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<document id="AB7CD798EBA46146F380495AB5CE9FE9" ID-DOI="10.5252/zoosystema2024v46a5" ID-ISSN="1638-9387" ID-Zenodo-Dep="10803267" ID-ZooBank="1B0779F3-FAE4-4105-8130-A26DE519721E" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1710145347001" checkinUser="plazi" docAuthor="Cardoso, Giovanna Monticelli, M, Diego de, Bento, edeiros &amp; Ferreira, Rodrigo Lopes" docDate="2024" docId="03C687BFC613FFD1101AF54D6E9BF856" docLanguage="en" docName="zoosystema.2024.46.5.pdf" docOrigin="Zoosystema 46 (5)" docSource="https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a5.pdf" docStyle="DocumentStyle:5CC576F744449C1860B4E72ABD884631.7:Zoosystema.2020-.journal_article" docStyleId="5CC576F744449C1860B4E72ABD884631" docStyleName="Zoosystema.2020-.journal_article" docStyleVersion="7" docTitle="Ctenorillo cearensis Cardoso &amp; Ferreira 2024, n. sp." docType="treatment" docUuid="29FFBB09-6896-4C9B-97BE-E50E44C7C180" docUuidSource="ZooBank" docVersion="2" lastPageNumber="116" masterDocId="FFFFFFC7C606FFC6136FF25C6C6BFFBA" masterDocTitle="Unveiling a hidden diversity: descriptions of nine new species of Ctenorillo Verhoeff, 1942 (Isopoda, Armadillidae) discovered in Brazilian caves and their importance for conservation" masterLastPageNumber="132" masterPageNumber="95" pageNumber="114" updateTime="1710145921992" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="C7373108A58627B8BE53F9F73AA5BAD7" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="AA51A8362DFE8C3C3551899F4E96C554">
<mods:title id="87B090678DF96CD10BA9B3C4E67F1D3C">Unveiling a hidden diversity: descriptions of nine new species of Ctenorillo Verhoeff, 1942 (Isopoda, Armadillidae) discovered in Brazilian caves and their importance for conservation</mods:title>
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<mods:name id="DB935AB739045AFD072AC63C62D215D1" type="personal">
<mods:role id="A4466B8268E7BCC6321ACA52B751FCD4">
<mods:roleTerm id="B9AE303EAA81B470F50503EE39523915">Author</mods:roleTerm>
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<mods:namePart id="AE703661F97CD2342D61B70A79374E45">Cardoso, Giovanna Monticelli</mods:namePart>
<mods:affiliation id="7BB5BD2228AA0E4A502530F1B0138109">Universidade Federal de Lavras (UFLA), Centro de Estudos em Biologia Subterrânea, Departamento de Biologia, programa de Pós-graduação em Ecologia Aplicada, Campus Universitário, CP 3037, 37200 - 000 Lavras, Minas Gerais (Brasil) gmcardoso. bio @ gmail. com (corresponding author)</mods:affiliation>
<mods:nameIdentifier id="96D8DD417D212CFDA9C487FBB6AF6DA5" type="email">bio@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="F669EB22525979ADD71661D12F5122C5" type="personal">
<mods:role id="FA6028D555F9B8F2A52955568810392B">
<mods:roleTerm id="217136F26A044D35A0102AD4BF95F04A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F3C8F4548499CC84BB0D5202627C8F25">M, Diego de</mods:namePart>
</mods:name>
<mods:name id="91B4A617F47DB17AA6C662F90BD427E7" type="personal">
<mods:role id="E2F8526B4A0E64609C5E6442CF5B6417">
<mods:roleTerm id="79D2321EC474950B9E90DD8058832E56">Author</mods:roleTerm>
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<mods:namePart id="DAEDFDF38B1AC0FE743B736A3556E610">Bento, edeiros</mods:namePart>
</mods:name>
<mods:name id="01C6A61F2BBFCCD8E9F8E713B8BD488C" type="personal">
<mods:role id="A74AB386CDC2CB263A98D7B61BDB6239">
<mods:roleTerm id="06A62B984B62660ED8B793810A63AA3D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9A9250E0EA1BE46E35C199229224E18D">Ferreira, Rodrigo Lopes</mods:namePart>
<mods:affiliation id="939303A0BB616E708D06290082426E18">Universidade Federal de Lavras (UFLA), Centro de Estudos em Biologia Subterrânea, Departamento de Biologia, programa de Pós-graduação em Ecologia Aplicada, Campus Universitário, CP 3037, 37200 - 000 Lavras, Minas Gerais (Brasil) drops @ ufla. br</mods:affiliation>
<mods:nameIdentifier id="9600D71CC75708A22BCA6A66A2BDB1D1" type="email">drops@ufla.br</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="A4BD2964034EB81929633617D1454CAB">text</mods:typeOfResource>
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<mods:title id="7E61BC94791BCB56DF680B4B3A84673A">Zoosystema</mods:title>
</mods:titleInfo>
<mods:part id="5B2C45B477DF0C4CFAA32FECBAF9BA30">
<mods:date id="F9034E6B6CEFA2B7D73DF5BA727BDF2A">2024</mods:date>
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<mods:number id="CA54B69A403C404E00869019F50124F0">2024-03-05</mods:number>
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<mods:detail id="F38ECEFFC2FA9BB7E01634CD0D3E828A" type="volume">
<mods:number id="5A5A395CBD12C7C557A49BE466602D87">46</mods:number>
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<mods:number id="C160BF012498FB90A6F92CBD9A42E0A1">5</mods:number>
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<mods:start id="BF4F35F4FDF9A10BA386B3E4DAD7E307">95</mods:start>
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<mods:url id="CDCA3B674CF5CB1B13328DFC864E9159">https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a5.pdf</mods:url>
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<subSubSection id="C3756522C613FFD3101AF54D690CF896" box="[885,1383,1809,1836]" pageId="21" pageNumber="114" type="nomenclature">
<paragraph id="8BD036A9C613FFD3101AF54D690CF896" blockId="21.[885,1383,1809,1868]" box="[885,1383,1809,1836]" pageId="21" pageNumber="114">
<heading id="D09881C5C613FFD3101AF54D690CF896" box="[885,1383,1809,1836]" centered="true" fontSize="11" level="2" pageId="21" pageNumber="114" reason="2">
<taxonomicName id="4C6F4D2AC613FFD3101AF54D6949F896" authority="Cardoso &amp; Ferreira" authorityName="Cardoso &amp; Ferreira" authorityYear="2024" box="[885,1314,1809,1836]" class="Malacostraca" family="Armadillidae" genus="Ctenorillo" kingdom="Animalia" order="Isopoda" pageId="21" pageNumber="114" phylum="Arthropoda" rank="species" species="cearensis" status="sp. nov.">
