387 lines
45 KiB
XML
387 lines
45 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1090.76292" ID-GBIF-Dataset="8bdbd00b-4927-4adb-9609-0cc35e1a7920" ID-PMC="PMC9005471" ID-Pensoft-Pub="1313-2970-1090-161" ID-Pensoft-UUID="1E6F228182BA5078A6144181DB93F0DE" ID-PubMed="35586844" ID-ZooBank="3B13C71EE11B49D2891C050DBD514872" ModsDocID="1313-2970-1090-161" checkinTime="1648647504971" checkinUser="pensoft" docAuthor="Nethupul, Hasitha, Stoehr, Sabine & Zhang, Haibin" docDate="2022" docId="07D71215F3FD5F089C88ACF1CFA74132" docLanguage="en" docName="ZooKeys 1090: 161-216" docOrigin="ZooKeys 1090" docPubDate="2022-03-30" docSource="http://dx.doi.org/10.3897/zookeys.1090.76292" docTitle="Asteroschema lissum H. L. Clark 1939" docType="treatment" docVersion="5" id="1E6F228182BA5078A6144181DB93F0DE" lastPageNumber="161" masterDocId="1E6F228182BA5078A6144181DB93F0DE" masterDocTitle="Order Euryalida (Echinodermata, Ophiuroidea), new species and new records from the South China Sea and the Northwest Pacific seamounts" masterLastPageNumber="216" masterPageNumber="161" pageNumber="161" updateTime="1668151560559" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Order Euryalida (Echinodermata, Ophiuroidea), new species and new records from the South China Sea and the Northwest Pacific seamounts</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Nethupul, Hasitha</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-0590-2719</mods:nameIdentifier>
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<mods:affiliation>Institute of Deep-sea Science and Engineering, Chinese Academy of Sciences, CAS, 57200 Sanya, China & University of Chinese Academy of Sciences, Beijing 100039, China</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Stoehr, Sabine</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-2586-7239</mods:nameIdentifier>
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<mods:affiliation>Swedish Museum of Natural History, Dept of Zoology, Box 50007, 10405 Stockholm, Sweden</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Zhang, Haibin</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-5429-9851</mods:nameIdentifier>
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<mods:affiliation>Institute of Deep-sea Science and Engineering, Chinese Academy of Sciences, CAS, 57200 Sanya, China</mods:affiliation>
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<mods:nameIdentifier type="email">hzhang@idsse.ac.cn</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-03-30</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1090</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>161</mods:start>
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<mods:end>216</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1090.76292</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1090.76292</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1090-161</mods:identifier>
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<mods:identifier type="ZooBank">3B13C71EE11B49D2891C050DBD514872</mods:identifier>
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<mods:identifier type="Pensoft-UUID">1E6F228182BA5078A6144181DB93F0DE</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="194310898" LSID="urn:lsid:plazi:treatment:07D71215F3FD5F089C88ACF1CFA74132" httpUri="http://treatment.plazi.org/id/07D71215F3FD5F089C88ACF1CFA74132" lastPageNumber="161" pageId="0" pageNumber="161">
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<subSubSection pageId="0" pageNumber="161" type="nomenclature">
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<paragraph pageId="0" pageNumber="161">
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<taxonomicName LSID="07D71215-F3FD-5F08-9C88-ACF1CFA74132" authority="H. L. Clark, 1939" authorityName="H. L. Clark" authorityYear="1939" class="Ophiuroidea" family="Euryalidae" genus="Asteroschema" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Asteroschema lissum" order="Euryalida" pageId="0" pageNumber="161" phylum="Echinodermata" rank="species" species="lissum">Asteroschema cf. lissum H. L. Clark, 1939</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="161" type="description">
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<paragraph pageId="0" pageNumber="161">
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<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Asteroschema cf. lissum H. L. Clark, 1939 (IDSSE-EEB-SW 0079) A dorsal view B ventral view C dorsal disc D skin on the disc E lateral disc F ventral disc G dorsal arm (middle) H lateral arm (middle) I ventral arm (proximal) J outer arm spine (proximal) K arm spines (middle) L arm spines (distal). Abbreviations: as adoral shield, ass adoral shield spine, ars arm spine, go granular ossicle, gs genital slit, iars inner arm spine, j jaw, oars outer arm spine, ots oral tentacle scale, rs radial shield, t teeth, tp tentacle pore. Scale bars: 2 mm (A, B, G); 1 mm (C, E, F, H, I); 500 µm (J, K); 200 µm (D, L)." figureDoi="10.3897/zookeys.1090.76292.figure16" httpUri="https://binary.pensoft.net/fig/665892" pageId="0" pageNumber="161">Figures 16</figureCitation>
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<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Asteroschema cf. lissum H. L. Clark, 1939 (IDSSE-EEB-SW 0079) A, B lateral arm plate (external, internal) C outer arm spine (middle) D arm spine (distal) E skin from dorsal arm base (granular ossicles) F-J vertebrae F proximal view G distal view H lateral view I dorsal view J ventral view. Abbreviations: d dorsal, dist distal, lac lateral ambulacral canals, mo muscle opening, no nerve opening, pb podial basin, prox proximal, st secondary teeth, v ventral. Scale bars: 800 µm (F-J); 500 µm (A, C); 300 µm (B, D); 200 µm (E)." figureDoi="10.3897/zookeys.1090.76292.figure17" httpUri="https://binary.pensoft.net/fig/665893" pageId="0" pageNumber="161">, 17</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="161" type="reference_group">
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<paragraph pageId="0" pageNumber="161">
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<taxonomicName authorityName="H. L. Clark" authorityYear="1939" class="Ophiuroidea" family="Euryalidae" genus="Asteroschema" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Asteroschema lissum" order="Euryalida" pageId="0" pageNumber="161" phylum="Echinodermata" rank="species" species="lissum">Asteroschema lissum</taxonomicName>
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H. L. Clark, 1939: 37-39, figs 1-3.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="161" type="materials_examined">
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<paragraph pageId="0" pageNumber="161">Material examined.</paragraph>
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<paragraph pageId="0" pageNumber="161">
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<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" country="China" location="China" specimenCount="2">
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<collectingCountry name="China">China</collectingCountry>
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•
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<specimenCount type="generic">2 specimens</specimenCount>
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; South
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</materialsCitation>
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<materialsCitation accessionNumber="OK044300" collectingDate="2020-03-30" country="China" elevation="1515" latitude="13.603333" location="Sea" longLatPrecision="13" longitude="113.56233" municipality="Zhongsha Islands" specimenCount="1">
