionTargetId="figure-0@32.[150,1112,159,1602]" captionTargetPageId="32" captionText="Figs 68-73: Median lobe of aedoeagus, dorsal aspect. Anthracus javaensis nov.sp. (68) HT and A. siamensis nov.sp. (69) PT, Thailand, Umphang, (70, 73) PT, Laos, Pakkading, (71) HT, (72) Thailand, Ban Khoun." figureDoi="http://doi.org/10.5281/zenodo.5194872" httpUri="https://zenodo.org/record/5194872/files/figure.png" pageId="12" pageNumber="1373">68</figureCitation>
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,
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<figureCitation id="4CCAA3A6A81DF11BFD42FEC4FD3A982B" box="[686,713,312,334]" captionStart="Fig 80" captionStartId="34.[178,210,909,931]" captionTargetBox="[266,997,212,896]" captionTargetId="figure-65@34.[266,997,212,896]" captionTargetPageId="34" captionText="Fig 80: Distribution: A. nesophilus (ANDREWES) (triangles), A. javaensis nov.sp. (white star), A. sumatraensis nov.sp. (square) A. philippinensis nov.sp. (circle), A. siamensis nov.sp. (black star)." figureDoi="http://doi.org/10.5281/zenodo.5193681" httpUri="https://zenodo.org/record/5193681/files/figure.png" pageId="12" pageNumber="1373">80</figureCitation>
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)
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="9CEBECA8A81DF11BFF5EFE9AFE0A98CD" pageId="12" pageNumber="1373" type="materials_examined">
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<paragraph id="D44EBF23A81DF11BFF5EFE9AFE0A98CD" blockId="12.[178,1086,357,425]" pageId="12" pageNumber="1373">
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Type material:
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<materialsCitation id="6499B57EA81DF11BFE6FFE99FE0A98CD" ID-GBIF-Occurrence="3766781347" collectingDate="1994-08-07" collectionCode="NHMW" collectorName="Schuh & B. Jaeger" country="Indonesia" elevation="1500" location="Bandung" municipality="Situ Lembang" pageId="12" pageNumber="1373" specimenCount="1" specimenCount-male="1" stateProvince="Java" typeStatus="holotype">
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<typeStatus id="0B4A0181A81DF11BFE6FFE99FE24981E" box="[387,471,357,379]" pageId="12" pageNumber="1373" type="holotype">Holotype</typeStatus>
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:
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<specimenCount id="C2F774AAA81DF11BFE08FE99FE04981F" box="[484,503,357,378]" pageId="12" pageNumber="1373" type="male">♂</specimenCount>
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(
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<collectionCode id="B2E027E6A81DF11BFDE8FE99FDA2981E" box="[516,593,357,379]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="12" pageNumber="1373" type="Museum">NHMW</collectionCode>
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) labelled "
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<collectingCountry id="ACE6FFB3A81DF11BFD58FE99FCC6981F" box="[692,821,357,378]" name="Indonesia" pageId="12" pageNumber="1373">INDONESIA</collectingCountry>
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: W
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<collectingRegion id="163571C1A81DF11BFCB6FE9AFC71981E" box="[858,898,358,379]" country="Indonesia" name="Jawa Timur" pageId="12" pageNumber="1373">Java</collectingRegion>
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/
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<collectingMunicipality id="342A2559A81DF11BFC7AFE99FBEA981E" box="[918,1049,357,379]" pageId="12" pageNumber="1373">Situ Lembang</collectingMunicipality>
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, ca.
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<quantity id="130912C6A81DF11BFF5EFE80FF0298F7" box="[178,241,380,402]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="12" pageNumber="1373" unit="m" value="1500.0">
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<elevation id="5FDC5810A81DF11BFF5EFE80FF0298F7" box="[178,241,380,402]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="12" pageNumber="1373" unit="m" value="1500.0">1500m</elevation>
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</quantity>
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/
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<locationDeviation id="C5FCA601A81DF11BFEE9FE80FEA498F4" bearing="0.0" box="[261,343,380,402]" distance="15000.0" pageId="12" pageNumber="1373">
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<quantity id="130912C6A81DF11BFEE9FE80FEB398F7" box="[261,320,380,402]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="12" pageNumber="1373" unit="km" value="15.0">15 km</quantity>
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N
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</locationDeviation>
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<location id="D12EE9F8A81DF11BFEB2FE80FE4398F7" LSID="urn:lsid:plazi:treatment:5C580E35A81DF11AFF5EFEC4FE699D28:D12EE9F8A81DF11BFEB2FE80FE4398F7" box="[350,432,380,402]" country="Indonesia" municipality="Situ Lembang" name="Bandung" pageId="12" pageNumber="1373" stateProvince="Java">Bandung</location>
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/ lg.
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<collectorName id="7904DAF5A81DF11BFE0FFE80FDEF98F7" box="[483,540,380,402]" pageId="12" pageNumber="1373">Schuh</collectorName>
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<date id="A04F99E3A81DF11BFDCEFE80FD8798F4" box="[546,628,380,401]" pageId="12" pageNumber="1373" value="1994-08-07">
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<collectingDate id="B00B600BA81DF11BFDCEFE80FD8798F4" box="[546,628,380,401]" pageId="12" pageNumber="1373" value="1994-08-07">7.8.1994</collectingDate>
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</date>
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" and "HOLOTYPE ♂ /
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<originalDetermination id="F7EF07DFA81DF11BFCB4FE80FF3798CD" pageId="12" pageNumber="1373">
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<taxonomicName id="13F1C4A0A81DF11BFCB4FE80FC4698F7" authorityName="Motschoulsky" authorityYear="1850" box="[856,949,380,402]" class="Insecta" family="Carabidae" genus="Anthracus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="1373" phylum="Arthropoda" rank="genus">Anthracus</taxonomicName>
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/ javaensis
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<taxonomicNameLabel id="FDB6DE4AA81DF11BFBCFFE80FF3798CD" pageId="12" pageNumber="1373" rank="species">sp. n.</taxonomicNameLabel>
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</originalDetermination>
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/ des.
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<collectorName id="7904DAF5A81DF11BFF13FE6FFEA098CD" box="[255,339,403,424]" pageId="12" pageNumber="1373">
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B.
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<bibRefCitation id="B060C2D2A81DF11BFEF5FE6FFEA098CD" author="JAEGER B" box="[281,339,403,424]" pageId="12" pageNumber="1373" refId="ref12895" refString="JAEGER B. (2015): Two new maculate species of the Anthracus annamensis group from Nepal and adjacent areas (Insecta: Coleoptera: Carabidae: Harpalini: Stenolophina). Pp. 339 - 345. - In: HARTMANN M. & J. WEIPERT (eds), Biodiversitat und Naturausstattung im Himalaya V. - Verein der Freunde und Forderer des Naturkundemuseums Erfurt e. V. Erfurt: 580 pp." type="book" year="2015">Jaeger</bibRefCitation>
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</collectorName>
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2015". [red label]
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="9CEBECA8A81DF11BFF5EFE4BFBFA98A8" box="[178,1033,439,461]" pageId="12" pageNumber="1373" type="etymology">
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<paragraph id="D44EBF23A81DF11BFF5EFE4BFBFA98A8" blockId="12.[178,1033,439,461]" box="[178,1033,439,461]" pageId="12" pageNumber="1373">Etymology: The species name refers to the known distribution of the species.</paragraph>
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</subSubSection>
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<subSubSection id="9CEBECA8A81DF11AFF5EFE20FDCA9B0D" lastPageId="13" lastPageNumber="1374" pageId="12" pageNumber="1373" type="description">
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<paragraph id="D44EBF23A81DF11BFF5EFE20FEA09B6A" blockId="12.[178,1085,476,527]" pageId="12" pageNumber="1373">
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Description: General appearance as figured (
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<figureCitation id="4CCAA3A6A81DF11BFD00FE20FCCE9897" box="[748,829,476,498]" captionStart="Figs 19-20" captionStartId="23.[178,218,1493,1515]" captionTargetBox="[178,1085,212,1573]" captionTargetId="figure-0@23.[178,1085,212,1573]" captionTargetPageId="23" captionText="Figs 19-20: Anthracus javaensis nov.sp. Habitus, head and pronotum. (19-20) HT." figureDoi="http://doi.org/10.5281/zenodo.5193673" httpUri="https://zenodo.org/record/5193673/files/figure.png" pageId="12" pageNumber="1373">Fig. 19</figureCitation>
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). Body length 4.0 mm; width
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<quantity id="130912C6A81DF11BFF1EFE05FEBC9B6A" box="[242,335,505,527]" metricMagnitude="-3" metricUnit="m" metricValue="1.55" pageId="12" pageNumber="1373" unit="mm" value="1.55">1.55 mm</quantity>
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.
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</paragraph>
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<paragraph id="D44EBF23A81DF11BFF5EFDE3FEB89A59" blockId="12.[178,1085,543,828]" pageId="12" pageNumber="1373">Shiny, pronotum weakly, elytra moderately iridescent. Head dark brown, with clypeus reddish brown, labrum and mandibles reddish yellow (inner margins and apices of the latter blackish). Pronotum darker than head, dark to blackish brown, with lateral furrows paler in basal half. Elytra dark reddish yellow, with each elytron having a very large almost blackish central macula, expanding laterally to interval 8, and leaving base, apex and first interval reddish yellow. Legs and palpi pale yellowish brown, antennae with first two antennomeres yellowish brown, remaining ones moderately infuscated. Ventral surface mainly brown to reddish brown, with proepisterna markedly darker brown, metepisterna and lateral parts of abdominal sternites moderately darker brown, epipleura reddish brown.</paragraph>
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<paragraph id="D44EBF23A81DF11BFF5EFCB7FE6D9D29" blockId="12.[178,1086,843,1100]" pageId="12" pageNumber="1373">
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Head (
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<figureCitation id="4CCAA3A6A81DF11BFF1AFCB7FE959A04" box="[246,358,843,865]" captionStart="Figs 19-20" captionStartId="23.[178,218,1493,1515]" captionTargetBox="[178,1085,212,1573]" captionTargetId="figure-0@23.[178,1085,212,1573]" captionTargetPageId="23" captionText="Figs 19-20: Anthracus javaensis nov.sp. Habitus, head and pronotum. (19-20) HT." figureDoi="http://doi.org/10.5281/zenodo.5193673" httpUri="https://zenodo.org/record/5193673/files/figure.png" pageId="12" pageNumber="1373">Figs 19-20</figureCitation>
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) including eyes 0.86 times as wide as pronotum, with eyes moderately prominent (head 1.61 times as wide as head between eyes). Labrum with apical margin linear but somewhat sloped down to the right side. Mandibles medium sized, left mandible rather short and somewhat obtuse at apex, but not thickened or truncate. Antennae moderately long, 2.46 times as long as pronotum and 0.86 times as long as elytra. Microsculpture on clypeus and labrum distinct and almost isodiametric, partly weakly transverse, on frons with very lightly impressed (partly obliterated) and on vertex with moderately impressed isodiametric meshes becoming weakly transverse in front of pronotal apical margin.
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</paragraph>
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<paragraph id="D44EBF23A81DF11BFF5EFBA7FC919CB5" blockId="12.[178,1086,1115,1488]" pageId="12" pageNumber="1373">
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Pronotum (
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<figureCitation id="4CCAA3A6A81DF11BFEC8FBA7FE679D14" box="[292,404,1115,1137]" captionStart="Figs 19-20" captionStartId="23.[178,218,1493,1515]" captionTargetBox="[178,1085,212,1573]" captionTargetId="figure-0@23.[178,1085,212,1573]" captionTargetPageId="23" captionText="Figs 19-20: Anthracus javaensis nov.sp. Habitus, head and pronotum. (19-20) HT." figureDoi="http://doi.org/10.5281/zenodo.5193673" httpUri="https://zenodo.org/record/5193673/files/figure.png" pageId="12" pageNumber="1373">Figs 19-20</figureCitation>
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) 1.30 times as wide as long, 1.27 times as wide as head, widest in second quarter, lateral seta inserted a little posterior to beginning of second quarter. Apical margin almost rectilinear, anterior angles narrowly rounded at tips, weakly projecting forward. Sides convex in anterior half and rectilinearly narrowed to posterior angles, which are obtuse and moderately rounded. Basal margin very weakly convex medially and moderately oblique to posterior angles,. Lateral furrows moderately wide in anterior half becoming only slightly wider in posterior quarter where they are fused with baso-lateral impressions. Baso-lateral impressions medium sized, distinctly delimited from the convex pronotal disc and the weakly depressed medial part of base, flattened to basal and lateral margins. Anterior transverse impression suggested. Microsculpture on disc with moderately impressed moderately transverse meshes, becoming weakly impressed and strongly transverse at sides, at baso-lateral impressions and lateral furrows with distinctly impressed isodiametric to weakly transverse meshes.
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</paragraph>
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<paragraph id="D44EBF23A81DF11AFF5EFA23FDCC984E" blockId="12.[178,1086,1503,1584]" lastBlockId="13.[178,1086,219,299]" lastPageId="13" lastPageNumber="1374" pageId="12" pageNumber="1373">
|
|
Elytra (
|
|
<figureCitation id="4CCAA3A6A81DF11BFF12FA23FEBB9C93" box="[254,328,1503,1526]" captionStart="Figs 19-20" captionStartId="23.[178,218,1493,1515]" captionTargetBox="[178,1085,212,1573]" captionTargetId="figure-0@23.[178,1085,212,1573]" captionTargetPageId="23" captionText="Figs 19-20: Anthracus javaensis nov.sp. Habitus, head and pronotum. (19-20) HT." figureDoi="http://doi.org/10.5281/zenodo.5193673" httpUri="https://zenodo.org/record/5193673/files/figure.png" pageId="12" pageNumber="1373">Fig. 19</figureCitation>
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|
) rather long, 1.59 times as long as wide, 2.84 times as long and 1.39 times as wide as pronotum. Sides weakly widened posteriad, widest just posterior to middle. Subapical sinuation weak. Elytral striae distinctly impressed and impunctate, intervals rather flat, becoming weakly narrowed and moderately convex at apex. Microsculpture on scutellum and around basal pore isodiametric, on elytral intervals only with traces of very lightly impressed transverse lines.
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</paragraph>
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|
<paragraph id="D44EBF23A81CF11AFF5EFEC7FCAC98EE" blockId="13.[178,1085,315,395]" pageId="13" pageNumber="1374">Metepisterna long, distinctly narrowed posteriad, at inner margin about 1.6 times longer than wide at basal margin. Prosternum medially with 5 and close to apical margin with a row of 8 medium long setae. Prosternal process with 1 distinct seta.</paragraph>
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|
<paragraph id="D44EBF23A81CF11AFF5EFE67FC16988E" blockId="13.[178,1085,410,491]" pageId="13" pageNumber="1374">Protarsomeres 1-4 of males markedly and mesotarsomeres 2-4 moderately dilated. Protarsomere 4 markedly and mesotarsomere 4 moderately bilobed. Pro- and mesotarsomeres 1-4 of males with biseriately arranged adhesive hairs on ventral surface.</paragraph>
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<paragraph id="D44EBF23A81CF11AFF5EFE06FDCA9B0D" blockId="13.[178,1085,506,616]" pageId="13" pageNumber="1374">
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Median lobe of aedeagus (
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|
<figureCitation id="4CCAA3A6A81CF11AFE2EFE06FDE19B75" box="[450,530,506,528]" captionStart="Figs 62-67" captionStartId="31.[178,218,1520,1542]" captionTargetBox="[150,1112,174,1495]" captionTargetId="figure-0@31.[150,1112,174,1617]" captionTargetPageId="31" captionText="Figs 62-67: Median lobe of aedoeagus, lateral aspect. Anthracus javaensis nov.sp. (62) HT and A. siamensis nov.sp. (63, 64, 67) PT, Laos, Pakkading, (65) HT, (66) Thailand, Ban Khoun." figureDoi="http://doi.org/10.5281/zenodo.6546849" httpUri="https://zenodo.org/record/6546849/files/figure.png" pageId="13" pageNumber="1374">Figs 62</figureCitation>
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,
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<figureCitation id="4CCAA3A6A81CF11AFDF1FE06FDC49B75" box="[541,567,506,528]" captionStart="Figs 68-73" captionStartId="32.[178,218,1420,1442]" captionTargetBox="[150,1112,159,1602]" captionTargetId="figure-0@32.[150,1112,159,1602]" captionTargetPageId="32" captionText="Figs 68-73: Median lobe of aedoeagus, dorsal aspect. Anthracus javaensis nov.sp. (68) HT and A. siamensis nov.sp. (69) PT, Thailand, Umphang, (70, 73) PT, Laos, Pakkading, (71) HT, (72) Thailand, Ban Khoun." figureDoi="http://doi.org/10.5281/zenodo.5194872" httpUri="https://zenodo.org/record/5194872/files/figure.png" pageId="13" pageNumber="1374">68</figureCitation>
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) large, with general appearance as figured. Apical lamella rather short and broad, not narrowed (dorsal aspect), the apex abruptly bent upward (lateral aspect). Internal structures composed of 3 larger subapical teeth, and three medium sized teeth in basal half.
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</paragraph>
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</subSubSection>
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<subSubSection id="9CEBECA8A81CF11AFF5EFD8BFDE19D6F" pageId="13" pageNumber="1374" type="discussion">
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<paragraph id="D44EBF23A81CF11AFF5EFD8BFDE19D6F" blockId="13.[178,1086,631,1034]" pageId="13" pageNumber="1374">
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Remarks and Comparisons:
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<taxonomicName id="13F1C4A0A81CF11AFD74FD84FCE19BE8" authority="Jaeger, 2015" authorityName="Jaeger" authorityYear="2015" box="[664,786,632,653]" class="Insecta" family="Carabidae" genus="Anthracus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="1374" phylum="Arthropoda" rank="species" species="javaensis" status="sp. nov.">
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<emphasis id="E6856331A81CF11AFD74FD84FCE19BE8" box="[664,786,632,653]" italics="true" pageId="13" pageNumber="1374">A. javaensis</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel id="FDB6DE4AA81CF11AFCF5FD84FC979BE8" box="[793,868,632,653]" pageId="13" pageNumber="1374" rank="species">nov.sp.</taxonomicNameLabel>
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,
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<taxonomicName id="13F1C4A0A81CF11AFC83FD84FC1E9BE8" authority="Jaeger, 2015" authorityName="Jaeger" authorityYear="2015" box="[879,1005,632,653]" class="Insecta" family="Carabidae" genus="Anthracus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="1374" phylum="Arthropoda" rank="species" species="siamensis" status="sp. nov.">
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|
<emphasis id="E6856331A81CF11AFC83FD84FC1E9BE8" box="[879,1005,632,653]" italics="true" pageId="13" pageNumber="1374">A. siamensis</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel id="FDB6DE4AA81CF11AFC18FD84FBCD9BE8" box="[1012,1086,632,653]" pageId="13" pageNumber="1374" rank="species">nov.sp.</taxonomicNameLabel>
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from
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|
<collectingCountry id="ACE6FFB3A81CF11AFF07FD69FEEF9BCF" box="[235,284,661,682]" name="Laos" pageId="13" pageNumber="1374">Laos</collectingCountry>
|
|
and
|
|
<collectingCountry id="ACE6FFB3A81CF11AFEBCFD69FE599BCE" box="[336,426,661,683]" name="Thailand" pageId="13" pageNumber="1374">Thailand</collectingCountry>
|
|
and
|
|
<taxonomicName id="13F1C4A0A81CF11AFE32FD69FD789BCF" authority="JAEGER" authorityName="JAEGER" authorityYear="2015" box="[478,651,661,682]" class="Insecta" family="Carabidae" genus="Anthracus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="1374" phylum="Arthropoda" rank="species" species="skalei">
|
|
<emphasis id="E6856331A81CF11AFE32FD69FDC49BCF" box="[478,567,661,682]" italics="true" pageId="13" pageNumber="1374">A. skalei</emphasis>
|
|
JAEGER
|
|
</taxonomicName>
|
|
from
|
|
<collectingCountry id="ACE6FFB3A81CF11AFD20FD69FCF89BCE" box="[716,779,661,683]" name="Nepal" pageId="13" pageNumber="1374">Nepal</collectingCountry>
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|
, NE
|
|
<collectingCountry id="ACE6FFB3A81CF11AFCACFD69FC849BCE" box="[832,887,661,683]" name="India" pageId="13" pageNumber="1374">India</collectingCountry>
|
|
,
|
|
<collectingCountry id="ACE6FFB3A81CF11AFC6EFD69FC389BCE" box="[898,971,661,683]" name="Bhutan" pageId="13" pageNumber="1374">Bhutan</collectingCountry>
|
|
and
|
|
<collectingCountry id="ACE6FFB3A81CF11AFC13FD69FF009BAD" name="Bangladesh" pageId="13" pageNumber="1374">Bangladesh</collectingCountry>
|
|
represent a complex of closely related, allopatrically distributed taxa which are characterized by a similar shape of the median lobe of the aedeagus with apex short in dorsal view and abruptly bent dorsad in lateral view. The species of this complex are partially very similar in general appearance, but can be distinguished by constant different internal structures of the aedeagus.
|
|
<taxonomicName id="13F1C4A0A81CF11AFDA5FCDBFD369A58" authority="Jaeger, 2015" authorityName="Jaeger" authorityYear="2015" box="[585,709,807,829]" class="Insecta" family="Carabidae" genus="Anthracus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="1374" phylum="Arthropoda" rank="species" species="javaensis" status="sp. nov.">
|
|
<emphasis id="E6856331A81CF11AFDA5FCDBFD369A58" box="[585,709,807,829]" italics="true" pageId="13" pageNumber="1374">A. javaensis</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel id="FDB6DE4AA81CF11AFD21FCD4FCE49A58" box="[717,791,808,829]" pageId="13" pageNumber="1374" rank="species">nov.sp.</taxonomicNameLabel>
|
|
is the largest species of this complex and differs from
|
|
<taxonomicName id="13F1C4A0A81CF11AFE24FCB9FDBA9A3F" authorityName="Jaeger" authorityYear="2015" box="[456,585,837,858]" class="Insecta" family="Carabidae" genus="Anthracus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="1374" phylum="Arthropoda" rank="species" species="siamensis">
|
|
<emphasis id="E6856331A81CF11AFE24FCB9FDBA9A3F" box="[456,585,837,858]" italics="true" pageId="13" pageNumber="1374">A. siamensis</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="13F1C4A0A81CF11AFD6DFCB9FD2F9A3C" authorityName="JAEGER" authorityYear="2015" box="[641,732,836,858]" class="Insecta" family="Carabidae" genus="Anthracus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="1374" phylum="Arthropoda" rank="species" species="skalei">
|
|
<emphasis id="E6856331A81CF11AFD6DFCB9FD2F9A3C" box="[641,732,836,858]" italics="true" pageId="13" pageNumber="1374">A. skalei</emphasis>
|
|
</taxonomicName>
|
|
by its large size, usually also by some body proportions (see HW/PW, EW/PW, EL/PL) and the different internal structures of the aedeagus (
|
|
<figureCitation id="4CCAA3A6A81CF11AFE48FC83FDEC9AF0" box="[420,543,895,917]" captionStart-0="Figs 62-67" captionStart-1="Figs 68-73" captionStart-2="Figs 74-79" captionStartId-0="31.[178,218,1520,1542]" captionStartId-1="32.[178,218,1420,1442]" captionStartId-2="33.[178,218,1494,1516]" captionTargetBox-0="[150,1112,174,1495]" captionTargetBox-1="[150,1112,159,1602]" captionTargetBox-2="[150,1107,181,1473]" captionTargetId-0="figure-0@31.[150,1112,174,1617]" captionTargetId-1="figure-0@32.[150,1112,159,1602]" captionTargetId-2="figure-0@33.[150,1112,181,1624]" captionTargetPageId-0="31" captionTargetPageId-1="32" captionTargetPageId-2="33" captionText-0="Figs 62-67: Median lobe of aedoeagus, lateral aspect. Anthracus javaensis nov.sp. (62) HT and A. siamensis nov.sp. (63, 64, 67) PT, Laos, Pakkading, (65) HT, (66) Thailand, Ban Khoun." captionText-1="Figs 68-73: Median lobe of aedoeagus, dorsal aspect. Anthracus javaensis nov.sp. (68) HT and A. siamensis nov.sp. (69) PT, Thailand, Umphang, (70, 73) PT, Laos, Pakkading, (71) HT, (72) Thailand, Ban Khoun." captionText-2="Figs 74-79: Median lobe of aedoeagus, lateral and dorsal aspect. Anthracus skalei JAEGER (74) PT, Bangladesh, Nasirabad, (75) PT, India, Kurseong, A. spec. 1 near A. biplagiatus (BOHEMAN) (76, 77) Bangladesh, Nasirabad, A. spec. 2 near A. annamensis (BATES) (78,79) Myanmar, Pegu." figureDoi-0="http://doi.org/10.5281/zenodo.6546849" figureDoi-1="http://doi.org/10.5281/zenodo.5194872" figureDoi-2="http://doi.org/10.5281/zenodo.6546851" httpUri-0="https://zenodo.org/record/6546849/files/figure.png" httpUri-1="https://zenodo.org/record/5194872/files/figure.png" httpUri-2="https://zenodo.org/record/6546851/files/figure.png" pageId="13" pageNumber="1374">Figs. 62-75</figureCitation>
|
|
). From
|
|
<taxonomicName id="13F1C4A0A81CF11AFD99FC83FCF29AF1" baseAuthorityName="ANDREWES" baseAuthorityYear="1936" box="[629,769,895,916]" class="Insecta" family="Carabidae" genus="Anthracus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="1374" phylum="Arthropoda" rank="species" species="nesophilus">
|
|
<emphasis id="E6856331A81CF11AFD99FC83FCF29AF1" box="[629,769,895,916]" italics="true" pageId="13" pageNumber="1374">A. nesophilus</emphasis>
|
|
</taxonomicName>
|
|
which occurs also in western Java it can be separated by characters given under this species. From
|
|
<taxonomicName id="13F1C4A0A81CF11AFC68FC60FBD49AD7" authorityName="Jaeger" authorityYear="2015" box="[900,1063,924,946]" class="Insecta" family="Carabidae" genus="Anthracus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="1374" phylum="Arthropoda" rank="species" species="sumatraensis">
|
|
<emphasis id="E6856331A81CF11AFC68FC60FBD49AD7" box="[900,1063,924,946]" italics="true" pageId="13" pageNumber="1374">A. sumatraensis</emphasis>
|
|
</taxonomicName>
|
|
it differs by the dark head and pronotum, larger size, some body proportions (HW/PW, EL/EW), and the different external shape and internal structures of the median lobe of the aedeagus (
|
|
<figureCitation id="4CCAA3A6A81CF11AFEAEFC08FE649D6F" box="[322,407,1012,1034]" captionStart="Figs 62-67" captionStartId="31.[178,218,1520,1542]" captionTargetBox="[150,1112,174,1495]" captionTargetId="figure-0@31.[150,1112,174,1617]" captionTargetPageId="31" captionText="Figs 62-67: Median lobe of aedoeagus, lateral aspect. Anthracus javaensis nov.sp. (62) HT and A. siamensis nov.sp. (63, 64, 67) PT, Laos, Pakkading, (65) HT, (66) Thailand, Ban Khoun." figureDoi="http://doi.org/10.5281/zenodo.6546849" httpUri="https://zenodo.org/record/6546849/files/figure.png" pageId="13" pageNumber="1374">Figs. 62</figureCitation>
|
|
,
|
|
<figureCitation id="4CCAA3A6A81CF11AFE4DFC08FE4F9D6F" box="[417,444,1012,1034]" captionStart="Figs 68-73" captionStartId="32.[178,218,1420,1442]" captionTargetBox="[150,1112,159,1602]" captionTargetId="figure-0@32.[150,1112,159,1602]" captionTargetPageId="32" captionText="Figs 68-73: Median lobe of aedoeagus, dorsal aspect. Anthracus javaensis nov.sp. (68) HT and A. siamensis nov.sp. (69) PT, Thailand, Umphang, (70, 73) PT, Laos, Pakkading, (71) HT, (72) Thailand, Ban Khoun." figureDoi="http://doi.org/10.5281/zenodo.5194872" httpUri="https://zenodo.org/record/5194872/files/figure.png" pageId="13" pageNumber="1374">68</figureCitation>
|
|
,
|
|
<figureCitation id="4CCAA3A6A81CF11AFE24FC08FDF59D6F" box="[456,518,1012,1034]" captionStart="Figs 50-55" captionStartId="29.[178,218,1492,1514]" captionTargetBox="[150,1110,191,1479]" captionTargetId="figure-0@29.[150,1112,189,1632]" captionTargetPageId="29" captionText="Figs 50-55: Anthracus sumatraensis nov.sp. Median lobe of aedoeagus, lateral and dorsal aspect. (50, 52, 53, 55) PT, (51, 54) HT." figureDoi="http://doi.org/10.5281/zenodo.6546843" httpUri="https://zenodo.org/record/6546843/files/figure.png" pageId="13" pageNumber="1374">50-55</figureCitation>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="9CEBECA8A81CF11AFF5EFBE5FE699D28" pageId="13" pageNumber="1374" type="distribution">
|
|
<paragraph id="D44EBF23A81CF11AFF5EFBE5FE699D28" blockId="13.[178,1085,1049,1101]" pageId="13" pageNumber="1374">
|
|
Distribution
|
|
<taxonomicName id="13F1C4A0A81CF11AFE70FBE6FDEA9D4A" authority="Jaeger, 2015" authorityName="Jaeger" authorityYear="2015" box="[412,537,1050,1071]" class="Insecta" family="Carabidae" genus="Anthracus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="1374" phylum="Arthropoda" rank="species" species="javaensis" status="sp. nov.">
|
|
<emphasis id="E6856331A81CF11AFE70FBE6FDEA9D4A" box="[412,537,1050,1071]" italics="true" pageId="13" pageNumber="1374">A. javaensis</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel id="FDB6DE4AA81CF11AFDCEFBE6FDA09D4A" box="[546,595,1050,1071]" pageId="13" pageNumber="1374" rank="species">sp.n.</taxonomicNameLabel>
|
|
is so far known only from the
|
|
<typeStatus id="0B4A0181A81CF11AFC4CFBE6FC3F9D4A" box="[928,972,1050,1071]" pageId="13" pageNumber="1374">type</typeStatus>
|
|
locality in western
|
|
<collectingRegion id="163571C1A81CF11AFEEBFBCBFEC79D29" box="[263,308,1079,1100]" country="Indonesia" name="Jawa Timur" pageId="13" pageNumber="1374">Java</collectingRegion>
|
|
(
|
|
<figureCitation id="4CCAA3A6A81CF11AFEAEFBCAFE7D9D28" box="[322,398,1078,1101]" captionStart="Fig 80" captionStartId="34.[178,210,909,931]" captionTargetBox="[266,997,212,896]" captionTargetId="figure-65@34.[266,997,212,896]" captionTargetPageId="34" captionText="Fig 80: Distribution: A. nesophilus (ANDREWES) (triangles), A. javaensis nov.sp. (white star), A. sumatraensis nov.sp. (square) A. philippinensis nov.sp. (circle), A. siamensis nov.sp. (black star)." figureDoi="http://doi.org/10.5281/zenodo.5193681" httpUri="https://zenodo.org/record/5193681/files/figure.png" pageId="13" pageNumber="1374">Fig. 80</figureCitation>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |