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<document id="E0EAAA2BBAAD0F7E4B15307811263756" ID-DOI="10.5281/zenodo.174076" ID-GBIF-Dataset="50e26ef8-c2f5-47ff-a6bd-b6b34323800b" ID-ISSN="1175­5326" ID-Zenodo-Dep="174076" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459789464809" checkinUser="plazi" docAuthor="Schwendinger, Peter J." docDate="2006" docId="03A5879D753CFF94FE985BDAD9F02E6B" docLanguage="en" docName="zt01325p266.pdf" docOrigin="Zootaxa 1325" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Martensiellus tenuipalpus Schwendinger, 2006, n. sp." docType="treatment" docVersion="9" lastPageNumber="262" masterDocId="FF9CFFE5753FFF93FF905E66DD6C2F5D" masterDocTitle="A taxonomic revision of the family Oncopodidae VI. Martensiellus, a new genus from Borneo, and the discovery of a tarsal pore organ in Oncopodidae (Opiliones: Laniatores)" masterLastPageNumber="266" masterPageNumber="255" pageNumber="258" updateTime="1698219045570" updateUser="plazi">
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<mods:title id="1690E2C07409E34A736FE773A62C299A">A taxonomic revision of the family Oncopodidae VI. Martensiellus, a new genus from Borneo, and the discovery of a tarsal pore organ in Oncopodidae (Opiliones: Laniatores)</mods:title>
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<mods:date id="37AD637114BE7A69B1DDA6705F2B49BC">2006</mods:date>
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<paragraph id="8BB3368B753CFF90FE985BDADFE12A8B" blockId="3.[264,653,1468,1529]" box="[264,653,1468,1494]" pageId="3" pageNumber="258">
<heading id="D0FB81E7753CFF90FE985BDADFE12A8B" bold="true" box="[264,653,1468,1494]" fontSize="11" level="1" pageId="3" pageNumber="258" reason="1">
<emphasis id="B978EA99753CFF90FE985BDADFE12A8B" bold="true" box="[264,653,1468,1494]" pageId="3" pageNumber="258">
<taxonomicName id="4C0C4D08753CFF90FE985BDADF2F2A8B" ID-CoL="7X64B" box="[264,579,1468,1494]" class="Arachnida" family="Sandokanidae" genus="Martensiellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="258" phylum="Arthropoda" rank="species" species="tenuipalpus" status="sp. nov.">
<emphasis id="B978EA99753CFF90FE985BDADF2F2A8B" bold="true" box="[264,579,1468,1494]" italics="true" pageId="3" pageNumber="258">Martensiellus tenuipalpus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B57E2753CFF90FDDB5BDADFE12A8B" box="[587,653,1468,1494]" pageId="3" pageNumber="258" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
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</paragraph>
<paragraph id="8BB3368B753CFF90FE985BB9DC102AA4" blockId="3.[264,653,1468,1529]" box="[264,380,1503,1529]" pageId="3" pageNumber="258">
<figureCitation id="13372A0E753CFF90FE985BB9DC102AA4" box="[264,380,1503,1529]" captionStart-0="FIGURES 1 – 3" captionStart-1="FIGURES 4 – 10" captionStart-2="FIGURES 11 – 17" captionStartId-0="2.[264,383,1797,1821]" captionStartId-1="5.[264,383,1011,1035]" captionStartId-2="6.[264,383,1444,1468]" captionTargetBox-0="[278,1303,404,1743]" captionTargetBox-1="[275,1314,521,997]" captionTargetBox-2="[278,1309,534,1410]" captionTargetId-0="figure@2.[264,1323,388,1769]" captionTargetId-1="figure@5.[264,1323,503,1005]" captionTargetId-2="figure@6.[264,1323,518,1422]" captionTargetPageId-0="2" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="FIGURES 1 – 3. Martensiellus tenuipalpus n. g., n. sp., male holotype, habitus (1 dorsal view, 2 ventral view, 3 lateral view). Scale bar: 1 mm." captionText-1="FIGURES 4 – 10. Martensiellus tenuipalpus n. g., n. sp., male (4 – 9 holotype, 10 paratype). 4. Chelicera, lateral view. 5. Palpus, lateral view. 6. Distal parts of legs, lateral view (6 leg I, 7 leg II, 8 leg III, 9 leg IV). 10. Palpus and anterior part of body, lateral view. Scale bars: 1 mm (4 – 9 same scale)." captionText-2="FIGURES 11 – 17. Martensiellus tenuipalpus n. g., n. sp., penis (11 – 14 holotype, 15 – 17 paratype). 11 – 12. Total penis (11 dorsal view, 12 lateral view). 13 – 15, 17. Apex of penis in resting position (13, 15 dorsal view, 14, 17 lateral view). 16. Apex of penis with expanded glans, dorsal view. Abbreviations: d. m. distal margin of truncus penis, l. s. lateral sclerite of glans, m. p. median plate of glans, m. t. membranous tubes of glans, s stylus. Scale bars: 0.5 mm (11 – 12), 0.1 mm (13 – 17)." httpUri-0="https://zenodo.org/record/174077/files/figure.png" httpUri-1="https://zenodo.org/record/174078/files/figure.png" httpUri-2="https://zenodo.org/record/174079/files/figure.png" pageId="3" pageNumber="258">Figs 1–17</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3166500753CFF90FE985842DEA2298C" pageId="3" pageNumber="258" type="description">
<paragraph id="8BB3368B753CFF90FE985842DE822966" blockId="3.[264,1006,1572,1595]" box="[264,1006,1572,1595]" pageId="3" pageNumber="258">
<taxonomicName id="4C0C4D08753CFF90FE985842DCFA2966" box="[264,406,1572,1595]" class="Arachnida" family="Oncopodidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="258" phylum="Arthropoda" rank="family">Oncopodidae</taxonomicName>
g. sp.: 
<bibRefCitation id="EF9D4B7A753CFF90FE745842DE282966" author="Schwendinger" box="[484,836,1572,1595]" pageId="3" pageNumber="266" refString="Schwendinger, P. J. &amp; Martens, J. (2002 b) Penis morphology in Oncopodidae (Opiliones, Laniatores): evolutionary trends and relationships. Journal of Arachnology, 30, 425 - 434." type="journal article" year="2002" yearSuffix="b">Schwendinger &amp; Martens 2002b</bibRefCitation>
: figs 50–51, 59.
</paragraph>
<paragraph id="8BB3368B753CFF90FE985800DEA2298C" blockId="3.[264,1323,1638,1905]" pageId="3" pageNumber="258">
<emphasis id="B978EA99753CFF90FE985800DC2229DD" bold="true" box="[264,334,1638,1664]" pageId="3" pageNumber="258">
<typeStatus id="54B78829753CFF90FE985800DC2229DD" box="[264,334,1638,1664]" pageId="3" pageNumber="258">Types</typeStatus>
</emphasis>
. Male 
<typeStatus id="54B78829753CFF90FE3A5801DF7C29DC" box="[426,528,1639,1665]" pageId="3" pageNumber="258" type="holotype">holotype</typeStatus>
(AMNH) and male 
<typeStatus id="54B78829753CFF90FC9B5801DE1C29DC" box="[779,880,1639,1665]" pageId="3" pageNumber="258" type="paratype">paratype</typeStatus>
(ventral scutum broken; MHNG; ex AMNH, donated by L. Prendini) from Eastern 
<collectingCountry id="F31B761B753CFF90FCAB58E9DEC629F4" box="[827,938,1679,1705]" name="Malaysia" pageId="3" pageNumber="258">Malaysia</collectingCountry>
, 
<collectingCountry id="F31B761B753CFF90FC2758E9D97329F4" box="[951,1055,1679,1705]" name="Malaysia" pageId="3" pageNumber="258">Sarawak</collectingCountry>
, Baleh River, north of Kapit, 
<quantity id="4CF49B6E753CFF90FEC858D1DCFA298C" box="[344,406,1719,1745]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="258" unit="m" value="40.0">40 m</quantity>
, collected 
<date id="FFB2104B753CFF90FD8358D1DFDC298C" box="[531,688,1719,1745]" pageId="3" pageNumber="258" value="1976-06-25">25 June 1976</date>
by Walter C. Sedgwick.
</paragraph>
</subSubSection>
<subSubSection id="C3166500753CFF90FEA858B8DCFA2814" pageId="3" pageNumber="258" type="etymology">
<paragraph id="8BB3368B753CFF90FEA858B8DCFA2814" blockId="3.[264,1323,1638,1905]" pageId="3" pageNumber="258">
<emphasis id="B978EA99753CFF90FEA858B8DCA829A5" bold="true" box="[312,452,1758,1784]" pageId="3" pageNumber="258">Etymology.</emphasis>
The specific epithet, a noun in apposition formed from the Latin 
<emphasis id="B978EA99753CFF90FB5358B9D86529A5" box="[1219,1289,1759,1784]" italics="true" pageId="3" pageNumber="258">tenuis</emphasis>
(= thin, slender) and 
<emphasis id="B978EA99753CFF90FE4A5961DF44287D" box="[474,552,1799,1824]" italics="true" pageId="3" pageNumber="258">palpus</emphasis>
(= feeler), refers to the unusually slender palpal tarsus of males in this species.
</paragraph>
</subSubSection>
<subSubSection id="C3166500753CFF97FEA85930DC892DC1" lastPageId="4" lastPageNumber="259" pageId="3" pageNumber="258" type="diagnosis">
<paragraph id="8BB3368B753CFF97FEA85930DC892DC1" blockId="3.[264,1323,1638,1905]" lastBlockId="4.[264,1324,284,1899]" lastPageId="4" lastPageNumber="259" pageId="3" pageNumber="258">
<emphasis id="B978EA99753CFF90FEA85930DCDC282D" bold="true" box="[312,432,1878,1904]" pageId="3" pageNumber="258">Diagnosis</emphasis>
. Externally similar to 
<taxonomicName id="4C0C4D08753CFF90FD575931DE82282D" box="[711,1006,1879,1904]" class="Arachnida" family="Sandokanidae" genus="Caenoncopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="258" phylum="Arthropoda" rank="species" species="cuspidatus">
<emphasis id="B978EA99753CFF90FD575931DE82282D" box="[711,1006,1879,1904]" italics="true" pageId="3" pageNumber="258">Caenoncopus cuspidatus</emphasis>
</taxonomicName>
(see 
<bibRefCitation id="EF9D4B7A753CFF90FBA55931D84B282C" author="Schwendinger" box="[1077,1319,1879,1905]" pageId="3" pageNumber="266" refString="Schwendinger, P. J. (1992) New Oncopodidae (Opiliones, Laniatores) from Southeast Asia. Revue suisse de Zoologie, 99 (1), 177 - 199." type="journal article" year="1992">Schwendinger 1992</bibRefCitation>
: 190–192, figs 76–80; 
<bibRefCitation id="EF9D4B7A753BFF97FD8C5F7ADECF2E6B" author="Martens" box="[540,931,284,310]" pageId="4" pageNumber="266" refString="Martens, J. &amp; Schwendinger, P. J. (1998) A taxonomic revision of the family Oncopodidae I. New genera and new species of Gnomulus Thorell (Opiliones, Laniatores). Revue suisse de Zoologie, 105 (3), 499 - 555." type="journal article" year="1998">Martens &amp; Schwendinger 1998</bibRefCitation>
: 507, figs 4c–f, 5–8), but with stronger pilosity; no carapace­opisthosoma bridge; eye mound (tubercle) distinctly elevated; carapace region fairly large; ventral scutal areas distinctly elevated; anteroproximal process on coxa III present; genital operculum small (
<figureCitation id="13372A0E753BFF97FBC75FF5D9AF2EF0" box="[1111,1219,403,429]" captionStart="FIGURES 1 – 3" captionStartId="2.[264,383,1797,1821]" captionTargetBox="[278,1303,404,1743]" captionTargetId="figure@2.[264,1323,388,1769]" captionTargetPageId="2" captionText="FIGURES 1 – 3. Martensiellus tenuipalpus n. g., n. sp., male holotype, habitus (1 dorsal view, 2 ventral view, 3 lateral view). Scale bar: 1 mm." httpUri="https://zenodo.org/record/174077/files/figure.png" pageId="4" pageNumber="259">Figs 1–3</figureCitation>
); palpus with distinct ventral process on trochanter, no process on femur, tarsus exceptionally thin in distal portion; leg tarsi/tarsomeres (
<figureCitation id="13372A0E753BFF97FD755F84DE0F2EA1" box="[741,867,482,508]" captionStart="FIGURES 4 – 10" captionStartId="5.[264,383,1011,1035]" captionTargetBox="[275,1314,521,997]" captionTargetId="figure@5.[264,1323,503,1005]" captionTargetPageId="5" captionText="FIGURES 4 – 10. Martensiellus tenuipalpus n. g., n. sp., male (4 – 9 holotype, 10 paratype). 4. Chelicera, lateral view. 5. Palpus, lateral view. 6. Distal parts of legs, lateral view (6 leg I, 7 leg II, 8 leg III, 9 leg IV). 10. Palpus and anterior part of body, lateral view. Scale bars: 1 mm (4 – 9 same scale)." httpUri="https://zenodo.org/record/174078/files/figure.png" pageId="4" pageNumber="259">Figs 4–10</figureCitation>
) much longer and larger than in 
<taxonomicName id="4C0C4D08753BFF97FA805F85DCE92D7E" class="Arachnida" family="Sandokanidae" genus="Caenoncopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="259" phylum="Arthropoda" rank="species" species="cuspidatus">
<emphasis id="B978EA99753BFF97FA805F85DCE92D7E" italics="true" pageId="4" pageNumber="259">C. cuspidatus</emphasis>
</taxonomicName>
. Penis similar to that of 
<taxonomicName id="4C0C4D08753BFF97FD2B5C6CD99D2D79" authority="Martens &amp; Schwendinger" authorityName="Martens &amp; Schwendinger" box="[699,1265,522,548]" class="Arachnida" family="Sandokanidae" genus="Biantoncopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="259" phylum="Arthropoda" rank="species" species="fuscus">
<emphasis id="B978EA99753BFF97FD2B5C6CDEDD2D7E" box="[699,945,522,547]" italics="true" pageId="4" pageNumber="259">Biantoncopus fuscus</emphasis>
Martens &amp; Schwendinger
</taxonomicName>
(see 
<bibRefCitation id="EF9D4B7A753BFF97FE985C54DFE02D11" author="Martens" box="[264,652,562,588]" pageId="4" pageNumber="266" refString="Martens, J. &amp; Schwendinger, P. J. (1998) A taxonomic revision of the family Oncopodidae I. New genera and new species of Gnomulus Thorell (Opiliones, Laniatores). Revue suisse de Zoologie, 105 (3), 499 - 555." type="journal article" year="1998">Martens &amp; Schwendinger 1998</bibRefCitation>
: 521–525, figs 58–70), but glans wider, situated more distally, with indistinct membranous socket, wider lateral sclerites and narrower median plate (
<figureCitation id="13372A0E753BFF97FEC05CE4DCB42DC1" box="[336,472,642,668]" captionStart="FIGURES 11 – 17" captionStartId="6.[264,383,1444,1468]" captionTargetBox="[278,1309,534,1410]" captionTargetId="figure@6.[264,1323,518,1422]" captionTargetPageId="6" captionText="FIGURES 11 – 17. Martensiellus tenuipalpus n. g., n. sp., penis (11 – 14 holotype, 15 – 17 paratype). 11 – 12. Total penis (11 dorsal view, 12 lateral view). 13 – 15, 17. Apex of penis in resting position (13, 15 dorsal view, 14, 17 lateral view). 16. Apex of penis with expanded glans, dorsal view. Abbreviations: d. m. distal margin of truncus penis, l. s. lateral sclerite of glans, m. p. median plate of glans, m. t. membranous tubes of glans, s stylus. Scale bars: 0.5 mm (11 – 12), 0.1 mm (13 – 17)." httpUri="https://zenodo.org/record/174079/files/figure.png" pageId="4" pageNumber="259">Figs 11–17</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3166500753BFF95FEA85CCFD87A282C" lastPageId="6" lastPageNumber="261" pageId="4" pageNumber="259" type="description">
<paragraph id="8BB3368B753BFF97FEA85CCFDC202C3F" blockId="4.[264,1324,284,1899]" pageId="4" pageNumber="259">
<emphasis id="B978EA99753BFF97FEA85CCFDCBC2D9E" bold="true" box="[312,464,681,707]" pageId="4" pageNumber="259">Description.</emphasis>
Male (
<typeStatus id="54B78829753BFF97FDB95CCFDFFB2D9E" box="[553,663,681,707]" pageId="4" pageNumber="259" type="holotype">holotype</typeStatus>
). Colouration (in alcohol) and surface texture: Ground colour light orange, with indistinct darker pattern on dorsal scutal elevations (probably lightened by long­time preservation in alcohol; 
<figureCitation id="13372A0E753BFF97FCCD5C9FDECB2C4E" box="[861,935,761,787]" captionStart="FIGURES 1 – 3" captionStartId="2.[264,383,1797,1821]" captionTargetBox="[278,1303,404,1743]" captionTargetId="figure@2.[264,1323,388,1769]" captionTargetPageId="2" captionText="FIGURES 1 – 3. Martensiellus tenuipalpus n. g., n. sp., male holotype, habitus (1 dorsal view, 2 ventral view, 3 lateral view). Scale bar: 1 mm." httpUri="https://zenodo.org/record/174077/files/figure.png" pageId="4" pageNumber="259">Fig. 1</figureCitation>
). A finely granular surface on prosoma, genital operculum, chelicera (except for fingers), legs and palpus (except for tarsi).
</paragraph>
<paragraph id="8BB3368B753BFF97FEA85D16D99A2B95" blockId="4.[264,1324,284,1899]" pageId="4" pageNumber="259">
Carapace region with rounded, slightly anteriad­inclined eye tubercle carrying a small pointed tip; carapace­opisthosoma bridge absent; no lateral tubercles in posterior part of carapace region. Dorsal scutal areas of opisthosoma region only indistinctly elevated; ventral scutal areas moderately swollen and pallid (
<figureCitation id="13372A0E753BFF97FC295D81D9602B5C" box="[953,1036,999,1025]" captionStart="FIGURES 1 – 3" captionStartId="2.[264,383,1797,1821]" captionTargetBox="[278,1303,404,1743]" captionTargetId="figure@2.[264,1323,388,1769]" captionTargetPageId="2" captionText="FIGURES 1 – 3. Martensiellus tenuipalpus n. g., n. sp., male holotype, habitus (1 dorsal view, 2 ventral view, 3 lateral view). Scale bar: 1 mm." httpUri="https://zenodo.org/record/174077/files/figure.png" pageId="4" pageNumber="259">Fig. 3</figureCitation>
). Ventral scutum and opisthosoma region of dorsal scutum densely covered by short hairs (
<figureCitation id="13372A0E753BFF97FBCA5A69D9AB2B74" box="[1114,1223,1039,1065]" captionStart="FIGURES 1 – 3" captionStartId="2.[264,383,1797,1821]" captionTargetBox="[278,1303,404,1743]" captionTargetId="figure@2.[264,1323,388,1769]" captionTargetPageId="2" captionText="FIGURES 1 – 3. Martensiellus tenuipalpus n. g., n. sp., male holotype, habitus (1 dorsal view, 2 ventral view, 3 lateral view). Scale bar: 1 mm." httpUri="https://zenodo.org/record/174077/files/figure.png" pageId="4" pageNumber="259">Figs 1–3</figureCitation>
). Palpal coxa with small ventral process; leg coxa I with indistinct anterolateral process; leg coxae II and III with distinct anteroproximal processes, no posteroproximal process on coxa II (
<figureCitation id="13372A0E753BFF97FE815AE0DC0C2BFD" box="[273,352,1158,1184]" captionStart="FIGURES 1 – 3" captionStartId="2.[264,383,1797,1821]" captionTargetBox="[278,1303,404,1743]" captionTargetId="figure@2.[264,1323,388,1769]" captionTargetPageId="2" captionText="FIGURES 1 – 3. Martensiellus tenuipalpus n. g., n. sp., male holotype, habitus (1 dorsal view, 2 ventral view, 3 lateral view). Scale bar: 1 mm." httpUri="https://zenodo.org/record/174077/files/figure.png" pageId="4" pageNumber="259">Fig. 2</figureCitation>
); dorsal side of leg coxa IV without tubercles. Genital operculum somewhat triangular, wider than long; posterior margin of stigmatic pit without tubercle (
<figureCitation id="13372A0E753BFF97FB305AC8D9842B95" box="[1184,1256,1198,1224]" captionStart="FIGURES 1 – 3" captionStartId="2.[264,383,1797,1821]" captionTargetBox="[278,1303,404,1743]" captionTargetId="figure@2.[264,1323,388,1769]" captionTargetPageId="2" captionText="FIGURES 1 – 3. Martensiellus tenuipalpus n. g., n. sp., male holotype, habitus (1 dorsal view, 2 ventral view, 3 lateral view). Scale bar: 1 mm." httpUri="https://zenodo.org/record/174077/files/figure.png" pageId="4" pageNumber="259">Fig. 2</figureCitation>
).
</paragraph>
<paragraph id="8BB3368B753BFF97FEA85AB0D95C2A4A" blockId="4.[264,1324,284,1899]" pageId="4" pageNumber="259">
Chelicera (
<figureCitation id="13372A0E753BFF97FE285AB0DF6C2BAD" box="[440,512,1238,1264]" captionStart="FIGURES 4 – 10" captionStartId="5.[264,383,1011,1035]" captionTargetBox="[275,1314,521,997]" captionTargetId="figure@5.[264,1323,503,1005]" captionTargetPageId="5" captionText="FIGURES 4 – 10. Martensiellus tenuipalpus n. g., n. sp., male (4 – 9 holotype, 10 paratype). 4. Chelicera, lateral view. 5. Palpus, lateral view. 6. Distal parts of legs, lateral view (6 leg I, 7 leg II, 8 leg III, 9 leg IV). 10. Palpus and anterior part of body, lateral view. Scale bars: 1 mm (4 – 9 same scale)." httpUri="https://zenodo.org/record/174078/files/figure.png" pageId="4" pageNumber="259">Fig. 4</figureCitation>
): Hand weak, with a few small hairs dorsally; proximal article with a higher dorsodistal and a lower dorsomedian boss, no ventral process.
</paragraph>
<paragraph id="8BB3368B753BFF97FEA85B43DCCB2AEB" blockId="4.[264,1324,284,1899]" pageId="4" pageNumber="259">
Palpus (
<figureCitation id="13372A0E753BFF97FE075B43DC8C2A62" box="[407,480,1317,1343]" captionStart="FIGURES 4 – 10" captionStartId="5.[264,383,1011,1035]" captionTargetBox="[275,1314,521,997]" captionTargetId="figure@5.[264,1323,503,1005]" captionTargetPageId="5" captionText="FIGURES 4 – 10. Martensiellus tenuipalpus n. g., n. sp., male (4 – 9 holotype, 10 paratype). 4. Chelicera, lateral view. 5. Palpus, lateral view. 6. Distal parts of legs, lateral view (6 leg I, 7 leg II, 8 leg III, 9 leg IV). 10. Palpus and anterior part of body, lateral view. Scale bars: 1 mm (4 – 9 same scale)." httpUri="https://zenodo.org/record/174078/files/figure.png" pageId="4" pageNumber="259">Fig. 5</figureCitation>
): Trochanter with slightly distad­inclined ventral process; femur short, with indistinct, broadly rounded ventroproximal boss, no process; tarsus cylindrical, exceptionally thin in distal half, there about as deep and wide as claw (possibly a sexual dimorphism).
</paragraph>
<paragraph id="8BB3368B753BFF97FEA85BA2DF5D2970" blockId="4.[264,1324,284,1899]" pageId="4" pageNumber="259">
Legs (
<figureCitation id="13372A0E753BFF97FE125BA2DC872A83" box="[386,491,1476,1502]" captionStart="FIGURES 4 – 10" captionStartId="5.[264,383,1011,1035]" captionTargetBox="[275,1314,521,997]" captionTargetId="figure@5.[264,1323,503,1005]" captionTargetPageId="5" captionText="FIGURES 4 – 10. Martensiellus tenuipalpus n. g., n. sp., male (4 – 9 holotype, 10 paratype). 4. Chelicera, lateral view. 5. Palpus, lateral view. 6. Distal parts of legs, lateral view (6 leg I, 7 leg II, 8 leg III, 9 leg IV). 10. Palpus and anterior part of body, lateral view. Scale bars: 1 mm (4 – 9 same scale)." httpUri="https://zenodo.org/record/174078/files/figure.png" pageId="4" pageNumber="259">Figs 6–9</figureCitation>
) 1342 (from shortest to longest), tarsal formula 1­1­2­2; tarsi I and II somewhat ovoid (
<figureCitation id="13372A0E753BFF97FE4F5B8ADF21295B" box="[479,589,1516,1542]" captionStart="FIGURES 4 – 10" captionStartId="5.[264,383,1011,1035]" captionTargetBox="[275,1314,521,997]" captionTargetId="figure@5.[264,1323,503,1005]" captionTargetPageId="5" captionText="FIGURES 4 – 10. Martensiellus tenuipalpus n. g., n. sp., male (4 – 9 holotype, 10 paratype). 4. Chelicera, lateral view. 5. Palpus, lateral view. 6. Distal parts of legs, lateral view (6 leg I, 7 leg II, 8 leg III, 9 leg IV). 10. Palpus and anterior part of body, lateral view. Scale bars: 1 mm (4 – 9 same scale)." httpUri="https://zenodo.org/record/174078/files/figure.png" pageId="4" pageNumber="259">Figs 6–7</figureCitation>
), with a deep pore dorsally; tarsus of leg II about 1.6 times longer than deep (
<figureCitation id="13372A0E753BFF97FE4B5875DF4F2970" box="[475,547,1555,1581]" captionStart="FIGURES 4 – 10" captionStartId="5.[264,383,1011,1035]" captionTargetBox="[275,1314,521,997]" captionTargetId="figure@5.[264,1323,503,1005]" captionTargetPageId="5" captionText="FIGURES 4 – 10. Martensiellus tenuipalpus n. g., n. sp., male (4 – 9 holotype, 10 paratype). 4. Chelicera, lateral view. 5. Palpus, lateral view. 6. Distal parts of legs, lateral view (6 leg I, 7 leg II, 8 leg III, 9 leg IV). 10. Palpus and anterior part of body, lateral view. Scale bars: 1 mm (4 – 9 same scale)." httpUri="https://zenodo.org/record/174078/files/figure.png" pageId="4" pageNumber="259">Fig. 7</figureCitation>
).
</paragraph>
<paragraph id="8BB3368B753BFF96FEA8585DD9EB2E88" blockId="4.[264,1324,284,1899]" lastBlockId="5.[264,1325,284,469]" lastPageId="5" lastPageNumber="260" pageId="4" pageNumber="259">
Penis (
<figureCitation id="13372A0E753BFF97FE1C585DDF762908" box="[396,538,1595,1621]" captionStart="FIGURES 11 – 17" captionStartId="6.[264,383,1444,1468]" captionTargetBox="[278,1309,534,1410]" captionTargetId="figure@6.[264,1323,518,1422]" captionTargetPageId="6" captionText="FIGURES 11 – 17. Martensiellus tenuipalpus n. g., n. sp., penis (11 – 14 holotype, 15 – 17 paratype). 11 – 12. Total penis (11 dorsal view, 12 lateral view). 13 – 15, 17. Apex of penis in resting position (13, 15 dorsal view, 14, 17 lateral view). 16. Apex of penis with expanded glans, dorsal view. Abbreviations: d. m. distal margin of truncus penis, l. s. lateral sclerite of glans, m. p. median plate of glans, m. t. membranous tubes of glans, s stylus. Scale bars: 0.5 mm (11 – 12), 0.1 mm (13 – 17)." httpUri="https://zenodo.org/record/174079/files/figure.png" pageId="4" pageNumber="259">Figs 11–14</figureCitation>
): Truncus fairly robust, slightly constricted midway, from there widening to a second moderate constriction below glans; distal margin of truncus widely rounded; several subapical setae laterally, two on each side separated from other setae and flanking proximal part of glans. Glans distad­directed, wider than truncus at that point, slightly projecting beyond distal margin of truncus; lateral sclerites wide, somewhat scoop­shaped, their outer part flat and their inner part projecting dorsad; median plate rectangular, longer than wide (visible only in expanded state; see 
<figureCitation id="13372A0E753BFF97FBB7594FD911281E" box="[1063,1149,1833,1859]" captionStart="FIGURES 11 – 17" captionStartId="6.[264,383,1444,1468]" captionTargetBox="[278,1309,534,1410]" captionTargetId="figure@6.[264,1323,518,1422]" captionTargetPageId="6" captionText="FIGURES 11 – 17. Martensiellus tenuipalpus n. g., n. sp., penis (11 – 14 holotype, 15 – 17 paratype). 11 – 12. Total penis (11 dorsal view, 12 lateral view). 13 – 15, 17. Apex of penis in resting position (13, 15 dorsal view, 14, 17 lateral view). 16. Apex of penis with expanded glans, dorsal view. Abbreviations: d. m. distal margin of truncus penis, l. s. lateral sclerite of glans, m. p. median plate of glans, m. t. membranous tubes of glans, s stylus. Scale bars: 0.5 mm (11 – 12), 0.1 mm (13 – 17)." httpUri="https://zenodo.org/record/174079/files/figure.png" pageId="4" pageNumber="259">Fig. 16</figureCitation>
for 
<typeStatus id="54B78829753BFF97FB25594FD873281E" box="[1205,1311,1833,1859]" pageId="4" pageNumber="259" type="paratype">paratype</typeStatus>
), lying dorsally of lateral sclerites (as in 
<taxonomicName id="4C0C4D08753BFF97FD7C5937DEE12837" box="[748,909,1873,1898]" class="Arachnida" family="Sandokanidae" genus="Biantoncopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="259" phylum="Arthropoda" rank="genus">
<emphasis id="B978EA99753BFF97FD7C5937DEE12837" box="[748,909,1873,1898]" italics="true" pageId="4" pageNumber="259">Biantoncopus</emphasis>
</taxonomicName>
, but not in 
<taxonomicName id="4C0C4D08753BFF97FBB35937D9A52837" box="[1059,1225,1873,1898]" class="Arachnida" family="Sandokanidae" genus="Palaeoncopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="259" phylum="Arthropoda" rank="genus">
<emphasis id="B978EA99753BFF97FBB35937D9A52837" box="[1059,1225,1873,1898]" italics="true" pageId="4" pageNumber="259">Palaeoncopus</emphasis>
</taxonomicName>
); stylus slender, with bifid tip but without pair of subapical teeth; membranous tubes wide, distally truncate and bent dorsad, lying between stylus and a somewhat pentagonal median knob. In expanded state (see 
<figureCitation id="13372A0E753AFF96FD8D5F0DDF1E2ED8" box="[541,626,363,389]" captionStart="FIGURES 11 – 17" captionStartId="6.[264,383,1444,1468]" captionTargetBox="[278,1309,534,1410]" captionTargetId="figure@6.[264,1323,518,1422]" captionTargetPageId="6" captionText="FIGURES 11 – 17. Martensiellus tenuipalpus n. g., n. sp., penis (11 – 14 holotype, 15 – 17 paratype). 11 – 12. Total penis (11 dorsal view, 12 lateral view). 13 – 15, 17. Apex of penis in resting position (13, 15 dorsal view, 14, 17 lateral view). 16. Apex of penis with expanded glans, dorsal view. Abbreviations: d. m. distal margin of truncus penis, l. s. lateral sclerite of glans, m. p. median plate of glans, m. t. membranous tubes of glans, s stylus. Scale bars: 0.5 mm (11 – 12), 0.1 mm (13 – 17)." httpUri="https://zenodo.org/record/174079/files/figure.png" pageId="5" pageNumber="260">Fig. 16</figureCitation>
for 
<typeStatus id="54B78829753AFF96FD385F0DDE632ED8" box="[680,783,363,389]" pageId="5" pageNumber="260" type="paratype">paratype</typeStatus>
) median plate and stylus of glans protruding distad, and membranous tubes folded posteriad, as in the penis of 
<taxonomicName id="4C0C4D08753AFF96FBA55FF5DF592E88" authority="Martens &amp; Schwendinger" authorityName="Martens &amp; Schwendinger" class="Arachnida" family="Sandokanidae" genus="Biantoncopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="260" phylum="Arthropoda" rank="species" species="fuscus">
<emphasis id="B978EA99753AFF96FBA55FF5D8472EF1" box="[1077,1323,403,428]" italics="true" pageId="5" pageNumber="260">Biantoncopus fuscus</emphasis>
Martens &amp; Schwendinger
</taxonomicName>
(see 
<bibRefCitation id="EF9D4B7A753AFF96FDE35FDDDE8A2E88" author="Martens" box="[627,998,443,469]" pageId="5" pageNumber="266" refString="Martens, J. &amp; Schwendinger, P. J. (1998) A taxonomic revision of the family Oncopodidae I. New genera and new species of Gnomulus Thorell (Opiliones, Laniatores). Revue suisse de Zoologie, 105 (3), 499 - 555." type="journal article" year="1998">Martens &amp; Schwendinger 1998</bibRefCitation>
: figs 64– 65).
</paragraph>
<caption id="DF736603753AFF96FE985D95DC212B2E" httpUri="https://zenodo.org/record/174078/files/figure.png" pageId="5" pageNumber="260" targetBox="[275,1314,521,997]" targetPageId="5">
<paragraph id="8BB3368B753AFF96FE985D95DC212B2E" blockId="5.[264,1323,1011,1139]" pageId="5" pageNumber="260">
<emphasis id="B978EA99753AFF96FE985D95DCAA2B56" bold="true" box="[264,454,1011,1035]" pageId="5" pageNumber="260">FIGURES 4–10.</emphasis>
<taxonomicName id="4C0C4D08753AFF96FE415D92DF872B56" box="[465,747,1012,1035]" class="Arachnida" family="Sandokanidae" genus="Martensiellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="260" phylum="Arthropoda" rank="species" species="tenuipalpus">
<emphasis id="B978EA99753AFF96FE415D92DF872B56" box="[465,747,1012,1035]" italics="true" pageId="5" pageNumber="260">Martensiellus tenuipalpus</emphasis>
</taxonomicName>
<emphasis id="B978EA99753AFF96FD665D95DE112B56" bold="true" box="[758,893,1011,1035]" pageId="5" pageNumber="260">n. g., n. sp.</emphasis>
, male (4–9 holotype, 10 paratype). 4. Chelicera, lateral view. 5. Palpus, lateral view. 6. Distal parts of legs, lateral view (6 leg I, 7 leg II, 8 leg III, 9 leg IV). 10. Palpus and anterior part of body, lateral view. Scale bars: 1 mm (4–9 same scale).
</paragraph>
</caption>
<paragraph id="8BB3368B753AFF96FEA85AC0DC532A4D" blockId="5.[264,1322,1190,1296]" pageId="5" pageNumber="260">
Measurements of male 
<typeStatus id="54B78829753AFF96FDCA5AC0DFAC2B9D" box="[602,704,1190,1216]" pageId="5" pageNumber="260" type="holotype">holotype</typeStatus>
(male 
<typeStatus id="54B78829753AFF96FC8C5AC0DEED2B9D" box="[796,897,1190,1216]" pageId="5" pageNumber="260" type="paratype">paratype</typeStatus>
in parentheses): Body 3.51 (3.47) long, 2.18 (2.18) wide; carapace region 1.04 (1.02) long, 1.30 (1.30) wide; palpus and legs:
</paragraph>
<paragraph id="8BB3368B753AFF96FE1D5B39D9952901" pageId="5" pageNumber="260">
<table id="F90CC42B753A006CFE9D5B39D86C2901" box="[269,1280,1375,1628]" gridcols="8" gridrows="6" pageId="5" pageNumber="260">
<tr id="353C34C9753A006CFE9D5B39D86C2A2B" box="[269,1280,1375,1398]" gridrow="0" pageId="5" pageNumber="260" rowspan-0="1">
<th id="76ED5DB5753A006CFE1D5B39DF6D2A2B" box="[397,513,1375,1398]" gridcol="1" gridrow="0" pageId="5" pageNumber="260">Trochanter</th>
<th id="76ED5DB5753A006CFDD35B39DFE52A2B" box="[579,649,1375,1398]" gridcol="2" gridrow="0" pageId="5" pageNumber="260">Femur</th>