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<collectingCountry name="China">China</collectingCountry>
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<location LSID="urn:lsid:plazi:treatment:07D71215F3FD5F089C88ACF1CFA74132:1ADF278E5E22D4F0DD4DBF428A901438" country="China" latitude="13.603333" longLatPrecision="13" longitude="113.56233" municipality="Zhongsha Islands" name="Sea">Sea</location>
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,
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<collectingMunicipality>Zhongsha Islands</collectingMunicipality>
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, seamount;
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<geoCoordinate degrees="13" direction="north" minutes="36.20" orientation="latitude" precision="9" value="13.603333">13°36.20'N</geoCoordinate>
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,
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<geoCoordinate degrees="113" direction="east" minutes="33.74" orientation="longitude" precision="9" value="113.56233">113°33.74'E</geoCoordinate>
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; depth
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<quantity metricMagnitude="3" metricUnit="m" metricValue="1.515" unit="m" value="1515.0">
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<elevation metricMagnitude="3" metricUnit="m" metricValue="1.515" unit="m" value="1515.0">1515 m</elevation>
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</quantity>
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;
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<collectingDate value="2020-03-30">30 Mar. 2020</collectingDate>
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; Collecting event: stn. SC025;
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<normalizedToken originalValue="‘Shenhaiyongshi’">'Shenhaiyongshi'</normalizedToken>
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msv leg; preserved in -80 °C; GenBank:
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<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/OK044300">OK044300</accessionNumber>
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; IDSSE-EEB-SW0079, IDSSE-EEB-SW080. Northwest Pacific •
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<specimenCount type="generic">1 specimen</specimenCount>
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; near Mariana Trench, Southeast of
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</materialsCitation>
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<materialsCitation accessionNumber="OK044299, OL712207" collectingDate="2019-09-03" country="Guam Island" elevation="1160" latitude="12.111167" location="Guam Island" longLatPrecision="13" longitude="141.62117" specimenCount="1">
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<collectingCountry name="Guam Island">Guam Island</collectingCountry>
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, seamount,
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<geoCoordinate degrees="12" direction="north" minutes="6.67" orientation="latitude" precision="9" value="12.111167">12°6.67'N</geoCoordinate>
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,
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<geoCoordinate degrees="141" direction="east" minutes="37.27" orientation="longitude" precision="9" value="141.62117">141°37.27'E</geoCoordinate>
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; depth
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<quantity metricMagnitude="3" metricUnit="m" metricValue="1.16" unit="m" value="1160.0">
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<elevation metricMagnitude="3" metricUnit="m" metricValue="1.16" unit="m" value="1160.0">1160 m</elevation>
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</quantity>
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;
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<collectingDate value="2019-09-03">03 Sep. 2019</collectingDate>
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; Collecting event: stn. SC033;
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<normalizedToken originalValue="‘Shenhaiyongshi’">'Shenhaiyongshi'</normalizedToken>
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msv leg; preserved in -80 °C; GenBank:
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<accessionNumber isEnumeration="true">OK044299, OL712207</accessionNumber>
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; IDSSE-EEB-SW0081
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="161" type="description">
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<paragraph pageId="0" pageNumber="161">Description.</paragraph>
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<paragraph pageId="0" pageNumber="161">
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IDSSE-EEB-SW0079: disc diameter 11.5 mm, length of arms 165 mm, arm base width 3-3.5 mm (Fig.
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<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Asteroschema cf. lissum H. L. Clark, 1939 (IDSSE-EEB-SW 0079) A dorsal view B ventral view C dorsal disc D skin on the disc E lateral disc F ventral disc G dorsal arm (middle) H lateral arm (middle) I ventral arm (proximal) J outer arm spine (proximal) K arm spines (middle) L arm spines (distal). Abbreviations: as adoral shield, ass adoral shield spine, ars arm spine, go granular ossicle, gs genital slit, iars inner arm spine, j jaw, oars outer arm spine, ots oral tentacle scale, rs radial shield, t teeth, tp tentacle pore. Scale bars: 2 mm (A, B, G); 1 mm (C, E, F, H, I); 500 µm (J, K); 200 µm (D, L)." figureDoi="10.3897/zookeys.1090.76292.figure16" httpUri="https://binary.pensoft.net/fig/665892" pageId="0" pageNumber="161">16</figureCitation>
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).
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</paragraph>
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<paragraph pageId="0" pageNumber="161">
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<emphasis bold="true" pageId="0" pageNumber="161">
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<emphasis italics="true" pageId="0" pageNumber="161">Disc</emphasis>
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.
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</emphasis>
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Disc flat, slightly raised above arms, swollen in center (Fig.
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<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Asteroschema cf. lissum H. L. Clark, 1939 (IDSSE-EEB-SW 0079) A dorsal view B ventral view C dorsal disc D skin on the disc E lateral disc F ventral disc G dorsal arm (middle) H lateral arm (middle) I ventral arm (proximal) J outer arm spine (proximal) K arm spines (middle) L arm spines (distal). Abbreviations: as adoral shield, ass adoral shield spine, ars arm spine, go granular ossicle, gs genital slit, iars inner arm spine, j jaw, oars outer arm spine, ots oral tentacle scale, rs radial shield, t teeth, tp tentacle pore. Scale bars: 2 mm (A, B, G); 1 mm (C, E, F, H, I); 500 µm (J, K); 200 µm (D, L)." figureDoi="10.3897/zookeys.1090.76292.figure16" httpUri="https://binary.pensoft.net/fig/665892" pageId="0" pageNumber="161">16A, B</figureCitation>
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). Disc covered with smooth, small granular ossicles (Fig.
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<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Asteroschema cf. lissum H. L. Clark, 1939 (IDSSE-EEB-SW 0079) A dorsal view B ventral view C dorsal disc D skin on the disc E lateral disc F ventral disc G dorsal arm (middle) H lateral arm (middle) I ventral arm (proximal) J outer arm spine (proximal) K arm spines (middle) L arm spines (distal). Abbreviations: as adoral shield, ass adoral shield spine, ars arm spine, go granular ossicle, gs genital slit, iars inner arm spine, j jaw, oars outer arm spine, ots oral tentacle scale, rs radial shield, t teeth, tp tentacle pore. Scale bars: 2 mm (A, B, G); 1 mm (C, E, F, H, I); 500 µm (J, K); 200 µm (D, L)." figureDoi="10.3897/zookeys.1090.76292.figure16" httpUri="https://binary.pensoft.net/fig/665892" pageId="0" pageNumber="161">16C, D</figureCitation>
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). Granular ossicles dense and small in disc center (six or eight grains in 1 mm), but slightly larger and separated at distal end of radial shield (five or six grains in 1 mm) (Fig.
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<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Asteroschema cf. lissum H. L. Clark, 1939 (IDSSE-EEB-SW 0079) A dorsal view B ventral view C dorsal disc D skin on the disc E lateral disc F ventral disc G dorsal arm (middle) H lateral arm (middle) I ventral arm (proximal) J outer arm spine (proximal) K arm spines (middle) L arm spines (distal). Abbreviations: as adoral shield, ass adoral shield spine, ars arm spine, go granular ossicle, gs genital slit, iars inner arm spine, j jaw, oars outer arm spine, ots oral tentacle scale, rs radial shield, t teeth, tp tentacle pore. Scale bars: 2 mm (A, B, G); 1 mm (C, E, F, H, I); 500 µm (J, K); 200 µm (D, L)." figureDoi="10.3897/zookeys.1090.76292.figure16" httpUri="https://binary.pensoft.net/fig/665892" pageId="0" pageNumber="161">16C</figureCitation>
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). Radial shields wide, similar in size, curved, slightly raised above disc but not meeting in center (Fig.
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<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Asteroschema cf. lissum H. L. Clark, 1939 (IDSSE-EEB-SW 0079) A dorsal view B ventral view C dorsal disc D skin on the disc E lateral disc F ventral disc G dorsal arm (middle) H lateral arm (middle) I ventral arm (proximal) J outer arm spine (proximal) K arm spines (middle) L arm spines (distal). Abbreviations: as adoral shield, ass adoral shield spine, ars arm spine, go granular ossicle, gs genital slit, iars inner arm spine, j jaw, oars outer arm spine, ots oral tentacle scale, rs radial shield, t teeth, tp tentacle pore. Scale bars: 2 mm (A, B, G); 1 mm (C, E, F, H, I); 500 µm (J, K); 200 µm (D, L)." figureDoi="10.3897/zookeys.1090.76292.figure16" httpUri="https://binary.pensoft.net/fig/665892" pageId="0" pageNumber="161">16A, C</figureCitation>
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). Radial shields clearly recognizable under thin skin embedded with ossicles (Fig.
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<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Asteroschema cf. lissum H. L. Clark, 1939 (IDSSE-EEB-SW 0079) A dorsal view B ventral view C dorsal disc D skin on the disc E lateral disc F ventral disc G dorsal arm (middle) H lateral arm (middle) I ventral arm (proximal) J outer arm spine (proximal) K arm spines (middle) L arm spines (distal). Abbreviations: as adoral shield, ass adoral shield spine, ars arm spine, go granular ossicle, gs genital slit, iars inner arm spine, j jaw, oars outer arm spine, ots oral tentacle scale, rs radial shield, t teeth, tp tentacle pore. Scale bars: 2 mm (A, B, G); 1 mm (C, E, F, H, I); 500 µm (J, K); 200 µm (D, L)." figureDoi="10.3897/zookeys.1090.76292.figure16" httpUri="https://binary.pensoft.net/fig/665892" pageId="0" pageNumber="161">16A, C</figureCitation>
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). Genital slits narrow, concealed by polygonal granular ossicles, and vertical on ventral interradii (Fig.
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<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Asteroschema cf. lissum H. L. Clark, 1939 (IDSSE-EEB-SW 0079) A dorsal view B ventral view C dorsal disc D skin on the disc E lateral disc F ventral disc G dorsal arm (middle) H lateral arm (middle) I ventral arm (proximal) J outer arm spine (proximal) K arm spines (middle) L arm spines (distal). Abbreviations: as adoral shield, ass adoral shield spine, ars arm spine, go granular ossicle, gs genital slit, iars inner arm spine, j jaw, oars outer arm spine, ots oral tentacle scale, rs radial shield, t teeth, tp tentacle pore. Scale bars: 2 mm (A, B, G); 1 mm (C, E, F, H, I); 500 µm (J, K); 200 µm (D, L)." figureDoi="10.3897/zookeys.1090.76292.figure16" httpUri="https://binary.pensoft.net/fig/665892" pageId="0" pageNumber="161">16E</figureCitation>
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). Jaws elongated, mostly naked without granular ossicles (Fig.
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<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Asteroschema cf. lissum H. L. Clark, 1939 (IDSSE-EEB-SW 0079) A dorsal view B ventral view C dorsal disc D skin on the disc E lateral disc F ventral disc G dorsal arm (middle) H lateral arm (middle) I ventral arm (proximal) J outer arm spine (proximal) K arm spines (middle) L arm spines (distal). Abbreviations: as adoral shield, ass adoral shield spine, ars arm spine, go granular ossicle, gs genital slit, iars inner arm spine, j jaw, oars outer arm spine, ots oral tentacle scale, rs radial shield, t teeth, tp tentacle pore. Scale bars: 2 mm (A, B, G); 1 mm (C, E, F, H, I); 500 µm (J, K); 200 µm (D, L)." figureDoi="10.3897/zookeys.1090.76292.figure16" httpUri="https://binary.pensoft.net/fig/665892" pageId="0" pageNumber="161">16F</figureCitation>
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). Flattened, spearhead-shaped teeth, and six to seven granular ossicles that resemble lateral oral papillae at apex of jaw (Fig.
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<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Asteroschema cf. lissum H. L. Clark, 1939 (IDSSE-EEB-SW 0079) A dorsal view B ventral view C dorsal disc D skin on the disc E lateral disc F ventral disc G dorsal arm (middle) H lateral arm (middle) I ventral arm (proximal) J outer arm spine (proximal) K arm spines (middle) L arm spines (distal). Abbreviations: as adoral shield, ass adoral shield spine, ars arm spine, go granular ossicle, gs genital slit, iars inner arm spine, j jaw, oars outer arm spine, ots oral tentacle scale, rs radial shield, t teeth, tp tentacle pore. Scale bars: 2 mm (A, B, G); 1 mm (C, E, F, H, I); 500 µm (J, K); 200 µm (D, L)." figureDoi="10.3897/zookeys.1090.76292.figure16" httpUri="https://binary.pensoft.net/fig/665892" pageId="0" pageNumber="161">16F</figureCitation>
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). Ventral disc covered with widely separated small granular ossicles (six or seven grains in 1 mm), but mostly covered by translucent thin skin (Fig.
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<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Asteroschema cf. lissum H. L. Clark, 1939 (IDSSE-EEB-SW 0079) A dorsal view B ventral view C dorsal disc D skin on the disc E lateral disc F ventral disc G dorsal arm (middle) H lateral arm (middle) I ventral arm (proximal) J outer arm spine (proximal) K arm spines (middle) L arm spines (distal). Abbreviations: as adoral shield, ass adoral shield spine, ars arm spine, go granular ossicle, gs genital slit, iars inner arm spine, j jaw, oars outer arm spine, ots oral tentacle scale, rs radial shield, t teeth, tp tentacle pore. Scale bars: 2 mm (A, B, G); 1 mm (C, E, F, H, I); 500 µm (J, K); 200 µm (D, L)." figureDoi="10.3897/zookeys.1090.76292.figure16" httpUri="https://binary.pensoft.net/fig/665892" pageId="0" pageNumber="161">16F</figureCitation>
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). Adoral shields large, distal edge convex, concealed by thin skin with scattered small, smooth granular ossicles (Fig.
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<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Asteroschema cf. lissum H. L. Clark, 1939 (IDSSE-EEB-SW 0079) A dorsal view B ventral view C dorsal disc D skin on the disc E lateral disc F ventral disc G dorsal arm (middle) H lateral arm (middle) I ventral arm (proximal) J outer arm spine (proximal) K arm spines (middle) L arm spines (distal). Abbreviations: as adoral shield, ass adoral shield spine, ars arm spine, go granular ossicle, gs genital slit, iars inner arm spine, j jaw, oars outer arm spine, ots oral tentacle scale, rs radial shield, t teeth, tp tentacle pore. Scale bars: 2 mm (A, B, G); 1 mm (C, E, F, H, I); 500 µm (J, K); 200 µm (D, L)." figureDoi="10.3897/zookeys.1090.76292.figure16" httpUri="https://binary.pensoft.net/fig/665892" pageId="0" pageNumber="161">16F</figureCitation>
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). Oral shields not discernible and oral tentacle pore naked without ossicles (Fig.
|
||
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Asteroschema cf. lissum H. L. Clark, 1939 (IDSSE-EEB-SW 0079) A dorsal view B ventral view C dorsal disc D skin on the disc E lateral disc F ventral disc G dorsal arm (middle) H lateral arm (middle) I ventral arm (proximal) J outer arm spine (proximal) K arm spines (middle) L arm spines (distal). Abbreviations: as adoral shield, ass adoral shield spine, ars arm spine, go granular ossicle, gs genital slit, iars inner arm spine, j jaw, oars outer arm spine, ots oral tentacle scale, rs radial shield, t teeth, tp tentacle pore. Scale bars: 2 mm (A, B, G); 1 mm (C, E, F, H, I); 500 µm (J, K); 200 µm (D, L)." figureDoi="10.3897/zookeys.1090.76292.figure16" httpUri="https://binary.pensoft.net/fig/665892" pageId="0" pageNumber="161">16F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.1090.76292.figure16" httpUri="https://binary.pensoft.net/fig/665892" pageId="0" pageNumber="161" start="Figure 16" startId="F16">
|
||
<paragraph pageId="0" pageNumber="161">
|
||
<emphasis bold="true" pageId="0" pageNumber="161">Figure 16.</emphasis>
|
||
<taxonomicName authorityName="H. L. Clark" authorityYear="1939" class="Ophiuroidea" family="Euryalidae" genus="Asteroschema" kingdom="Animalia" lsidName="Asteroschema lissum" order="Euryalida" pageId="0" pageNumber="161" phylum="Echinodermata" rank="species" species="lissum">Asteroschema cf. lissum</taxonomicName>
|
||
H. L. Clark, 1939 (IDSSE-EEB-SW0079)
|
||
<emphasis bold="true" pageId="0" pageNumber="161">A</emphasis>
|
||
dorsal view
|
||
<emphasis bold="true" pageId="0" pageNumber="161">B</emphasis>
|
||
ventral view
|
||
<emphasis bold="true" pageId="0" pageNumber="161">C</emphasis>
|
||
dorsal disc
|
||
<emphasis bold="true" pageId="0" pageNumber="161">D</emphasis>
|
||
skin on the disc
|
||
<emphasis bold="true" pageId="0" pageNumber="161">E</emphasis>
|
||
lateral disc
|
||
<emphasis bold="true" pageId="0" pageNumber="161">F</emphasis>
|
||
ventral disc
|
||
<emphasis bold="true" pageId="0" pageNumber="161">G</emphasis>
|
||
dorsal arm (middle)
|
||
<emphasis bold="true" pageId="0" pageNumber="161">H</emphasis>
|
||
lateral arm (middle)
|
||
<emphasis bold="true" pageId="0" pageNumber="161">I</emphasis>
|
||
ventral arm (proximal)
|
||
<emphasis bold="true" pageId="0" pageNumber="161">J</emphasis>
|
||
outer arm spine (proximal)
|
||
<emphasis bold="true" pageId="0" pageNumber="161">K</emphasis>
|
||
arm spines (middle)
|
||
<emphasis bold="true" pageId="0" pageNumber="161">L</emphasis>
|
||
arm spines (distal). Abbreviations:
|
||
<emphasis bold="true" pageId="0" pageNumber="161">as</emphasis>
|
||
adoral shield,
|
||
<emphasis bold="true" pageId="0" pageNumber="161">ass</emphasis>
|
||
adoral shield spine,
|
||
<emphasis bold="true" pageId="0" pageNumber="161">ars</emphasis>
|
||
arm spine,
|
||
<emphasis bold="true" pageId="0" pageNumber="161">go</emphasis>
|
||
granular ossicle,
|
||
<emphasis bold="true" pageId="0" pageNumber="161">gs</emphasis>
|
||
genital slit,
|
||
<emphasis bold="true" pageId="0" pageNumber="161">iars</emphasis>
|
||
inner arm spine,
|
||
<emphasis bold="true" pageId="0" pageNumber="161">j</emphasis>
|
||
jaw,
|
||
<emphasis bold="true" pageId="0" pageNumber="161">oars</emphasis>
|
||
outer arm spine,
|
||
<emphasis bold="true" pageId="0" pageNumber="161">ots</emphasis>
|
||
oral tentacle scale,
|
||
<emphasis bold="true" pageId="0" pageNumber="161">rs</emphasis>
|
||
radial shield,
|
||
<emphasis bold="true" pageId="0" pageNumber="161">t</emphasis>
|
||
teeth,
|
||
<emphasis bold="true" pageId="0" pageNumber="161">tp</emphasis>
|
||
tentacle pore. Scale bars: 2 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="161">A, B, G</emphasis>
|
||
); 1 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="161">C, E, F, H, I</emphasis>
|
||
); 500
|
||
<normalizedToken originalValue="µm">µm</normalizedToken>
|
||
(
|
||
<emphasis bold="true" pageId="0" pageNumber="161">J, K</emphasis>
|
||
); 200
|
||
<normalizedToken originalValue="µm">µm</normalizedToken>
|
||
(
|
||
<emphasis bold="true" pageId="0" pageNumber="161">D, L</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="161">
|
||
<emphasis bold="true" pageId="0" pageNumber="161">
|
||
<emphasis italics="true" pageId="0" pageNumber="161">Arms</emphasis>
|
||
.
|
||
</emphasis>
|
||
Arms slightly arched, circular, from middle to distal half narrower and more cylindrical (Fig.
|
||
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Asteroschema cf. lissum H. L. Clark, 1939 (IDSSE-EEB-SW 0079) A dorsal view B ventral view C dorsal disc D skin on the disc E lateral disc F ventral disc G dorsal arm (middle) H lateral arm (middle) I ventral arm (proximal) J outer arm spine (proximal) K arm spines (middle) L arm spines (distal). Abbreviations: as adoral shield, ass adoral shield spine, ars arm spine, go granular ossicle, gs genital slit, iars inner arm spine, j jaw, oars outer arm spine, ots oral tentacle scale, rs radial shield, t teeth, tp tentacle pore. Scale bars: 2 mm (A, B, G); 1 mm (C, E, F, H, I); 500 µm (J, K); 200 µm (D, L)." figureDoi="10.3897/zookeys.1090.76292.figure16" httpUri="https://binary.pensoft.net/fig/665892" pageId="0" pageNumber="161">16G</figureCitation>
|
||
). Dorsal arm base covered with granular ossicles similar to dorsal disc (six or eight grains in 1 mm), increasingly separated and decreasing in size along the middle segments of the arm (seven or eight grains in 1 mm), distally widely separated (five or six grains in 1 mm) (Fig.
|
||
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Asteroschema cf. lissum H. L. Clark, 1939 (IDSSE-EEB-SW 0079) A dorsal view B ventral view C dorsal disc D skin on the disc E lateral disc F ventral disc G dorsal arm (middle) H lateral arm (middle) I ventral arm (proximal) J outer arm spine (proximal) K arm spines (middle) L arm spines (distal). Abbreviations: as adoral shield, ass adoral shield spine, ars arm spine, go granular ossicle, gs genital slit, iars inner arm spine, j jaw, oars outer arm spine, ots oral tentacle scale, rs radial shield, t teeth, tp tentacle pore. Scale bars: 2 mm (A, B, G); 1 mm (C, E, F, H, I); 500 µm (J, K); 200 µm (D, L)." figureDoi="10.3897/zookeys.1090.76292.figure16" httpUri="https://binary.pensoft.net/fig/665892" pageId="0" pageNumber="161">16G, H</figureCitation>
|
||
). On lateral arm plates, granular ossicles widely separated but continuing to base of arm (Fig.
|
||
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Asteroschema cf. lissum H. L. Clark, 1939 (IDSSE-EEB-SW 0079) A dorsal view B ventral view C dorsal disc D skin on the disc E lateral disc F ventral disc G dorsal arm (middle) H lateral arm (middle) I ventral arm (proximal) J outer arm spine (proximal) K arm spines (middle) L arm spines (distal). Abbreviations: as adoral shield, ass adoral shield spine, ars arm spine, go granular ossicle, gs genital slit, iars inner arm spine, j jaw, oars outer arm spine, ots oral tentacle scale, rs radial shield, t teeth, tp tentacle pore. Scale bars: 2 mm (A, B, G); 1 mm (C, E, F, H, I); 500 µm (J, K); 200 µm (D, L)." figureDoi="10.3897/zookeys.1090.76292.figure16" httpUri="https://binary.pensoft.net/fig/665892" pageId="0" pageNumber="161">16H, K</figureCitation>
|
||
). Only one or three ventral arm segments near arm base covered with few granular ossicles similar to ventral disc (six or seven grains in 1 mm), then completely naked along the arm (Fig.
|
||
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Asteroschema cf. lissum H. L. Clark, 1939 (IDSSE-EEB-SW 0079) A dorsal view B ventral view C dorsal disc D skin on the disc E lateral disc F ventral disc G dorsal arm (middle) H lateral arm (middle) I ventral arm (proximal) J outer arm spine (proximal) K arm spines (middle) L arm spines (distal). Abbreviations: as adoral shield, ass adoral shield spine, ars arm spine, go granular ossicle, gs genital slit, iars inner arm spine, j jaw, oars outer arm spine, ots oral tentacle scale, rs radial shield, t teeth, tp tentacle pore. Scale bars: 2 mm (A, B, G); 1 mm (C, E, F, H, I); 500 µm (J, K); 200 µm (D, L)." figureDoi="10.3897/zookeys.1090.76292.figure16" httpUri="https://binary.pensoft.net/fig/665892" pageId="0" pageNumber="161">16I, J</figureCitation>
|
||
). First arm spine appears at second arm segment, second arm spine at ninth or thirteenth segment (Fig.
|
||
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Asteroschema cf. lissum H. L. Clark, 1939 (IDSSE-EEB-SW 0079) A dorsal view B ventral view C dorsal disc D skin on the disc E lateral disc F ventral disc G dorsal arm (middle) H lateral arm (middle) I ventral arm (proximal) J outer arm spine (proximal) K arm spines (middle) L arm spines (distal). Abbreviations: as adoral shield, ass adoral shield spine, ars arm spine, go granular ossicle, gs genital slit, iars inner arm spine, j jaw, oars outer arm spine, ots oral tentacle scale, rs radial shield, t teeth, tp tentacle pore. Scale bars: 2 mm (A, B, G); 1 mm (C, E, F, H, I); 500 µm (J, K); 200 µm (D, L)." figureDoi="10.3897/zookeys.1090.76292.figure16" httpUri="https://binary.pensoft.net/fig/665892" pageId="0" pageNumber="161">16I-K</figureCitation>
|
||
). Inner arm spine initially short, thick with thorny pointed tip, at middle arm cylindrical, taller, one and a half arm segment in length, with flattened thorny tip (Fig.
|
||
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Asteroschema cf. lissum H. L. Clark, 1939 (IDSSE-EEB-SW 0079) A dorsal view B ventral view C dorsal disc D skin on the disc E lateral disc F ventral disc G dorsal arm (middle) H lateral arm (middle) I ventral arm (proximal) J outer arm spine (proximal) K arm spines (middle) L arm spines (distal). Abbreviations: as adoral shield, ass adoral shield spine, ars arm spine, go granular ossicle, gs genital slit, iars inner arm spine, j jaw, oars outer arm spine, ots oral tentacle scale, rs radial shield, t teeth, tp tentacle pore. Scale bars: 2 mm (A, B, G); 1 mm (C, E, F, H, I); 500 µm (J, K); 200 µm (D, L)." figureDoi="10.3897/zookeys.1090.76292.figure16" httpUri="https://binary.pensoft.net/fig/665892" pageId="0" pageNumber="161">16J, K</figureCitation>
|
||
). Outer arm spine half as long as inner spine in middle region, with smooth to thorny tip (Fig.
|
||
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Asteroschema cf. lissum H. L. Clark, 1939 (IDSSE-EEB-SW 0079) A dorsal view B ventral view C dorsal disc D skin on the disc E lateral disc F ventral disc G dorsal arm (middle) H lateral arm (middle) I ventral arm (proximal) J outer arm spine (proximal) K arm spines (middle) L arm spines (distal). Abbreviations: as adoral shield, ass adoral shield spine, ars arm spine, go granular ossicle, gs genital slit, iars inner arm spine, j jaw, oars outer arm spine, ots oral tentacle scale, rs radial shield, t teeth, tp tentacle pore. Scale bars: 2 mm (A, B, G); 1 mm (C, E, F, H, I); 500 µm (J, K); 200 µm (D, L)." figureDoi="10.3897/zookeys.1090.76292.figure16" httpUri="https://binary.pensoft.net/fig/665892" pageId="0" pageNumber="161">16K</figureCitation>
|
||
). Both arm spines equal in length at distal end of arm, and compound hook with five or six secondary teeth (Fig.
|
||
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Asteroschema cf. lissum H. L. Clark, 1939 (IDSSE-EEB-SW 0079) A dorsal view B ventral view C dorsal disc D skin on the disc E lateral disc F ventral disc G dorsal arm (middle) H lateral arm (middle) I ventral arm (proximal) J outer arm spine (proximal) K arm spines (middle) L arm spines (distal). Abbreviations: as adoral shield, ass adoral shield spine, ars arm spine, go granular ossicle, gs genital slit, iars inner arm spine, j jaw, oars outer arm spine, ots oral tentacle scale, rs radial shield, t teeth, tp tentacle pore. Scale bars: 2 mm (A, B, G); 1 mm (C, E, F, H, I); 500 µm (J, K); 200 µm (D, L)." figureDoi="10.3897/zookeys.1090.76292.figure16" httpUri="https://binary.pensoft.net/fig/665892" pageId="0" pageNumber="161">16L</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="161">
|
||
<emphasis bold="true" pageId="0" pageNumber="161">
|
||
<emphasis italics="true" pageId="0" pageNumber="161">Color</emphasis>
|
||
.
|
||
</emphasis>
|
||
In live specimen, reddish brown but radial shields slightly lighter in color due to thin skin (Fig.
|
||
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Asteroschema cf. lissum H. L. Clark, 1939 (IDSSE-EEB-SW 0079) A dorsal view B ventral view C dorsal disc D skin on the disc E lateral disc F ventral disc G dorsal arm (middle) H lateral arm (middle) I ventral arm (proximal) J outer arm spine (proximal) K arm spines (middle) L arm spines (distal). Abbreviations: as adoral shield, ass adoral shield spine, ars arm spine, go granular ossicle, gs genital slit, iars inner arm spine, j jaw, oars outer arm spine, ots oral tentacle scale, rs radial shield, t teeth, tp tentacle pore. Scale bars: 2 mm (A, B, G); 1 mm (C, E, F, H, I); 500 µm (J, K); 200 µm (D, L)." figureDoi="10.3897/zookeys.1090.76292.figure16" httpUri="https://binary.pensoft.net/fig/665892" pageId="0" pageNumber="161">16</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="161" type="ossicle morphology">
|
||
<paragraph pageId="0" pageNumber="161">Ossicle morphology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="161">
|
||
Lateral arm plate with two arm spine articular structures, with large muscle and nerve openings (Fig.
|
||
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Asteroschema cf. lissum H. L. Clark, 1939 (IDSSE-EEB-SW 0079) A, B lateral arm plate (external, internal) C outer arm spine (middle) D arm spine (distal) E skin from dorsal arm base (granular ossicles) F-J vertebrae F proximal view G distal view H lateral view I dorsal view J ventral view. Abbreviations: d dorsal, dist distal, lac lateral ambulacral canals, mo muscle opening, no nerve opening, pb podial basin, prox proximal, st secondary teeth, v ventral. Scale bars: 800 µm (F-J); 500 µm (A, C); 300 µm (B, D); 200 µm (E)." figureDoi="10.3897/zookeys.1090.76292.figure17" httpUri="https://binary.pensoft.net/fig/665893" pageId="0" pageNumber="161">17A, B</figureCitation>
|
||
). Inner arm spine from proximal and middle half of arm cylindrical, with thorny tip (Fig.
|
||
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Asteroschema cf. lissum H. L. Clark, 1939 (IDSSE-EEB-SW 0079) A, B lateral arm plate (external, internal) C outer arm spine (middle) D arm spine (distal) E skin from dorsal arm base (granular ossicles) F-J vertebrae F proximal view G distal view H lateral view I dorsal view J ventral view. Abbreviations: d dorsal, dist distal, lac lateral ambulacral canals, mo muscle opening, no nerve opening, pb podial basin, prox proximal, st secondary teeth, v ventral. Scale bars: 800 µm (F-J); 500 µm (A, C); 300 µm (B, D); 200 µm (E)." figureDoi="10.3897/zookeys.1090.76292.figure17" httpUri="https://binary.pensoft.net/fig/665893" pageId="0" pageNumber="161">17C</figureCitation>
|
||
). Distally, arm spine turns into compound hook with secondary teeth (Fig.
|
||
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Asteroschema cf. lissum H. L. Clark, 1939 (IDSSE-EEB-SW 0079) A, B lateral arm plate (external, internal) C outer arm spine (middle) D arm spine (distal) E skin from dorsal arm base (granular ossicles) F-J vertebrae F proximal view G distal view H lateral view I dorsal view J ventral view. Abbreviations: d dorsal, dist distal, lac lateral ambulacral canals, mo muscle opening, no nerve opening, pb podial basin, prox proximal, st secondary teeth, v ventral. Scale bars: 800 µm (F-J); 500 µm (A, C); 300 µm (B, D); 200 µm (E)." figureDoi="10.3897/zookeys.1090.76292.figure17" httpUri="https://binary.pensoft.net/fig/665893" pageId="0" pageNumber="161">17D</figureCitation>
|
||
). Arm and disc concealed by less dense, wider, and short granular ossicles (Fig.
|
||
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Asteroschema cf. lissum H. L. Clark, 1939 (IDSSE-EEB-SW 0079) A, B lateral arm plate (external, internal) C outer arm spine (middle) D arm spine (distal) E skin from dorsal arm base (granular ossicles) F-J vertebrae F proximal view G distal view H lateral view I dorsal view J ventral view. Abbreviations: d dorsal, dist distal, lac lateral ambulacral canals, mo muscle opening, no nerve opening, pb podial basin, prox proximal, st secondary teeth, v ventral. Scale bars: 800 µm (F-J); 500 µm (A, C); 300 µm (B, D); 200 µm (E)." figureDoi="10.3897/zookeys.1090.76292.figure17" httpUri="https://binary.pensoft.net/fig/665893" pageId="0" pageNumber="161">17E</figureCitation>
|
||
). Vertebrae with streptospondylous articulation, dorsally a median longitudinal furrow, ventrally with deep median longitudinal groove with lateral ambulacral canals, podial basins relatively small (Fig.
|
||
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Asteroschema cf. lissum H. L. Clark, 1939 (IDSSE-EEB-SW 0079) A, B lateral arm plate (external, internal) C outer arm spine (middle) D arm spine (distal) E skin from dorsal arm base (granular ossicles) F-J vertebrae F proximal view G distal view H lateral view I dorsal view J ventral view. Abbreviations: d dorsal, dist distal, lac lateral ambulacral canals, mo muscle opening, no nerve opening, pb podial basin, prox proximal, st secondary teeth, v ventral. Scale bars: 800 µm (F-J); 500 µm (A, C); 300 µm (B, D); 200 µm (E)." figureDoi="10.3897/zookeys.1090.76292.figure17" httpUri="https://binary.pensoft.net/fig/665893" pageId="0" pageNumber="161">17F-J</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.1090.76292.figure17" httpUri="https://binary.pensoft.net/fig/665893" pageId="0" pageNumber="161" start="Figure 17" startId="F17">
|
||
<paragraph pageId="0" pageNumber="161">
|
||
<emphasis bold="true" pageId="0" pageNumber="161">Figure 17.</emphasis>
|
||
<taxonomicName authorityName="H. L. Clark" authorityYear="1939" class="Ophiuroidea" family="Euryalidae" genus="Asteroschema" kingdom="Animalia" lsidName="Asteroschema lissum" order="Euryalida" pageId="0" pageNumber="161" phylum="Echinodermata" rank="species" species="lissum">Asteroschema cf. lissum</taxonomicName>
|
||
H. L. Clark, 1939 (IDSSE-EEB-SW0079)
|
||
<emphasis bold="true" pageId="0" pageNumber="161">A, B</emphasis>
|
||
lateral arm plate (external, internal)
|
||
<emphasis bold="true" pageId="0" pageNumber="161">C</emphasis>
|
||
outer arm spine (middle)
|
||
<emphasis bold="true" pageId="0" pageNumber="161">D</emphasis>
|
||
arm spine (distal)
|
||
<emphasis bold="true" pageId="0" pageNumber="161">E</emphasis>
|
||
skin from dorsal arm base (granular ossicles)
|
||
<emphasis bold="true" pageId="0" pageNumber="161">F-J</emphasis>
|
||
vertebrae
|
||
<emphasis bold="true" pageId="0" pageNumber="161">F</emphasis>
|
||
proximal view
|
||
<emphasis bold="true" pageId="0" pageNumber="161">G</emphasis>
|
||
distal view
|
||
<emphasis bold="true" pageId="0" pageNumber="161">H</emphasis>
|
||
lateral view
|
||
<emphasis bold="true" pageId="0" pageNumber="161">I</emphasis>
|
||
dorsal view
|
||
<emphasis bold="true" pageId="0" pageNumber="161">J</emphasis>
|
||
ventral view. Abbreviations:
|
||
<emphasis bold="true" pageId="0" pageNumber="161">d</emphasis>
|
||
dorsal,
|
||
<emphasis bold="true" pageId="0" pageNumber="161">dist</emphasis>
|
||
distal,
|
||
<emphasis bold="true" pageId="0" pageNumber="161">lac</emphasis>
|
||
lateral ambulacral canals,
|
||
<emphasis bold="true" pageId="0" pageNumber="161">mo</emphasis>
|
||
muscle opening,
|
||
<emphasis bold="true" pageId="0" pageNumber="161">no</emphasis>
|
||
nerve opening,
|
||
<emphasis bold="true" pageId="0" pageNumber="161">pb</emphasis>
|
||
podial basin,
|
||
<emphasis bold="true" pageId="0" pageNumber="161">prox</emphasis>
|
||
proximal,
|
||
<emphasis bold="true" pageId="0" pageNumber="161">st</emphasis>
|
||
secondary teeth,
|
||
<emphasis bold="true" pageId="0" pageNumber="161">v</emphasis>
|
||
ventral. Scale bars: 800
|
||
<normalizedToken originalValue="µm">µm</normalizedToken>
|
||
(
|
||
<emphasis bold="true" pageId="0" pageNumber="161">F-J</emphasis>
|
||
); 500
|
||
<normalizedToken originalValue="µm">µm</normalizedToken>
|
||
(
|
||
<emphasis bold="true" pageId="0" pageNumber="161">A, C</emphasis>
|
||
); 300
|
||
<normalizedToken originalValue="µm">µm</normalizedToken>
|
||
(
|
||
<emphasis bold="true" pageId="0" pageNumber="161">B, D</emphasis>
|
||
); 200
|
||
<normalizedToken originalValue="µm">µm</normalizedToken>
|
||
(
|
||
<emphasis bold="true" pageId="0" pageNumber="161">E</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="161" type="distribution">
|
||
<paragraph pageId="0" pageNumber="161">Distribution.</paragraph>
|
||
<paragraph pageId="0" pageNumber="161">797-1515 m depth. Maldives, South China Sea, Northwest Pacific.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="161" type="remarks">
|
||
<paragraph pageId="0" pageNumber="161">Remarks.</paragraph>
|
||
<paragraph pageId="0" pageNumber="161">
|
||
<taxonomicName authorityName="H. L. Clark" authorityYear="1939" class="Ophiuroidea" family="Euryalidae" genus="Asteroschema" kingdom="Animalia" lsidName="Asteroschema lissum" order="Euryalida" pageId="0" pageNumber="161" phylum="Echinodermata" rank="species" species="lissum">
|
||
<emphasis italics="true" pageId="0" pageNumber="161">Asteroschema lissum</emphasis>
|
||
</taxonomicName>
|
||
was first described by H. L.
|
||
<bibRefCitation author="Clark, HL" journalOrPublisher="Bulletin of the Museum of Comparative Zoology at Harvard" pageId="0" pageNumber="161" refId="B12" refString="Clark, HL, 1939. Ophiuroidea. Scientific Reports from the John Murray Exp. 1933-34 6: 29-136." title="Ophiuroidea. Scientific Reports from the John Murray Exp. 1933 - 34 6: 29 - 136." year="1939">Clark (1939)</bibRefCitation>
|
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, with type locality Maldives waters in the Indian Ocean. This is the first redescription after the original description. The specimens from our collection were close to H. L.
|
||
<normalizedToken originalValue="Clark’s">Clark's</normalizedToken>
|
||
description but we noticed some differences, such as: characters of radial shields, and granular ossicles at ventral disc and base of arm. We hesitate to fully associate our specimens with
|
||
<taxonomicName authorityName="H. L. Clark" authorityYear="1939" class="Ophiuroidea" family="Euryalidae" genus="Asteroschema" kingdom="Animalia" lsidName="Asteroschema lissum" order="Euryalida" pageId="0" pageNumber="161" phylum="Echinodermata" rank="species" species="lissum">
|
||
<emphasis italics="true" pageId="0" pageNumber="161">Asteroschema lissum</emphasis>
|
||
</taxonomicName>
|
||
or propose a new species, due to uncertainty of these morphological variations. Some of these variations may be affected by size, maturity, and environment (holotype disc diameter 7.5 mm).
|
||
<taxonomicName authorityName="H. L. Clark" authorityYear="1939" class="Ophiuroidea" family="Euryalidae" genus="Asteroschema" kingdom="Animalia" lsidName="Asteroschema lissum" order="Euryalida" pageId="0" pageNumber="161" phylum="Echinodermata" rank="species" species="lissum">
|
||
<emphasis italics="true" pageId="0" pageNumber="161">Asteroschema lissum</emphasis>
|
||
</taxonomicName>
|
||
strongly resembles
|
||
<taxonomicName lsidName="A. hemigymnum" pageId="0" pageNumber="161" rank="species" species="hemigymnum">
|
||
<emphasis italics="true" pageId="0" pageNumber="161">A. hemigymnum</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName lsidName="A. intectum" pageId="0" pageNumber="161" rank="species" species="intectum">
|
||
<emphasis italics="true" pageId="0" pageNumber="161">A. intectum</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName lsidName="A. sublaeve" pageId="0" pageNumber="161" rank="species" species="sublaeve">
|
||
<emphasis italics="true" pageId="0" pageNumber="161">A. sublaeve</emphasis>
|
||
</taxonomicName>
|
||
by having similar granular density, and almost naked ventral disc and arms but differs in characters of the radial shields, start of second arm spine, granulation pattern on the disc and arm (Table
|
||
<tableCitation captionStart="Table 2" captionStartId="T2" captionText="Table 2. Tabular key to the species of Asteroschema and Ophiocreas. Abbreviations: ASS arm segment, ARS arm spine, GO granular ossicles, RS radial shield, CO conical ossicles, PO plate-like ossicles, TP terminal projection, GS genital slits." httpUri="http://table.plazi.org/id/FB8AE5BBB9BA374FFA6C03C1361A84BF" pageId="0" pageNumber="161" tableUuid="FB8AE5BBB9BA374FFA6C03C1361A84BF">2</tableCitation>
|
||
). This is the first record of
|
||
<taxonomicName lsidName="A. lissum" pageId="0" pageNumber="161" rank="species" species="lissum">
|
||
<emphasis italics="true" pageId="0" pageNumber="161">A. lissum</emphasis>
|
||
</taxonomicName>
|
||
from the South China Sea and the North Pacific Ocean, if it is indeed this species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |