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<document id="442BEB1FE7B44A08F9C8D80D5814C0DA" ID-DOI="10.11646/zootaxa.3954.1.1" ID-GBIF-Dataset="44878647-9450-4895-ad32-d1fc79eb30ea" ID-ISSN="1175-5326" ID-Zenodo-Dep="288397" ID-ZooBank="AA3DB71F-AD9A-4205-889B-FB212E367A37" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461048726863" checkinUser="plazi" docAuthor="Hippa, Heikki, Steenis, Jeroen Van &amp; Mutin, Valeri A." docDate="2015" docId="03E04D5BFF9CFF97FF71D14966A7FBDF" docLanguage="en" docName="zt03954p067.pdf" docOrigin="Zootaxa 3954 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Sphegina (Asiosphegina) raduloides Hippa, Steenis &amp; Mutin, 2015, sp. n." docType="treatment" docUuid="A3317818-AABC-4EE8-BEE6-45137ABA0A3E" docUuidSource="ZooBank" docVersion="7" lastPageNumber="30" masterDocId="FFD93523FF80FF8AFFE6D30E6410FF87" masterDocTitle="The genus Sphegina Meigen (Diptera, Syrphidae) in a biodiversity hotspot: the thirty-six sympatric species in Kambaiti, Myanmar" masterLastPageNumber="67" masterPageNumber="1" pageNumber="29" updateTime="1698577992032" updateUser="plazi">
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<mods:title id="1CCEBD4361EC83FC665E29C7DA3CD407">The genus Sphegina Meigen (Diptera, Syrphidae) in a biodiversity hotspot: the thirty-six sympatric species in Kambaiti, Myanmar</mods:title>
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<mods:name id="C3C8418BA0CFDEEFCC2A5A7A360EA357" type="personal">
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<mods:namePart id="170B413F9C78700D776001EACD140308">Hippa, Heikki</mods:namePart>
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<mods:namePart id="BBF6043FF1B13059994A3675C930D09B">Steenis, Jeroen Van</mods:namePart>
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<mods:name id="5882A4087A3578EBF50A32919EE9658F" type="personal">
<mods:role id="3469699002A4644FA5A79135B85530F1">
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<mods:namePart id="B2B26D0585A32E73597EC884E2E689B6">Mutin, Valeri A.</mods:namePart>
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<mods:title id="37F41C5E255B77E6530CF89F019954B4">Zootaxa</mods:title>
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<mods:date id="C1B4658803B75450309A6E0186A5ABAB">2015</mods:date>
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<mods:number id="2A707241826AAD1F57B6BC0118D4DD62">3954</mods:number>
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<mods:classification id="ED88F2D372E81E94B3C66F42A5F86D1B">journal article</mods:classification>
<mods:identifier id="11F9E97D6C00EAA8B8734D5ED7ED3D8C" type="DOI">10.11646/zootaxa.3954.1.1</mods:identifier>
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<mods:identifier id="CBB58BC0429DB375646EEBBE84D5BB1A" type="Zenodo-Dep">288397</mods:identifier>
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<treatment id="03E04D5BFF9CFF97FF71D14966A7FBDF" ID-DOI="http://doi.org/10.5281/zenodo.5692189" ID-GBIF-Taxon="127691271" ID-Zenodo-Dep="5692189" LSID="urn:lsid:zoobank.org:act:A3317818-AABC-4EE8-BEE6-45137ABA0A3E" httpUri="http://treatment.plazi.org/id/03E04D5BFF9CFF97FF71D14966A7FBDF" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29">
<subSubSection id="C353AFC6FF9CFF96FF71D1496530FD04" pageId="28" pageNumber="29" type="nomenclature">
<paragraph id="8BF6FC4DFF9CFF96FF71D1496686FDE5" blockId="28.[151,662,583,643]" box="[151,662,583,610]" pageId="28" pageNumber="29">
<heading id="D0BE4B21FF9CFF96FF71D1496686FDE5" bold="true" box="[151,662,583,610]" fontSize="11" level="1" pageId="28" pageNumber="29" reason="1">
<emphasis id="B93D205FFF9CFF96FF71D1496686FDE5" bold="true" box="[151,662,583,610]" pageId="28" pageNumber="29">
<taxonomicName id="4C4987CEFF9CFF96FF71D149665DFDE6" ID-CoL="8F3ZG" box="[151,589,583,609]" class="Insecta" family="Syrphidae" genus="Sphegina" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="raduloides" status="sp. nov." subGenus="Asiosphegina">
<emphasis id="B93D205FFF9CFF96FF71D149665DFDE6" bold="true" box="[151,589,583,609]" italics="true" pageId="28" pageNumber="29">Sphegina (Asiosphegina) raduloides</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20E9D24FF9CFF96FDB2D1466686FDE5" box="[596,662,584,610]" pageId="28" pageNumber="29" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BF6FC4DFF9CFF96FF71D1646530FD04" blockId="28.[151,662,583,643]" box="[151,288,618,643]" pageId="28" pageNumber="29">
<figureCitation id="1372E0C8FF9CFF96FF71D16464F7FD04" box="[151,231,618,643]" captionStart="FIGURE 15" captionStartId="28.[151,250,1913,1935]" captionTargetBox="[194,1388,921,1881]" captionTargetId="figure@28.[188,1398,911,1891]" captionTargetPageId="28" captionText="FIGURE 15. Sphegina (Asiosphegina) raduloides sp. n. (A paratype, B, C, D holotype). A. Posterior part of sternite IV, ventral view; the unmodified pilosity omitted. B. Genitalia, lateral view. C. Left surstylus, lateral view. D. Left superior lobe, lateral view. Scale 0.5 mm." httpUri="https://zenodo.org/record/288411/files/figure.png" pageId="28" pageNumber="29">Figs 15</figureCitation>
AD
</paragraph>
</subSubSection>
<subSubSection id="C353AFC6FF9CFF97FF71D1BC666EFD2F" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29" type="description">
<paragraph id="8BF6FC4DFF9CFF97FF71D1BC666EFD2F" blockId="28.[151,1436,690,859]" lastBlockId="29.[151,1437,151,1112]" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29">
<emphasis id="B93D205FFF9CFF96FF71D1BC64E2FD4C" bold="true" box="[151,242,690,715]" pageId="28" pageNumber="29">MALE.</emphasis>
Body length
<quantity id="4CB151A8FF9CFF96FE7CD1BC65E7FD4D" box="[410,503,690,715]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="28" pageNumber="29" unit="mm" value="6.5">6.5 mm</quantity>
, wing length
<quantity id="4CB151A8FF9CFF96FD79D1BC66EEFD4D" box="[671,766,690,715]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="28" pageNumber="29" unit="mm" value="5.6">5.6 mm</quantity>
.
<emphasis id="B93D205FFF9CFF96FCE8D1BC6745FD4C" bold="true" box="[782,853,690,715]" pageId="28" pageNumber="29">Head.</emphasis>
Face strongly concave, weakly projected anteroventrally; frontal prominence weakly developed (similar to Fig. 1F). Width of vertex at anterior ocellus:width of head 1:4.1; depth of occipital fossa ca. 1/4 of the width of an eye in dorsal view [1:3.9]. Width of face:width of head 1:4.6. Face yellow, densely pale pollinose. Gena shiny yellow. Frons and vertex shiny black; lunula shiny brown, a narrow pale-pollinose cross band just posterior of the frontal prominence; the pile very short, erect and pale. Occiput dull black. Antenna brownish; basoflagellomere oval [length:width 1:0.8], paler baso-ventrally; arista pilose.
<emphasis id="B93D205FFF9DFF97FECCD3B2659AFF52" bold="true" box="[298,394,188,213]" pageId="29" pageNumber="30">Thorax.</emphasis>
Colour dull black; postpronotum pale pollinose; scutum with short, adpressed densely placed pale pile; scutellum semicircular, with adpressed pale pile which becomes longer and stronger towards the apex and with a pair of long pale setae at apical margin.
<emphasis id="B93D205FFF9DFF97FD03D20A673DFE9A" bold="true" box="[741,813,260,285]" pageId="29" pageNumber="30">Wing.</emphasis>
Hyaline, stigma pale.
<emphasis id="B93D205FFF9DFF97FBD6D20A607FFE9A" bold="true" box="[1072,1135,260,285]" pageId="29" pageNumber="30">Legs.</emphasis>
Pro- and mesoleg yellow; tarsomeres darker toward apex, sometimes 4 and 5 brown. Metaleg: coxa brownish; trochanter brownish, simple; femur brownish, the basal 1/3 yellow, the yellow basal part with 23 small black and yellow setulae ventrally, well separated from the black setae on the dark part of the femur; metatibia without apico-ventral tooth, brownish, the basal 1/3 and an annulus on the apical 1/2 yellow; tarsus black or brown (in paler specimens).
<emphasis id="B93D205FFF9DFF97FB2FD29A6154FE2A" bold="true" box="[1225,1348,404,429]" pageId="29" pageNumber="30">Abdomen.</emphasis>
Length ratio of tergites I, II,
<date id="FFF7DA8DFF9DFF97FE64D2B66614FE57" box="[386,516,439,464]" pageId="29" pageNumber="30">III and IV 1</date>
:3.3:1.7:1.3. Colour of tergite black, anterior 1/3 of tergite III reddish, pile mainly pale, short and adpressed, becoming longer laterally; tergite I with 3 black (sometimes the first one yellow), strong, well separated setae at lateral margin; sternite IV (
<figureCitation id="1372E0C8FF9DFF97FD32D2F16734FD9F" box="[724,804,511,536]" captionStart="FIGURE 15" captionStartId="28.[151,250,1913,1935]" captionTargetBox="[194,1388,921,1881]" captionTargetId="figure@28.[188,1398,911,1891]" captionTargetPageId="28" captionText="FIGURE 15. Sphegina (Asiosphegina) raduloides sp. n. (A paratype, B, C, D holotype). A. Posterior part of sternite IV, ventral view; the unmodified pilosity omitted. B. Genitalia, lateral view. C. Left surstylus, lateral view. D. Left superior lobe, lateral view. Scale 0.5 mm." httpUri="https://zenodo.org/record/288411/files/figure.png" pageId="29" pageNumber="30">Fig. 15</figureCitation>
A) black, posterior margin behind setae yellow, the pile pale, the stronger setae and bristles black, the number and arrangement of the setae and bristles slightly variable; sternites VIVIII simple, black, the pile dense, long and pale.
<emphasis id="B93D205FFF9DFF97FC91D14967FDFDE7" bold="true" box="[887,1005,583,608]" pageId="29" pageNumber="30">Genitalia,</emphasis>
<figureCitation id="1372E0C8FF9DFF97FC1ED149605EFDE7" box="[1016,1102,583,608]" captionStart="FIGURE 15" captionStartId="28.[151,250,1913,1935]" captionTargetBox="[194,1388,921,1881]" captionTargetId="figure@28.[188,1398,911,1891]" captionTargetPageId="28" captionText="FIGURE 15. Sphegina (Asiosphegina) raduloides sp. n. (A paratype, B, C, D holotype). A. Posterior part of sternite IV, ventral view; the unmodified pilosity omitted. B. Genitalia, lateral view. C. Left surstylus, lateral view. D. Left superior lobe, lateral view. Scale 0.5 mm." httpUri="https://zenodo.org/record/288411/files/figure.png" pageId="29" pageNumber="30">Figs 15</figureCitation>
BD. Note the asymmetrical surstyli and superior lobes and the concavity on the dorsal margin of the dorsal lobe of the left side surstylus giving it an angulate shape.
<emphasis id="B93D205FFF9DFF97FE64D1816612FD2F" bold="true" box="[386,514,655,680]" pageId="29" pageNumber="30">FEMALE.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<caption id="DF36ACC5FF9CFF96FF71D4776575F84A" httpUri="https://zenodo.org/record/288411/files/figure.png" pageId="28" pageNumber="29" targetBox="[194,1388,921,1881]" targetPageId="28">
<paragraph id="8BF6FC4DFF9CFF96FF71D4776575F84A" blockId="28.[151,1435,1913,1997]" pageId="28" pageNumber="29">
<emphasis id="B93D205FFF9CFF96FF71D477650FF809" bold="true" box="[151,287,1913,1935]" pageId="28" pageNumber="29">FIGURE 15.</emphasis>
<taxonomicName id="4C4987CEFF9CFF96FEC2D4776684F808" box="[292,660,1913,1935]" class="Insecta" family="Syrphidae" genus="Sphegina" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="raduloides" status="sp. nov." subGenus="Asiosphegina">
<emphasis id="B93D205FFF9CFF96FEC2D4776684F808" box="[292,660,1913,1935]" italics="true" pageId="28" pageNumber="29">Sphegina (Asiosphegina) raduloides</emphasis>
</taxonomicName>
<emphasis id="B93D205FFF9CFF96FD7FD47466C2F808" bold="true" box="[665,722,1914,1935]" pageId="28" pageNumber="29">
<taxonomicNameLabel id="A20E9D24FF9CFF96FD7FD47466C2F808" box="[665,722,1914,1935]" pageId="28" pageNumber="29" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(A paratype, B, C, D holotype).
<emphasis id="B93D205FFF9CFF96FBFCD4776023F809" bold="true" box="[1050,1075,1913,1934]" pageId="28" pageNumber="29">A.</emphasis>
Posterior part of sternite IV, ventral view; the unmodified pilosity omitted.
<emphasis id="B93D205FFF9CFF96FDCFD4966650F82A" bold="true" box="[553,576,1944,1965]" pageId="28" pageNumber="29">B.</emphasis>
Genitalia, lateral view.
<emphasis id="B93D205FFF9CFF96FCD1D4966740F829" bold="true" box="[823,848,1944,1966]" pageId="28" pageNumber="29">C.</emphasis>
Left surstylus, lateral view.
<emphasis id="B93D205FFF9CFF96FB90D496609FF82A" bold="true" box="[1142,1167,1944,1965]" pageId="28" pageNumber="29">D.</emphasis>
Left superior lobe, lateral view. Scale 0.5 mm.
</paragraph>
</caption>
<subSubSection id="C353AFC6FF9DFF97FF21D1BA654EFC93" pageId="29" pageNumber="30" type="materials_examined">
<paragraph id="8BF6FC4DFF9DFF97FF21D1BA654EFC93" blockId="29.[151,1437,151,1112]" pageId="29" pageNumber="30">
<emphasis id="B93D205FFF9DFF97FF21D1BA656EFD4A" bold="true" box="[199,382,692,717]" pageId="29" pageNumber="30">
<typeStatus id="54F242EFFF9DFF97FF21D1BA6512FD4A" box="[199,258,692,717]" pageId="29" pageNumber="30">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54F242EFFF9DFF97FE68D1BA6632FD4B" box="[398,546,692,716]" pageId="29" pageNumber="30" type="holotype">HOLOTYPE</typeStatus>
. ♂, N.E.
<collectingCountry id="F35EBCDDFF9DFF97FD4FD1BB66E8FD4B" box="[681,760,693,716]" name="Myanmar" pageId="29" pageNumber="30">Burma</collectingCountry>
, Kambaiti,
<quantity id="4CB151A8FF9DFF97FC6AD1BA67FCFD4B" box="[908,1004,692,716]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="29" pageNumber="30" unit="m" value="2000.0">2000 m</quantity>
,
<date id="FFF7DA8DFF9DFF97FBE7D1BA609AFD4B" box="[1025,1162,692,717]" pageId="29" pageNumber="30" value="1934-04-26">26.iv. 1934</date>
, R.
<taxonomicName id="4C4987CEFF9DFF97FB2ED1BA6131FD4A" box="[1224,1313,692,717]" class="Insecta" family="Syrphidae" genus="Sphegina" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="19" phylum="Arthropoda" rank="species" species="malaise">Malaise</taxonomicName>
(
<collectionCode id="ED586488FF9DFF97FADFD1BA6180FD4B" box="[1337,1424,692,716]" httpUri="http://grbio.org/cool/8z6v-f8is" name="Saskatchewan Museum of Natural History" pageId="29" pageNumber="30">SMNH</collectionCode>
).
<typeStatus id="54F242EFFF9DFF97FF71D1D6653FFD77" box="[151,303,728,752]" pageId="29" pageNumber="30" type="paratype">PARATYPES</typeStatus>
.
<specimenCount id="9D4F37C4FF9DFF97FED9D1D66572FD68" box="[319,354,728,752]" pageId="29" pageNumber="30" type="generic">2♂</specimenCount>
with same data as
<typeStatus id="54F242EFFF9DFF97FDD5D1D96685FD77" box="[563,661,727,752]" pageId="29" pageNumber="30" type="holotype">holotype</typeStatus>
except
<quantity id="4CB151A8FF9DFF97FD0DD1D6672DFD77" box="[747,829,727,752]" metricMagnitude="3" metricUnit="m" metricValue="2.1336" pageId="29" pageNumber="30" unit="ft" value="7000.0">7000 ft</quantity>
,
<date id="FFF7DA8DFF9DFF97FCAED1D6676EFD77" box="[840,894,728,752]" pageId="29" pageNumber="30">14.v.</date>
(
<collectionCode id="ED586488FF9DFF97FC6BD1D667F2FD77" box="[909,994,728,752]" httpUri="http://grbio.org/cool/8z6v-f8is" name="Saskatchewan Museum of Natural History" pageId="29" pageNumber="30">SMNH</collectionCode>
,
<collectionCode id="ED586488FF9DFF97FC0AD1D6603AFD77" box="[1004,1066,728,752]" pageId="29" pageNumber="30">NBC</collectionCode>
);
<specimenCount id="9D4F37C4FF9DFF97FBD8D1D66071FD68" box="[1086,1121,728,752]" pageId="29" pageNumber="30" type="generic">1♂</specimenCount>
with same data except
<quantity id="4CB151A8FF9DFF97FA82D1D664BBFC92" metricMagnitude="3" metricUnit="m" metricValue="2.1336" pageId="29" pageNumber="30" unit="ft" value="7000.0">7000 ft</quantity>
,
<date id="FFF7DA8DFF9DFF97FF50D1F264FCFC93" box="[182,236,764,788]" pageId="29" pageNumber="30">23.v.</date>
(
<collectionCode id="ED586488FF9DFF97FF1DD1F26542FC93" box="[251,338,764,788]" httpUri="http://grbio.org/cool/8z6v-f8is" name="Saskatchewan Museum of Natural History" pageId="29" pageNumber="30">SMNH</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C353AFC6FF9DFF97FF21D01166F9FCDA" pageId="29" pageNumber="30" type="etymology">
<paragraph id="8BF6FC4DFF9DFF97FF21D01166F9FCDA" blockId="29.[151,1437,151,1112]" pageId="29" pageNumber="30">
<emphasis id="B93D205FFF9DFF97FF21D011655EFCBF" bold="true" box="[199,334,799,824]" pageId="29" pageNumber="30">Etymology.</emphasis>
The name is formed from the name of
<taxonomicName id="4C4987CEFF9DFF97FCD1D02E67B4FCBF" box="[823,932,800,824]" class="Insecta" family="Syrphidae" genus="Sphegina" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="radula">
<emphasis id="B93D205FFF9DFF97FCD1D02E67B4FCBF" box="[823,932,800,824]" italics="true" pageId="29" pageNumber="30">S. radula</emphasis>
</taxonomicName>
by adding the Latin suffix -
<emphasis id="B93D205FFF9DFF97FB19D02E612AFCBF" box="[1279,1338,800,824]" italics="true" pageId="29" pageNumber="30">oides</emphasis>
, like or resembling, referring to the similarity to that species.
</paragraph>
</subSubSection>
<subSubSection id="C353AFC6FF9DFF97FF21D06966A7FBDF" pageId="29" pageNumber="30" type="discussion">
<paragraph id="8BF6FC4DFF9DFF97FF21D06966A7FBDF" blockId="29.[151,1437,151,1112]" pageId="29" pageNumber="30">
<emphasis id="B93D205FFF9DFF97FF21D069655CFC07" bold="true" box="[199,332,871,896]" pageId="29" pageNumber="30">Discussion.</emphasis>
<taxonomicName id="4C4987CEFF9DFF97FEB5D066662AFC07" box="[339,570,872,896]" class="Insecta" family="Syrphidae" genus="Sphegina" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="raduloides">
<emphasis id="B93D205FFF9DFF97FEB5D066662AFC07" box="[339,570,872,896]" italics="true" pageId="29" pageNumber="30">Sphegina raduloides</emphasis>
</taxonomicName>
is similar to
<emphasis id="B93D205FFF9DFF97FD2DD06667DCFC07" box="[715,972,872,896]" italics="true" pageId="29" pageNumber="30">
<taxonomicName id="4C4987CEFF9DFF97FD2DD0666724FC07" box="[715,820,872,896]" class="Insecta" family="Syrphidae" genus="Sphegina" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="radula">S. radula</taxonomicName>
,
<taxonomicName id="4C4987CEFF9DFF97FCD9D06667DCFC07" box="[831,972,872,896]" class="Insecta" family="Syrphidae" genus="Sphegina" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="subradula">S. subradula</taxonomicName>
</emphasis>
and
<taxonomicName id="4C4987CEFF9DFF97FBE4D0666093FC07" box="[1026,1155,872,896]" class="Insecta" family="Syrphidae" genus="Sphegina" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="trichaeta">
<emphasis id="B93D205FFF9DFF97FBE4D0666093FC07" box="[1026,1155,872,896]" italics="true" pageId="29" pageNumber="30">S. trichaeta</emphasis>
</taxonomicName>
. Except for the character mentioned in the key it differs from
<taxonomicName id="4C4987CEFF9DFF97FDA7D08266D4FC23" box="[577,708,908,932]" class="Insecta" family="Syrphidae" genus="Sphegina" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="trichaeta">
<emphasis id="B93D205FFF9DFF97FDA7D08266D4FC23" box="[577,708,908,932]" italics="true" pageId="29" pageNumber="30">S. trichaeta</emphasis>
</taxonomicName>
by lacking three very strong setae on left side of the posterior margin of male sternite IV, greatly deviating from the other setosity, and from all the discussed species by having an unusual concavity on the dorsal margin of the dorsal lobe of the left side surstylus which makes the margin angulate, and by having the right side superior lobe posteriorly bi-lobed. The ventral one of these lobes possibly belongs to the ventral part of sternite IX which in that case is unusually pushed posteriorly. The aedeagus resembles that of
<taxonomicName id="4C4987CEFF9DFF97FF00D74E655DFBDF" box="[230,333,1088,1112]" class="Insecta" family="Syrphidae" genus="Sphegina" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="radula">
<emphasis id="B93D205FFF9DFF97FF00D74E655DFBDF" box="[230,333,1088,1112]" italics="true" pageId="29" pageNumber="30">S. radula</emphasis>
</taxonomicName>
by having the lateral lobe short.
</paragraph>
</subSubSection>
</treatment>
</document>