treatments-xml/data/03/C6/87/03C687C8FFFC1826954C9F83B224C026.xml

<document id="D1FC2F6E2145D4B8BEC88FA9D5192492" ID-CLB-Dataset="22068" ID-DOI="10.5852/ejt.2017.335" ID-GBIF-Dataset="f3382433-ea2c-4ec8-b1fe-44320ac0893b" ID-ISSN="2118-9773" ID-Zenodo-Dep="3832422" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589830608528" checkinUser="carolina" docAuthor="Framenau, Volker W. &amp; Hudson, Peter" docDate="2017" docId="03C687C8FFFC1826954C9F83B224C026" docLanguage="en" docName="ejt-2017-335.pdf.imf" docOrigin="European Journal of Taxonomy 335" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Tetralycosa caudex Framenau &amp; Hudson 2017, sp. nov." docType="treatment" docUuid="5390F04B-FA2D-40FD-BA4C-78777A071684" docUuidSource="ZooBank" docVersion="7" lastPageNumber="27" masterDocId="FFFFFFB0FFE4183C97399D48B752C779" masterDocTitle="Taxonomy, systematics and biology of the Australian halotolerant wolf spider genus Tetralycosa (Araneae: Lycosidae: Artoriinae)" masterLastPageNumber="72" masterPageNumber="1" pageNumber="25" updateTime="1698840103139" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="8C0A38241AE84DDBE325D6890003DD33">Taxonomy, systematics and biology of the Australian halotolerant wolf spider genus Tetralycosa (Araneae: Lycosidae: Artoriinae)</mods:title>
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<mods:namePart id="123F82574FEC650A7B6DF5EAA008FF53">Framenau, Volker W.</mods:namePart>
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<mods:namePart id="2D1105004004A1359FFF463460EEA8B3">Hudson, Peter</mods:namePart>
<mods:affiliation id="F0EC540682DA4CF55D5CC7D982049632">South Australian Museum, North Terrace, Adelaide, South Australia 5000, Australia. &amp; Email: Peter. Hudson @ samuseum. sa. gov. au</mods:affiliation>
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<mods:title id="19FA4738FCEE70519C87E9F90F29AC78">European Journal of Taxonomy</mods:title>
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<mods:date id="74A7FD460B8F221E0B6047BC7E6CE665">2017</mods:date>
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<treatment id="03C687C8FFFC1826954C9F83B224C026" ID-DOI="http://doi.org/10.5281/zenodo.3850580" ID-GBIF-Taxon="164167100" ID-Zenodo-Dep="3850580" LSID="urn:lsid:zoobank.org:act:5390F04B-FA2D-40FD-BA4C-78777A071684" httpUri="http://treatment.plazi.org/id/03C687C8FFFC1826954C9F83B224C026" lastPageId="26" lastPageNumber="27" pageId="24" pageNumber="25">
<subSubSection id="C3756555FFFC1824954C9F83B4ECC59F" box="[629,958,715,742]" pageId="24" pageNumber="25" type="nomenclature">
<paragraph id="8BD036DEFFFC1824954C9F83B4ECC59F" blockId="24.[368,1220,715,824]" box="[629,958,715,742]" pageId="24" pageNumber="25">
<heading id="D09881B2FFFC1824954C9F83B4ECC59F" box="[629,958,715,742]" centered="true" fontSize="11" level="2" pageId="24" pageNumber="25" reason="2">
<taxonomicName id="4C6F4D5DFFFC1824954C9F83B40EC59F" ID-CoL="55P4D" authority="Framenau &amp; Hudson, 2017" authorityName="Framenau &amp; Hudson" authorityYear="2017" box="[629,860,715,742]" class="Arachnida" family="Lycosidae" genus="Tetralycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="caudex" status="sp. nov.">
<emphasis id="B91BEACCFFFC1824954C9F83B40EC59F" bold="true" box="[629,860,715,742]" italics="true" pageId="24" pageNumber="25">Tetralycosa caudex</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22857B7FFFC1824945D9F84B4ECC59F" box="[868,958,716,742]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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urn:lsid:zoobank.org:act: 
<uuid id="FFC90C0BFFFC182495BE9FBCB396C476" box="[647,1220,756,783]" pageId="24" pageNumber="25">
5390F04B-FA2D-40FD-BA4C-78777A 
<date id="FFD1101EFFFC182493519FBCB396C476" box="[1128,1220,756,783]" pageId="24" pageNumber="25" value="84-07-16">071684</date>
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</paragraph>
<paragraph id="8BD036DEFFFC182495939E55B4D8C44E" blockId="24.[368,1220,715,824]" box="[682,906,797,824]" pageId="24" pageNumber="25">
<figureCitation id="13542A5BFFFC182495939E55B5A0C441" box="[682,754,797,824]" captionStart="Fig" captionStartId="20.[189,232,1859,1885]" captionTargetBox="[331,1256,273,1831]" captionTargetId="figure@20.[325,1262,265,1837]" captionTargetPageId="20" captionText="Fig. 5. Tetralycosa Roewer, 1960, microscopic photographs showing dorsal and ventral views. A–B. T. oraria (L. Koch, 1876), ♂ (WAM T93541). C–D. T. oraria (L. Koch, 1876), ♀ (WAM T93944). E–F. T. arabanae (Framenau, Gotch &amp; Austin, 2006), ♂ (WAM T47298). G–H. T. arabanae (Framenau, Gotch &amp; Austin, 2006), ♀ (WAM T47298). I–J. T. caudex sp. nov., ♀ (WAM T55576). Scale bars: A–J = 2.00 mm." figureDoi="http://doi.org/10.5281/zenodo.3832433" httpUri="https://zenodo.org/record/3832433/files/figure.png" pageId="24" pageNumber="25">Figs 5</figureCitation>
I–J; 8A–B; 9
</paragraph>
</subSubSection>
<subSubSection id="C3756555FFFC182497849E2BB329C4CA" box="[189,1147,867,947]" pageId="24" pageNumber="25" type="diagnosis">
<paragraph id="8BD036DEFFFC182497849E2BB664C404" blockId="24.[189,310,867,893]" box="[189,310,867,893]" pageId="24" pageNumber="25">
<heading id="D09881B2FFFC182497849E2BB664C404" bold="true" box="[189,310,867,893]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
<emphasis id="B91BEACCFFFC182497849E2BB664C404" bold="true" box="[189,310,867,893]" pageId="24" pageNumber="25">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BD036DEFFFC182497849ED0B329C4CA" blockId="24.[189,1147,920,947]" box="[189,1147,920,947]" pageId="24" pageNumber="25">
<taxonomicName id="4C6F4D5DFFFC182497849ED0B6CDC4CB" authority="Framenau &amp; Hudson, 2017" authorityName="Framenau &amp; Hudson" authorityYear="2017" box="[189,415,920,946]" class="Arachnida" family="Lycosidae" genus="Tetralycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="caudex" status="sp. nov.">
<emphasis id="B91BEACCFFFC182497849ED0B6CDC4CB" box="[189,415,920,946]" italics="true" pageId="24" pageNumber="25">Tetralycosa caudex</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22857B7FFFC1824969E9ED0B553C4CB" box="[423,513,920,946]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
is very similar to 
<taxonomicName id="4C6F4D5DFFFC182495EC9ED0B430C4CB" baseAuthorityName="Framenau, Gotch &amp; Austin" baseAuthorityYear="2006" box="[725,866,920,946]" class="Arachnida" family="Lycosidae" genus="Tetralycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="arabanae">
<emphasis id="B91BEACCFFFC182495EC9ED0B430C4CB" box="[725,866,920,946]" italics="true" pageId="24" pageNumber="25">T. arabanae</emphasis>
</taxonomicName>
(see ‘Diagnosis’ there).
</paragraph>
</subSubSection>
<subSubSection id="C3756555FFFC182497849E97B513C32C" pageId="24" pageNumber="25" type="etymology">
<paragraph id="8BD036DEFFFC182497849E97B616C480" blockId="24.[189,324,991,1017]" box="[189,324,991,1017]" pageId="24" pageNumber="25">
<heading id="D09881B2FFFC182497849E97B616C480" bold="true" box="[189,324,991,1017]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
<emphasis id="B91BEACCFFFC182497849E97B616C480" bold="true" box="[189,324,991,1017]" pageId="24" pageNumber="25">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BD036DEFFFC18249784995AB513C32C" blockId="24.[189,1399,1042,1110]" pageId="24" pageNumber="25">
The specific epithet is a noun in apposition, from the Latin ‘ 
<taxonomicName id="4C6F4D5DFFFC182494BE995AB488C355" authorityName="Framenau &amp; Hudson" authorityYear="2017" box="[903,986,1042,1068]" class="Arachnida" family="Lycosidae" genus="Tetralycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="caudex">
<emphasis id="B91BEACCFFFC182494BE995AB488C355" box="[903,986,1042,1068]" italics="true" pageId="24" pageNumber="25">caudex</emphasis>
</taxonomicName>
’, wooden stem, in reference to the 
<typeStatus id="54D4887CFFFC182497849974B7A2C32F" box="[189,240,1084,1110]" pageId="24" pageNumber="25">type</typeStatus>
locality, Woodstock Station.
</paragraph>
</subSubSection>
<subSubSection id="C3756555FFFC1825978499CAB430C60E" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25" type="materials_examined">
<paragraph id="8BD036DEFFFC1824978499CAB63EC3E5" blockId="24.[189,364,1154,1180]" box="[189,364,1154,1180]" pageId="24" pageNumber="25">
<heading id="D09881B2FFFC1824978499CAB63EC3E5" bold="true" box="[189,364,1154,1180]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
<emphasis id="B91BEACCFFFC1824978499CAB63EC3E5" bold="true" box="[189,364,1154,1180]" pageId="24" pageNumber="25">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="8BD036DEFFFC182497D399FEB60EC3A9" blockId="24.[189,1399,1206,1314]" box="[234,348,1206,1232]" pageId="24" pageNumber="25">
<emphasis id="B91BEACCFFFC182497D399FEB60EC3A9" bold="true" box="[234,348,1206,1232]" pageId="24" pageNumber="25">
<typeStatus id="54D4887CFFFC182497D399FEB60EC3A9" box="[234,348,1206,1232]" pageId="24" pageNumber="25" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BD036DEFFFC182497849997B485C25B" blockId="24.[189,1399,1206,1314]" pageId="24" pageNumber="25">
<materialsCitation id="3B073C83FFFC182497849997B485C25B" ID-GBIF-Occurrence="2620516301" collectingDate="1988-03-27" collectingMethod="pitfall trap" collectionCode="WAM" collectorName="W. F. Humphreys" country="Australia" latitude="-21.616667" location="Woodstock Station" longLatPrecision="21" longitude="119.02334" pageId="24" pageNumber="25" specimenCode="WAM T55570" specimenCount="1" specimenCount-female="1" stateProvince="Western Australia" typeStatus="holotype">
<collectingCountry id="F378764EFFFC182497849997B63AC380" box="[189,360,1247,1273]" name="Australia" pageId="24" pageNumber="25">AUSTRALIA</collectingCountry>
: 
<specimenCount id="9D69FD57FFFC1824964C9997B6D9C380" box="[373,395,1247,1273]" pageId="24" pageNumber="25" type="female">♀</specimenCount>
, 
<collectingRegion id="49ABF83CFFFC182496A29997B523C381" box="[411,625,1246,1273]" country="Australia" name="Western Australia" pageId="24" pageNumber="25">Western Australia</collectingRegion>
, 
<location id="8EB06005FFFC182495479996B433C381" LSID="urn:lsid:plazi:treatment:03C687C8FFFC1826954C9F83B224C026:8EB06005FFFC182495479996B433C381" box="[638,865,1246,1272]" country="Australia" latitude="-21.616667" longLatPrecision="21" longitude="119.02334" name="Woodstock Station" pageId="24" pageNumber="25" stateProvince="Western Australia">Woodstock Station</location>
, 
<geoCoordinate id="EE5B5019FFFC182494569995B353C380" box="[879,1025,1245,1273]" degrees="21" direction="south" minutes="37" orientation="latitude" pageId="24" pageNumber="25" precision="15" seconds="00" value="-21.616667">21°37′00″ S</geoCoordinate>
, 
<geoCoordinate id="EE5B5019FFFC182493299995B3E3C380" box="[1040,1201,1245,1273]" degrees="119" direction="east" minutes="01" orientation="longitude" pageId="24" pageNumber="25" precision="15" seconds="24" value="119.02334">119°01′24″ E</geoCoordinate>
, 
<collectingMethod id="522E4EC9FFFC182493F99996B212C380" box="[1216,1344,1246,1273]" pageId="24" pageNumber="25">pitfall trap</collectingMethod>
, site WS9, trap #3, 
<date id="FFD1101EFFFC1824965E984FB555C258" box="[359,519,1287,1314]" pageId="24" pageNumber="25" value="1988-03-27">
<collectingDate id="EF95E9F6FFFC1824965E984FB555C258" box="[359,519,1287,1314]" pageId="24" pageNumber="25" value="1988-03-27">27 Mar. 1988</collectingDate>
</date>
, 
<collectorName id="269A5308FFFC1824952A9840B58BC258" box="[531,729,1287,1314]" pageId="24" pageNumber="25">W.F. Humphreys</collectorName>
leg. (
<specimenCode id="DBC99EA5FFFC18249423984FB499C25B" box="[794,971,1287,1314]" pageId="24" pageNumber="25">WAM T55570</specimenCode>
).
</materialsCitation>
</paragraph>
<caption id="DF106656FFFC182497849AE4B4D5C093" ID-DOI="http://doi.org/10.5281/zenodo.3832441" ID-Zenodo-Dep="3832441" httpUri="https://zenodo.org/record/3832441/files/figure.png" pageId="24" pageNumber="25" startId="24.[189,232,1964,1990]" targetBox="[189,1387,1359,1919]" targetPageId="24">
<paragraph id="8BD036DEFFFC182497849AE4B4D5C093" blockId="24.[189,1398,1964,2026]" pageId="24" pageNumber="25">
<emphasis id="B91BEACCFFFC182497849AE4B659C0BF" bold="true" box="[189,267,1964,1990]" pageId="24" pageNumber="25">Fig. 8.</emphasis>
<taxonomicName id="4C6F4D5DFFFC1824962B9AE4B6A7C0BF" authority="Framenau &amp; Hudson, 2017" authorityName="Framenau &amp; Hudson" authorityYear="2017" box="[274,501,1964,1990]" class="Arachnida" family="Lycosidae" genus="Tetralycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="caudex" status="sp. nov.">
<emphasis id="B91BEACCFFFC1824962B9AE4B6A7C0BF" box="[274,501,1964,1990]" italics="true" pageId="24" pageNumber="25">Tetralycosa caudex</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22857B7FFFC182496C49AE5B50AC0BE" box="[509,600,1965,1991]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B91BEACCFFFC182495669AE4B526C0BF" bold="true" box="[607,628,1964,1990]" pageId="24" pageNumber="25">A</emphasis>
. Epigyne, dorsal view (holotype, ♀, WAM T55570). 
<emphasis id="B91BEACCFFFC182493D29AE5B3ADC0BE" bold="true" box="[1259,1279,1965,1991]" pageId="24" pageNumber="25">B</emphasis>
. Epigyne, ventral view (♀, WAM T62497). Scale bar: A–B = 0.47 mm.
</paragraph>
</caption>
<paragraph id="8BD036DEFFFD182597849C43B6ABC65C" blockId="25.[189,505,267,293]" box="[189,505,267,293]" pageId="25" pageNumber="26">
<heading id="D09881B2FFFD182597849C43B6ABC65C" bold="true" box="[189,505,267,293]" fontSize="11" level="3" pageId="25" pageNumber="26" reason="3">
<emphasis id="B91BEACCFFFD182597849C43B6ABC65C" bold="true" box="[189,505,267,293]" pageId="25" pageNumber="26">Other material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BD036DEFFFD182597849C72B430C60E" blockId="25.[189,1399,312,375]" pageId="25" pageNumber="26">
<collectingCountry id="F378764EFFFD182597849C72B634C62D" box="[189,358,314,340]" name="Australia" pageId="25" pageNumber="26">AUSTRALIA</collectingCountry>
, 
<collectingRegion id="49ABF83CFFFD182596569C72B510C62A" box="[367,578,313,340]" country="Australia" name="Western Australia" pageId="25" pageNumber="26">Western Australia</collectingRegion>
: 
<specimenCount id="9D69FD57FFFD182595729C72B5C9C62D" box="[587,667,314,340]" pageId="25" pageNumber="26" type="female">10 ♀♀</specimenCount>
, 
<specimenCount id="9D69FD57FFFD1825959F9C72B582C62D" box="[678,720,314,340]" pageId="25" pageNumber="26" type="female">1 ♀</specimenCount>
with spiderlings, 23 juvs, Woodstock Station, 
<geoCoordinate id="EE5B5019FFFD182593DA9C70B223C62D" box="[1251,1393,312,340]" degrees="21" direction="south" minutes="37" orientation="latitude" pageId="25" pageNumber="26" precision="15" seconds="00" value="-21.616667">21°37′00″ S</geoCoordinate>
, 
<geoCoordinate id="EE5B5019FFFD182597849C13B608C60E" box="[189,346,347,375]" degrees="119" direction="east" minutes="01" orientation="longitude" pageId="25" pageNumber="26" precision="15" seconds="24" value="119.02334">119°01′24″ E</geoCoordinate>
(
<collectionCode id="ED7EAE1BFFFD182596509C15B6E1C60E" box="[361,435,349,375]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="25" pageNumber="26" type="Museum">WAM</collectionCode>
T55576, T62493, T62497, T62611).
</paragraph>
</subSubSection>
<subSubSection id="C3756555FFFD182697849CD6B604C226" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26" type="description">
<paragraph id="8BD036DEFFFD182597849CD6B61CC6C1" blockId="25.[189,334,414,440]" box="[189,334,414,440]" pageId="25" pageNumber="26">
<heading id="D09881B2FFFD182597849CD6B61CC6C1" bold="true" box="[189,334,414,440]" fontSize="11" level="3" pageId="25" pageNumber="26" reason="3">
<emphasis id="B91BEACCFFFD182597849CD6B61CC6C1" bold="true" box="[189,334,414,440]" pageId="25" pageNumber="26">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BD036DEFFFD182597D39C85B678C69E" blockId="25.[189,313,461,522]" box="[234,298,461,487]" pageId="25" pageNumber="26">
<emphasis id="B91BEACCFFFD182597D39C85B678C69E" bold="true" box="[234,298,461,487]" pageId="25" pageNumber="26">Male</emphasis>
</paragraph>
<paragraph id="8BD036DEFFFD182597849CB8B66BC573" blockId="25.[189,313,461,522]" box="[189,313,496,522]" pageId="25" pageNumber="26">Unknown.</paragraph>
<paragraph id="8BD036DEFFFD182597D39F79B5BCC535" blockId="25.[189,1399,561,658]" box="[234,750,561,588]" pageId="25" pageNumber="26">
<emphasis id="B91BEACCFFFD182597D39F79B617C532" bold="true" box="[234,325,561,587]" pageId="25" pageNumber="26">Female</emphasis>
(based on 
<typeStatus id="54D4887CFFFD182596FC9F79B57DC532" box="[453,559,561,587]" pageId="25" pageNumber="26" type="holotype">holotype</typeStatus>
, WAM T55570)
</paragraph>
<paragraph id="8BD036DEFFFD182597849F1DB225C5EB" blockId="25.[189,1399,561,658]" pageId="25" pageNumber="26">
MEASUREMENTS. 
<specimenCount id="9D69FD57FFFD182596459F1DB6C0C516" box="[380,402,597,623]" pageId="25" pageNumber="26" type="female">♀</specimenCount>
<typeStatus id="54D4887CFFFD182596A29F1CB557C517" box="[411,517,596,622]" pageId="25" pageNumber="26" type="holotype">holotype</typeStatus>
, WAM T53393: TL 7.80, CL 3.83, CW 2.28. Eyes: AME 0.20, ALE 0.16, PME 0.33, PLE 0.23. Row of eyes: AE 0.80, PME 0.74, PLE 0.97. Sternum length/width: 1.73/1.43.
</paragraph>
<caption id="DF106656FFFD182597849A10B5F0C0EC" ID-DOI="http://doi.org/10.5281/zenodo.3832443" ID-Zenodo-Dep="3832443" httpUri="https://zenodo.org/record/3832443/files/figure.png" pageId="25" pageNumber="26" startId="25.[189,232,1880,1906]" targetBox="[202,1386,729,1834]" targetPageId="25">
<paragraph id="8BD036DEFFFD182597849A10B5F0C0EC" blockId="25.[189,1398,1880,1942]" pageId="25" pageNumber="26">
<emphasis id="B91BEACCFFFD182597849A10B65AC00B" bold="true" box="[189,264,1880,1906]" pageId="25" pageNumber="26">Fig. 9.</emphasis>
<taxonomicName id="4C6F4D5DFFFD182596379A10B5F5C00A" authority="(L. Koch, 1876)" baseAuthorityName="L. Koch" baseAuthorityYear="1876" box="[270,679,1880,1907]" class="Arachnida" family="Lycosidae" genus="Tetralycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="oraria">
<emphasis id="B91BEACCFFFD182596379A10B6B4C00B" box="[270,486,1880,1906]" italics="true" pageId="25" pageNumber="26">Tetralycosa oraria</emphasis>
(L. 
<bibRefCitation id="EFFE4B2FFFFD1825952A9A10B5CCC00A" author="Koch" box="[531,670,1880,1907]" firstAuthor="Koch" pageId="25" pageNumber="26" refId="ref38160" refString="Koch L. 1876. Die Arachniden Australiens nach der Natur beschrieben und abgebildet. 1. Theil. 6. Lieferung. Verlag von Bauer und Raspe, Nurnberg." type="book" year="1876">Koch, 1876</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="4C6F4D5DFFFD182595889A11B381C00A" authority="(Framenau, Gotch &amp; Austin, 1976)" baseAuthorityName="Framenau, Gotch &amp; Austin" baseAuthorityYear="1976" box="[689,1235,1880,1907]" class="Arachnida" family="Lycosidae" genus="Tetralycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="arabanae">
<emphasis id="B91BEACCFFFD182595889A11B469C00B" box="[689,827,1880,1906]" italics="true" pageId="25" pageNumber="26">T. arabanae</emphasis>
(Framenau, Gotch &amp;Austin, 1976)
</taxonomicName>
and 
<taxonomicName id="4C6F4D5DFFFD182592309A11B224C00B" authority="Framenau &amp; Hudson, 2017" authorityName="Framenau &amp; Hudson" authorityYear="2017" box="[1289,1398,1880,1906]" class="Arachnida" family="Lycosidae" genus="Tetralycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="caudex" status="sp. nov.">
<emphasis id="B91BEACCFFFD182592309A11B224C00B" box="[1289,1398,1880,1906]" italics="true" pageId="25" pageNumber="26">T. caudex</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22857B7FFFD182597849A34B648C0EF" box="[189,282,1916,1942]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
, distribution records in Australia.
</paragraph>
</caption>
<paragraph id="8BD036DEFFFE182697849C43B218C615" blockId="26.[189,1399,267,365]" pageId="26" pageNumber="27">Labium length/width: 0.57/0.53. AL 3.98, AW 3.00. Legs: Lengths of segments (femur + patella/tibia + metatarsus + tarsus = total length): Pedipalp 1.49+1.52+ – + 0.92 = 3.93, I 2.80+3.38+2.17+1.06 = 9.41, II 2.57+3.09+2.15+1.06 = 8.87, III 2.55+2.89+2.37+1.26 = 9.07, IV 3.20+4.35+3.03+1.54 = 12.12.</paragraph>
<paragraph id="8BD036DEFFFE182697849CDCB68AC6A9" blockId="26.[189,1398,403,465]" pageId="26" pageNumber="27">
VARIATION. 
<specimenCount id="9D69FD57FFFE1826967D9CDCB608C6D7" box="[324,346,404,430]" pageId="26" pageNumber="27" type="female">♀</specimenCount>
(range, mean ± SD): TL 5.85–9.00, 7.18 ± 1.07; CL 3.00–4.20, 3.56 ± 0.47; CW 2.25– 3.15, 2.70 ± 0.36; n = 8.
</paragraph>
<paragraph id="8BD036DEFFFE182697849CB0B477C505" blockId="26.[189,1399,504,636]" pageId="26" pageNumber="27">
CARAPACE (
<figureCitation id="13542A5BFFFE182696079CB0B6DCC56A" box="[318,398,504,531]" captionStart="Fig" captionStartId="20.[189,232,1859,1885]" captionTargetBox="[331,1256,273,1831]" captionTargetId="figure@20.[325,1262,265,1837]" captionTargetPageId="20" captionText="Fig. 5. Tetralycosa Roewer, 1960, microscopic photographs showing dorsal and ventral views. A–B. T. oraria (L. Koch, 1876), ♂ (WAM T93541). C–D. T. oraria (L. Koch, 1876), ♀ (WAM T93944). E–F. T. arabanae (Framenau, Gotch &amp; Austin, 2006), ♂ (WAM T47298). G–H. T. arabanae (Framenau, Gotch &amp; Austin, 2006), ♀ (WAM T47298). I–J. T. caudex sp. nov., ♀ (WAM T55576). Scale bars: A–J = 2.00 mm." figureDoi="http://doi.org/10.5281/zenodo.3832433" httpUri="https://zenodo.org/record/3832433/files/figure.png" pageId="26" pageNumber="27">Fig. 5I</figureCitation>
). Dorsal line straight in lateral view; yellow-brown with lighter radial pattern; cephalic area slightly darker; indistinct grey pigmentation anteriorly of fovea, few white setae along carapace margins and laterally of PE; brown macrosetae in eye region and between eye region and fovea; six long bristles below AME, one long bristle between AME.
</paragraph>
<paragraph id="8BD036DEFFFE182697849FEBB4A7C5C4" blockId="26.[189,1013,675,701]" box="[189,1013,675,701]" pageId="26" pageNumber="27">EYES. Row of AE slightly wider than row of PME; row of AE straight.</paragraph>
<paragraph id="8BD036DEFFFE182697849FACB40EC458" blockId="26.[189,1398,740,801]" pageId="26" pageNumber="27">CHELICERAE. Brown; covered with brown setae and macrosetae. Two promarginal teeth, with the apical largest, three retromarginal teeth, with the basal smallest.</paragraph>
<paragraph id="8BD036DEFFFE182697849E01B496C41A" blockId="26.[189,964,841,868]" box="[189,964,841,868]" pageId="26" pageNumber="27">
STERNUM (
<figureCitation id="13542A5BFFFE1826960C9E01B6DAC41D" box="[309,392,841,868]" captionStart="Fig" captionStartId="20.[189,232,1859,1885]" captionTargetBox="[331,1256,273,1831]" captionTargetId="figure@20.[325,1262,265,1837]" captionTargetPageId="20" captionText="Fig. 5. Tetralycosa Roewer, 1960, microscopic photographs showing dorsal and ventral views. A–B. T. oraria (L. Koch, 1876), ♂ (WAM T93541). C–D. T. oraria (L. Koch, 1876), ♀ (WAM T93944). E–F. T. arabanae (Framenau, Gotch &amp; Austin, 2006), ♂ (WAM T47298). G–H. T. arabanae (Framenau, Gotch &amp; Austin, 2006), ♀ (WAM T47298). I–J. T. caudex sp. nov., ♀ (WAM T55576). Scale bars: A–J = 2.00 mm." figureDoi="http://doi.org/10.5281/zenodo.3832433" httpUri="https://zenodo.org/record/3832433/files/figure.png" pageId="26" pageNumber="27">Fig. 5J</figureCitation>
). Yellow-brown; covered with long brown setae.
</paragraph>
<paragraph id="8BD036DEFFFE182697849EC3B497C4DD" blockId="26.[189,965,906,933]" box="[189,965,906,933]" pageId="26" pageNumber="27">LABIUM. Dark brown, basally darker; front end truncate and white.</paragraph>
<paragraph id="8BD036DEFFFE182697849E84B5A0C354" blockId="26.[189,1399,972,1069]" pageId="26" pageNumber="27">
ABDOMEN. Dorsally olive-brown with pairs of irregular lighter patches; indistinct brown lanceolate heart mark; mainly brown and fewer white setae, few brown macrosetae (
<figureCitation id="13542A5BFFFE182694E29EA7B37CC373" box="[987,1070,1007,1034]" captionStart="Fig" captionStartId="20.[189,232,1859,1885]" captionTargetBox="[331,1256,273,1831]" captionTargetId="figure@20.[325,1262,265,1837]" captionTargetPageId="20" captionText="Fig. 5. Tetralycosa Roewer, 1960, microscopic photographs showing dorsal and ventral views. A–B. T. oraria (L. Koch, 1876), ♂ (WAM T93541). C–D. T. oraria (L. Koch, 1876), ♀ (WAM T93944). E–F. T. arabanae (Framenau, Gotch &amp; Austin, 2006), ♂ (WAM T47298). G–H. T. arabanae (Framenau, Gotch &amp; Austin, 2006), ♀ (WAM T47298). I–J. T. caudex sp. nov., ♀ (WAM T55576). Scale bars: A–J = 2.00 mm." figureDoi="http://doi.org/10.5281/zenodo.3832433" httpUri="https://zenodo.org/record/3832433/files/figure.png" pageId="26" pageNumber="27">Fig. 5I</figureCitation>
). Venter yellow-brown with brown setae. Spinnerets yellow-brown (
<figureCitation id="13542A5BFFFE182695A9995AB5B7C354" box="[656,741,1042,1069]" captionStart="Fig" captionStartId="20.[189,232,1859,1885]" captionTargetBox="[331,1256,273,1831]" captionTargetId="figure@20.[325,1262,265,1837]" captionTargetPageId="20" captionText="Fig. 5. Tetralycosa Roewer, 1960, microscopic photographs showing dorsal and ventral views. A–B. T. oraria (L. Koch, 1876), ♂ (WAM T93541). C–D. T. oraria (L. Koch, 1876), ♀ (WAM T93944). E–F. T. arabanae (Framenau, Gotch &amp; Austin, 2006), ♂ (WAM T47298). G–H. T. arabanae (Framenau, Gotch &amp; Austin, 2006), ♀ (WAM T47298). I–J. T. caudex sp. nov., ♀ (WAM T55576). Scale bars: A–J = 2.00 mm." figureDoi="http://doi.org/10.5281/zenodo.3832433" httpUri="https://zenodo.org/record/3832433/files/figure.png" pageId="26" pageNumber="27">Fig. 5J</figureCitation>
).
</paragraph>
<paragraph id="8BD036DEFFFE18269784991CB353C3CD" blockId="26.[189,1399,1108,1205]" pageId="26" pageNumber="27">
EPIGYNE. Ventral view (
<figureCitation id="13542A5BFFFE182696D9991CB517C317" box="[480,581,1108,1134]" captionStart="Fig" captionStartId="24.[189,232,1964,1990]" captionTargetBox="[189,1387,1359,1919]" captionTargetId="figure@24.[189,1399,1359,1931]" captionTargetPageId="24" captionText="Fig. 8. Tetralycosa caudex sp. nov. A. Epigyne, dorsal view (holotype, ♀, WAM T55570). B. Epigyne, ventral view (♀, WAM T62497). Scale bar: A–B = 0.47 mm." figureDoi="http://doi.org/10.5281/zenodo.3832441" httpUri="https://zenodo.org/record/3832441/files/figure.png" pageId="26" pageNumber="27">Fig. 8A</figureCitation>
): median septum inverted T-shaped with rounded anterior curves; anterior margin continuous and nearly straight. Dorsal view (
<figureCitation id="13542A5BFFFE18269489993FB340C3E8" box="[944,1042,1143,1169]" captionStart="Fig" captionStartId="24.[189,232,1964,1990]" captionTargetBox="[189,1387,1359,1919]" captionTargetId="figure@24.[189,1399,1359,1931]" captionTargetPageId="24" captionText="Fig. 8. Tetralycosa caudex sp. nov. A. Epigyne, dorsal view (holotype, ♀, WAM T55570). B. Epigyne, ventral view (♀, WAM T62497). Scale bar: A–B = 0.47 mm." figureDoi="http://doi.org/10.5281/zenodo.3832441" httpUri="https://zenodo.org/record/3832441/files/figure.png" pageId="26" pageNumber="27">Fig. 8B</figureCitation>
): large bulbous spermathecal heads; spermathecal stalks slim and attach posteriorly to spermathecae.
</paragraph>
<paragraph id="8BD036DEFFFE182697849994B604C226" blockId="26.[189,1398,1243,1375]" pageId="26" pageNumber="27">LEGS. Leg formula IV&gt;I&gt;III&gt;II. Light brown, some grey pigmentation; basal two thirds of femora yellow-brown. Pedipalp with very long claw. Spination of leg I: femur: three dorsal, one apicoprolateral, two retrolateral; tibia: two ventral pairs, two prolateral; metatarsus: two ventral pairs, one prolateral, one apicoventral.</paragraph>
</subSubSection>
<subSubSection id="C3756555FFFE1826978498CFB598C125" pageId="26" pageNumber="27" type="discussion">
<paragraph id="8BD036DEFFFE1826978498CFB67DC2D8" blockId="26.[189,303,1415,1441]" box="[189,303,1415,1441]" pageId="26" pageNumber="27">
<heading id="D09881B2FFFE1826978498CFB67DC2D8" bold="true" box="[189,303,1415,1441]" fontSize="11" level="3" pageId="26" pageNumber="27" reason="3">
<emphasis id="B91BEACCFFFE1826978498CFB67DC2D8" bold="true" box="[189,303,1415,1441]" pageId="26" pageNumber="27">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BD036DEFFFE1826978498FDB598C125" blockId="26.[189,1399,1461,1628]" pageId="26" pageNumber="27">
Although the male of 
<taxonomicName id="4C6F4D5DFFFE1826968798FEB57FC2B6" authority="Framenau &amp; Hudson, 2017" authorityName="Framenau &amp; Hudson" authorityYear="2017" box="[446,557,1461,1487]" class="Arachnida" family="Lycosidae" genus="Tetralycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="caudex" status="sp. nov.">
<emphasis id="B91BEACCFFFE1826968798FEB57FC2B6" box="[446,557,1461,1487]" italics="true" pageId="26" pageNumber="27">T. caudex</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22857B7FFFE1826950D98FEB5DDC2A9" box="[564,655,1462,1488]" pageId="26" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
is unknown, female somatic and genital morphology match the diagnosis of the genus and our phylogenetic analysis places it here. Some characters are unusual within the 
<taxonomicName id="4C6F4D5DFFFE182697DE98B3B63DC16C" authorityName="Roewer" authorityYear="1960" box="[231,367,1531,1557]" class="Arachnida" family="Lycosidae" genus="Tetralycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEACCFFFE182697DE98B3B63DC16C" box="[231,367,1531,1557]" italics="true" pageId="26" pageNumber="27">Tetralycosa</emphasis>
</taxonomicName>
; for example, 
<taxonomicName id="4C6F4D5DFFFE1826952E98B4B5D7C16C" authority="Framenau &amp; Hudson, 2017" authorityName="Framenau &amp; Hudson" authorityYear="2017" box="[535,645,1531,1557]" class="Arachnida" family="Lycosidae" genus="Tetralycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="caudex" status="sp. nov.">
<emphasis id="B91BEACCFFFE1826952E98B4B5D7C16C" box="[535,645,1531,1557]" italics="true" pageId="26" pageNumber="27">T. caudex</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22857B7FFFE182695B598B4B5B7C16F" box="[652,741,1532,1558]" pageId="26" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
is the only species within the genus in which leg II is the shortest. In addition, the 
<typeStatus id="54D4887CFFFE182696E49B57B542C140" box="[477,528,1567,1593]" pageId="26" pageNumber="27">type</typeStatus>
and only locality of this species in northern 
<collectingRegion id="49ABF83CFFFE1826932B9B57B3B1C140" box="[1042,1251,1567,1593]" country="Australia" name="Western Australia" pageId="26" pageNumber="27">Western Australia</collectingRegion>
falls outside the general distribution of the genus (
<figureCitation id="13542A5BFFFE1826954D9B0AB5EEC125" box="[628,700,1602,1628]" captionStart="Fig" captionStartId="25.[189,232,1880,1906]" captionTargetBox="[202,1386,729,1834]" captionTargetId="figure@25.[189,1398,717,1846]" captionTargetPageId="25" captionText="Fig. 9. Tetralycosa oraria (L. Koch, 1876), T. arabanae (Framenau, Gotch &amp;Austin, 1976) and T. caudex sp. nov., distribution records in Australia." figureDoi="http://doi.org/10.5281/zenodo.3832443" httpUri="https://zenodo.org/record/3832443/files/figure.png" pageId="26" pageNumber="27">Fig. 9</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3756555FFFE182697849BCBB377C196" pageId="26" pageNumber="27" type="biology_ecology">
<paragraph id="8BD036DEFFFE182697849BCBB52CC1E4" blockId="26.[189,638,1667,1693]" box="[189,638,1667,1693]" pageId="26" pageNumber="27">
<heading id="D09881B2FFFE182697849BCBB52CC1E4" bold="true" box="[189,638,1667,1693]" fontSize="11" level="3" pageId="26" pageNumber="27" reason="3">
<emphasis id="B91BEACCFFFE182697849BCBB52CC1E4" bold="true" box="[189,638,1667,1693]" pageId="26" pageNumber="27">Life history and habitat preferences</emphasis>
</heading>
</paragraph>
<paragraph id="8BD036DEFFFE182697849BFAB377C196" blockId="26.[189,1399,1714,1776]" pageId="26" pageNumber="27">
Only a limited number of female 
<taxonomicName id="4C6F4D5DFFFE1826957A9BFBB5E1C1B5" authority="Framenau &amp; Hudson, 2017" authorityName="Framenau &amp; Hudson" authorityYear="2017" box="[579,691,1714,1740]" class="Arachnida" family="Lycosidae" genus="Tetralycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="caudex" status="sp. nov.">
<emphasis id="B91BEACCFFFE1826957A9BFBB5E1C1B5" box="[579,691,1714,1740]" italics="true" pageId="26" pageNumber="27">T. caudex</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22857B7FFFE182695839BFAB446C1B5" box="[698,788,1714,1740]" pageId="26" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
have been found, some of them in granite with sand near a creek. Most records are from May, one female was found in March.
</paragraph>
</subSubSection>
<subSubSection id="C3756555FFFE182697849A5EB224C026" box="[189,1398,1814,1888]" pageId="26" pageNumber="27" type="distribution">
<paragraph id="8BD036DEFFFE182697849A5EB604C049" blockId="26.[189,342,1814,1840]" box="[189,342,1814,1840]" pageId="26" pageNumber="27">
<heading id="D09881B2FFFE182697849A5EB604C049" bold="true" box="[189,342,1814,1840]" fontSize="11" level="3" pageId="26" pageNumber="27" reason="3">
<emphasis id="B91BEACCFFFE182697849A5EB604C049" bold="true" box="[189,342,1814,1840]" pageId="26" pageNumber="27">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BD036DEFFFE182697849A0DB224C026" blockId="26.[189,1398,1861,1888]" box="[189,1398,1861,1888]" pageId="26" pageNumber="27">
Only known from the 
<typeStatus id="54D4887CFFFE182696829A0EB6BCC019" box="[443,494,1862,1888]" pageId="26" pageNumber="27">type</typeStatus>
locality, Woodstock Station in the Pilbara region of 
<collectingRegion id="49ABF83CFFFE1826937D9A0EB246C026" box="[1092,1300,1861,1888]" country="Australia" name="Western Australia" pageId="26" pageNumber="27">Western Australia</collectingRegion>
(
<figureCitation id="13542A5BFFFE182692189A0DB23AC026" box="[1313,1384,1861,1887]" captionStart="Fig" captionStartId="25.[189,232,1880,1906]" captionTargetBox="[202,1386,729,1834]" captionTargetId="figure@25.[189,1398,717,1846]" captionTargetPageId="25" captionText="Fig. 9. Tetralycosa oraria (L. Koch, 1876), T. arabanae (Framenau, Gotch &amp;Austin, 1976) and T. caudex sp. nov., distribution records in Australia." figureDoi="http://doi.org/10.5281/zenodo.3832443" httpUri="https://zenodo.org/record/3832443/files/figure.png" pageId="26" pageNumber="27">Fig. 9</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>