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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.114.1490" ID-GBIF-Dataset="fcb1d660-b485-4a65-9bdf-479ef9762383" ID-PMC="PMC3130342" ID-Pensoft-Pub="1313-2970-114-1" ID-PubMed="21976993" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1313-2970-114-1" ModsDocOrigin="ZooKeys 114" ModsDocTitle="Two new cave rnicolous genera of Julidae (Diplopoda, Julida), with notes on the tribe Brachyiulini and on julid subanal hooks and anchors" checkinTime="1451250225350" checkinUser="pensoft" docAuthor="Akkari, Nesrine, Stoev, Pavel &amp; Enghoff, Henrik" docDate="2011" docId="7E174013E9BC22ADB3901BC9F457ACD8" docLanguage="en" docName="ZooKeys 114: 1-14" docOrigin="ZooKeys 114" docSource="http://dx.doi.org/10.3897/zookeys.114.1490" docTitle="Titanophyllum spiliarum Akkari, Stoev &amp; Enghoff, 2011, sp. n." docType="treatment" docVersion="4" lastPageNumber="4" masterDocId="FF98FF9CFFD7FFBB9318FFED383DCD5F" masterDocTitle="Two new cave rnicolous genera of Julidae (Diplopoda, Julida), with notes on the tribe Brachyiulini and on julid subanal hooks and anchors" masterLastPageNumber="14" masterPageNumber="1" pageNumber="3" updateTime="1668151782143" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Two new cave rnicolous genera of Julidae (Diplopoda, Julida), with notes on the tribe Brachyiulini and on julid subanal hooks and anchors</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Akkari, Nesrine</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Stoev, Pavel</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Enghoff, Henrik</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2011</mods:date>
<mods:detail type="volume">
<mods:number>114</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>14</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.114.1490</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.114.1490</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-114-1</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152030072" LSID="urn:lsid:zoobank.org:act:C4B8738C-470F-4F54-A4F4-B69741C20EF4" httpUri="http://treatment.plazi.org/id/7E174013E9BC22ADB3901BC9F457ACD8" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
<subSubSection pageId="2" pageNumber="3" type="nomenclature">
<paragraph pageId="2" pageNumber="3">
<taxonomicName LSID="urn:lsid:zoobank.org:act:C4B8738C-470F-4F54-A4F4-B69741C20EF4" class="Diplopoda" family="Julidae" genus="Titanophyllum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Titanophyllum spiliarum" order="Julida" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="spiliarum">Titanophyllum spiliarum</taxonomicName>
<taxonomicNameLabel pageId="2" pageNumber="3">sp. n.</taxonomicNameLabel>
Figs 91014
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="3" type="material examined">
<paragraph pageId="2" pageNumber="3">Material examined.</paragraph>
<paragraph pageId="2" pageNumber="3">Holotype: adult ♂, Greece, Magnesia, Othris Mts., village of Kofi, Titanospilia (Cave of Titans), 13.VII.2003, P. Beron leg. (National Museum of Natural History Sofia NMNHS); Paratypes: 4 ♂♂, 5 ♀♀, same locality, date and collector (ZMUC); 12 adult ♂♂, 11 adult ♀♀, 2 subadult ♂♂, 2 subadult ♀♀, same locality, date and collector (NMNHS).</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="3" type="description of locality">
<paragraph pageId="2" pageNumber="3">Description of locality.</paragraph>
<paragraph pageId="2" pageNumber="3">Titanospilia is an approximately 100 m long vertical cave composed of a single voluminous hall. All material was collected at the bottom of the shaft (P. Beron, pers. comm.).</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="3" type="etymology">
<paragraph pageId="2" pageNumber="3">Etymology.</paragraph>
<paragraph pageId="2" pageNumber="3">The names means &quot;of caves&quot; in Greek and emphasizes the troglomorphic character of the species.</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="3" type="description">
<paragraph pageId="2" pageNumber="3">General description.</paragraph>
<paragraph pageId="2" pageNumber="3">
(all measurements in mm). Body uniformly pale to yellowish, legs brownish, metazonites with a slightly darker posterior band; length: 17.333.5 mm, vertical body diameter (height, H) 1-1.2 (♂) and 1-1.4 (♀); length/height ratio 16 (♂). Head: ocelli absent (Fig. 9); frons with 2 setae; gnathochilarium with 3 setae in apical part of each stipites, and with a long seta on each lamella lingualis; 4 supralabral setae and a row of ca 8 labral setae; mandibular stipites not expanded; male first leg-pair reduced and hook-shaped; antennal length about 1.6
<normalizedToken originalValue="×H">xH</normalizedToken>
. Body rings with more or less dense striation and a whorl of moderately dense long setae; 50-61 (♂) and 47-54 (♀) podous rings, 1-2 apodous rings + telson. Defense glands visible as dark spots opening on the suture. Length of legs ca 0.83
<normalizedToken originalValue="×H">xH</normalizedToken>
. Preanal ring dorsally only slightly protruding beyond anal margin; subanal scale with a small hook pointing anteriad (Fig. 14); anal valves pilose, with long setae. Male 7th body ring with well developed ventral lobes.
</paragraph>
</subSubSection>
<subSubSection lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3" type="gonopods">
<paragraph pageId="2" pageNumber="3">Gonopods.</paragraph>
<paragraph lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
Gonopods protruding from the 7th body ring. Anterior gonopod (promerite, p) much shorter than posterior gonopod (Figs 10, 11), uniformly broad along its length, slightly expanded at midlength; apex incised, with a lower angular
<pageBreakToken pageId="3" pageNumber="4" start="start">process</pageBreakToken>
mesally and a higher triangular one laterally (Fig. 11). Posterior side of promerite at middle with a quite high, semitriangular ridge (r) pointing postero-laterad (Fig. 11). Flagellum (f) (Figs 10, 11) moderately long, falcate, pointing ventrad, emerging from the base of ridge.
</paragraph>
<paragraph pageId="3" pageNumber="4">Posterior gonopod (Figs 12, 13) unipartite, long, broad at base and midlength, gradually tapering distally; with a proximal lobe (lo) laterally (Fig. 12) and a subbasal fold (fo) mesally (Fig. 10), the latter giving rise to a mesal groove (g) running proximal and distal (Figs 10, 13) and ending in a subapical opening (op) (Fig. 12). Gonopodal apex (t) with a pointed tip (Figs 12, 13).</paragraph>
<caption pageId="3" pageNumber="4">
<paragraph pageId="3" pageNumber="4">
Figure 9.
<taxonomicName class="Diplopoda" family="Julidae" genus="Titanophyllum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Titanophyllum spiliarum" order="Julida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="spiliarum">Titanophyllum spiliarum</taxonomicName>
gen. n., sp. n., habitus. Scale bar: 2mm.
</paragraph>
</caption>
<caption pageId="3" pageNumber="4">
<paragraph pageId="3" pageNumber="4">
Figures 1013.
<taxonomicName class="Diplopoda" family="Julidae" genus="Titanophyllum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Titanophyllum spiliarum" order="Julida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="spiliarum">Titanophyllum spiliarum</taxonomicName>
gen. n., sp. n., gonopods: 10 right gonopod, mesal view 11 left promerite, posterior view 12 right posterior gonopod, lateral view 13 close up of the tip of posterior gonopods, mesal view. Abbreviations: f: flagellum, fo: fold, g: groove, lo: lobe, op: opening, p: promerite, r: ridge, t: tip.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>