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<document id="DE874F392673577F156F22B1A4A44EAA" ID-DOI="10.5281/zenodo.5460634" ID-ISSN="2345-7600" ID-Zenodo-Dep="5460634" ID-ZooBank="1E7214C5-24E8-4860-BF13-203FD93A2C19" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630239335276" checkinUser="valdenar" docAuthor="Pholyotha, Arthit, Sutcharit, Chirasak &amp; Panha, Somsak" docDate="2018" docId="03A587FA7827A03DFF49BFAC30935323" docLanguage="en" docName="66rbz763-781.pdf" docOrigin="Raffles Bulletin of Zoology 66" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Macrochlamys coleus Pholyotha &amp; Sutcharit &amp; Panha 2018, new species" docType="treatment" docUuid="52C0D943-69AB-4BC7-8850-9A15AFBBF7D2" docUuidSource="ZooBank" docVersion="4" lastPageNumber="769" masterDocId="FF9CFF827821A03BFFC7BF1034285476" masterDocTitle="The land snail genus Macrochlamys Gray, 1847 from Thailand, with descriptions of five new species (Pulmonata: Ariophantidae)" masterLastPageNumber="781" masterPageNumber="763" pageNumber="769" updateTime="1699195389588" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="C1F51B41A6D48F9A027A7186AF138174">The land snail genus Macrochlamys Gray, 1847 from Thailand, with descriptions of five new species (Pulmonata: Ariophantidae)</mods:title>
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<mods:namePart id="7CD362B8D970A97F176E50455A3709D8">Pholyotha, Arthit</mods:namePart>
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<mods:namePart id="E26F8ECE4408B0990BDC8CC933D81563">Sutcharit, Chirasak</mods:namePart>
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<mods:namePart id="6BF5724A1C6BAC169BE28D1ABA0B3891">Panha, Somsak</mods:namePart>
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<mods:title id="3C3F2EA8802D5BB3E95C98C922436D9E">Raffles Bulletin of Zoology</mods:title>
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<mods:date id="93EB658D22686D423A21A6F804059E3D">2018</mods:date>
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<mods:number id="5991E44565FA704357401A46D536132D">2018-12-03</mods:number>
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<mods:number id="26AE8462533BE49A1E13879491661632">66</mods:number>
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<mods:identifier id="A2BE5F58F26BF2EEFCB85596D1046726" type="ISSN">2345-7600</mods:identifier>
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<treatment id="03A587FA7827A03DFF49BFAC30935323" ID-DOI="http://doi.org/10.5281/zenodo.5458063" ID-Zenodo-Dep="5458063" LSID="urn:lsid:zoobank.org:act:52C0D943-69AB-4BC7-8850-9A15AFBBF7D2" httpUri="http://treatment.plazi.org/id/03A587FA7827A03DFF49BFAC30935323" lastPageNumber="769" pageId="6" pageNumber="769">
<subSubSection id="C31665677827A03DFF49BFAC372954A3" box="[142,769,188,213]" pageId="6" pageNumber="769" type="nomenclature">
<paragraph id="8BB336EC7827A03DFF49BFAC372954A3" blockId="6.[142,769,188,245]" box="[142,769,188,213]" pageId="6" pageNumber="769">
<heading id="D0FB81807827A03DFF49BFAC372954A3" bold="true" box="[142,769,188,213]" centered="true" fontSize="10" level="2" pageId="6" pageNumber="769" reason="2">
<emphasis id="B978EAFE7827A03DFF49BFAC372954A3" bold="true" box="[142,769,188,213]" pageId="6" pageNumber="769">
<taxonomicName id="4C0C4D6F7827A03DFF49BFAC364554A3" ID-CoL="3X42Z" authority="Pholyotha &amp; Panha" authorityName="Pholyotha &amp; Sutcharit &amp; Panha" authorityYear="2018" box="[142,621,188,213]" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="769" phylum="Mollusca" rank="species" species="coleus" status="sp. nov.">
<emphasis id="B978EAFE7827A03DFF49BFAC35A854A2" bold="true" box="[142,384,188,212]" italics="true" pageId="6" pageNumber="769">Macrochlamys coleus</emphasis>
Pholyotha &amp; Panha
</taxonomicName>
,
<taxonomicNameLabel id="A24B57857827A03DFDBCBFAD372954A3" box="[635,769,189,213]" pageId="6" pageNumber="769" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31665677827A03DFEE6BFCD36455483" box="[289,621,221,245]" pageId="6" pageNumber="769" type="description">
<paragraph id="8BB336EC7827A03DFEE6BFCD36455483" blockId="6.[142,769,188,245]" box="[289,621,221,245]" pageId="6" pageNumber="769">
(
<figureCitation id="13372A697827A03DFEEEBFCD35A15483" box="[297,393,221,245]" captionStart="Fig" captionStartId="2.[135,170,1546,1568]" captionTargetBox="[136,1474,192,1529]" captionTargetId="figure-7@2.[136,1474,189,1531]" captionTargetPageId="2" captionText="Fig. 2. Living snails of Macrochlamys spp. from Thailand. A, Macrochlamys aurantia new species, paratype (CUMZ 7103), Thong Pha Phum, Kanchanaburi, shell width about 15 mm; B, M. coleus new species, paratype (CUMZ 7106), Sai Yok, Kanchanaburi, shell width about 15 mm; C,M. caverna new species, paratype (CUMZ 7109), Tha Wung, Lopburi, shell width about 17 mm; D, M. lemma new species, paratype (CUMZ 7116), Ban Mi, Lopburi, shell width about 15 mm; E, M. tanymentula new species, paratype (CUMZ 7119), Thong Pha Phum, Kanchanaburi, shell width about 10 mm; F, M. kelantanensis (CUMZ 7125), Lam Kaen, Phang-Nga, shell width about 20 mm." figureDoi="http://doi.org/10.5281/zenodo.5460640" httpUri="https://zenodo.org/record/5460640/files/figure.png" pageId="6" pageNumber="769">Figs. 2B</figureCitation>
,
<figureCitation id="13372A697827A03DFE52BFCD35F15482" box="[405,473,221,245]" captionStart="Fig" captionStartId="3.[113,148,1896,1918]" captionTargetBox="[113,1451,328,1881]" captionTargetId="figure-9@3.[113,1451,328,1881]" captionTargetPageId="3" captionText="Fig. 3. Shells of Macrochlamys aurantia new species (A, B), M. coleus new species (C, D), and M. caverna new species (E, F). A, Macrochlamys aurantia holotype (CUMZ 7101); B, paratype (CUMZ 7102); C, M. coleus holotype (CUMZ 7104); D, paratype (CUMZ 7105); E, M. caverna holotype (CUMZ 7107); F, paratype (CUMZ 7108)." figureDoi="http://doi.org/10.5281/zenodo.5460642" httpUri="https://zenodo.org/record/5460642/files/figure.png" pageId="6" pageNumber="769">3C, D</figureCitation>
,
<figureCitation id="13372A697827A03DFE20BFCD36345483" box="[487,540,221,245]" captionStart="Fig" captionStartId="5.[113,148,1838,1860]" captionTargetBox="[113,1451,270,1823]" captionTargetId="figure-9@5.[113,1451,270,1823]" captionTargetPageId="5" captionText="Fig. 4. Genitalia of Macrochlamys aurantia new species (A–E), and M. coleus new species (F–I). A–E, Macrochlamys aurantia paratype (CUMZ 7103): A, whole reproductive system; B, internal structure of penis with penial verge; C, spermatophore; D, detail of spines between sperm sac and tail filament; E, spines on the end of tail filament. F–I, Macrochlamys coleus paratype (CUMZ 7106): F, whole reproductive system; G, internal structure of penis with penial verge; H, spermatophore; I, head filament. Abbreviations: at, atrium; da, dart apparatus; drm, dart retractor muscle; e, epiphallus; ec, epiphallic caecum; fl, flagellum; fo, free oviduct; gd, gametolytic duct; gs, gametolytic sac; hf, head filament; ov, oviduct; p, penis; pc, penial caecum; pg, prostate gland; pp, penial pilaster; prm, penial retractor muscle; pv, penial verge; ss, sperm sac; tf, tail filament; v, vagina and vd, vas deferens. Asterisk indicates the tissue casting." figureDoi="http://doi.org/10.5281/zenodo.5460644" httpUri="https://zenodo.org/record/5460644/files/figure.png" pageId="6" pageNumber="769">4F–I</figureCitation>
,
<figureCitation id="13372A697827A03DFDEFBFCD364F5483" box="[552,615,221,245]" captionStart="Fig" captionStartId="10.[136,171,1215,1237]" captionTargetBox="[136,1474,189,1200]" captionTargetId="figure-7@10.[136,1474,189,1200]" captionTargetPageId="10" captionText="Fig. 6. SEM images of the radula of Macrochlamys aurantia new species (A–C) (paratype; CUMZ 7103), M. coleus new species (D–F) (paratype; CUMZ 7106), and M. caverna new species (G–I) (paratype; CUMZ 7109). Central tooth (indicated by ‘C’) with lateral teeth (A, D, G); lateral teeth with tricuspid marginal teeth transition (B, E, H); bicuspid marginal teeth (C, F, I)." figureDoi="http://doi.org/10.5281/zenodo.5460650" httpUri="https://zenodo.org/record/5460650/files/figure.png" pageId="6" pageNumber="769">6D–F</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31665677827A03DFF4FBE0D36045643" pageId="6" pageNumber="769" type="materials_examined">
<paragraph id="8BB336EC7827A03DFF4FBE0D36045643" blockId="6.[136,775,285,565]" pageId="6" pageNumber="769">
<emphasis id="B978EAFE7827A03DFF4FBE0D35465543" bold="true" box="[136,366,285,309]" pageId="6" pageNumber="769">Material examined.</emphasis>
<materialsCitation id="3B643CB17827A03DFEBEBE0D350855A2" collectionCode="CUMZ" country="Thailand" latitude="14.533778" location="Wat Sunantha Wanaram" longLatPrecision="1" longitude="98.837166" municipality="Sai Yok District" pageId="6" pageNumber="769" specimenCode="CUMZ 7104" specimenCount="1" stateProvince="Kanchanaburi" typeStatus="holotype">
<typeStatus id="54B7884E7827A03DFEBEBE0D35F65543" box="[377,478,285,309]" pageId="6" pageNumber="769" type="holotype">Holotype</typeStatus>
(
<figureCitation id="13372A697827A03DFE37BE0D36615543" box="[496,585,285,309]" captionStart="Fig" captionStartId="3.[113,148,1896,1918]" captionTargetBox="[113,1451,328,1881]" captionTargetId="figure-9@3.[113,1451,328,1881]" captionTargetPageId="3" captionText="Fig. 3. Shells of Macrochlamys aurantia new species (A, B), M. coleus new species (C, D), and M. caverna new species (E, F). A, Macrochlamys aurantia holotype (CUMZ 7101); B, paratype (CUMZ 7102); C, M. coleus holotype (CUMZ 7104); D, paratype (CUMZ 7105); E, M. caverna holotype (CUMZ 7107); F, paratype (CUMZ 7108)." figureDoi="http://doi.org/10.5281/zenodo.5460642" httpUri="https://zenodo.org/record/5460642/files/figure.png" pageId="6" pageNumber="769">Fig. 3C</figureCitation>
; shell height
<quantity id="4CF49B097827A03DFD22BE0D349C5522" metricMagnitude="-3" metricUnit="m" metricValue="6.6" pageId="6" pageNumber="769" unit="mm" value="6.6">6.6 mm</quantity>
, shell width 13.0 mm, aperture height 5.0 mm, aperture width
<quantity id="4CF49B097827A03DFF13BE4D351A5502" box="[212,306,349,373]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="6" pageNumber="769" unit="mm" value="6.2">6.2 mm</quantity>
, 5½ whorls) (
<specimenCode id="DBAA9E977827A03DFE1DBE4D36585503" box="[474,624,349,373]" collectionCode="CUMZ" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" pageId="6" pageNumber="769">CUMZ 7104</specimenCode>
), limestone outcrops at
<location id="8ED360377827A03DFED6BE6D360E55E2" LSID="urn:lsid:plazi:treatment:03A587FA7827A03DFF49BFAC30935323:8ED360377827A03DFED6BE6D360E55E2" box="[273,550,381,405]" country="Thailand" latitude="14.533778" longLatPrecision="1" longitude="98.837166" municipality="Sai Yok District" name="Wat Sunantha Wanaram" pageId="6" pageNumber="769" stateProvince="Kanchanaburi">Wat Sunantha Wanaram</location>
(Temple),
<collectingMunicipality id="6BD7AC967827A03DFD6DBE6D34F355C3" pageId="6" pageNumber="769">Sai Yok District</collectingMunicipality>
,
<collectingRegion id="49C8F80E7827A03DFF2FBE8D35C155C3" box="[232,489,413,437]" country="Thailand" name="Kanchanaburi" pageId="6" pageNumber="769">Kanchanaburi Province</collectingRegion>
,
<collectingCountry id="F31B767C7827A03DFE32BE8D367F55C3" box="[501,599,413,437]" name="Thailand" pageId="6" pageNumber="769">Thailand</collectingCountry>
,
<geoCoordinate id="EE38502B7827A03DFDA3BE8D372B55C2" box="[612,771,411,437]" degrees="14" direction="north" minutes="32" orientation="latitude" pageId="6" pageNumber="769" precision="1" seconds="01.6" value="14.533778">14° 32′01.6″N</geoCoordinate>
,
<geoCoordinate id="EE38502B7827A03DFF4FBEAD350855A2" box="[136,288,443,469]" degrees="98" direction="east" minutes="50" orientation="longitude" pageId="6" pageNumber="769" precision="1" seconds="13.8" value="98.837166">98° 50′13.8″E</geoCoordinate>
</materialsCitation>
.
<materialsCitation id="3B643CB17827A03DFEEEBEAD36015643" collectionCode="CUMZ" country="Thailand" latitude="14.533778" location="Wat Sunantha Wanaram" longLatPrecision="1" longitude="98.837166" municipality="Sai Yok District" pageId="6" pageNumber="769" specimenCode="CUMZ 7105, CUMZ 7106" specimenCount="10" stateProvince="Kanchanaburi" typeStatus="paratype">
<typeStatus id="54B7884E7827A03DFEEEBEAD35BD55A3" box="[297,405,445,469]" pageId="6" pageNumber="769" type="paratype">Paratypes</typeStatus>
: one shell (
<figureCitation id="13372A697827A03DFDCFBEAD367455A3" box="[520,604,445,469]" captionStart="Fig" captionStartId="3.[113,148,1896,1918]" captionTargetBox="[113,1451,328,1881]" captionTargetId="figure-9@3.[113,1451,328,1881]" captionTargetPageId="3" captionText="Fig. 3. Shells of Macrochlamys aurantia new species (A, B), M. coleus new species (C, D), and M. caverna new species (E, F). A, Macrochlamys aurantia holotype (CUMZ 7101); B, paratype (CUMZ 7102); C, M. coleus holotype (CUMZ 7104); D, paratype (CUMZ 7105); E, M. caverna holotype (CUMZ 7107); F, paratype (CUMZ 7108)." figureDoi="http://doi.org/10.5281/zenodo.5460642" httpUri="https://zenodo.org/record/5460642/files/figure.png" pageId="6" pageNumber="769">Fig. 3D</figureCitation>
; shell height 6.0 mm, shell width
<quantity id="4CF49B097827A03DFEFBBECD35B55582" box="[316,413,477,501]" metricMagnitude="-2" metricUnit="m" metricValue="1.23" pageId="6" pageNumber="769" unit="mm" value="12.3">12.3 mm</quantity>
, aperture height
<quantity id="4CF49B097827A03DFD97BECD368C5582" box="[592,676,477,501]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="6" pageNumber="769" unit="mm" value="4.5">4.5 mm</quantity>
, aperture width
<quantity id="4CF49B097827A03DFF17BEED35015662" box="[208,297,509,533]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" pageId="6" pageNumber="769" unit="mm" value="6.1">6.1 mm</quantity>
, 5¼ whorls) (
<specimenCode id="DBAA9E977827A03DFE00BEED36735663" box="[455,603,509,533]" collectionCode="CUMZ" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" pageId="6" pageNumber="769">CUMZ 7105</specimenCode>
),
<specimenCount id="9D0AFD657827A03DFDB6BEED372E5663" box="[625,774,509,533]" pageId="6" pageNumber="769" type="generic">10 specimens</specimenCount>
(
<specimenCode id="DBAA9E977827A03DFF48BD0D35085643" box="[143,288,541,565]" collectionCode="CUMZ" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" pageId="6" pageNumber="769">CUMZ 7106</specimenCode>
), same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31665677827A03DFF4FBD4D36B156C2" pageId="6" pageNumber="769" type="etymology">
<paragraph id="8BB336EC7827A03DFF4FBD4D36B156C2" blockId="6.[136,775,605,693]" pageId="6" pageNumber="769">
<emphasis id="B978EAFE7827A03DFF4FBD4D35235603" bold="true" box="[136,267,605,629]" pageId="6" pageNumber="769">Etymology.</emphasis>
The specific epithet “
<taxonomicName id="4C0C4D6F7827A03DFDC3BD4D36615603" authorityName="Pholyotha &amp; Sutcharit &amp; Panha" authorityYear="2018" box="[516,585,605,629]" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="769" phylum="Mollusca" rank="species" species="coleus">
<emphasis id="B978EAFE7827A03DFDC3BD4D36615603" box="[516,585,605,629]" italics="true" pageId="6" pageNumber="769">coleus</emphasis>
</taxonomicName>
”, from the Latin word meaning “sheath”, refers to the well-developed tissue covering between the vagina and dart apparatus.
</paragraph>
</subSubSection>
<subSubSection id="C31665677827A03DFF4FBDCD37D85683" pageId="6" pageNumber="769" type="description">
<paragraph id="8BB336EC7827A03DFF4FBDCD358C5042" blockId="6.[134,776,733,1077]" pageId="6" pageNumber="769">
<emphasis id="B978EAFE7827A03DFF4FBDCD353A5683" bold="true" box="[136,274,733,757]" pageId="6" pageNumber="769">Description.</emphasis>
Shell medium-sized (
<figureCitation id="13372A697827A03DFE33BDCD364C5682" box="[500,612,733,757]" captionStart="Fig" captionStartId="3.[113,148,1896,1918]" captionTargetBox="[113,1451,328,1881]" captionTargetId="figure-9@3.[113,1451,328,1881]" captionTargetPageId="3" captionText="Fig. 3. Shells of Macrochlamys aurantia new species (A, B), M. coleus new species (C, D), and M. caverna new species (E, F). A, Macrochlamys aurantia holotype (CUMZ 7101); B, paratype (CUMZ 7102); C, M. coleus holotype (CUMZ 7104); D, paratype (CUMZ 7105); E, M. caverna holotype (CUMZ 7107); F, paratype (CUMZ 7108)." figureDoi="http://doi.org/10.5281/zenodo.5460642" httpUri="https://zenodo.org/record/5460642/files/figure.png" pageId="6" pageNumber="769">Fig. 3C, D</figureCitation>
; shell height up to
<quantity id="4CF49B097827A03DFF64BDED34D25762" box="[163,250,765,789]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" pageId="6" pageNumber="769" unit="mm" value="6.6">6.6 mm</quantity>
; shell width up to 13.0 mm), dextral, depressed, slightly thick, translucent brown in colour. Shell surface nearly smooth, with very fine growth lines. Embryonic shell surface smooth, with about 2½ whorls. Whorls 5–6, increasing regularly. Suture wide and shallow, channel-shaped. Spire depressed, apex slightly raised with spire angle of about 148–152°. Last whorl well rounded. Aperture crescent ovoid shaped, obliquely open, with simple lip. Columellar margin little developed and slightly reflected near umbilicus. Umbilicus narrowly open.
</paragraph>
<paragraph id="8BB336EC7827A03DFF4FBB4D358351E3" blockId="6.[135,776,1117,1429]" pageId="6" pageNumber="769">
<emphasis id="B978EAFE7827A03DFF4FBB4D35165003" box="[136,318,1117,1141]" italics="true" pageId="6" pageNumber="769">Genital organs</emphasis>
. Atrium (at) short. Penis prolonged, cylindrical-shaped with rounded bulge corresponding to penial verge. Penial caecum (pc) slightly prolonged. Epiphallus (e) slender tube about two times longer than penis. Epiphallic caecum (ec) coiled about one circle, located at middle of epiphallus and attached with thick penial retractor muscle (prm). Flagellum (fl) short, cylindrical shaped and stretched around half of epiphallus length. Vas deference (vd) a thin tube connected between distal end of epiphallus and free oviduct (
<figureCitation id="13372A697827A03DFE8CBA6D35B751E3" box="[331,415,1405,1429]" captionStart="Fig" captionStartId="5.[113,148,1838,1860]" captionTargetBox="[113,1451,270,1823]" captionTargetId="figure-9@5.[113,1451,270,1823]" captionTargetPageId="5" captionText="Fig. 4. Genitalia of Macrochlamys aurantia new species (A–E), and M. coleus new species (F–I). A–E, Macrochlamys aurantia paratype (CUMZ 7103): A, whole reproductive system; B, internal structure of penis with penial verge; C, spermatophore; D, detail of spines between sperm sac and tail filament; E, spines on the end of tail filament. F–I, Macrochlamys coleus paratype (CUMZ 7106): F, whole reproductive system; G, internal structure of penis with penial verge; H, spermatophore; I, head filament. Abbreviations: at, atrium; da, dart apparatus; drm, dart retractor muscle; e, epiphallus; ec, epiphallic caecum; fl, flagellum; fo, free oviduct; gd, gametolytic duct; gs, gametolytic sac; hf, head filament; ov, oviduct; p, penis; pc, penial caecum; pg, prostate gland; pp, penial pilaster; prm, penial retractor muscle; pv, penial verge; ss, sperm sac; tf, tail filament; v, vagina and vd, vas deferens. Asterisk indicates the tissue casting." figureDoi="http://doi.org/10.5281/zenodo.5460644" httpUri="https://zenodo.org/record/5460644/files/figure.png" pageId="6" pageNumber="769">Fig. 4F</figureCitation>
).
</paragraph>
<paragraph id="8BB336EC7827A03DFF4FBAAD36985243" blockId="6.[135,775,1469,1589]" pageId="6" pageNumber="769">
Inner penial wall supported proximally with small and irregular wrinkled penial pilasters (pp), distally with thin longitudinal and wrinkled folds. Penial verge (pv) very small and short, situated at distal end of penis (
<figureCitation id="13372A697827A03DFD8CB90D368C5243" box="[587,676,1565,1589]" captionStart="Fig" captionStartId="5.[113,148,1838,1860]" captionTargetBox="[113,1451,270,1823]" captionTargetId="figure-9@5.[113,1451,270,1823]" captionTargetPageId="5" captionText="Fig. 4. Genitalia of Macrochlamys aurantia new species (A–E), and M. coleus new species (F–I). A–E, Macrochlamys aurantia paratype (CUMZ 7103): A, whole reproductive system; B, internal structure of penis with penial verge; C, spermatophore; D, detail of spines between sperm sac and tail filament; E, spines on the end of tail filament. F–I, Macrochlamys coleus paratype (CUMZ 7106): F, whole reproductive system; G, internal structure of penis with penial verge; H, spermatophore; I, head filament. Abbreviations: at, atrium; da, dart apparatus; drm, dart retractor muscle; e, epiphallus; ec, epiphallic caecum; fl, flagellum; fo, free oviduct; gd, gametolytic duct; gs, gametolytic sac; hf, head filament; ov, oviduct; p, penis; pc, penial caecum; pg, prostate gland; pp, penial pilaster; prm, penial retractor muscle; pv, penial verge; ss, sperm sac; tf, tail filament; v, vagina and vd, vas deferens. Asterisk indicates the tissue casting." figureDoi="http://doi.org/10.5281/zenodo.5460644" httpUri="https://zenodo.org/record/5460644/files/figure.png" pageId="6" pageNumber="769">Fig. 4G</figureCitation>
).
</paragraph>
<paragraph id="8BB336EC7827A03DFF4FB94D34D853E3" blockId="6.[135,776,1629,1941]" pageId="6" pageNumber="769">
Vagina (v) slightly long, almost same length as penis and cylindrical. Dart apparatus (da) long, enlarged cylindrical shape, located at proximal end of vagina, with thick connective tissue encircled at proximal end (asterisk in
<figureCitation id="13372A697827A03DFD1AB9AD34805283" captionStart="Fig" captionStartId="5.[113,148,1838,1860]" captionTargetBox="[113,1451,270,1823]" captionTargetId="figure-9@5.[113,1451,270,1823]" captionTargetPageId="5" captionText="Fig. 4. Genitalia of Macrochlamys aurantia new species (A–E), and M. coleus new species (F–I). A–E, Macrochlamys aurantia paratype (CUMZ 7103): A, whole reproductive system; B, internal structure of penis with penial verge; C, spermatophore; D, detail of spines between sperm sac and tail filament; E, spines on the end of tail filament. F–I, Macrochlamys coleus paratype (CUMZ 7106): F, whole reproductive system; G, internal structure of penis with penial verge; H, spermatophore; I, head filament. Abbreviations: at, atrium; da, dart apparatus; drm, dart retractor muscle; e, epiphallus; ec, epiphallic caecum; fl, flagellum; fo, free oviduct; gd, gametolytic duct; gs, gametolytic sac; hf, head filament; ov, oviduct; p, penis; pc, penial caecum; pg, prostate gland; pp, penial pilaster; prm, penial retractor muscle; pv, penial verge; ss, sperm sac; tf, tail filament; v, vagina and vd, vas deferens. Asterisk indicates the tissue casting." figureDoi="http://doi.org/10.5281/zenodo.5460644" httpUri="https://zenodo.org/record/5460644/files/figure.png" pageId="6" pageNumber="769">Fig. 4F</figureCitation>
). Gametolytic sac (gs) elongate and bulbous. Gametolytic duct (gd) long and cylindrical (one spermatophore in
<figureCitation id="13372A697827A03DFD1CB9ED34805343" captionStart="Fig" captionStartId="5.[113,148,1838,1860]" captionTargetBox="[113,1451,270,1823]" captionTargetId="figure-9@5.[113,1451,270,1823]" captionTargetPageId="5" captionText="Fig. 4. Genitalia of Macrochlamys aurantia new species (A–E), and M. coleus new species (F–I). A–E, Macrochlamys aurantia paratype (CUMZ 7103): A, whole reproductive system; B, internal structure of penis with penial verge; C, spermatophore; D, detail of spines between sperm sac and tail filament; E, spines on the end of tail filament. F–I, Macrochlamys coleus paratype (CUMZ 7106): F, whole reproductive system; G, internal structure of penis with penial verge; H, spermatophore; I, head filament. Abbreviations: at, atrium; da, dart apparatus; drm, dart retractor muscle; e, epiphallus; ec, epiphallic caecum; fl, flagellum; fo, free oviduct; gd, gametolytic duct; gs, gametolytic sac; hf, head filament; ov, oviduct; p, penis; pc, penial caecum; pg, prostate gland; pp, penial pilaster; prm, penial retractor muscle; pv, penial verge; ss, sperm sac; tf, tail filament; v, vagina and vd, vas deferens. Asterisk indicates the tissue casting." figureDoi="http://doi.org/10.5281/zenodo.5460644" httpUri="https://zenodo.org/record/5460644/files/figure.png" pageId="6" pageNumber="769">Fig. 4F</figureCitation>
). Free oviduct (fo) short, about similar length to flagellum, proximal end encircled with brownish tissue. Oviduct (ov) large lobules with prostate gland (pg) running alongside (
<figureCitation id="13372A697827A03DFF57B86D34CC53E3" box="[144,228,1917,1941]" captionStart="Fig" captionStartId="5.[113,148,1838,1860]" captionTargetBox="[113,1451,270,1823]" captionTargetId="figure-9@5.[113,1451,270,1823]" captionTargetPageId="5" captionText="Fig. 4. Genitalia of Macrochlamys aurantia new species (A–E), and M. coleus new species (F–I). A–E, Macrochlamys aurantia paratype (CUMZ 7103): A, whole reproductive system; B, internal structure of penis with penial verge; C, spermatophore; D, detail of spines between sperm sac and tail filament; E, spines on the end of tail filament. F–I, Macrochlamys coleus paratype (CUMZ 7106): F, whole reproductive system; G, internal structure of penis with penial verge; H, spermatophore; I, head filament. Abbreviations: at, atrium; da, dart apparatus; drm, dart retractor muscle; e, epiphallus; ec, epiphallic caecum; fl, flagellum; fo, free oviduct; gd, gametolytic duct; gs, gametolytic sac; hf, head filament; ov, oviduct; p, penis; pc, penial caecum; pg, prostate gland; pp, penial pilaster; prm, penial retractor muscle; pv, penial verge; ss, sperm sac; tf, tail filament; v, vagina and vd, vas deferens. Asterisk indicates the tissue casting." figureDoi="http://doi.org/10.5281/zenodo.5460644" httpUri="https://zenodo.org/record/5460644/files/figure.png" pageId="6" pageNumber="769">Fig. 4F</figureCitation>
).
</paragraph>
<paragraph id="8BB336EC7827A03DFF4FB8AD31E85543" blockId="6.[135,775,1981,2101]" lastBlockId="6.[834,1476,189,309]" pageId="6" pageNumber="769">
Spermatophore long, needle-shaped and translucent (
<figureCitation id="13372A697827A03DFD1CB8AD34855383" captionStart="Fig" captionStartId="5.[113,148,1838,1860]" captionTargetBox="[113,1451,270,1823]" captionTargetId="figure-9@5.[113,1451,270,1823]" captionTargetPageId="5" captionText="Fig. 4. Genitalia of Macrochlamys aurantia new species (A–E), and M. coleus new species (F–I). A–E, Macrochlamys aurantia paratype (CUMZ 7103): A, whole reproductive system; B, internal structure of penis with penial verge; C, spermatophore; D, detail of spines between sperm sac and tail filament; E, spines on the end of tail filament. F–I, Macrochlamys coleus paratype (CUMZ 7106): F, whole reproductive system; G, internal structure of penis with penial verge; H, spermatophore; I, head filament. Abbreviations: at, atrium; da, dart apparatus; drm, dart retractor muscle; e, epiphallus; ec, epiphallic caecum; fl, flagellum; fo, free oviduct; gd, gametolytic duct; gs, gametolytic sac; hf, head filament; ov, oviduct; p, penis; pc, penial caecum; pg, prostate gland; pp, penial pilaster; prm, penial retractor muscle; pv, penial verge; ss, sperm sac; tf, tail filament; v, vagina and vd, vas deferens. Asterisk indicates the tissue casting." figureDoi="http://doi.org/10.5281/zenodo.5460644" httpUri="https://zenodo.org/record/5460644/files/figure.png" pageId="6" pageNumber="769">Fig. 4H</figureCitation>
). Head filament (hf) short, gradually tapering to a point (
<figureCitation id="13372A697827A03DFF57B8ED34F15C63" box="[144,217,2045,2069]" captionStart="Fig" captionStartId="5.[113,148,1838,1860]" captionTargetBox="[113,1451,270,1823]" captionTargetId="figure-9@5.[113,1451,270,1823]" captionTargetPageId="5" captionText="Fig. 4. Genitalia of Macrochlamys aurantia new species (A–E), and M. coleus new species (F–I). A–E, Macrochlamys aurantia paratype (CUMZ 7103): A, whole reproductive system; B, internal structure of penis with penial verge; C, spermatophore; D, detail of spines between sperm sac and tail filament; E, spines on the end of tail filament. F–I, Macrochlamys coleus paratype (CUMZ 7106): F, whole reproductive system; G, internal structure of penis with penial verge; H, spermatophore; I, head filament. Abbreviations: at, atrium; da, dart apparatus; drm, dart retractor muscle; e, epiphallus; ec, epiphallic caecum; fl, flagellum; fo, free oviduct; gd, gametolytic duct; gs, gametolytic sac; hf, head filament; ov, oviduct; p, penis; pc, penial caecum; pg, prostate gland; pp, penial pilaster; prm, penial retractor muscle; pv, penial verge; ss, sperm sac; tf, tail filament; v, vagina and vd, vas deferens. Asterisk indicates the tissue casting." figureDoi="http://doi.org/10.5281/zenodo.5460644" httpUri="https://zenodo.org/record/5460644/files/figure.png" pageId="6" pageNumber="769">Fig. 4I</figureCitation>
). Sperm sac (ss) enlarged, cylindrical capsule-shaped with sperm mass. Tail filament (tf) long, curved with many spines. Region close to sperm sac bear a single trichotomous spine divided into three pointed tips. Middle region with a row of five, long curved spines. Region furthest away from sperm sac contains small cluster of short spines (
<figureCitation id="13372A697827A03DFA9BBE0D319C5543" box="[1372,1460,285,309]" captionStart="Fig" captionStartId="5.[113,148,1838,1860]" captionTargetBox="[113,1451,270,1823]" captionTargetId="figure-9@5.[113,1451,270,1823]" captionTargetPageId="5" captionText="Fig. 4. Genitalia of Macrochlamys aurantia new species (A–E), and M. coleus new species (F–I). A–E, Macrochlamys aurantia paratype (CUMZ 7103): A, whole reproductive system; B, internal structure of penis with penial verge; C, spermatophore; D, detail of spines between sperm sac and tail filament; E, spines on the end of tail filament. F–I, Macrochlamys coleus paratype (CUMZ 7106): F, whole reproductive system; G, internal structure of penis with penial verge; H, spermatophore; I, head filament. Abbreviations: at, atrium; da, dart apparatus; drm, dart retractor muscle; e, epiphallus; ec, epiphallic caecum; fl, flagellum; fo, free oviduct; gd, gametolytic duct; gs, gametolytic sac; hf, head filament; ov, oviduct; p, penis; pc, penial caecum; pg, prostate gland; pp, penial pilaster; prm, penial retractor muscle; pv, penial verge; ss, sperm sac; tf, tail filament; v, vagina and vd, vas deferens. Asterisk indicates the tissue casting." figureDoi="http://doi.org/10.5281/zenodo.5460644" httpUri="https://zenodo.org/record/5460644/files/figure.png" pageId="6" pageNumber="769">Fig. 4H</figureCitation>
).
</paragraph>
<paragraph id="8BB336EC7827A03DFC84BE4D318155A3" blockId="6.[835,1474,349,469]" pageId="6" pageNumber="769">
<emphasis id="B978EAFE7827A03DFC84BE4D37B85503" box="[835,912,349,373]" italics="true" pageId="6" pageNumber="769">Radula</emphasis>
. Each row contains about 92 teeth with formula (45- (12-11)-1-(11-12)-46). Central tooth symmetric tricuspid. Lateral teeth asymmetric tricuspid. Marginal teeth obliquely bicuspid starting from tooth number 11–12 (
<figureCitation id="13372A697827A03DFAEFBEAD31B555A3" box="[1320,1437,445,469]" captionStart="Fig" captionStartId="10.[136,171,1215,1237]" captionTargetBox="[136,1474,189,1200]" captionTargetId="figure-7@10.[136,1474,189,1200]" captionTargetPageId="10" captionText="Fig. 6. SEM images of the radula of Macrochlamys aurantia new species (A–C) (paratype; CUMZ 7103), M. coleus new species (D–F) (paratype; CUMZ 7106), and M. caverna new species (G–I) (paratype; CUMZ 7109). Central tooth (indicated by ‘C’) with lateral teeth (A, D, G); lateral teeth with tricuspid marginal teeth transition (B, E, H); bicuspid marginal teeth (C, F, I)." figureDoi="http://doi.org/10.5281/zenodo.5460650" httpUri="https://zenodo.org/record/5460650/files/figure.png" pageId="6" pageNumber="769">Fig. 6D–F</figureCitation>
).
</paragraph>
<paragraph id="8BB336EC7827A03DFC84BEED37D85683" blockId="6.[834,1476,509,757]" pageId="6" pageNumber="769">
<emphasis id="B978EAFE7827A03DFC84BEED37D65663" box="[835,1022,509,533]" italics="true" pageId="6" pageNumber="769">External features</emphasis>
. Animal with reticulated skin. Head, eye stalks and body dark grey, and tail orange (
<figureCitation id="13372A697827A03DFAC8BD0D314B5643" box="[1295,1379,541,565]" captionStart="Fig" captionStartId="2.[135,170,1546,1568]" captionTargetBox="[136,1474,192,1529]" captionTargetId="figure-7@2.[136,1474,189,1531]" captionTargetPageId="2" captionText="Fig. 2. Living snails of Macrochlamys spp. from Thailand. A, Macrochlamys aurantia new species, paratype (CUMZ 7103), Thong Pha Phum, Kanchanaburi, shell width about 15 mm; B, M. coleus new species, paratype (CUMZ 7106), Sai Yok, Kanchanaburi, shell width about 15 mm; C,M. caverna new species, paratype (CUMZ 7109), Tha Wung, Lopburi, shell width about 17 mm; D, M. lemma new species, paratype (CUMZ 7116), Ban Mi, Lopburi, shell width about 15 mm; E, M. tanymentula new species, paratype (CUMZ 7119), Thong Pha Phum, Kanchanaburi, shell width about 10 mm; F, M. kelantanensis (CUMZ 7125), Lam Kaen, Phang-Nga, shell width about 20 mm." figureDoi="http://doi.org/10.5281/zenodo.5460640" httpUri="https://zenodo.org/record/5460640/files/figure.png" pageId="6" pageNumber="769">Fig. 2B</figureCitation>
). Caudal foss (cf) long and narrow. Caudal horn (ch) large and dark grey in colour. Mantle edge (shell lobes and dorsal lobes) well developed, orange with black pigmented spots spread all over. Shell lobes and dorsal lobes similar in morphology to
<taxonomicName id="4C0C4D6F7827A03DFC99BDAE37F456A3" authority="Pholyotha &amp; Sutcharit &amp; Panha, 2018" authorityName="Pholyotha &amp; Sutcharit &amp; Panha" authorityYear="2018" box="[862,988,702,725]" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="769" phylum="Mollusca" rank="species" species="aurantia" status="sp. nov.">
<emphasis id="B978EAFE7827A03DFC99BDAE37F456A3" box="[862,988,702,725]" italics="true" pageId="6" pageNumber="769">M. aurantia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B57857827A03DFC25BDAE304C56A3" box="[994,1124,701,725]" pageId="6" pageNumber="769" rank="species">new species</taxonomicNameLabel>
. Snails secrete light orange slime when disturbed.
</paragraph>
</subSubSection>
<subSubSection id="C31665677827A03DFC84BC0D305657C3" pageId="6" pageNumber="769" type="distribution">
<paragraph id="8BB336EC7827A03DFC84BC0D305657C3" blockId="6.[835,1474,797,949]" pageId="6" pageNumber="769">
<emphasis id="B978EAFE7827A03DFC84BC0D30455743" bold="true" box="[835,1133,797,821]" pageId="6" pageNumber="769">Distribution and habitat.</emphasis>
This new species is currently known only from the
<typeStatus id="54B7884E7827A03DFBF9BC2E30455720" box="[1086,1133,830,854]" pageId="6" pageNumber="769">type</typeStatus>
locality, an area of limestone outcrops with a relatively low population density. The snails were mostly found on decaying leaf litter or sometimes climbing on limestone walls.
</paragraph>
</subSubSection>
<subSubSection id="C31665677827A03DFC84BCCD30935323" pageId="6" pageNumber="769" type="discussion">
<paragraph id="8BB336EC7827A03DFC84BCCD314D5083" blockId="6.[834,1474,989,1270]" pageId="6" pageNumber="769">
<emphasis id="B978EAFE7827A03DFC84BCCD37995783" bold="true" box="[835,945,989,1013]" pageId="6" pageNumber="769">Remarks.</emphasis>
<taxonomicName id="4C0C4D6F7827A03DFC72BCCD30B15783" authority="Pholyotha &amp; Sutcharit &amp; Panha, 2018" authorityName="Pholyotha &amp; Sutcharit &amp; Panha" authorityYear="2018" box="[949,1177,989,1013]" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="769" phylum="Mollusca" rank="species" species="coleus" status="sp. nov.">
<emphasis id="B978EAFE7827A03DFC72BCCD30B15783" box="[949,1177,989,1013]" italics="true" pageId="6" pageNumber="769">Macrochlamys coleus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B57857827A03DFB59BCCE31335783" box="[1182,1307,989,1013]" pageId="6" pageNumber="769" rank="species">new species</taxonomicNameLabel>
is distinguished by a depressed shell, dark brown in colour, and rather shallowly channelled suture. The genitalia have one circle of coiled epiphallic caecum, and dart apparatus with a thick connective tissue encircling the proximal end. Its spermatophore has a single trichotomously branched spine at the proximal end, five long spines in the middle, and a cluster of small spines at the distal end of tail filaments. Its radula morphology is very similar to
<taxonomicName id="4C0C4D6F7827A03DFB19BBCE31765083" authorityName="Pholyotha &amp; Sutcharit &amp; Panha" authorityYear="2018" box="[1246,1374,1246,1269]" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="769" phylum="Mollusca" rank="species" species="aurantia">
<emphasis id="B978EAFE7827A03DFB19BBCE31765083" box="[1246,1374,1246,1269]" italics="true" pageId="6" pageNumber="769">M. aurantia</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BB336EC7827A03DFC84BA0D30935323" blockId="6.[834,1476,1309,1877]" pageId="6" pageNumber="769">
This new species is close to
<taxonomicName id="4C0C4D6F7827A03DFBA0BA0E30CD5143" authority="Pholyotha &amp; Sutcharit &amp; Panha, 2018" authorityName="Pholyotha &amp; Sutcharit &amp; Panha" authorityYear="2018" box="[1127,1253,1310,1333]" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="769" phylum="Mollusca" rank="species" species="aurantia" status="sp. nov.">
<emphasis id="B978EAFE7827A03DFBA0BA0E30CD5143" box="[1127,1253,1310,1333]" italics="true" pageId="6" pageNumber="769">M. aurantia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B57857827A03DFB2BBA0E31455143" box="[1260,1389,1309,1333]" pageId="6" pageNumber="769" rank="species">new species</taxonomicNameLabel>
, but can be differentiated by its small last whorl and shallow channelshaped suture. Anatomically, the genitalia have a short flagellum and well-developed connective tissue surrounding the proximal end of the dart apparatus; characters missing in
<taxonomicName id="4C0C4D6F7827A03DFCA7BAAE37C851A3" authority="Pholyotha &amp; Sutcharit &amp; Panha, 2018" authorityName="Pholyotha &amp; Sutcharit &amp; Panha" authorityYear="2018" box="[864,992,1470,1493]" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="769" phylum="Mollusca" rank="species" species="aurantia" status="sp. nov.">
<emphasis id="B978EAFE7827A03DFCA7BAAE37C851A3" box="[864,992,1470,1493]" italics="true" pageId="6" pageNumber="769">M. aurantia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B57857827A03DFC2FBAAE304451A3" box="[1000,1132,1469,1493]" pageId="6" pageNumber="769" rank="species">new species</taxonomicNameLabel>
. In addition, the spermatophore of this new species has a trichotomous spine on the tail filament close to the sperm sac and a row of long and curved spines on the middle part of the tail filament, while
<taxonomicName id="4C0C4D6F7827A03DFC84B92E37E05223" authority="Pholyotha &amp; Sutcharit &amp; Panha, 2018" authorityName="Pholyotha &amp; Sutcharit &amp; Panha" authorityYear="2018" box="[835,968,1598,1621]" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="769" phylum="Mollusca" rank="species" species="aurantia" status="sp. nov.">
<emphasis id="B978EAFE7827A03DFC84B92E37E05223" box="[835,968,1598,1621]" italics="true" pageId="6" pageNumber="769">M. aurantia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B57857827A03DFC15B92E30705223" box="[978,1112,1597,1621]" pageId="6" pageNumber="769" rank="species">new species</taxonomicNameLabel>
has one dichotomous spine and two simple spines on the tail filament close to the sperm sac. This new species differs from
<taxonomicName id="4C0C4D6F7827A03DFB7CB96E37AB52C3" authority="(Blanford, 1865)" baseAuthorityName="Blanford" baseAuthorityYear="1865" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="769" phylum="Mollusca" rank="species" species="hypoleuca">
<emphasis id="B978EAFE7827A03DFB7CB96E316552E3" box="[1211,1357,1661,1685]" italics="true" pageId="6" pageNumber="769">M. hypoleuca</emphasis>
(
<bibRefCitation id="EF9D4B1D7827A03DFA9CB96D375352C3" author="Blanford WT" pageId="6" pageNumber="769" pagination="66 - 68" refId="ref12581" refString="Blanford WT (1865) Contributions to Indian Malacology, No. V. Descriptions of new land shells from Arakan, Pegu, and Ava; with notes on the distribution of described species. Journal of the Asiatic Society of Bengal, 34: 66 - 68." type="journal article" year="1865">Blanford, 1865</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4C0C4D6F7827A03DFC70B98E30DE52C3" authority="(Blanford, 1865)" baseAuthorityName="Blanford" baseAuthorityYear="1865" box="[951,1270,1693,1717]" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="769" phylum="Mollusca" rank="species" species="nebulosa">
<emphasis id="B978EAFE7827A03DFC70B98E301352C3" box="[951,1083,1693,1717]" italics="true" pageId="6" pageNumber="769">M. nebulosa</emphasis>
(
<bibRefCitation id="EF9D4B1D7827A03DFB8DB98D30C752C3" author="Blanford WT" box="[1098,1263,1693,1717]" pageId="6" pageNumber="769" pagination="66 - 68" refId="ref12581" refString="Blanford WT (1865) Contributions to Indian Malacology, No. V. Descriptions of new land shells from Arakan, Pegu, and Ava; with notes on the distribution of described species. Journal of the Asiatic Society of Bengal, 34: 66 - 68." type="journal article" year="1865">Blanford, 1865</bibRefCitation>
)
</taxonomicName>
from
<collectingCountry id="F31B767C7827A03DFAFEB98E318952C0" box="[1337,1441,1694,1718]" name="Myanmar" pageId="6" pageNumber="769">Myanmar</collectingCountry>
by its depressed shell with a broad last whorl and dark brown shell colour. In comparison,
<taxonomicName id="4C0C4D6F7827A03DFBBAB9CE31385283" baseAuthorityName="Blanford" baseAuthorityYear="1865" box="[1149,1296,1757,1781]" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="769" phylum="Mollusca" rank="species" species="hypoleuca">
<emphasis id="B978EAFE7827A03DFBBAB9CE31385283" box="[1149,1296,1757,1781]" italics="true" pageId="6" pageNumber="769">M. hypoleuca</emphasis>
</taxonomicName>
has a two-toned shell that is brown above the periphery and white below, while
<taxonomicName id="4C0C4D6F7827A03DFC4FB80E30275343" baseAuthorityName="Blanford" baseAuthorityYear="1865" box="[904,1039,1821,1845]" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="769" phylum="Mollusca" rank="species" species="nebulosa">
<emphasis id="B978EAFE7827A03DFC4FB80E30275343" box="[904,1039,1821,1845]" italics="true" pageId="6" pageNumber="769">M. nebulosa</emphasis>
</taxonomicName>
has a conoidal, depressed, shell that is bluntly angulated at the periphery.
</paragraph>
</subSubSection>
</treatment>
</document>
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