<emphasis id="B91BEABBC613FFD3101AF54D682DF896" bold="true" box="[885,1094,1809,1836]" italics="true" pageId="21" pageNumber="114">Ctenorillo cearensis</emphasis>
Cardoso &amp; Ferreira
</taxonomicName>
,
<taxonomicNameLabel id="A22857C0C613FFD31643F54E690CF896" box="[1324,1383,1810,1836]" pageId="21" pageNumber="114" rank="species">n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3756522C613FFD31763F56D69FEF8C5" pageId="21" pageNumber="114" type="description">
<paragraph id="8BD036A9C613FFD31763F56D68A4F8F6" blockId="21.[885,1383,1809,1868]" box="[1036,1231,1841,1868]" pageId="21" pageNumber="114">
(
<figureCitation id="13542A2CC613FFD3177CF56D680EF8F6" box="[1043,1125,1841,1868]" captionStart="FIG" captionStartId="21.[132,143,1668,1686]" captionTargetBox="[132,1455,215,1619]" captionTargetId="figure-128@21.[132,1455,215,1619]" captionTargetPageId="21" captionText="FIG. 14. — Ctenorillo cearensis Cardoso &amp; Ferreira,n. sp.,holotype (male,4 mm,ISLA96021):A, habitus,lateral view;B, disposition of dorsal tubercles;C, cephalon, frontal view; D, cephalon, dorsal view; E, epimera 1-3 ventral view; F, pleonites 3-5, uropods and telson, dorsal view; G, antennula; H, antenna; I, right mandible; J, left mandible; K, maxillula; L, maxilla; M, maxilliped. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10803297" httpUri="https://zenodo.org/record/10803297/files/figure.png" pageId="21" pageNumber="114">Figs 14</figureCitation>
;
<figureCitation id="13542A2CC613FFD3171EF56D68E4F8F6" box="[1137,1167,1841,1868]" captionStart="FIG" captionStartId="22.[132,143,1289,1307]" captionTargetBox="[134,1395,216,1241]" captionTargetId="figure-425@22.[132,1455,216,1247]" captionTargetPageId="22" captionText="FIG. 15. — Ctenorillo cearensis Cardoso &amp; Ferreira,n. sp., paratype (male, 4 mm, ISLA96021):A, uropod,B, pereopod 1; C, pereopod 7; D, pleopod 1 and genital papilla; E, pleopod 2; F, pleopod 3 exopod; G, pleopod 4 exopod; H, pleopod 5 exopod." figureDoi="http://doi.org/10.5281/zenodo.10803299" httpUri="https://zenodo.org/record/10803299/files/figure.png" pageId="21" pageNumber="114">15</figureCitation>
;
<figureCitation id="13542A2CC613FFD317F4F56E68ACF8F6" box="[1179,1223,1842,1868]" captionStart="FIG" captionStartId="35.[132,143,1225,1243]" captionTargetBox="[132,1455,216,1183]" captionTargetId="figure-480@35.[699,983,655,744]" captionTargetPageId="35" captionText="FIG. 27. — Habitus of the species of the genus Ctenorillo Verhoeff, 1942 discussed in the present paper:A, Ctenorillo ferrarai Campos-Filho, Araujo &amp; Taiti,2014; B, Ctenorillo pelado Cardoso &amp; Ferreira, n. sp.; C, Ctenorillo araguaia Cardoso &amp; Ferreira, n. sp.; D, Ctenorillo intertidalis Cardoso &amp; Ferreira, n. sp.; E, Ctenorillo ubajarensis Cardoso &amp; Ferreira,n. sp.; F, Ctenorillo cearensis Cardoso &amp; Ferreira,n. sp.; G, Ctenorillo quiteriensis Cardoso &amp; Ferreira,n. sp.; H, Ctenorillo potiguar Cardoso &amp; Ferreira, n. sp.; I, Ctenorillo iuiuensis n. sp.; J, Ctenorillo jequitinhonha Cardoso &amp; Ferreira, n. sp. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10803325" httpUri="https://zenodo.org/record/10803325/files/figure.png" pageId="21" pageNumber="114">27F</figureCitation>
)
</paragraph>
<paragraph id="8BD036A9C613FFD31028F53169FEF8C5" blockId="21.[839,1429,1901,1919]" box="[839,1429,1901,1919]" pageId="21" pageNumber="114">
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urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
</subSubSection>
<subSubSection id="C3756522C613FFD01042F5C26D69F9F2" lastPageId="22" lastPageNumber="115" pageId="21" pageNumber="114" type="materials_examined">
<paragraph id="8BD036A9C613FFD31042F5C2692AF851" blockId="21.[813,1455,1950,2027]" pageId="21" pageNumber="114">
<materialsCitation id="3B073CF4C613FFD31042F5C2692AF851" collectingDate="2021-03-10" collectingDateMax="2021-03-18" collectingDateMin="2021-03-10" collectionCode="TYPE, MATERIAL" collectorName="Brazil &amp; Ceara &amp; Santa Quiteria &amp; Carste" country="Iceland" latitude="-4.5647225" location="ISLA" longLatPrecision="21" longitude="-39.776665" pageId="21" pageNumber="114" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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. —
<emphasis id="B91BEABBC613FFD31081F5C268FFF80D" bold="true" box="[1006,1172,1950,1975]" pageId="21" pageNumber="114">
<typeStatus id="54D4880BC613FFD31081F5C26824F80C" box="[1006,1103,1950,1974]" pageId="21" pageNumber="114" type="holotype">Holotype</typeStatus>
.
<collectorName id="269A537FC613FFD31735F5C368FFF80D" box="[1114,1172,1951,1975]" pageId="21" pageNumber="114">Brazil</collectorName>
</emphasis>
•
<specimenCount id="9D69FD20C613FFD317C4F5C268A5F80F" box="[1195,1230,1950,1974]" count="1" pageId="21" pageNumber="114" type="male">1♂</specimenCount>
(
<quantity id="4C979B4CC613FFD317B3F5C26971F80F" box="[1244,1306,1950,1974]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="21" pageNumber="114" unit="mm" value="4.0">4 mm</quantity>
; parts in slide);
<collectorName id="269A537FC613FFD31042F5E46F0DF86A" box="[813,870,1976,2000]" pageId="21" pageNumber="114">Ceará</collectorName>
state;
<collectorName id="269A537FC613FFD310C8F5E4685DF86A" box="[935,1078,1976,2000]" pageId="21" pageNumber="114">Santa Quitéria</collectorName>
municipality; W_04 cave;
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,
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;
<collectorName id="269A537FC613FFD310DBF58F6F99F851" box="[948,1010,2003,2027]" pageId="21" pageNumber="114">Carste</collectorName>
leg.;
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</date>
;
<collectingCountry id="F3787639C613FFD317A7F58F696BF851" box="[1224,1280,2003,2027]" name="Iceland" pageId="21" pageNumber="114">ISLA</collectingCountry>
96021.
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</paragraph>
<caption id="DF106621C610FFD013EBF7556F0CFA88" ID-DOI="http://doi.org/10.5281/zenodo.10803299" ID-Zenodo-Dep="10803299" httpUri="https://zenodo.org/record/10803299/files/figure.png" pageId="22" pageNumber="115" startId="22.[132,143,1289,1307]" targetBox="[134,1395,216,1241]" targetPageId="22" targetType="figure">
<paragraph id="8BD036A9C610FFD013EBF7556F0CFA88" blockId="22.[132,1455,1289,1330]" pageId="22" pageNumber="115">
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. 15. —
<taxonomicName id="4C6F4D2AC610FFD013B2F7556E49FAA1" authority="Cardoso &amp; Ferreira" authorityName="Cardoso &amp; Ferreira" authorityYear="2024" box="[221,546,1289,1307]" class="Malacostraca" family="Armadillidae" genus="Ctenorillo" kingdom="Animalia" order="Isopoda" pageId="22" pageNumber="115" phylum="Arthropoda" rank="species" species="cearensis" status="sp. nov.">
<emphasis id="B91BEABBC610FFD013B2F7556DEBFAA1" box="[221,384,1289,1307]" italics="true" pageId="22" pageNumber="115">Ctenorillo cearensis</emphasis>
Cardoso &amp; Ferreira
</taxonomicName>
,
<taxonomicNameLabel id="A22857C0C610FFD01146F7556E30FAA1" box="[553,603,1289,1307]" pageId="22" pageNumber="115" rank="species">n. sp.</taxonomicNameLabel>
, paratype (male, 4 mm, ISLA96021):
<emphasis id="B91BEABBC610FFD010E8F7556FFFFAA1" bold="true" box="[903,916,1289,1307]" pageId="22" pageNumber="115">A</emphasis>
, uropod,
<emphasis id="B91BEABBC610FFD0108CF7556F9BFAA1" bold="true" box="[995,1008,1289,1307]" pageId="22" pageNumber="115">B</emphasis>
, pereopod 1;
<emphasis id="B91BEABBC610FFD0170CF755681AFAA1" bold="true" box="[1123,1137,1289,1307]" pageId="22" pageNumber="115">C</emphasis>
, pereopod 7;
<emphasis id="B91BEABBC610FFD0178BF7556899FAA1" bold="true" box="[1252,1266,1289,1307]" pageId="22" pageNumber="115">D</emphasis>
, pleopod 1 and genital papilla;
<emphasis id="B91BEABBC610FFD013AAF77C6CBAFA88" bold="true" box="[197,209,1312,1330]" pageId="22" pageNumber="115">E</emphasis>
, pleopod 2;
<emphasis id="B91BEABBC610FFD01255F77C6D2EFA88" bold="true" box="[314,325,1312,1330]" pageId="22" pageNumber="115">F</emphasis>
, pleopod 3 exopod;
<emphasis id="B91BEABBC610FFD0129CF77C6E6AFA88" bold="true" box="[499,513,1312,1330]" pageId="22" pageNumber="115">G</emphasis>
, pleopod 4 exopod;
<emphasis id="B91BEABBC610FFD011C0F77C6ED6FA88" bold="true" box="[687,701,1312,1330]" pageId="22" pageNumber="115">H</emphasis>
, pleopod 5 exopod.
</paragraph>
</caption>
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. —
<emphasis id="B91BEABBC610FFD0121DF7256DDAFA2B" bold="true" box="[370,433,1401,1425]" pageId="22" pageNumber="115">
<collectingCountry id="F3787639C610FFD0121DF7256DDAFA2B" box="[370,433,1401,1425]" name="Brazil" pageId="22" pageNumber="115">Brazil</collectingCountry>
</emphasis>
•
<specimenCount id="9D69FD20C610FFD012A2F72B6D9FFA34" box="[461,500,1399,1423]" count="1" pageId="22" pageNumber="115" type="male">1 ♂</specimenCount>
; SQ_26 cave;
<geoCoordinate id="EE5B506EC610FFD011FAF72B6F69FA35" box="[661,770,1399,1423]" degrees="4" direction="south" minutes="33" orientation="latitude" pageId="22" pageNumber="115" precision="15" seconds="38" value="-4.560556">4°33’38”S</geoCoordinate>
,
<geoCoordinate id="EE5B506EC610FFD013EBF7CE6D63FA10" box="[132,264,1426,1450]" degrees="39" direction="west" minutes="46" orientation="longitude" pageId="22" pageNumber="115" precision="15" seconds="51" value="-39.780834">39°46’51”W</geoCoordinate>
;
<collectorName id="269A537FC610FFD01278F7CE6D31FA10" box="[279,346,1426,1450]" pageId="22" pageNumber="115">Carste</collectorName>
leg.;
<date id="FFD11069C610FFD012F8F7CE6E53FA10" box="[407,568,1426,1450]" pageId="22" pageNumber="115" value="2021-03-10" valueMax="2021-03-18" valueMin="2021-03-10">
<collectingDate id="EF95E981C610FFD012F8F7CE6E53FA10" box="[407,568,1426,1450]" pageId="22" pageNumber="115" value="2021-03-10" valueMax="2021-03-18" valueMin="2021-03-10">10-18.III.2021</collectingDate>
</date>
; ISLA96020
</materialsCitation>
•
<specimenCount id="9D69FD20C610FFD011B3F7CE6F6BFA13" box="[732,768,1426,1450]" count="1" pageId="22" pageNumber="115" type="female">1 ♀</specimenCount>
; W_16 cave;
<geoCoordinate id="EE5B506EC610FFD01391F7F06D0DFA7E" box="[254,358,1452,1476]" degrees="4" direction="south" minutes="33" orientation="latitude" pageId="22" pageNumber="115" precision="15" seconds="43" value="-4.5619445">4°33’43”S</geoCoordinate>
,
<geoCoordinate id="EE5B506EC610FFD01200F7F06D87FA7E" box="[367,492,1452,1476]" degrees="39" direction="west" minutes="46" orientation="longitude" pageId="22" pageNumber="115" precision="15" seconds="44" value="-39.77889">39°46’44”W</geoCoordinate>
; Carste leg.;
<date id="FFD11069C610FFD01106F7F06F69FA7E" box="[617,770,1452,1476]" pageId="22" pageNumber="115" value="2021-03-10" valueMax="2021-03-18" valueMin="2021-03-10">10-18.III.2021</date>
; ISLA96026 •
<specimenCount id="9D69FD20C610FFD0127FF79B6D5FFA64" box="[272,308,1479,1503]" count="1" pageId="22" pageNumber="115" type="female">1 ♀</specimenCount>
; SQ_03 cave;
<geoCoordinate id="EE5B506EC610FFD012A7F79A6E5AFA64" box="[456,561,1478,1502]" degrees="4" direction="south" minutes="33" orientation="latitude" pageId="22" pageNumber="115" precision="15" seconds="53" value="-4.5647225">4°33’53”S</geoCoordinate>
,
<geoCoordinate id="EE5B506EC610FFD01154F79B6ED1FA65" box="[571,698,1479,1503]" degrees="39" direction="west" minutes="46" orientation="longitude" pageId="22" pageNumber="115" precision="15" seconds="40" value="-39.77778">39°46’40”W</geoCoordinate>
; Carste leg.;
<date id="FFD11069C610FFD013DCF7BD6D25FA43" box="[179,334,1505,1529]" pageId="22" pageNumber="115" value="2021-03-10" valueMax="2021-03-18" valueMin="2021-03-10">10-18.III.2021</date>
; ISLA96033 •
<specimenCount id="9D69FD20C610FFD0128CF7BD6E63FA42" box="[483,520,1505,1529]" count="2" pageId="22" pageNumber="115" type="male">2 ♂</specimenCount>
; SQ_P2 cave;
<geoCoordinate id="EE5B506EC610FFD011F5F7BD6F68FA43" box="[666,771,1505,1529]" degrees="4" direction="south" minutes="33" orientation="latitude" pageId="22" pageNumber="115" precision="15" seconds="37" value="-4.560278">4°33’37”S</geoCoordinate>
,
<geoCoordinate id="EE5B506EC610FFD013EBF7A76D6CF9A9" box="[132,263,1531,1555]" degrees="39" direction="west" minutes="45" orientation="longitude" pageId="22" pageNumber="115" precision="15" seconds="57" value="-39.765835">39°45’57”W</geoCoordinate>
; Carste leg.;
<date id="FFD11069C610FFD012F8F7A06E53F9AE" box="[407,568,1532,1556]" pageId="22" pageNumber="115" value="2021-03-10" valueMax="2021-03-18" valueMin="2021-03-10">10-18.III.2021</date>
; ISLA96962 •
<specimenCount id="9D69FD20C610FFD011B3F7A06F6BF9A9" box="[732,768,1532,1556]" count="3" pageId="22" pageNumber="115" type="female">3 ♀</specimenCount>
; SQ_03 cave;
<geoCoordinate id="EE5B506EC610FFD01264F44A6D03F994" box="[267,360,1558,1582]" degrees="4" direction="south" minutes="34" orientation="latitude" pageId="22" pageNumber="115" precision="15" seconds="4" value="-4.5677776">4°34’4”S</geoCoordinate>
,
<geoCoordinate id="EE5B506EC610FFD0121DF44A6D88F994" box="[370,483,1558,1582]" degrees="39" direction="west" minutes="47" orientation="longitude" pageId="22" pageNumber="115" precision="15" seconds="0" value="-39.783333">39°47’0”W</geoCoordinate>
; Carste leg.;
<date id="FFD11069C610FFD01107F44A6F68F994" box="[616,771,1558,1582]" pageId="22" pageNumber="115" value="2021-03-10" valueMax="2021-03-18" valueMin="2021-03-10">10-18.III.2021</date>
; ISLA96025.
</paragraph>
</subSubSection>
<subSubSection id="C3756522C610FFD013EBF4306DA3F969" pageId="22" pageNumber="115" type="etymology">
<paragraph id="8BD036A9C610FFD013EBF4306DA3F969" blockId="22.[132,777,1643,1747]" pageId="22" pageNumber="115">
<smallCapsWord id="8D36A075C610FFD013EBF4306C95F938" baselines="1661,1662" box="[132,254,1644,1667]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Etymology" pageId="22" pageNumber="115">ETYMOLOGY</smallCapsWord>
. — The new species name,
<taxonomicName id="4C6F4D2AC610FFD01176F4376E10F939" authorityName="Cardoso &amp; Ferreira" authorityYear="2024" box="[537,635,1643,1667]" class="Malacostraca" family="Armadillidae" genus="Ctenorillo" kingdom="Animalia" order="Isopoda" pageId="22" pageNumber="115" phylum="Arthropoda" rank="species" species="cearensis">“cearensis</taxonomicName>
”, refers to the people who are born in the State of
<collectingRegion id="49ABF84BC610FFD0128CF4DA6E75F924" box="[483,542,1670,1694]" country="Brazil" name="Ceara" pageId="22" pageNumber="115">Ceará</collectingRegion>
,
<collectingCountry id="F3787639C610FFD01147F4DA6E0FF924" box="[552,612,1670,1694]" name="Brazil" pageId="22" pageNumber="115">Brazil</collectingCountry>
. The word
<collectingRegion id="49ABF84BC610FFD011BFF4DA6F62F924" box="[720,777,1670,1694]" country="Brazil" name="Ceara" pageId="22" pageNumber="115">Ceará</collectingRegion>
comes from the tupy language that means the macaw’s singing (o canto dajanaia in Portuguese).
</paragraph>
</subSubSection>
<subSubSection id="C3756522C610FFD013EBF4AA6DD9F816" pageId="22" pageNumber="115" type="diagnosis">
<paragraph id="8BD036A9C610FFD013EBF4AA6DD9F816" blockId="22.[131,777,1782,1964]" pageId="22" pageNumber="115">
<smallCapsWord id="8D36A075C610FFD013EBF4AA6C84F8B6" baselines="1799,1800" box="[132,239,1782,1805]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Diagnosis" pageId="22" pageNumber="115">DIAGNOSIS</smallCapsWord>
.— Dorsum covered by weakly developed tubercles and ribs with three rows on cephalon (4, 4, 4); three rows on pereonite 1 (2, 8, 9); two rows on pereonites 2-6 (4 + 11) and on pereonite 7 (4 + 7); one row of four tubercles on pleonites 3-5; and two paramedian tubercles on telson. Pleopod 1 exopod with round distal portion, outer margin straight; endopod with distal portion straight, four times longer than exopod.
</paragraph>
</subSubSection>
<subSubSection id="C3756522C610FFD013EBF58C6EA0F85D" box="[132,715,1999,2024]" pageId="22" pageNumber="115" type="distribution">
<paragraph id="8BD036A9C610FFD013EBF58C6EA0F85D" blockId="22.[132,715,1999,2024]" box="[132,715,1999,2024]" pageId="22" pageNumber="115">
<smallCapsWord id="8D36A075C610FFD013EBF58C6D79F85C" baselines="2017,2018" box="[132,274,2000,2023]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Distribution" pageId="22" pageNumber="115">DISTRIBUTION</smallCapsWord>
. — Santa Quitéria municipality,
<collectingRegion id="49ABF84BC610FFD01136F58C6EACF85D" box="[601,711,2000,2024]" country="Brazil" name="Ceara" pageId="22" pageNumber="115">Ceará state</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C3756522C610FFD11042F7286D56F916" lastPageId="23" lastPageNumber="116" pageId="22" pageNumber="115" type="description">
<paragraph id="8BD036A9C610FFD01042F7286FD6FA36" blockId="22.[811,1457,1396,2028]" box="[813,957,1396,1422]" pageId="22" pageNumber="115">
<smallCapsWord id="8D36A075C610FFD01042F7286FD6FA36" baselines="1415,1415" box="[813,957,1396,1422]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Description" pageId="22" pageNumber="115">DESCRIPTION</smallCapsWord>
</paragraph>
<paragraph id="8BD036A9C610FFD11042F7CF6DDFFE0A" blockId="22.[811,1457,1396,2028]" lastBlockId="23.[132,777,215,432]" lastPageId="23" lastPageNumber="116" pageId="22" pageNumber="115">
Maximum size:
<specimenCount id="9D69FD20C610FFD010B9F7CA6F87FA16" box="[982,1004,1430,1452]" count="1" pageId="22" pageNumber="115" type="male">♂</specimenCount>
and
<specimenCount id="9D69FD20C610FFD0174FF7CA6858FA16" box="[1056,1075,1430,1452]" count="1" pageId="22" pageNumber="115" type="female">♀</specimenCount>
,
<quantity id="4C979B4CC610FFD01750F7CF68E8FA14" box="[1087,1155,1427,1454]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="22" pageNumber="115" unit="mm" value="4.0">4 mm</quantity>
. Color grey (
<figureCitation id="13542A2CC610FFD01665F7CF6903FA17" box="[1290,1384,1427,1453]" captionStart="FIG" captionStartId="35.[132,143,1225,1243]" captionTargetBox="[132,1455,216,1183]" captionTargetId="figure-480@35.[699,983,655,744]" captionTargetPageId="35" captionText="FIG. 27. — Habitus of the species of the genus Ctenorillo Verhoeff, 1942 discussed in the present paper:A, Ctenorillo ferrarai Campos-Filho, Araujo &amp; Taiti,2014; B, Ctenorillo pelado Cardoso &amp; Ferreira, n. sp.; C, Ctenorillo araguaia Cardoso &amp; Ferreira, n. sp.; D, Ctenorillo intertidalis Cardoso &amp; Ferreira, n. sp.; E, Ctenorillo ubajarensis Cardoso &amp; Ferreira,n. sp.; F, Ctenorillo cearensis Cardoso &amp; Ferreira,n. sp.; G, Ctenorillo quiteriensis Cardoso &amp; Ferreira,n. sp.; H, Ctenorillo potiguar Cardoso &amp; Ferreira, n. sp.; I, Ctenorillo iuiuensis n. sp.; J, Ctenorillo jequitinhonha Cardoso &amp; Ferreira, n. sp. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10803325" httpUri="https://zenodo.org/record/10803325/files/figure.png" pageId="22" pageNumber="115">Fig. 27F</figureCitation>
). Dorsum covered by weakly developed tubercles and ribs (
<figureCitation id="13542A2CC610FFD01621F7EF6F54FA57" captionStart="FIG" captionStartId="21.[132,143,1668,1686]" captionTargetBox="[132,1455,215,1619]" captionTargetId="figure-128@21.[132,1455,215,1619]" captionTargetPageId="21" captionText="FIG. 14. — Ctenorillo cearensis Cardoso &amp; Ferreira,n. sp.,holotype (male,4 mm,ISLA96021):A, habitus,lateral view;B, disposition of dorsal tubercles;C, cephalon, frontal view; D, cephalon, dorsal view; E, epimera 1-3 ventral view; F, pleonites 3-5, uropods and telson, dorsal view; G, antennula; H, antenna; I, right mandible; J, left mandible; K, maxillula; L, maxilla; M, maxilliped. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10803297" httpUri="https://zenodo.org/record/10803297/files/figure.png" pageId="22" pageNumber="115">Fig. 14A, B</figureCitation>
): cephalon with three rows (4, 4, and 4 from front to back of vertex); pereonite 1 with
<quantity id="4C979B4CC610FFD0170CF7AE68F5F9B7" box="[1123,1182,1522,1549]" metricMagnitude="-1" metricUnit="m" metricValue="3.81" pageId="22" pageNumber="115" unit="in" value="15.0">15 in</quantity>
three rows (2, 8 and 9); pereonites 2-6 with 15 (4+11); pereonite 7 with 11 (4 +7); pleonites 3-5 with one row of four tubercles; telson with two paramedian tubercles. Dorsal cuticle with short triangular scale setae; pereonites 1-7 bearing one line of
<emphasis id="B91BEABBC610FFD01678F42E69DAF936" box="[1303,1457,1650,1676]" italics="true" pageId="22" pageNumber="115">noduli lateralis</emphasis>
per side on outer surface of posterior tubercle of second line of tubercle. Cephalon with frontal shield protruding above vertex; eye consisting of eight ommatidia (
<figureCitation id="13542A2CC610FFD0167CF48E69C8F956" box="[1299,1443,1746,1772]" captionStart="FIG" captionStartId="21.[132,143,1668,1686]" captionTargetBox="[132,1455,215,1619]" captionTargetId="figure-128@21.[132,1455,215,1619]" captionTargetPageId="21" captionText="FIG. 14. — Ctenorillo cearensis Cardoso &amp; Ferreira,n. sp.,holotype (male,4 mm,ISLA96021):A, habitus,lateral view;B, disposition of dorsal tubercles;C, cephalon, frontal view; D, cephalon, dorsal view; E, epimera 1-3 ventral view; F, pleonites 3-5, uropods and telson, dorsal view; G, antennula; H, antenna; I, right mandible; J, left mandible; K, maxillula; L, maxilla; M, maxilliped. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10803297" httpUri="https://zenodo.org/record/10803297/files/figure.png" pageId="22" pageNumber="115">Fig. 14C, D</figureCitation>
). Pereonite 1 inner lobe of schisma rounded, extending beyond posterior margin of outer lobe; pereonite 2 with triangular ventral tooth reaching posterior margin of epimera (
<figureCitation id="13542A2CC610FFD01627F56D69CEF8F6" box="[1352,1445,1841,1868]" captionStart="FIG" captionStartId="21.[132,143,1668,1686]" captionTargetBox="[132,1455,215,1619]" captionTargetId="figure-128@21.[132,1455,215,1619]" captionTargetPageId="21" captionText="FIG. 14. — Ctenorillo cearensis Cardoso &amp; Ferreira,n. sp.,holotype (male,4 mm,ISLA96021):A, habitus,lateral view;B, disposition of dorsal tubercles;C, cephalon, frontal view; D, cephalon, dorsal view; E, epimera 1-3 ventral view; F, pleonites 3-5, uropods and telson, dorsal view; G, antennula; H, antenna; I, right mandible; J, left mandible; K, maxillula; L, maxilla; M, maxilliped. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10803297" httpUri="https://zenodo.org/record/10803297/files/figure.png" pageId="22" pageNumber="115">Fig. 14E</figureCitation>
); pereonites 2-7 with wide quadrangular epimera (
<figureCitation id="13542A2CC610FFD01651F50D69CAF8D6" box="[1342,1441,1873,1900]" captionStart="FIG" captionStartId="21.[132,143,1668,1686]" captionTargetBox="[132,1455,215,1619]" captionTargetId="figure-128@21.[132,1455,215,1619]" captionTargetPageId="21" captionText="FIG. 14. — Ctenorillo cearensis Cardoso &amp; Ferreira,n. sp.,holotype (male,4 mm,ISLA96021):A, habitus,lateral view;B, disposition of dorsal tubercles;C, cephalon, frontal view; D, cephalon, dorsal view; E, epimera 1-3 ventral view; F, pleonites 3-5, uropods and telson, dorsal view; G, antennula; H, antenna; I, right mandible; J, left mandible; K, maxillula; L, maxilla; M, maxilliped. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10803297" httpUri="https://zenodo.org/record/10803297/files/figure.png" pageId="22" pageNumber="115">Fig. 14A</figureCitation>
). Telson (
<figureCitation id="13542A2CC610FFD01010F52D6FB2F831" box="[895,985,1905,1931]" captionStart="FIG" captionStartId="21.[132,143,1668,1686]" captionTargetBox="[132,1455,215,1619]" captionTargetId="figure-128@21.[132,1455,215,1619]" captionTargetPageId="21" captionText="FIG. 14. — Ctenorillo cearensis Cardoso &amp; Ferreira,n. sp.,holotype (male,4 mm,ISLA96021):A, habitus,lateral view;B, disposition of dorsal tubercles;C, cephalon, frontal view; D, cephalon, dorsal view; E, epimera 1-3 ventral view; F, pleonites 3-5, uropods and telson, dorsal view; G, antennula; H, antenna; I, right mandible; J, left mandible; K, maxillula; L, maxilla; M, maxilliped. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10803297" httpUri="https://zenodo.org/record/10803297/files/figure.png" pageId="22" pageNumber="115">Fig. 14F</figureCitation>
) hourglass-shaped, proximal portion broader than distal portion. Antennula (
<figureCitation id="13542A2CC610FFD017E6F5CD6885F811" box="[1161,1262,1937,1963]" captionStart="FIG" captionStartId="21.[132,143,1668,1686]" captionTargetBox="[132,1455,215,1619]" captionTargetId="figure-128@21.[132,1455,215,1619]" captionTargetPageId="21" captionText="FIG. 14. — Ctenorillo cearensis Cardoso &amp; Ferreira,n. sp.,holotype (male,4 mm,ISLA96021):A, habitus,lateral view;B, disposition of dorsal tubercles;C, cephalon, frontal view; D, cephalon, dorsal view; E, epimera 1-3 ventral view; F, pleonites 3-5, uropods and telson, dorsal view; G, antennula; H, antenna; I, right mandible; J, left mandible; K, maxillula; L, maxilla; M, maxilliped. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10803297" httpUri="https://zenodo.org/record/10803297/files/figure.png" pageId="22" pageNumber="115">Fig. 14G</figureCitation>
) of three articles, second article shorter than first and third, third article with six apical aesthetascs. Antenna (
<figureCitation id="13542A2CC610FFD017E5F58D689BF851" box="[1162,1264,2001,2027]" captionStart="FIG" captionStartId="21.[132,143,1668,1686]" captionTargetBox="[132,1455,215,1619]" captionTargetId="figure-128@21.[132,1455,215,1619]" captionTargetPageId="21" captionText="FIG. 14. — Ctenorillo cearensis Cardoso &amp; Ferreira,n. sp.,holotype (male,4 mm,ISLA96021):A, habitus,lateral view;B, disposition of dorsal tubercles;C, cephalon, frontal view; D, cephalon, dorsal view; E, epimera 1-3 ventral view; F, pleonites 3-5, uropods and telson, dorsal view; G, antennula; H, antenna; I, right mandible; J, left mandible; K, maxillula; L, maxilla; M, maxilliped. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10803297" httpUri="https://zenodo.org/record/10803297/files/figure.png" pageId="22" pageNumber="115">Fig. 14H</figureCitation>
) short and stout, flagellum shorter than fifth article of peduncle; second article of flagellum about three times as long as first. Buccal pieces as
<taxonomicName id="4C6F4D2AC611FFD11386F34B6E43FE8B" authority="Cardoso &amp; Ferreira" authorityName="Cardoso &amp; Ferreira" authorityYear="2024" box="[233,552,279,305]" class="Malacostraca" family="Armadillidae" genus="Ctenorillo" kingdom="Animalia" order="Isopoda" pageId="23" pageNumber="116" phylum="Arthropoda" rank="species" species="pelado" status="n. sp.">
<emphasis id="B91BEABBC611FFD11386F34B6D27FE8B" box="[233,332,279,305]" italics="true" pageId="23" pageNumber="116">C. pelado</emphasis>
Cardoso &amp; Ferreira
</taxonomicName>
,
<taxonomicNameLabel id="A22857C0C611FFD1115CF3446E05FE88" box="[563,622,280,306]" pageId="23" pageNumber="116" rank="species">n. sp.</taxonomicNameLabel>
(
<figureCitation id="13542A2CC611FFD11112F34B6EA0FE8B" box="[637,715,279,305]" captionStart="FIG" captionStartId="21.[132,143,1668,1686]" captionTargetBox="[132,1455,215,1619]" captionTargetId="figure-128@21.[132,1455,215,1619]" captionTargetPageId="21" captionText="FIG. 14. — Ctenorillo cearensis Cardoso &amp; Ferreira,n. sp.,holotype (male,4 mm,ISLA96021):A, habitus,lateral view;B, disposition of dorsal tubercles;C, cephalon, frontal view; D, cephalon, dorsal view; E, epimera 1-3 ventral view; F, pleonites 3-5, uropods and telson, dorsal view; G, antennula; H, antenna; I, right mandible; J, left mandible; K, maxillula; L, maxilla; M, maxilliped. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10803297" httpUri="https://zenodo.org/record/10803297/files/figure.png" pageId="23" pageNumber="116">Fig. 14</figureCitation>
I-M). All pleopod exopods with monospiracular covered lungs. Uropod (
<figureCitation id="13542A2CC611FFD11381F30B6D38FEC8" box="[238,339,343,370]" captionStart="FIG" captionStartId="22.[132,143,1289,1307]" captionTargetBox="[134,1395,216,1241]" captionTargetId="figure-425@22.[132,1455,216,1247]" captionTargetPageId="22" captionText="FIG. 15. — Ctenorillo cearensis Cardoso &amp; Ferreira,n. sp., paratype (male, 4 mm, ISLA96021):A, uropod,B, pereopod 1; C, pereopod 7; D, pleopod 1 and genital papilla; E, pleopod 2; F, pleopod 3 exopod; G, pleopod 4 exopod; H, pleopod 5 exopod." figureDoi="http://doi.org/10.5281/zenodo.10803299" httpUri="https://zenodo.org/record/10803299/files/figure.png" pageId="23" pageNumber="116">Fig. 15A</figureCitation>
) protopod flattened, enlarged on basal portion; exopod outward, short, inserted dorsally close to medial margin of protopod.
</paragraph>
<paragraph id="8BD036A9C611FFD113EBF38A6CD2FE4A" blockId="23.[132,776,470,688]" box="[132,185,470,496]" pageId="23" pageNumber="116">
<emphasis id="B91BEABBC611FFD113EBF38A6CD2FE4A" box="[132,185,470,496]" italics="true" pageId="23" pageNumber="116">Male</emphasis>
</paragraph>
<paragraph id="8BD036A9C611FFD113EBF3AA6D49FD0A" blockId="23.[132,776,470,688]" pageId="23" pageNumber="116">
Pereopods without particular modifications (
<figureCitation id="13542A2CC611FFD1111DF3AA6E92FDAA" box="[626,761,502,529]" captionStart="FIG" captionStartId="22.[132,143,1289,1307]" captionTargetBox="[134,1395,216,1241]" captionTargetId="figure-425@22.[132,1455,216,1247]" captionTargetPageId="22" captionText="FIG. 15. — Ctenorillo cearensis Cardoso &amp; Ferreira,n. sp., paratype (male, 4 mm, ISLA96021):A, uropod,B, pereopod 1; C, pereopod 7; D, pleopod 1 and genital papilla; E, pleopod 2; F, pleopod 3 exopod; G, pleopod 4 exopod; H, pleopod 5 exopod." figureDoi="http://doi.org/10.5281/zenodo.10803299" httpUri="https://zenodo.org/record/10803299/files/figure.png" pageId="23" pageNumber="116">Fig. 15B, C</figureCitation>
). Pleopod 1 exopod (
<figureCitation id="13542A2CC611FFD11236F04A6DD7FD8B" box="[345,444,534,561]" captionStart="FIG" captionStartId="22.[132,143,1289,1307]" captionTargetBox="[134,1395,216,1241]" captionTargetId="figure-425@22.[132,1455,216,1247]" captionTargetPageId="22" captionText="FIG. 15. — Ctenorillo cearensis Cardoso &amp; Ferreira,n. sp., paratype (male, 4 mm, ISLA96021):A, uropod,B, pereopod 1; C, pereopod 7; D, pleopod 1 and genital papilla; E, pleopod 2; F, pleopod 3 exopod; G, pleopod 4 exopod; H, pleopod 5 exopod." figureDoi="http://doi.org/10.5281/zenodo.10803299" httpUri="https://zenodo.org/record/10803299/files/figure.png" pageId="23" pageNumber="116">Fig. 15D</figureCitation>
) wider than long, round distal portion, outer margin straight; endopod with distal portion straight, four times longer than exopod. Pleopod 2 endopod longer than exopod (
<figureCitation id="13542A2CC611FFD11207F02A6DA1FD2B" box="[360,458,630,657]" captionStart="FIG" captionStartId="22.[132,143,1289,1307]" captionTargetBox="[134,1395,216,1241]" captionTargetId="figure-425@22.[132,1455,216,1247]" captionTargetPageId="22" captionText="FIG. 15. — Ctenorillo cearensis Cardoso &amp; Ferreira,n. sp., paratype (male, 4 mm, ISLA96021):A, uropod,B, pereopod 1; C, pereopod 7; D, pleopod 1 and genital papilla; E, pleopod 2; F, pleopod 3 exopod; G, pleopod 4 exopod; H, pleopod 5 exopod." figureDoi="http://doi.org/10.5281/zenodo.10803299" httpUri="https://zenodo.org/record/10803299/files/figure.png" pageId="23" pageNumber="116">Fig. 15E</figureCitation>
). Pleopods 3-5 exopods as in
<figureCitation id="13542A2CC611FFD113EBF0CA6C80FD0A" box="[132,235,661,688]" captionStart="FIG" captionStartId="22.[132,143,1289,1307]" captionTargetBox="[134,1395,216,1241]" captionTargetId="figure-425@22.[132,1455,216,1247]" captionTargetPageId="22" captionText="FIG. 15. — Ctenorillo cearensis Cardoso &amp; Ferreira,n. sp., paratype (male, 4 mm, ISLA96021):A, uropod,B, pereopod 1; C, pereopod 7; D, pleopod 1 and genital papilla; E, pleopod 2; F, pleopod 3 exopod; G, pleopod 4 exopod; H, pleopod 5 exopod." figureDoi="http://doi.org/10.5281/zenodo.10803299" httpUri="https://zenodo.org/record/10803299/files/figure.png" pageId="23" pageNumber="116">Figure 15</figureCitation>
F-H.
</paragraph>
<paragraph id="8BD036A9C611FFD113EBF08A6C8EFD54" blockId="23.[131,777,726,1708]" box="[132,229,726,752]" pageId="23" pageNumber="116">
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</paragraph>
<paragraph id="8BD036A9C611FFD113EBF0A96D56F916" blockId="23.[131,777,726,1708]" pageId="23" pageNumber="116">
Specimens of
<taxonomicName id="4C6F4D2AC611FFD1124FF0AA6EEAFCB5" authority="Cardoso &amp; Ferreira" authorityName="Cardoso &amp; Ferreira" authorityYear="2024" box="[288,641,757,784]" class="Malacostraca" family="Armadillidae" genus="Ctenorillo" kingdom="Animalia" order="Isopoda" pageId="23" pageNumber="116" phylum="Arthropoda" rank="species" species="cearensis" status="n. sp.">
<emphasis id="B91BEABBC611FFD1124FF0AA6DF7FCB5" box="[288,412,758,784]" italics="true" pageId="23" pageNumber="116">C. cearensis</emphasis>
Cardoso &amp; Ferreira
</taxonomicName>
,
<taxonomicNameLabel id="A22857C0C611FFD111E1F0AA6EA7FCAA" box="[654,716,758,784]" pageId="23" pageNumber="116" rank="species">n. sp.</taxonomicNameLabel>
were found in six limestone caves inserted in the caatinga biome in northern
<collectingRegion id="49ABF84BC611FFD11261F1696DE0FCEA" box="[270,395,821,848]" country="Brazil" name="Ceara" pageId="23" pageNumber="116">Ceará state</collectingRegion>
. The region experiences a Tropical Hot Semi-arid climate (BSh according to Koppen), with an average annual rainfall of
<quantity id="4C979B4CC611FFD112F1F1296E79FC35" box="[414,530,885,911]" metricMagnitude="-1" metricUnit="m" metricValue="7.997999999999999" pageId="23" pageNumber="116" unit="mm" value="799.8">799.8 mm</quantity>
concentrated between February and April (
<bibRefCitation id="EFFE4B58C611FFD11208F1CA6E5AFC15" author="ALVARES C. A. &amp; STAPE J. L. &amp; SENTELHAS P. C. &amp; MORAES G. &amp; DE &amp; LEONARDO J. &amp; SPAROVEK G." box="[359,561,917,946]" pageId="23" pageNumber="116" pagination="711 - 728" refId="ref31163" refString="ALVARES C. A., STAPE J. L., SENTELHAS P. C., MORAES G. DE, LEONARDO J. &amp; SPAROVEK G. 2013. - Koppen's climate classification map for Brazil. Meteorologische Zeitschrift 22 (6): 711 - 728. https: // doi. org / 10.1127 / 0941 - 2948 / 2013 / 0507" type="journal article" year="2013">
Álvares
<emphasis id="B91BEABBC611FFD112D3F1CA6D9AFC15" box="[444,497,917,943]" italics="true" pageId="23" pageNumber="116">et al.</emphasis>
2013
</bibRefCitation>
). Average temperatures in the region hover around 27°C, with approximately 3°C of thermal amplitude (
<bibRefCitation id="EFFE4B58C611FFD112C2F1896E51FC55" author="IPECE" box="[429,570,981,1007]" pageId="23" pageNumber="116" refId="ref33190" refString="IPECE 2011. - Instituto de Pesquisa e Estrategia Economica no Estado do Ceara. http: // www. ipece. ce. gov. br" type="url" year="2011">IPECE 2011</bibRefCitation>
). The area features