<th id="76ED5DB5753A006CFD2B5B39DE682A2B" box="[699,772,1375,1398]" gridcol="3" gridrow="0" pageId="5" pageNumber="260">Patella</th>
<th id="76ED5DB5753A006CFCA65B39DE012A2B" box="[822,877,1375,1398]" gridcol="4" gridrow="0" pageId="5" pageNumber="260">Tibia</th>
<th id="76ED5DB5753A006CFC0C5B39D97D2A2B" box="[924,1041,1375,1398]" gridcol="5" gridrow="0" pageId="5" pageNumber="260">Metatarsus</th>
<th id="76ED5DB5753A006CFBC25B39D9FB2A2B" box="[1106,1175,1375,1398]" gridcol="6" gridrow="0" pageId="5" pageNumber="260">Tarsus</th>
<th id="76ED5DB5753A006CFB5A5B39D86C2A2B" box="[1226,1280,1375,1398]" gridcol="7" gridrow="0" pageId="5" pageNumber="260">Total</th>
</tr>
<tr id="353C34C9753A006CFE9D5BE8D86C2AFB" box="[269,1280,1422,1446]" gridrow="1" pageId="5" pageNumber="260">
<th id="76ED5DB5753A006CFE9D5BE8DC362AFB" box="[269,346,1422,1446]" gridcol="0" gridrow="1" pageId="5" pageNumber="260">Palpus</th>
<td id="76ED5DB5753A006CFE1D5BE8DF6D2AFB" box="[397,513,1422,1446]" gridcol="1" gridrow="1" pageId="5" pageNumber="260">0.51</td>
<td id="76ED5DB5753A006CFDD35BE8DFE52AFB" box="[579,649,1422,1446]" gridcol="2" gridrow="1" pageId="5" pageNumber="260">0.63</td>
<td id="76ED5DB5753A006CFD2B5BE8DE682AFB" box="[699,772,1422,1446]" gridcol="3" gridrow="1" pageId="5" pageNumber="260">0.47</td>
<td id="76ED5DB5753A006CFCA65BE8DE012AFB" box="[822,877,1422,1446]" gridcol="4" gridrow="1" pageId="5" pageNumber="260">0.44</td>
<td id="76ED5DB5753A006CFC0C5BE8D97D2AFB" box="[924,1041,1422,1446]" gridcol="5" gridrow="1" pageId="5" pageNumber="260">
<emphasis id="B978EA99753AFF96FC0C5BE8DEC92AFB" bold="true" box="[924,933,1422,1446]" pageId="5" pageNumber="260">­</emphasis>
</td>
<td id="76ED5DB5753A006CFBC25BE8D9FB2AFB" box="[1106,1175,1422,1446]" gridcol="6" gridrow="1" pageId="5" pageNumber="260">1.17</td>
<td id="76ED5DB5753A006CFB5A5BE8D86C2AFB" box="[1226,1280,1422,1446]" gridcol="7" gridrow="1" pageId="5" pageNumber="260">3.22</td>
</tr>
<tr id="353C34C9753A006CFE9D5BDAD86C2A8E" box="[269,1280,1468,1491]" gridrow="2" pageId="5" pageNumber="260">
<th id="76ED5DB5753A006CFE9D5BDADC362A8E" box="[269,346,1468,1491]" gridcol="0" gridrow="2" pageId="5" pageNumber="260">Leg I</th>
<td id="76ED5DB5753A006CFE1D5BDADF6D2A8E" box="[397,513,1468,1491]" gridcol="1" gridrow="2" pageId="5" pageNumber="260">0.38</td>
<td id="76ED5DB5753A006CFDD35BDADFE52A8E" box="[579,649,1468,1491]" gridcol="2" gridrow="2" pageId="5" pageNumber="260">0.85</td>
<td id="76ED5DB5753A006CFD2B5BDADE682A8E" box="[699,772,1468,1491]" gridcol="3" gridrow="2" pageId="5" pageNumber="260">0.57</td>
<td id="76ED5DB5753A006CFCA65BDADE012A8E" box="[822,877,1468,1491]" gridcol="4" gridrow="2" pageId="5" pageNumber="260">0.47</td>
<td id="76ED5DB5753A006CFC0C5BDAD97D2A8E" box="[924,1041,1468,1491]" gridcol="5" gridrow="2" pageId="5" pageNumber="260">0.70</td>
<td id="76ED5DB5753A006CFBC25BDAD9FB2A8E" box="[1106,1175,1468,1491]" gridcol="6" gridrow="2" pageId="5" pageNumber="260">0.40</td>
<td id="76ED5DB5753A006CFB5A5BDAD86C2A8E" box="[1226,1280,1468,1491]" gridcol="7" gridrow="2" pageId="5" pageNumber="260">3.37</td>
</tr>
<tr id="353C34C9753A006CFE9D5B8CD86C295C" box="[269,1280,1514,1537]" gridrow="3" pageId="5" pageNumber="260">
<th id="76ED5DB5753A006CFE9D5B8CDC36295C" box="[269,346,1514,1537]" gridcol="0" gridrow="3" pageId="5" pageNumber="260">Leg II</th>
<td id="76ED5DB5753A006CFE1D5B8CDF6D295C" box="[397,513,1514,1537]" gridcol="1" gridrow="3" pageId="5" pageNumber="260">0.51</td>
<td id="76ED5DB5753A006CFDD35B8CDFE5295C" box="[579,649,1514,1537]" gridcol="2" gridrow="3" pageId="5" pageNumber="260">1.17</td>
<td id="76ED5DB5753A006CFD2B5B8CDE68295C" box="[699,772,1514,1537]" gridcol="3" gridrow="3" pageId="5" pageNumber="260">0.82</td>
<td id="76ED5DB5753A006CFCA65B8CDE01295C" box="[822,877,1514,1537]" gridcol="4" gridrow="3" pageId="5" pageNumber="260">0.73</td>
<td id="76ED5DB5753A006CFC0C5B8CD97D295C" box="[924,1041,1514,1537]" gridcol="5" gridrow="3" pageId="5" pageNumber="260">1.14</td>
<td id="76ED5DB5753A006CFBC25B8CD9FB295C" box="[1106,1175,1514,1537]" gridcol="6" gridrow="3" pageId="5" pageNumber="260">0.44</td>
<td id="76ED5DB5753A006CFB5A5B8CD86C295C" box="[1226,1280,1514,1537]" gridcol="7" gridrow="3" pageId="5" pageNumber="260">4.81</td>
</tr>
<tr id="353C34C9753A006CFE9D5871D86C2973" box="[269,1280,1559,1582]" gridrow="4" pageId="5" pageNumber="260">
<th id="76ED5DB5753A006CFE9D5871DC362973" box="[269,346,1559,1582]" gridcol="0" gridrow="4" pageId="5" pageNumber="260">Leg III</th>
<td id="76ED5DB5753A006CFE1D5871DF6D2973" box="[397,513,1559,1582]" gridcol="1" gridrow="4" pageId="5" pageNumber="260">0.38</td>
<td id="76ED5DB5753A006CFDD35871DFE52973" box="[579,649,1559,1582]" gridcol="2" gridrow="4" pageId="5" pageNumber="260">0.81</td>
<td id="76ED5DB5753A006CFD2B5871DE682973" box="[699,772,1559,1582]" gridcol="3" gridrow="4" pageId="5" pageNumber="260">0.63</td>
<td id="76ED5DB5753A006CFCA65871DE012973" box="[822,877,1559,1582]" gridcol="4" gridrow="4" pageId="5" pageNumber="260">0.51</td>
<td id="76ED5DB5753A006CFC0C5871D97D2973" box="[924,1041,1559,1582]" gridcol="5" gridrow="4" pageId="5" pageNumber="260">0.88</td>
<td id="76ED5DB5753A006CFBC25871D9FB2973" box="[1106,1175,1559,1582]" gridcol="6" gridrow="4" pageId="5" pageNumber="260">0.35</td>
<td id="76ED5DB5753A006CFB5A5871D86C2973" box="[1226,1280,1559,1582]" gridcol="7" gridrow="4" pageId="5" pageNumber="260">3.56</td>
</tr>
<tr id="353C34C9753A006CFE9D5823D86C2901" box="[269,1280,1605,1628]" gridrow="5" pageId="5" pageNumber="260">
<th id="76ED5DB5753A006CFE9D5823DC362901" box="[269,346,1605,1628]" gridcol="0" gridrow="5" pageId="5" pageNumber="260">Leg IV</th>
<td id="76ED5DB5753A006CFE1D5823DF6D2901" box="[397,513,1605,1628]" gridcol="1" gridrow="5" pageId="5" pageNumber="260">0.47</td>
<td id="76ED5DB5753A006CFDD35823DFE52901" box="[579,649,1605,1628]" gridcol="2" gridrow="5" pageId="5" pageNumber="260">1.01</td>
<td id="76ED5DB5753A006CFD2B5823DE682901" box="[699,772,1605,1628]" gridcol="3" gridrow="5" pageId="5" pageNumber="260">0.79</td>
<td id="76ED5DB5753A006CFCA65823DE012901" box="[822,877,1605,1628]" gridcol="4" gridrow="5" pageId="5" pageNumber="260">0.70</td>
<td id="76ED5DB5753A006CFC0C5823D97D2901" box="[924,1041,1605,1628]" gridcol="5" gridrow="5" pageId="5" pageNumber="260">1.23</td>
<td id="76ED5DB5753A006CFBC25823D9FB2901" box="[1106,1175,1605,1628]" gridcol="6" gridrow="5" pageId="5" pageNumber="260">0.38</td>
<td id="76ED5DB5753A006CFB5A5823D86C2901" box="[1226,1280,1605,1628]" gridcol="7" gridrow="5" pageId="5" pageNumber="260">4.58</td>
</tr>
</table>
</paragraph>
<paragraph id="8BB3368B753AFF96FEA858CADF74299B" blockId="5.[264,1323,1708,1894]" box="[312,536,1708,1734]" pageId="5" pageNumber="260">Female. Unknown.</paragraph>
<paragraph id="8BB3368B753AFF95FEA858B2DF632E8B" blockId="5.[264,1323,1708,1894]" lastBlockId="6.[264,1323,284,470]" lastPageId="6" lastPageNumber="261" pageId="5" pageNumber="260">
<emphasis id="B978EA99753AFF96FEA858B2DCC729B3" bold="true" box="[312,427,1748,1774]" pageId="5" pageNumber="260">Variation</emphasis>
. The male 
<typeStatus id="54B78829753AFF96FDD658B2DFC729B3" box="[582,683,1748,1774]" pageId="5" pageNumber="260" type="paratype">paratype</typeStatus>
is more strongly pigmented than the 
<typeStatus id="54B78829753AFF96FB0258B2D99029B3" box="[1170,1276,1748,1774]" pageId="5" pageNumber="260" type="holotype">holotype</typeStatus>
. Its ground colour is amber, with dark brown horizontal bands on dorsal scutum and with a dark reticulation in the carapace region; leg femora to metatarsi are darkened, leg tarsi and palpal tarsus light brown; its ventral body side is mostly amber, only the lateral margin of the ventral scutum and patches behind coxa IV and genital operculum are darkened. The ventral process on the palpal trochanter of the 
<typeStatus id="54B788297539FF95FCB95F22DEE22E03" box="[809,910,324,350]" pageId="6" pageNumber="261" type="paratype">paratype</typeStatus>
is ventrad­directed (
<figureCitation id="13372A0E7539FF95FB105F22D9BA2E03" box="[1152,1238,324,350]" captionStart="FIGURES 4 – 10" captionStartId="5.[264,383,1011,1035]" captionTargetBox="[275,1314,521,997]" captionTargetId="figure@5.[264,1323,503,1005]" captionTargetPageId="5" captionText="FIGURES 4 – 10. Martensiellus tenuipalpus n. g., n. sp., male (4 – 9 holotype, 10 paratype). 4. Chelicera, lateral view. 5. Palpus, lateral view. 6. Distal parts of legs, lateral view (6 leg I, 7 leg II, 8 leg III, 9 leg IV). 10. Palpus and anterior part of body, lateral view. Scale bars: 1 mm (4 – 9 same scale)." httpUri="https://zenodo.org/record/174078/files/figure.png" pageId="6" pageNumber="261">Fig. 10</figureCitation>
) rather than slightly distad­directed as in the 
<typeStatus id="54B788297539FF95FD405F0ADE5B2EDB" box="[720,823,364,390]" pageId="6" pageNumber="261" type="holotype">holotype</typeStatus>
(
<figureCitation id="13372A0E7539FF95FCD95F0ADEF82EDB" box="[841,916,364,390]" captionStart="FIGURES 4 – 10" captionStartId="5.[264,383,1011,1035]" captionTargetBox="[275,1314,521,997]" captionTargetId="figure@5.[264,1323,503,1005]" captionTargetPageId="5" captionText="FIGURES 4 – 10. Martensiellus tenuipalpus n. g., n. sp., male (4 – 9 holotype, 10 paratype). 4. Chelicera, lateral view. 5. Palpus, lateral view. 6. Distal parts of legs, lateral view (6 leg I, 7 leg II, 8 leg III, 9 leg IV). 10. Palpus and anterior part of body, lateral view. Scale bars: 1 mm (4 – 9 same scale)." httpUri="https://zenodo.org/record/174078/files/figure.png" pageId="6" pageNumber="261">Fig. 5</figureCitation>
). The penis of the 
<typeStatus id="54B788297539FF95FBED5F0AD98E2EDB" box="[1149,1250,364,390]" pageId="6" pageNumber="261" type="paratype">paratype</typeStatus>
has a slightly more rounded distal margin (
<figureCitation id="13372A0E7539FF95FD2A5FF2DE2D2EF3" box="[698,833,404,430]" captionStart="FIGURES 11 – 17" captionStartId="6.[264,383,1444,1468]" captionTargetBox="[278,1309,534,1410]" captionTargetId="figure@6.[264,1323,518,1422]" captionTargetPageId="6" captionText="FIGURES 11 – 17. Martensiellus tenuipalpus n. g., n. sp., penis (11 – 14 holotype, 15 – 17 paratype). 11 – 12. Total penis (11 dorsal view, 12 lateral view). 13 – 15, 17. Apex of penis in resting position (13, 15 dorsal view, 14, 17 lateral view). 16. Apex of penis with expanded glans, dorsal view. Abbreviations: d. m. distal margin of truncus penis, l. s. lateral sclerite of glans, m. p. median plate of glans, m. t. membranous tubes of glans, s stylus. Scale bars: 0.5 mm (11 – 12), 0.1 mm (13 – 17)." httpUri="https://zenodo.org/record/174079/files/figure.png" pageId="6" pageNumber="261">Figs 15–16</figureCitation>
), but otherwise it is very similar to that of the 
<typeStatus id="54B788297539FF95FEA35FDADCF62E8B" box="[307,410,444,470]" pageId="6" pageNumber="261" type="holotype">holotype</typeStatus>
(
<figureCitation id="13372A0E7539FF95FE3A5FDADF6C2E8B" box="[426,512,444,470]" captionStart="FIGURES 11 – 17" captionStartId="6.[264,383,1444,1468]" captionTargetBox="[278,1309,534,1410]" captionTargetId="figure@6.[264,1323,518,1422]" captionTargetPageId="6" captionText="FIGURES 11 – 17. Martensiellus tenuipalpus n. g., n. sp., penis (11 – 14 holotype, 15 – 17 paratype). 11 – 12. Total penis (11 dorsal view, 12 lateral view). 13 – 15, 17. Apex of penis in resting position (13, 15 dorsal view, 14, 17 lateral view). 16. Apex of penis with expanded glans, dorsal view. Abbreviations: d. m. distal margin of truncus penis, l. s. lateral sclerite of glans, m. p. median plate of glans, m. t. membranous tubes of glans, s stylus. Scale bars: 0.5 mm (11 – 12), 0.1 mm (13 – 17)." httpUri="https://zenodo.org/record/174079/files/figure.png" pageId="6" pageNumber="261">Fig. 13</figureCitation>
).
</paragraph>
<caption id="DF7366037539FF95FE985BC2D99E2966" httpUri="https://zenodo.org/record/174079/files/figure.png" pageId="6" pageNumber="261" targetBox="[278,1309,534,1410]" targetPageId="6">
<paragraph id="8BB3368B7539FF95FE985BC2D99E2966" blockId="6.[264,1324,1444,1596]" pageId="6" pageNumber="261">
<emphasis id="B978EA997539FF95FE985BC2DCA12AE1" bold="true" box="[264,461,1444,1468]" pageId="6" pageNumber="261">FIGURES 11–17.</emphasis>
<taxonomicName id="4C0C4D087539FF95FE445BC3DF872AE1" box="[468,747,1445,1468]" class="Arachnida" family="Sandokanidae" genus="Martensiellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="261" phylum="Arthropoda" rank="species" species="tenuipalpus">
<emphasis id="B978EA997539FF95FE445BC3DF872AE1" box="[468,747,1445,1468]" italics="true" pageId="6" pageNumber="261">Martensiellus tenuipalpus</emphasis>
</taxonomicName>
<emphasis id="B978EA997539FF95FD625BC2DE002AE1" bold="true" box="[754,876,1444,1468]" pageId="6" pageNumber="261">n. g., n. sp.</emphasis>
, penis (11–14 holotype, 15–17 paratype). 11–12. Total penis (11 dorsal view, 12 lateral view). 13–15, 17. Apex of penis in resting position (13, 15 dorsal view, 14, 17 lateral view). 16. Apex of penis with expanded glans, dorsal view. Abbreviations: 
<emphasis id="B978EA997539FF95FE3D5863DCB02941" box="[429,476,1541,1564]" italics="true" pageId="6" pageNumber="261">d.m.</emphasis>
distal margin of truncus penis, 
<emphasis id="B978EA997539FF95FCBB5863DE262941" box="[811,842,1541,1564]" italics="true" pageId="6" pageNumber="261">l.s.</emphasis>
lateral sclerite of glans, 
<emphasis id="B978EA997539FF95FBC05863D9132941" box="[1104,1151,1541,1564]" italics="true" pageId="6" pageNumber="261">m.p.</emphasis>
median plate of glans, 
<emphasis id="B978EA997539FF95FEDE5843DC1A2961" box="[334,374,1573,1596]" italics="true" pageId="6" pageNumber="261">m.t.</emphasis>
membranous tubes of glans, 
<emphasis id="B978EA997539FF95FD3E5843DFD42961" box="[686,696,1573,1596]" italics="true" pageId="6" pageNumber="261">s</emphasis>
stylus. Scale bars: 0.5 mm (11–12), 0.1 mm (13–17).
</paragraph>
</caption>
<paragraph id="8BB3368B7539FF95FEA85801D87A282C" blockId="6.[264,1323,1639,1905]" pageId="6" pageNumber="261">
<emphasis id="B978EA997539FF95FEA85801DC8D29DC" bold="true" box="[312,481,1639,1665]" pageId="6" pageNumber="261">Relationships</emphasis>
. The presence of a distinctly derived penis with a distad­directed, expandable glans in which the median plate lies dorsally of the lateral sclerites, shows that 
<taxonomicName id="4C0C4D087539FF95FE9858DEDCC5298C" box="[264,425,1720,1745]" class="Arachnida" family="Sandokanidae" genus="Martensiellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="261" phylum="Arthropoda" rank="genus">
<emphasis id="B978EA997539FF95FE9858DEDCC5298C" box="[264,425,1720,1745]" italics="true" pageId="6" pageNumber="261">Martensiellus</emphasis>
</taxonomicName>
<emphasis id="B978EA997539FF95FE2658D1DC9C298C" bold="true" box="[438,496,1719,1745]" pageId="6" pageNumber="261">n. g.</emphasis>
from Borneo is phylogenetically closest to 
<taxonomicName id="4C0C4D087539FF95FB8458DED9D9298C" box="[1044,1205,1720,1745]" class="Arachnida" family="Sandokanidae" genus="Biantoncopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="261" phylum="Arthropoda" rank="genus">
<emphasis id="B978EA997539FF95FB8458DED9D9298C" box="[1044,1205,1720,1745]" italics="true" pageId="6" pageNumber="261">Biantoncopus</emphasis>
</taxonomicName>
from the 
<collectingCountry id="F31B761B7539FF95FE9858B9DCE029A4" box="[264,396,1759,1785]" name="Philippines" pageId="6" pageNumber="261">Philippines</collectingCountry>
(see 
<bibRefCitation id="EF9D4B7A7539FF95FE5958B9DE2729A4" author="Schwendinger" box="[457,843,1759,1785]" pageId="6" pageNumber="266" refString="Schwendinger, P. J. &amp; Martens, J. (2002 b) Penis morphology in Oncopodidae (Opiliones, Laniatores): evolutionary trends and relationships. Journal of Arachnology, 30, 425 - 434." type="journal article" year="2002" yearSuffix="b">Schwendinger &amp; Martens 2002b</bibRefCitation>
: fig. 59; 
<taxonomicName id="4C0C4D087539FF95FC215886D98E29A4" box="[945,1250,1760,1785]" class="Arachnida" family="Sandokanidae" genus="Martensiellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="261" phylum="Arthropoda" rank="species" species="tenuipalpus" status="sp. nov.">
<emphasis id="B978EA997539FF95FC215886D98E29A4" box="[945,1250,1760,1785]" italics="true" pageId="6" pageNumber="261">Martensiellus tenuipalpus</emphasis>
</taxonomicName>
<emphasis id="B978EA997539FF95FB7958B9D84729A4" bold="true" box="[1257,1323,1759,1785]" pageId="6" pageNumber="261">
<taxonomicNameLabel id="A24B57E27539FF95FB7958B9D84729A4" box="[1257,1323,1759,1785]" pageId="6" pageNumber="261" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
under 
<taxonomicName id="4C0C4D087539FF95FEC45961DC9C287C" box="[340,496,1799,1825]" class="Arachnida" family="Oncopodidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="261" phylum="Arthropoda" rank="family">Oncopodidae</taxonomicName>
g. sp.). In its somatic morphology however, the new genus is rather conservative and more similar to 
<taxonomicName id="4C0C4D087539FF95FD335956DE252814" box="[675,841,1840,1865]" class="Arachnida" family="Sandokanidae" genus="Palaeoncopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="261" phylum="Arthropoda" rank="genus">
<emphasis id="B978EA997539FF95FD335956DE252814" box="[675,841,1840,1865]" italics="true" pageId="6" pageNumber="261">Palaeoncopus</emphasis>
</taxonomicName>
(with primitive penis morphology) and 
<taxonomicName id="4C0C4D087539FF95FE98593EDCCB282C" box="[264,423,1880,1905]" class="Arachnida" family="Sandokanidae" genus="Caenoncopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="261" phylum="Arthropoda" rank="genus">
<emphasis id="B978EA997539FF95FE98593EDCCB282C" box="[264,423,1880,1905]" italics="true" pageId="6" pageNumber="261">Caenoncopus</emphasis>
</taxonomicName>
(with extremely derived penis morphology), which both occur in Sumatra.
</paragraph>
</subSubSection>
<subSubSection id="C31665007538FF94FEA85F7DD9F02E6B" box="[312,1180,283,310]" pageId="7" pageNumber="262" type="distribution">
<paragraph id="8BB3368B7538FF94FEA85F7DD9F02E6B" blockId="7.[312,1180,283,310]" box="[312,1180,283,310]" pageId="7" pageNumber="262">
<emphasis id="B978EA997538FF94FEA85F7DDCBD2E68" bold="true" box="[312,465,283,309]" pageId="7" pageNumber="262">Distribution</emphasis>
. Known only from the environs of Kapit in central 
<collectingCountry id="F31B761B7538FF94FBBE5F7AD9FB2E6B" box="[1070,1175,284,310]" name="Malaysia" pageId="7" pageNumber="262">Sarawak</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>