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<document id="EF3124AA55146CCAAB479E463124768D" ID-CLB-Dataset="42416" ID-DOI="10.11646/zootaxa.3905.2.7" ID-GBIF-Dataset="d499b3b8-1297-49ad-a192-5f1bc960fd93" ID-ISSN="1175-5326" ID-Zenodo-Dep="236505" ID-ZooBank="E9B24CA7-D364-4A5D-8FF6-C44C01393D1D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460930526977" checkinUser="plazi" docAuthor="Trýzna, Miloš &amp; Baňař, Petr" docDate="2015" docId="4A02A864FFDF3342FF5DFBABFA351308" docLanguage="en" docName="zt03905p272.pdf" docOrigin="Zootaxa 3905 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tophoderes lidmilae Tryzna &amp; Banar, sp. nov." docType="treatment" docVersion="10" lastPageNumber="270" masterDocId="B63BD01CFFDE3344FFCAFFA6FFA9120C" masterDocTitle="A new species of Tophoderes Dejean from northern Madagascar with a checklist of the species (Coleoptera: Anthribidae)" masterLastPageNumber="272" masterPageNumber="264" pageNumber="265" updateTime="1698582135455" updateUser="ExternalLinkService">
<mods:mods id="CD063C7098CF09E486BDCA4471C3445C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B40AA810634EF7156519ED794398DF08">
<mods:title id="58AA0C9F579980C863EEFBC4475B6D52">A new species of Tophoderes Dejean from northern Madagascar with a checklist of the species (Coleoptera: Anthribidae)</mods:title>
</mods:titleInfo>
<mods:name id="E092180BA5088F04DB84A90696A79BF6" type="personal">
<mods:role id="057843DE44A331A1DFE9C4786483B5C5">
<mods:roleTerm id="92294D2DA4026D73704CC0BD06B15DDD">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F5BBA05DF4A04EDCA535AF71D471859E">Trýzna, Miloš</mods:namePart>
</mods:name>
<mods:name id="7C13699E7BDAA276F5544D3A80801827" type="personal">
<mods:role id="B624753ACD1858C617FA9992B5787D6E">
<mods:roleTerm id="BF998D27CCD6862A2389DBDCC4E38740">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5AC58B66B891247B455CE5772AA792C9">Baňař, Petr</mods:namePart>
</mods:name>
<mods:typeOfResource id="4F1CEB2B34BF3806D7FBD6D97F95A969">text</mods:typeOfResource>
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<mods:title id="6EE72B7D8201ED8C6FC25A55A102CB75">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="B852FF0FDBDA5257BE9508579640C9AE">
<mods:date id="FA9B33BF12056BD8D64A6F0136413DD3">2015</mods:date>
<mods:detail id="456271A8231C178F4052AA18C12CC7F3" type="volume">
<mods:number id="C0107D7A669909CD1327461A14DB3865">3905</mods:number>
</mods:detail>
<mods:detail id="E7FF01557DB4D8B69D19B3DBC56E1EAA" type="issue">
<mods:number id="E8D2CA8F099B5930C6BF2676CEFB6C4A">2</mods:number>
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<mods:classification id="E8734E5E427E0829C0198F12796D2E17">journal article</mods:classification>
<mods:identifier id="B7FB3463BFE039925153A83F5E27C9E6" type="CLB-Dataset">42416</mods:identifier>
<mods:identifier id="BD638E2BFB899D449EA5CD73441CD2A9" type="DOI">10.11646/zootaxa.3905.2.7</mods:identifier>
<mods:identifier id="61ABA78625B516EF7753A6C74B7A0FAB" type="GBIF-Dataset">d499b3b8-1297-49ad-a192-5f1bc960fd93</mods:identifier>
<mods:identifier id="386AC5DEEF9127FBA836F0054965D217" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="ABDDC9A9A18FA83296D75494543168CB" type="Zenodo-Dep">236505</mods:identifier>
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<treatment id="4A02A864FFDF3342FF5DFBABFA351308" ID-DOI="http://doi.org/10.5281/zenodo.6111827" ID-GBIF-Taxon="119590201" ID-Zenodo-Dep="6111827" LSID="urn:lsid:plazi:treatment:4A02A864FFDF3342FF5DFBABFA351308" httpUri="http://treatment.plazi.org/id/4A02A864FFDF3342FF5DFBABFA351308" lastPageId="6" lastPageNumber="270" pageId="1" pageNumber="265">
<subSubSection id="8AB14AF9FFDF3345FF5DFBABFEB61644" pageId="1" pageNumber="265" type="nomenclature">
<paragraph id="C2141972FFDF3345FF5DFBABFD6C162B" blockId="1.[151,709,1037,1096]" box="[151,709,1037,1063]" pageId="1" pageNumber="265">
<heading id="995CAE1EFFDF3345FF5DFBABFD6C162B" bold="true" box="[151,709,1037,1063]" fontSize="11" level="1" pageId="1" pageNumber="265" reason="1">
<emphasis id="F0DFC560FFDF3345FF5DFBABFD6C162B" bold="true" box="[151,709,1037,1063]" pageId="1" pageNumber="265">
<taxonomicName id="05AB62F1FFDF3345FF5DFBABFDC9162B" authority="Tryzna &amp; Banar" authorityName="Tryzna &amp; Banar" box="[151,608,1037,1063]" class="Insecta" family="Anthribidae" genus="Tophoderes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="265" phylum="Arthropoda" rank="species" species="lidmilae" status="sp. nov.">
<emphasis id="F0DFC560FFDF3345FF5DFBABFE22162B" bold="true" box="[151,395,1037,1063]" italics="true" pageId="1" pageNumber="265">Tophoderes lidmilae</emphasis>
Trýzna &amp; Baňař
</taxonomicName>
<taxonomicNameLabel id="EBEC781BFFDF3345FDADFBABFD6C162B" box="[615,709,1037,1063]" pageId="1" pageNumber="265" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="C2141972FFDF3345FF5DFB89FEB61644" blockId="1.[151,709,1037,1096]" box="[151,287,1071,1096]" pageId="1" pageNumber="265">