<specimenCount id="9D69FD20C611FFD113EAF1A86C96FBB5" box="[133,253,1012,1039]" count="3" pageId="23" pageNumber="116" type="generic" typeStatus="types">three types</specimenCount>
of vegetation: Tropical Steppe, located in both the residual massifs and dissected areas of the Sertaneja Depression; Thorny Deciduous Forest and Tropical Rainforest. Unfortunately, the biologists who collected the specimens did not provide any data on the caves or the conservation status of the surrounding region. However, at least 74 caves were inventoried in the region, and specimens of
<taxonomicName id="4C6F4D2AC611FFD111FEF6E86D37FB54" authority="Cardoso &amp; Ferreira" authorityName="Cardoso &amp; Ferreira" authorityYear="2024" class="Malacostraca" family="Armadillidae" genus="Ctenorillo" kingdom="Animalia" order="Isopoda" pageId="23" pageNumber="116" phylum="Arthropoda" rank="species" species="cearensis" status="n. sp.">
<emphasis id="B91BEABBC611FFD111FEF6E86F6CFB74" box="[657,775,1204,1230]" italics="true" pageId="23" pageNumber="116">C. cearensis</emphasis>
Cardoso &amp; Ferreira
</taxonomicName>
,
<taxonomicNameLabel id="A22857C0C611FFD11207F6886DCFFB54" box="[360,420,1236,1262]" pageId="23" pageNumber="116" rank="species">n. sp.</taxonomicNameLabel>
were found only in six of them. As mentioned for other species herein described, external inventories were not conducted, hence it is not possible to determine the actual distribution of this species. Nonetheless, the species does not present any troglomorphic traits, which would suggest its restriction to subterranean habitats. Importantly, the cave inventories were conducted to rank the importance of the caves in the area, as required by Brazilian environmental agencies for evaluating projects that may impact the environment. The potential installation of quarries in the area is a cause for concern since the limestone caves in the region are located close to uranium deposits (
<bibRefCitation id="EFFE4B58C611FFD111CBF46E6D4BF9D6" author="JUNIOR J. T. G. &amp; DE SOUZA M. J. N." pageId="23" pageNumber="116" pagination="1368 - 1377" refId="ref33217" refString="JUNIOR J. T. G. &amp; DE SOUZA M. J. N. 2012. - Caracterizacao ambiental de Santa Quiteria, Ceara: a nova cidade uranifera do Brasil. Revista GeoNorte 3 (5): 1368 - 1377." type="journal article" year="2012">Júnior &amp; de Souza 2012</bibRefCitation>
), and such enterprises could destroy the caves where the species occurs and severely alter the surrounding external habitats.
</paragraph>
</subSubSection>
<subSubSection id="C3756522C611FFD113EBF48E6E9BF856" pageId="23" pageNumber="116" type="discussion">
<paragraph id="8BD036A9C611FFD113EBF48E6C83F950" blockId="23.[132,775,1746,2028]" box="[132,232,1746,1772]" pageId="23" pageNumber="116">
<smallCapsWord id="8D36A075C611FFD113EBF48E6C83F950" baselines="1765,1765" box="[132,232,1746,1772]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Remarks" pageId="23" pageNumber="116">REMARKS</smallCapsWord>
</paragraph>
<paragraph id="8BD036A9C611FFD113EBF4AE6E9BF856" blockId="23.[132,775,1746,2028]" pageId="23" pageNumber="116">
As
<taxonomicName id="4C6F4D2AC611FFD113C6F4AE6E47F8B6" authority="Cardoso &amp; Ferreira" authorityName="Cardoso &amp; Ferreira" authorityYear="2024" box="[169,556,1778,1804]" class="Malacostraca" family="Armadillidae" genus="Ctenorillo" kingdom="Animalia" order="Isopoda" pageId="23" pageNumber="116" phylum="Arthropoda" rank="species" species="ubajarensis" status="n. sp.">
<emphasis id="B91BEABBC611FFD113C6F4AE6D28F8B6" box="[169,323,1778,1804]" italics="true" pageId="23" pageNumber="116">C. ubajarensis</emphasis>
Cardoso &amp; Ferreira
</taxonomicName>
,
<taxonomicNameLabel id="A22857C0C611FFD11155F4AE6E10F8B6" box="[570,635,1778,1804]" pageId="23" pageNumber="116" rank="species">n. sp.</taxonomicNameLabel>
,
<taxonomicName id="4C6F4D2AC611FFD111E6F4AE6D0BF896" authority="Cardoso &amp; Ferreira" authorityName="Cardoso &amp; Ferreira" authorityYear="2024" class="Malacostraca" family="Armadillidae" genus="Ctenorillo" kingdom="Animalia" order="Isopoda" pageId="23" pageNumber="116" phylum="Arthropoda" rank="species" species="cearensis" status="n. sp.">
<emphasis id="B91BEABBC611FFD111E6F4AE6F6DF8B6" box="[649,774,1778,1804]" italics="true" pageId="23" pageNumber="116">C. cearensis</emphasis>
Cardoso &amp; Ferreira
</taxonomicName>
,
<taxonomicNameLabel id="A22857C0C611FFD11201F54E6DC0F896" box="[366,427,1810,1836]" pageId="23" pageNumber="116" rank="species">n. sp.</taxonomicNameLabel>
presents the less developed tubercles, however the tubercles are more prominent than in
<taxonomicName id="4C6F4D2AC611FFD113E9F50E6DBEF8D1" authority="Cardoso &amp; Ferreira" authorityName="Cardoso &amp; Ferreira" authorityYear="2024" box="[134,469,1873,1900]" class="Malacostraca" family="Armadillidae" genus="Ctenorillo" kingdom="Animalia" order="Isopoda" pageId="23" pageNumber="116" phylum="Arthropoda" rank="species" species="cearensis" status="n. sp.">
<emphasis id="B91BEABBC611FFD113E9F50E6C96F8D1" box="[134,253,1874,1900]" italics="true" pageId="23" pageNumber="116">C. cearensis</emphasis>
Cardoso &amp; Ferreira
</taxonomicName>
,
<taxonomicNameLabel id="A22857C0C611FFD112B0F50E6E72F8D6" box="[479,537,1874,1900]" pageId="23" pageNumber="116" rank="species">n. sp.</taxonomicNameLabel>
As the other Brazilian species,
<taxonomicName id="4C6F4D2AC611FFD113B2F52E6E5EF831" authority="Cardoso &amp; Ferreira" authorityName="Cardoso &amp; Ferreira" authorityYear="2024" box="[221,565,1905,1932]" class="Malacostraca" family="Armadillidae" genus="Ctenorillo" kingdom="Animalia" order="Isopoda" pageId="23" pageNumber="116" phylum="Arthropoda" rank="species" species="cearensis" status="n. sp.">
<emphasis id="B91BEABBC611FFD113B2F52E6D3EF831" box="[221,341,1906,1932]" italics="true" pageId="23" pageNumber="116">C. cearensis</emphasis>
Cardoso &amp; Ferreira
</taxonomicName>
,
<taxonomicNameLabel id="A22857C0C611FFD1112EF52E6E16F836" box="[577,637,1906,1932]" pageId="23" pageNumber="116" rank="species">n. sp.</taxonomicNameLabel>
presents the pleonites 3-5 with four tubercles each and differs from them by the number of tubercles on pereon, with 19 tubercles on pereonite 1, 15 on pereonites 2-6, and 11 on pereonite 7.
</paragraph>
</subSubSection>
</treatment>
</document>
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