(
<figureCitation id="5A9005F7FFDF3345FF55FB89FEBE1644" box="[159,279,1071,1096]" captionStart-0="FIGURES 5 – 8" captionStart-1="FIGURES 9 – 10" captionStart-2="FIGURES 11 – 14. 11, 13" captionStart-3="FIGURE 15" captionStartId-0="3.[151,264,1953,1976]" captionStartId-1="4.[151,264,1230,1253]" captionStartId-2="5.[151,264,1679,1702]" captionStartId-3="6.[151,250,1392,1414]" captionTargetBox-0="[193,1369,200,1907]" captionTargetBox-1="[257,1342,205,1184]" captionTargetBox-2="[187,1399,541,1642]" captionTargetId-0="figure@3.[187,1399,193,1933]" captionTargetId-1="figure@4.[232,1355,193,1209]" captionTargetId-2="figure@5.[171,1415,531,1659]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURES 5 – 8. Tophoderes lidmilae sp. nov. Trýzna &amp; Baňař; 5, male holotype dorsal habitus; 6, female allotype dorsal habitus; 7, pronotum, male; 8, head dorsal, male. Scale bars in mm." captionText-1="FIGURES 9 – 10. Tophoderes lidmilae sp. nov. Trýzna &amp; Baňař, holotype male; 9, aedeagus, lateral view; 10, tectum, dorsal view. Scale bars in mm." captionText-2="FIGURES 11 – 14. 11, 13; Tophoderes frenatus (Klug, 1833) male; 12, 14, Tophoderes lidmilae sp. nov. Trýzna &amp; Baňař, holotype male; 11, 12, tectum, dorsal view; 13, 14, apex of tegmen, dorsal view. Scale bars in mm." captionText-3="FIGURE 15. Type locality of Tophoderes lidmilae sp. nov. Trýzna &amp; Baňař." httpUri-0="https://zenodo.org/record/236507/files/figure.png" httpUri-1="https://zenodo.org/record/236508/files/figure.png" httpUri-2="https://zenodo.org/record/236509/files/figure.png" pageId="1" pageNumber="265">Figs. 5–15</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="8AB14AF9FFDF3345FF5DFBD1FC2B1764" pageId="1" pageNumber="265" type="materials_examined">
<paragraph id="C2141972FFDF3345FF5DFBD1FBC4169C" blockId="1.[151,1436,1143,1960]" box="[151,1133,1143,1168]" pageId="1" pageNumber="265">
<emphasis id="F0DFC560FFDF3345FF5DFBD1FE9C169C" bold="true" box="[151,309,1143,1168]" pageId="1" pageNumber="265">
<typeStatus id="1D10A7D0FFDF3345FF5DFBD1FF7B169C" box="[151,210,1143,1168]" pageId="1" pageNumber="265">Type</typeStatus>
locality.
</emphasis>
North
<collectingCountry id="BABC59E2FFDF3345FE4EFBD1FDA4169C" box="[388,525,1143,1168]" name="Madagascar" pageId="1" pageNumber="265">Madagascar</collectingCountry>
, Antsiranana province, Montagne d´Ambre Nat. Park.
</paragraph>
<paragraph id="C2141972FFDF3345FF0DFB3AFC2B1764" blockId="1.[151,1436,1143,1960]" pageId="1" pageNumber="265">
<emphasis id="F0DFC560FFDF3345FF0DFB3AFB6B16BF" bold="true" box="[199,1218,1180,1205]" pageId="1" pageNumber="265">
<typeStatus id="1D10A7D0FFDF3345FF0DFB3AFEAB16B9" box="[199,258,1180,1205]" pageId="1" pageNumber="265">Type</typeStatus>
material.
<typeStatus id="1D10A7D0FFDF3345FE5FFB3AFDAB16B9" box="[405,514,1180,1205]" pageId="1" pageNumber="265" type="holotype">Holotype</typeStatus>
(male): N
<collectingCountry id="BABC59E2FFDF3345FD6CFB3AFCDC16B9" box="[678,885,1180,1205]" name="Madagascar" pageId="1" pageNumber="265">MADAGASCAR</collectingCountry>
, ANTSIRANANA PROVINCE
</emphasis>
: ‘N
<collectingCountry id="BABC59E2FFDF3345FAC5FB3AFA3116B9" box="[1295,1432,1180,1205]" name="Madagascar" pageId="1" pageNumber="265">Madagascar</collectingCountry>
, Ambohitra env. / Montagne d´Ambre N.P. /
<geoCoordinate id="A79F7FB5FFDF3345FD7AFB66FCC916D4" box="[688,864,1216,1240]" direction="south" orientation="latitude" pageId="1" pageNumber="265" precision="1" value="-12.476305">S 12°28´34,7´´</geoCoordinate>
;
<geoCoordinate id="A79F7FB5FFDF3345FCB8FB66FB8616D4" box="[882,1071,1215,1240]" direction="east" orientation="longitude" pageId="1" pageNumber="265" precision="1" value="49.21875">E 049°13´07,5´´</geoCoordinate>
/ J. Vybíral leg., 1.–
<date id="B6153FB2FFDF3345FAE0FB66FA3E16D4" box="[1322,1431,1216,1240]" pageId="1" pageNumber="265" value="2007-01-03">3.1.2007</date>
’ (
<collectionCode id="A4BA81B7FFDF3345FF55FB42FF5716F0" box="[159,254,1252,1276]" pageId="1" pageNumber="265">BSNPC</collectionCode>
).
<emphasis id="F0DFC560FFDF3345FEDBFB42FE4116F1" bold="true" box="[273,488,1252,1277]" pageId="1" pageNumber="265">Allotype (female):</emphasis>
‘
<collectingCountry id="BABC59E2FFDF3345FE3EFB42FD2916F1" box="[500,640,1252,1277]" name="Madagascar" pageId="1" pageNumber="265">Madagascar</collectingCountry>
N / Antsiranana prov. / Ambohitra env. /
<date id="B6153FB2FFDF3345FB9DFB42FA8916F0" box="[1111,1312,1252,1277]" pageId="1" pageNumber="265" value="2002-12-20" valueMax="2002-12-26" valueMin="2002-12-20">20.–26. XII. 2002</date>
/ Ivo Jeniš leg.’ (
<collectionCode id="A4BA81B7FFDF3345FF15FAAEFE9F172C" box="[223,310,1288,1312]" pageId="1" pageNumber="265">MTDC</collectionCode>
).
<emphasis id="F0DFC560FFDF3345FE85FAA1FD9D172C" bold="true" box="[335,564,1287,1312]" pageId="1" pageNumber="265">
<typeStatus id="1D10A7D0FFDF3345FE85FAA1FE15172C" box="[335,444,1287,1312]" pageId="1" pageNumber="265" type="paratype">Paratype</typeStatus>
(female):
</emphasis>
‘N
<collectingCountry id="BABC59E2FFDF3345FDA6FAA1FD5C172C" box="[620,757,1287,1312]" name="Madagascar" pageId="1" pageNumber="265">Madagascar</collectingCountry>
,
<date id="B6153FB2FFDF3345FCCDFAAEFCDE172C" box="[775,887,1288,1312]" pageId="1" pageNumber="265" value="2010-10-28">28.x.2010</date>
/ Mt. d´Ambre Nat. Park,
<quantity id="0553B497FFDF3345FB0CFAAEFA8B1713" box="[1222,1314,1288,1312]" metricMagnitude="3" metricUnit="m" metricValue="1.048" pageId="1" pageNumber="265" unit="m" value="1048.0">1048 m</quantity>
a.s.l., /
<geoCoordinate id="A79F7FB5FFDF3345FA46FAAEFE831748" direction="south" orientation="latitude" pageId="1" pageNumber="265" precision="1" value="-12.534472">S 12°32´04.1´´</geoCoordinate>
;
<geoCoordinate id="A79F7FB5FFDF3345FEFCFA8BFE451748" box="[310,492,1324,1348]" direction="east" orientation="longitude" pageId="1" pageNumber="265" precision="1" value="49.176113">E 049°10´34.0´´</geoCoordinate>
/ M. Trýzna leg.’ (
<collectionCode id="A4BA81B7FFDF3345FD75FA8AFCBF1748" box="[703,790,1324,1348]" pageId="1" pageNumber="265">MTDC</collectionCode>
). Red label [p]
<typeStatus id="1D10A7D0FFDF3345FC0BFA8AFBFC1748" box="[961,1109,1324,1348]" pageId="1" pageNumber="265" type="holotype">HOLOTYPE</typeStatus>
/ ALLOTYPE /
<typeStatus id="1D10A7D0FFDF3345FADBFA8AFA351748" box="[1297,1436,1324,1348]" pageId="1" pageNumber="265" type="paratype">PARATYPE</typeStatus>
/
<emphasis id="F0DFC560FFDF3345FF6FFAE9FE311764" box="[165,408,1359,1384]" italics="true" pageId="1" pageNumber="265">
<taxonomicName id="05AB62F1FFDF3345FF6FFAE9FE8F1764" box="[165,294,1359,1384]" class="Insecta" family="Anthribidae" genus="Tophoderes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="265" phylum="Arthropoda" rank="genus">Tophoderes</taxonomicName>
/
<taxonomicName id="05AB62F1FFDF3345FEF6FAE9FE311764" box="[316,408,1359,1384]" class="Insecta" family="Anthribidae" genus="Tophoderes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="265" phylum="Arthropoda" rank="species" species="lidmilae" status="sp. nov.">lidmilae</taxonomicName>
</emphasis>
<emphasis id="F0DFC560FFDF3345FE55FAE9FE511764" bold="true" box="[415,504,1359,1384]" pageId="1" pageNumber="265">
<taxonomicNameLabel id="EBEC781BFFDF3345FE55FAE9FE511764" box="[415,504,1359,1384]" pageId="1" pageNumber="265" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
/ M. Trýzna &amp; P. Baňař det., 2014.
</paragraph>
</subSubSection>
<subSubSection id="8AB14AF9FFDF3341FF0DFAD2FBAD1369" lastPageId="5" lastPageNumber="269" pageId="1" pageNumber="265" type="description">
<paragraph id="C2141972FFDF3345FF0DFAD2FC781468" blockId="1.[151,1436,1143,1960]" pageId="1" pageNumber="265">
<emphasis id="F0DFC560FFDF3345FF0DFAD2FEF01781" bold="true" box="[199,345,1396,1421]" pageId="1" pageNumber="265">Description.</emphasis>
Male
<typeStatus id="1D10A7D0FFDF3345FE6CFAD2FDA21781" box="[422,523,1396,1421]" pageId="1" pageNumber="265" type="holotype">holotype</typeStatus>
, (female
<typeStatus id="1D10A7D0FFDF3345FDB0FAD2FD491780" box="[634,736,1396,1420]" pageId="1" pageNumber="265" type="paratype">paratype</typeStatus>
).
<emphasis id="F0DFC560FFDF3345FD3FFAD3FC3E1780" box="[757,919,1397,1420]" italics="true" pageId="1" pageNumber="265">Measurements</emphasis>
(in mm): Total body length—26.20 (21.20). Head: total length—5.10 (4.70); length of rostrum—3.50 (2.10); maximum width of rostrum—3.40 (2.80); length of eye—1.60 (1.20); maximum width across eyes—3.85 (3.10); minimum distance between eyes—1.50 (1.50). Antenna: length of segments: II 1.00 (0.60), III—1.60 (0.80), IV—1.50 (0.70), V—1.25 (0.55), VI—1.10 (0.50), VII—1.10 (0.50), VIII—1.00 (0.55), IX—1.00 (0.75), X—0.80 (0.55), XI—0.75 (0.60). Pronotum: maximum length—6.10 (4.20); width at carina—7.70 (5.70); maximum width—8.40 (6.00); minimum width—4.00 (3.30). Elytra: maximum length—15.00 (12.30); maximum width—10.30 (8.70).
</paragraph>
<paragraph id="C2141972FFDF3345FF0DF9C9FA951484" blockId="1.[151,1436,1143,1960]" box="[199,1340,1647,1672]" pageId="1" pageNumber="265">
<emphasis id="F0DFC560FFDF3345FF0DF9C9FEF11484" bold="true" box="[199,344,1647,1672]" pageId="1" pageNumber="265">Colouration</emphasis>
of cuticle of entire body black, only distal part of tarsomere V dark brown in both sexes.
</paragraph>
<paragraph id="C2141972FFDF3340FF0DF932FF71146F" blockId="1.[151,1436,1143,1960]" lastBlockId="4.[151,1437,1323,2031]" lastPageId="4" lastPageNumber="268" pageId="1" pageNumber="265">
<emphasis id="F0DFC560FFDF3345FF0DF932FE9314A1" bold="true" box="[199,314,1684,1709]" pageId="1" pageNumber="265">Vestiture.</emphasis>
<emphasis id="F0DFC560FFDF3345FE8EF932FE2814A1" box="[324,385,1684,1709]" italics="true" pageId="1" pageNumber="265">Head</emphasis>
predominantly with black appressed setae. On dorsal part of head two distinct yellowish longitudinal stripes located from vertex along inner margins of eyes to dorsal part of rostrum, but not reaching half of its length. These stripes composed of light yellow appressed setae, weakly mixed with brown setae.
<emphasis id="F0DFC560FFDF3345FAE7F97BFA3514F8" box="[1325,1436,1757,1780]" italics="true" pageId="1" pageNumber="265">Pronotum</emphasis>
with five more or less distinct longitudinal spots of light yellow setae, two on distal margin and three on proximal third of pronotum, rest of pronotum covered with black appressed setae.
<emphasis id="F0DFC560FFDF3345FC31F882FBE71531" box="[1019,1102,1828,1853]" italics="true" pageId="1" pageNumber="265">Elytron</emphasis>
with one distinct transverse narrow band on distal third and one short indistinct oblique pinstripe in humerus which blends in with adjacent similarly coloured spots. Rest of elytra speckled, covered by mixed yellow and brown appressed setae (more noticeable in male than in female). Scutellum with yellowish setae.
<emphasis id="F0DFC560FFDF3345FC44F836FC5115AB" box="[910,1016,1936,1959]" italics="true" pageId="1" pageNumber="265">Antennae</emphasis>
with black pubescence, distal part of antennomeres III–VIII in male and VI–VIII in female covered by longer conspicuous yellowish appressed setae, remaining antennomeres and primarily antennomeres IX–XI (club) only with very short, soft, dense setae in both sexes. Legs covered with dense, fine black setae, all femora with indistinct light distal part, with yellowish stripe on the distal quarter. Tibiae unicolorous, covered with dense distinct yellow setae appressed in proximal part and subdecumbent in distal part, apical part of tibiae with only black setae in both sexes. All tarsomeres with black setae; tarsomere I with sparse yellowish to brown indistinct setae in male and tarsomere V with distinct yellow distinct setae in both sexes. Lateral parts of venter of thorax and abdominal sternites with dense black pubescence mixed with yellow and brown setae. Distal part of prosternum with one pair of rusty (male) or yellow (female) spots.
</paragraph>
<caption id="96D449FAFFDC3346FF5DF839FE7C15D8" httpUri="https://zenodo.org/record/236506/files/figure.png" pageId="2" pageNumber="266" targetBox="[214,1365,198,1917]" targetPageId="2">
<paragraph id="C2141972FFDC3346FF5DF839FE7C15D8" blockId="2.[151,1436,1951,2004]" pageId="2" pageNumber="266">
<emphasis id="F0DFC560FFDC3346FF5DF839FE9315B8" bold="true" box="[151,314,1951,1974]" pageId="2" pageNumber="266">FIGURES 1–4.</emphasis>
<taxonomicName id="05AB62F1FFDC3346FE8AF839FD3015B9" authority="Klug, 1833" authorityName="Klug" authorityYear="1833" box="[320,665,1951,1973]" class="Insecta" family="Anthribidae" genus="Tophoderes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="266" phylum="Arthropoda" rank="species" species="frenatus">
<emphasis id="F0DFC560FFDC3346FE8AF839FDA415B9" box="[320,525,1951,1973]" italics="true" pageId="2" pageNumber="266">Tophoderes frenatus</emphasis>
(Klug, 1833)
</taxonomicName>
; 1, male dorsal habitus; 2, female dorsal habitus; 3, pronotum, male; 4, head dorsal, male. Scale bars in mm.
</paragraph>
</caption>
<caption id="96D449FAFFDD3347FF5DF807FCEB15DA" httpUri="https://zenodo.org/record/236507/files/figure.png" pageId="3" pageNumber="267" targetBox="[193,1369,200,1907]" targetPageId="3">
<paragraph id="C2141972FFDD3347FF5DF807FCEB15DA" blockId="3.[151,1436,1953,2006]" pageId="3" pageNumber="267">
<emphasis id="F0DFC560FFDD3347FF5DF807FE9715BB" bold="true" box="[151,318,1953,1976]" pageId="3" pageNumber="267">FIGURES 5–8.</emphasis>
<taxonomicName id="05AB62F1FFDD3347FE8DF804FDB115B4" box="[327,536,1954,1976]" class="Insecta" family="Anthribidae" genus="Tophoderes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="267" phylum="Arthropoda" rank="species" species="lidmilae" status="sp. nov.">
<emphasis id="F0DFC560FFDD3347FE8DF804FDB115B4" box="[327,536,1954,1976]" italics="true" pageId="3" pageNumber="267">Tophoderes lidmilae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EBEC781BFFDD3347FDE8F804FDDF15BB" box="[546,630,1954,1975]" pageId="3" pageNumber="267" rank="species">
<emphasis id="F0DFC560FFDD3347FDE8F804FDDF15BB" bold="true" box="[546,630,1954,1975]" pageId="3" pageNumber="267">sp. nov.</emphasis>
</taxonomicNameLabel>
Trýzna &amp; Baňař; 5, male holotype dorsal habitus; 6, female allotype dorsal habitus; 7, pronotum, male; 8, head dorsal, male. Scale bars in mm.
</paragraph>
</caption>
<caption id="96D449FAFFDA3340FF5DFB68FE20170F" httpUri="https://zenodo.org/record/236508/files/figure.png" pageId="4" pageNumber="268" targetBox="[257,1342,205,1184]" targetPageId="4">
<paragraph id="C2141972FFDA3340FF5DFB68FE20170F" blockId="4.[151,1436,1230,1283]" pageId="4" pageNumber="268">
<emphasis id="F0DFC560FFDA3340FF5DFB68FEE016E8" bold="true" box="[151,329,1230,1253]" pageId="4" pageNumber="268">FIGURES 9–10.</emphasis>
<taxonomicName id="05AB62F1FFDA3340FE9AFB69FDB616E9" box="[336,543,1231,1253]" class="Insecta" family="Anthribidae" genus="Tophoderes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="268" phylum="Arthropoda" rank="species" species="lidmilae" status="sp. nov.">
<emphasis id="F0DFC560FFDA3340FE9AFB69FDB616E9" box="[336,543,1231,1253]" italics="true" pageId="4" pageNumber="268">Tophoderes lidmilae</emphasis>
</taxonomicName>
<emphasis id="F0DFC560FFDA3340FDEDFB69FDD016E8" bold="true" box="[551,633,1231,1252]" pageId="4" pageNumber="268">
<taxonomicNameLabel id="EBEC781BFFDA3340FDEDFB69FDD016E8" box="[551,633,1231,1252]" pageId="4" pageNumber="268" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Trýzna &amp; Baňař, holotype male; 9, aedeagus, lateral view; 10, tectum, dorsal view. Scale bars in mm.
</paragraph>
</caption>
<paragraph id="C2141972FFDA3341FF0DF9C8FC7B1311" blockId="4.[151,1437,1323,2031]" lastBlockId="5.[151,1436,151,501]" lastPageId="5" lastPageNumber="269" pageId="4" pageNumber="268">
<emphasis id="F0DFC560FFDA3340FF0DF9C8FE96148B" bold="true" box="[199,319,1646,1671]" pageId="4" pageNumber="268">Structure.</emphasis>
<emphasis id="F0DFC560FFDA3340FE83F9C8FE2F148B" box="[329,390,1646,1671]" italics="true" pageId="4" pageNumber="268">Head</emphasis>
relatively long, rostrum extended apically (
<figureCitation id="5A9005F7FFDA3340FC4AF9C8FC6F148B" box="[896,966,1646,1671]" captionStart="FIGURES 5 – 8" captionStartId="3.[151,264,1953,1976]" captionTargetBox="[193,1369,200,1907]" captionTargetId="figure@3.[187,1399,193,1933]" captionTargetPageId="3" captionText="FIGURES 5 – 8. Tophoderes lidmilae sp. nov. Trýzna &amp; Baňař; 5, male holotype dorsal habitus; 6, female allotype dorsal habitus; 7, pronotum, male; 8, head dorsal, male. Scale bars in mm." httpUri="https://zenodo.org/record/236507/files/figure.png" pageId="4" pageNumber="268">Fig. 8</figureCitation>
), not depressed in middle, with one fine longitudinal carina reaching between eyes in both sexes. Ratio of rostrum length to maximum width
<quantity id="0553B497FFDA3340FAC3F935FAFE14A0" box="[1289,1367,1683,1708]" metricMagnitude="-2" metricUnit="m" metricValue="2.6162" pageId="4" pageNumber="268" unit="in" value="1.03">1.03 in</quantity>
male,
<quantity id="0553B497FFDA3340FF5DF910FF4D14C3" box="[151,228,1718,1743]" metricMagnitude="-2" metricUnit="m" metricValue="1.905" pageId="4" pageNumber="268" unit="in" value="0.75">0.75 in</quantity>
female. Eyes spherical, not emarginate, dorsal ocular index
<quantity id="0553B497FFDA3340FC4EF911FC7B14C3" box="[900,978,1718,1743]" metricMagnitude="-2" metricUnit="m" metricValue="3.2512" pageId="4" pageNumber="268" unit="in" value="1.28">1.28 in</quantity>
male,
<quantity id="0553B497FFDA3340FBD6F911FBC314C3" box="[1052,1130,1718,1743]" metricMagnitude="-2" metricUnit="m" metricValue="4.7498" pageId="4" pageNumber="268" unit="in" value="1.87">1.87 in</quantity>
female. Ratio of maximum width across eyes to maximum width of rostrum
<quantity id="0553B497FFDA3340FD13F97DFC8214F8" box="[729,811,1755,1780]" metricMagnitude="-2" metricUnit="m" metricValue="2.8701999999999996" pageId="4" pageNumber="268" unit="in" value="1.13">1.13 in</quantity>
male,
<quantity id="0553B497FFDA3340FCB4F97DFC6714F8" box="[894,974,1755,1780]" metricMagnitude="-2" metricUnit="m" metricValue="2.8194" pageId="4" pageNumber="268" unit="in" value="1.11">1.11 in</quantity>
female. Antennae reaching to posterior margin of pronotum in male, females antennae shorter. Funicle thick, club moderately robust, antennomere IX longer than wide in male.
<emphasis id="F0DFC560FFDA3340FE0AF882FD861537" box="[448,559,1828,1851]" italics="true" pageId="4" pageNumber="268">Pronotum</emphasis>
slightly transverse (
<figureCitation id="5A9005F7FFDA3340FCDEF885FCF01537" box="[788,857,1827,1852]" captionStart="FIGURES 5 – 8" captionStartId="3.[151,264,1953,1976]" captionTargetBox="[193,1369,200,1907]" captionTargetId="figure@3.[187,1399,193,1933]" captionTargetPageId="3" captionText="FIGURES 5 – 8. Tophoderes lidmilae sp. nov. Trýzna &amp; Baňař; 5, male holotype dorsal habitus; 6, female allotype dorsal habitus; 7, pronotum, male; 8, head dorsal, male. Scale bars in mm." httpUri="https://zenodo.org/record/236507/files/figure.png" pageId="4" pageNumber="268">Fig. 7</figureCitation>
), ratio of its length to width at carina
<quantity id="0553B497FFDA3340FAC2F885FAFF1530" box="[1288,1366,1827,1852]" metricMagnitude="-2" metricUnit="m" metricValue="2.0066" pageId="4" pageNumber="268" unit="in" value="0.79">0.79 in</quantity>
male,
<quantity id="0553B497FFDA3340FF5DF8E1FF4C1553" box="[151,229,1862,1887]" metricMagnitude="-2" metricUnit="m" metricValue="1.8796" pageId="4" pageNumber="268" unit="in" value="0.74">0.74 in</quantity>
female, gradually extended anteriorly to half of its length, here widest, then narrowed to distal part. Disk of pronotum with three distinct humps and two considerable horns protruded obliquely forward from line of pronotum in both sexes. Dorsal transverse carina bisinuate with indistinct arch in middle. Lateral carina of pronotum welldeveloped, sharp, reaching beyond basal half of pronotum in lateral view, meeting with dorsal transverse carina at right-angle.
<emphasis id="F0DFC560FFDA3340FED5F870FECC15E3" box="[287,357,2006,2031]" italics="true" pageId="4" pageNumber="268">Elytra</emphasis>
broadly suboval (
<figureCitation id="5A9005F7FFDA3340FDE4F870FD3315E3" box="[558,666,2006,2031]" captionStart="FIGURES 5 – 8" captionStartId="3.[151,264,1953,1976]" captionTargetBox="[193,1369,200,1907]" captionTargetId="figure@3.[187,1399,193,1933]" captionTargetPageId="3" captionText="FIGURES 5 – 8. Tophoderes lidmilae sp. nov. Trýzna &amp; Baňař; 5, male holotype dorsal habitus; 6, female allotype dorsal habitus; 7, pronotum, male; 8, head dorsal, male. Scale bars in mm." httpUri="https://zenodo.org/record/236507/files/figure.png" pageId="4" pageNumber="268">Figs. 5–6</figureCitation>
), ratio of maximum length to maximum width of elytra
<quantity id="0553B497FFDA3340FAC0F870FAF115E3" box="[1290,1368,2006,2031]" metricMagnitude="-2" metricUnit="m" metricValue="3.7084" pageId="4" pageNumber="268" unit="in" value="1.46">1.46 in</quantity>
male,
<quantity id="0553B497FFDB3341FF5DFF3EFF4112BC" box="[151,232,151,176]" metricMagnitude="-2" metricUnit="m" metricValue="3.5814" pageId="5" pageNumber="269" unit="in" value="1.41">1.41 in</quantity>
female.
<emphasis id="F0DFC560FFDB3341FE85FF31FE1012BC" box="[335,441,151,176]" italics="true" pageId="5" pageNumber="269">Abdomen</emphasis>
as long as wide.
<emphasis id="F0DFC560FFDB3341FD40FF31FC9312BC" box="[650,826,151,176]" italics="true" pageId="5" pageNumber="269">Male genitalia.</emphasis>
Aedeagus robust (
<figureCitation id="5A9005F7FFDB3341FBDEFF31FBF212BC" box="[1044,1115,151,176]" captionStart="FIGURES 9 – 10" captionStartId="4.[151,264,1230,1253]" captionTargetBox="[257,1342,205,1184]" captionTargetId="figure@4.[232,1355,193,1209]" captionTargetPageId="4" captionText="FIGURES 9 – 10. Tophoderes lidmilae sp. nov. Trýzna &amp; Baňař, holotype male; 9, aedeagus, lateral view; 10, tectum, dorsal view. Scale bars in mm." httpUri="https://zenodo.org/record/236508/files/figure.png" pageId="5" pageNumber="269">Fig. 9</figureCitation>
), internal sac, consisting of ventral lobe only, very wide and long, reaching the apices of apodemes. Tectum very wide, its sclerotized plate broadly oval, almost egg-shaped (
<figureCitation id="5A9005F7FFDB3341FDDDFF79FDDC12F4" box="[535,629,223,248]" captionStart="FIGURES 9 – 10" captionStartId="4.[151,264,1230,1253]" captionTargetBox="[257,1342,205,1184]" captionTargetId="figure@4.[232,1355,193,1209]" captionTargetPageId="4" captionText="FIGURES 9 – 10. Tophoderes lidmilae sp. nov. Trýzna &amp; Baňař, holotype male; 9, aedeagus, lateral view; 10, tectum, dorsal view. Scale bars in mm." httpUri="https://zenodo.org/record/236508/files/figure.png" pageId="5" pageNumber="269">Figs. 10</figureCitation>
,
<figureCitation id="5A9005F7FFDB3341FD48FF46FD0812F4" box="[642,673,224,248]" captionStart="FIGURES 11 – 14. 11, 13" captionStartId="5.[151,264,1679,1702]" captionTargetBox="[187,1399,541,1642]" captionTargetId="figure@5.[171,1415,531,1659]" captionTargetPageId="5" captionText="FIGURES 11 – 14. 11, 13; Tophoderes frenatus (Klug, 1833) male; 12, 14, Tophoderes lidmilae sp. nov. Trýzna &amp; Baňař, holotype male; 11, 12, tectum, dorsal view; 13, 14, apex of tegmen, dorsal view. Scale bars in mm." httpUri="https://zenodo.org/record/236509/files/figure.png" pageId="5" pageNumber="269">12</figureCitation>
). Apodemal bridge irregularly rounded, widest in middle (
<figureCitation id="5A9005F7FFDB3341FAF0FF79FA2612F4" box="[1338,1423,223,248]" captionStart="FIGURES 11 – 14. 11, 13" captionStartId="5.[151,264,1679,1702]" captionTargetBox="[187,1399,541,1642]" captionTargetId="figure@5.[171,1415,531,1659]" captionTargetPageId="5" captionText="FIGURES 11 – 14. 11, 13; Tophoderes frenatus (Klug, 1833) male; 12, 14, Tophoderes lidmilae sp. nov. Trýzna &amp; Baňař, holotype male; 11, 12, tectum, dorsal view; 13, 14, apex of tegmen, dorsal view. Scale bars in mm." httpUri="https://zenodo.org/record/236509/files/figure.png" pageId="5" pageNumber="269">Fig. 12</figureCitation>
). Tegmen robust, its apex subrectangular (
<figureCitation id="5A9005F7FFDB3341FD97FEA2FD061310" box="[605,687,260,285]" captionStart="FIGURES 11 – 14. 11, 13" captionStartId="5.[151,264,1679,1702]" captionTargetBox="[187,1399,541,1642]" captionTargetId="figure@5.[171,1415,531,1659]" captionTargetPageId="5" captionText="FIGURES 11 – 14. 11, 13; Tophoderes frenatus (Klug, 1833) male; 12, 14, Tophoderes lidmilae sp. nov. Trýzna &amp; Baňař, holotype male; 11, 12, tectum, dorsal view; 13, 14, apex of tegmen, dorsal view. Scale bars in mm." httpUri="https://zenodo.org/record/236509/files/figure.png" pageId="5" pageNumber="269">Fig. 14</figureCitation>
), preapical flange simple.
</paragraph>
<paragraph id="C2141972FFDB3341FF0DFE81FBAD1369" blockId="5.[151,1436,151,501]" pageId="5" pageNumber="269">
<emphasis id="F0DFC560FFDB3341FF0DFE81FE14134C" bold="true" box="[199,445,295,320]" pageId="5" pageNumber="269">Sexual dimorphism.</emphasis>
Male antennae are more robust and moderately longer, reaching approximately to posterior margin of pronotum, abdominal ventrites are more slightly flattened.
</paragraph>
</subSubSection>
<subSubSection id="8AB14AF9FFDB3341FF0DFEC9FA261384" box="[199,1423,367,392]" pageId="5" pageNumber="269" type="etymology">
<paragraph id="C2141972FFDB3341FF0DFEC9FA261384" blockId="5.[151,1436,151,501]" box="[199,1423,367,392]" pageId="5" pageNumber="269">
<emphasis id="F0DFC560FFDB3341FF0DFEC9FEE71384" bold="true" box="[199,334,367,392]" pageId="5" pageNumber="269">Etymology.</emphasis>
Patronym, dedicated to the memory of the mother of the first author, Mgr. Lidmila Trýznová (†).
</paragraph>
</subSubSection>
<subSubSection id="8AB14AF9FFDB3342FF0DFE32FA351308" lastPageId="6" lastPageNumber="270" pageId="5" pageNumber="269" type="distribution">
<paragraph id="C2141972FFDB3341FF0DFE32FB6B13F9" blockId="5.[151,1436,151,501]" pageId="5" pageNumber="269">
<emphasis id="F0DFC560FFDB3341FF0DFE32FEC913A1" bold="true" box="[199,352,404,429]" pageId="5" pageNumber="269">Distribution.</emphasis>
Three specimens were collected in northern
<collectingCountry id="BABC59E2FFDB3341FC4FFE32FBA713A1" box="[901,1038,404,429]" name="Madagascar" pageId="5" pageNumber="269">Madagascar</collectingCountry>
, Antsiranana province, Montagne d´Ambre National Park, which is covered by a volcanic massif composed mainly of basaltic rocks. The
<typeStatus id="1D10A7D0FFDB3341FAF6FE1EFA3513DC" box="[1340,1436,440,464]" pageId="5" pageNumber="269" type="paratype">paratype</typeStatus>
was collected by the authors in tropical moist forest near a crater lake at 1048 meters elevation.
</paragraph>
<caption id="96D449FAFFDB3341FF5DF929FBD614C8" httpUri="https://zenodo.org/record/236509/files/figure.png" pageId="5" pageNumber="269" targetBox="[187,1399,541,1642]" targetPageId="5">
<paragraph id="C2141972FFDB3341FF5DF929FBD614C8" blockId="5.[151,1436,1679,1732]" pageId="5" pageNumber="269">
<emphasis id="F0DFC560FFDB3341FF5DF929FEFF14A9" bold="true" box="[151,342,1679,1702]" pageId="5" pageNumber="269">FIGURES 11–14.</emphasis>
11, 13;
<taxonomicName id="05AB62F1FFDB3341FE78F936FCBB14AA" authority="Klug, 1833" authorityName="Klug" authorityYear="1833" box="[434,786,1680,1702]" class="Insecta" family="Anthribidae" genus="Tophoderes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="269" phylum="Arthropoda" rank="species" species="frenatus">
<emphasis id="F0DFC560FFDB3341FE78F936FD2B14AA" box="[434,642,1680,1702]" italics="true" pageId="5" pageNumber="269">Tophoderes frenatus</emphasis>
(Klug, 1833)
</taxonomicName>
male; 12, 14,
<taxonomicName id="05AB62F1FFDB3341FC78F936FB2A14AA" box="[946,1155,1680,1702]" class="Insecta" family="Anthribidae" genus="Tophoderes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="269" phylum="Arthropoda" rank="species" species="lidmilae" status="sp. nov.">
<emphasis id="F0DFC560FFDB3341FC78F936FB2A14AA" box="[946,1155,1680,1702]" italics="true" pageId="5" pageNumber="269">Tophoderes lidmilae</emphasis>
</taxonomicName>
<emphasis id="F0DFC560FFDB3341FB46F936FB4814A9" bold="true" box="[1164,1249,1680,1701]" pageId="5" pageNumber="269">
<taxonomicNameLabel id="EBEC781BFFDB3341FB46F936FB4814A9" box="[1164,1249,1680,1701]" pageId="5" pageNumber="269" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Trýzna &amp; Baňař, holotype male; 11, 12, tectum, dorsal view; 13, 14, apex of tegmen, dorsal view. Scale bars in mm.
</paragraph>
</caption>
<paragraph id="C2141972FFDB3341FF0DF94AFC8C1525" blockId="5.[151,1436,1772,1833]" pageId="5" pageNumber="269">
<emphasis id="F0DFC560FFDB3341FF0DF94AFE6C1509" bold="true" box="[199,453,1772,1797]" pageId="5" pageNumber="269">Diferential diagnosis.</emphasis>
<taxonomicName id="05AB62F1FFDB3341FE07F94AFD1B1509" box="[461,690,1772,1797]" class="Insecta" family="Anthribidae" genus="Tophoderes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="269" phylum="Arthropoda" rank="species" species="lidmilae" status="sp. nov.">
<emphasis id="F0DFC560FFDB3341FE07F94AFD1B1509" box="[461,690,1772,1797]" italics="true" pageId="5" pageNumber="269">Tophoderes lidmilae</emphasis>
</taxonomicName>
<emphasis id="F0DFC560FFDB3341FD70F94AFCBC1509" bold="true" box="[698,789,1772,1797]" pageId="5" pageNumber="269">
<taxonomicNameLabel id="EBEC781BFFDB3341FD70F94AFCBC1509" box="[698,789,1772,1797]" pageId="5" pageNumber="269" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is very similar to
<taxonomicName id="05AB62F1FFDB3341FC2DF94BFB511509" authority="Klug, 1833" authorityName="Klug" authorityYear="1833" box="[999,1272,1772,1797]" class="Insecta" family="Anthribidae" genus="Tophoderes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="269" phylum="Arthropoda" rank="species" species="frenatus">
<emphasis id="F0DFC560FFDB3341FC2DF94BFBF61509" box="[999,1119,1773,1797]" italics="true" pageId="5" pageNumber="269">T. frenatus</emphasis>
(
<bibRefCitation id="A63A6483FFDB3341FBA4F94AFB591509" author="Klug" box="[1134,1264,1772,1797]" pageId="5" pageNumber="269" refString="Klug, F. (1833) Bericht uber eine auf Madagascar veranstaltete Sammlung von Insecten aus der Ordnung Coleoptera. Abhandlungen der Koniglich-Preussischen Akademie der Wissenschaft, Berlin, 1, 91 - 223." type="journal article" year="1833">Klug, 1833</bibRefCitation>
)
</taxonomicName>
from which it can be distinguished by characters given in the key below:
</paragraph>
<paragraph id="C2141972FFDB3341FF5DF8FEFA3515EE" blockId="5.[151,1436,1880,2018]" pageId="5" pageNumber="269">
(1) Pair of longitudinal stripes on head extended from vertex to only half of length of rostrum; pronotum without pair of wellmarked longitudinal stripes, with only five small longitudinal spots; humerus with only indistinct smudgy oblique stripes; surface of elytra mottled, covered with mix of yellow and brown setae; sclerotized plate of tectum broad, almost egg-shaped (
<figureCitation id="5A9005F7FFDB3341FABFF834FF5215C9" captionStart="FIGURES 11 – 14. 11, 13" captionStartId="5.[151,264,1679,1702]" captionTargetBox="[187,1399,541,1642]" captionTargetId="figure@5.[171,1415,531,1659]" captionTargetPageId="5" captionText="FIGURES 11 – 14. 11, 13; Tophoderes frenatus (Klug, 1833) male; 12, 14, Tophoderes lidmilae sp. nov. Trýzna &amp; Baňař, holotype male; 11, 12, tectum, dorsal view; 13, 14, apex of tegmen, dorsal view. Scale bars in mm." httpUri="https://zenodo.org/record/236509/files/figure.png" pageId="5" pageNumber="269">Fig. 12</figureCitation>
), apodemal bridge irregularly rounded (
<figureCitation id="5A9005F7FFDB3341FDB0F816FD6815C9" box="[634,705,1968,1989]" captionStart="FIGURES 11 – 14. 11, 13" captionStartId="5.[151,264,1679,1702]" captionTargetBox="[187,1399,541,1642]" captionTargetId="figure@5.[171,1415,531,1659]" captionTargetPageId="5" captionText="FIGURES 11 – 14. 11, 13; Tophoderes frenatus (Klug, 1833) male; 12, 14, Tophoderes lidmilae sp. nov. Trýzna &amp; Baňař, holotype male; 11, 12, tectum, dorsal view; 13, 14, apex of tegmen, dorsal view. Scale bars in mm." httpUri="https://zenodo.org/record/236509/files/figure.png" pageId="5" pageNumber="269">Fig. 12</figureCitation>
), preapical flange of tectum simple (
<figureCitation id="5A9005F7FFDB3341FBEAF816FBCF15C9" box="[1056,1126,1968,1989]" captionStart="FIGURES 11 – 14. 11, 13" captionStartId="5.[151,264,1679,1702]" captionTargetBox="[187,1399,541,1642]" captionTargetId="figure@5.[171,1415,531,1659]" captionTargetPageId="5" captionText="FIGURES 11 – 14. 11, 13; Tophoderes frenatus (Klug, 1833) male; 12, 14, Tophoderes lidmilae sp. nov. Trýzna &amp; Baňař, holotype male; 11, 12, tectum, dorsal view; 13, 14, apex of tegmen, dorsal view. Scale bars in mm." httpUri="https://zenodo.org/record/236509/files/figure.png" pageId="5" pageNumber="269">Fig. 14</figureCitation>
).................................................................................................
<taxonomicName id="05AB62F1FFDB3341FB88F86BFAE115EE" authority="Tryzna &amp; Banar" authorityName="Tryzna &amp; Banar" box="[1090,1352,1997,2018]" class="Insecta" family="Anthribidae" genus="Tophoderes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="269" phylum="Arthropoda" rank="species" species="lidmilae" status="sp. nov.">
<emphasis id="F0DFC560FFDB3341FB88F86BFB0E15EE" box="[1090,1191,1997,2018]" italics="true" pageId="5" pageNumber="269">T. lidmilae</emphasis>
Trýzna &amp; Baňař
</taxonomicName>
<emphasis id="F0DFC560FFDB3341FA85F86AFA3515EE" bold="true" box="[1359,1436,1996,2018]" pageId="5" pageNumber="269">
<taxonomicNameLabel id="EBEC781BFFDB3341FA85F86AFA3515EE" box="[1359,1436,1996,2018]" pageId="5" pageNumber="269" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="C2141972FFD83342FF5DFF31FA351308" blockId="6.[151,1436,151,260]" pageId="6" pageNumber="270">
(2) Pair of longitudinal stripes on head extended from vertex to distinctly beyond half of length of rostrum; pronotum with pair of well-marked longitudinal stripes subsequent to head and short humeral stripes; humerus with distinct oblique stripes; surface of elytra covered with brightly bordered spots with yellow setae; sclerotized plate of tectum narrow, longitudinal (
<figureCitation id="5A9005F7FFD83342FA8DFF74FA2612EB" box="[1351,1423,210,231]" captionStart="FIGURES 11 – 14. 11, 13" captionStartId="5.[151,264,1679,1702]" captionTargetBox="[187,1399,541,1642]" captionTargetId="figure@5.[171,1415,531,1659]" captionTargetPageId="5" captionText="FIGURES 11 – 14. 11, 13; Tophoderes frenatus (Klug, 1833) male; 12, 14, Tophoderes lidmilae sp. nov. Trýzna &amp; Baňař, holotype male; 11, 12, tectum, dorsal view; 13, 14, apex of tegmen, dorsal view. Scale bars in mm." httpUri="https://zenodo.org/record/236509/files/figure.png" pageId="6" pageNumber="270">Fig. 11</figureCitation>
), apodemal bridge regularly rounded (
<figureCitation id="5A9005F7FFD83342FDF5FF49FD2C1308" box="[575,645,239,260]" captionStart="FIGURES 11 – 14. 11, 13" captionStartId="5.[151,264,1679,1702]" captionTargetBox="[187,1399,541,1642]" captionTargetId="figure@5.[171,1415,531,1659]" captionTargetPageId="5" captionText="FIGURES 11 – 14. 11, 13; Tophoderes frenatus (Klug, 1833) male; 12, 14, Tophoderes lidmilae sp. nov. Trýzna &amp; Baňař, holotype male; 11, 12, tectum, dorsal view; 13, 14, apex of tegmen, dorsal view. Scale bars in mm." httpUri="https://zenodo.org/record/236509/files/figure.png" pageId="6" pageNumber="270">Fig. 11</figureCitation>
), preapical flange of tectum broadly flattened (
<figureCitation id="5A9005F7FFD83342FB82FF49FB241308" box="[1096,1165,239,260]" captionStart="FIGURES 11 – 14. 11, 13" captionStartId="5.[151,264,1679,1702]" captionTargetBox="[187,1399,541,1642]" captionTargetId="figure@5.[171,1415,531,1659]" captionTargetPageId="5" captionText="FIGURES 11 – 14. 11, 13; Tophoderes frenatus (Klug, 1833) male; 12, 14, Tophoderes lidmilae sp. nov. Trýzna &amp; Baňař, holotype male; 11, 12, tectum, dorsal view; 13, 14, apex of tegmen, dorsal view. Scale bars in mm." httpUri="https://zenodo.org/record/236509/files/figure.png" pageId="6" pageNumber="270">Fig. 13</figureCitation>
)...
<taxonomicName id="05AB62F1FFD83342FB7BFF49FA351308" authority="Klug, 1833" authorityName="Klug" authorityYear="1833" box="[1201,1436,239,260]" class="Insecta" family="Anthribidae" genus="Tophoderes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="270" phylum="Arthropoda" rank="species" species="frenatus">
<emphasis id="F0DFC560FFD83342FB7BFF49FAB01308" box="[1201,1305,239,260]" italics="true" pageId="6" pageNumber="270">T. frenatus</emphasis>
(
<bibRefCitation id="A63A6483FFD83342FAEDFF49FA3C1308" author="Klug" box="[1319,1429,239,260]" pageId="6" pageNumber="270" refString="Klug, F. (1833) Bericht uber eine auf Madagascar veranstaltete Sammlung von Insecten aus der Ordnung Coleoptera. Abhandlungen der Koniglich-Preussischen Akademie der Wissenschaft, Berlin, 1, 91 - 223." type="journal article" year="1833">Klug, 1833</bibRefCitation>
)
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>
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