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<document ID-DOI="http://doi.org/10.5281/zenodo.6870843" ID-ISBN="978-84-16728-08-4" ID-Zenodo-Dep="6870843" approvalRequired="304" approvalRequired_for_illustrations="45" approvalRequired_for_matCits="75" approvalRequired_for_taxonomicNames="40" approvalRequired_for_textStreams="75" approvalRequired_for_treatments="69" checkinTime="1658335596803" checkinUser="felipe" docAuthor="Russell A. Mittermeier &amp; Don E. Wilson" docDate="2018" docId="3D474A54A010877DFAF7A73C16AEF95E" docLanguage="en" docName="hbmw_8_Soricidae_0332.pdf.imf" docOrigin="Handbook of the Mammals of the World Volume 8 Insectivores, Sloths and Colugos, Barcelona: Lynx Edicions" docTitle="Soriculus nigrescens" docType="treatment" docVersion="3" lastPageNumber="452" masterDocId="C17E322CA0288744FF8DAB47125EFFF9" masterDocTitle="Soricidae" masterLastPageNumber="551" masterPageNumber="332" pageNumber="451" updateTime="1658350298165" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Soricidae</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Russell A. Mittermeier</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Don E. Wilson</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:originInfo>
<mods:dateIssued>2018</mods:dateIssued>
<mods:dateOther type="pubDate">2018-07-31</mods:dateOther>
<mods:publisher>Lynx Edicions</mods:publisher>
<mods:place>
<mods:placeTerm>Barcelona</mods:placeTerm>
</mods:place>
</mods:originInfo>
<mods:titleInfo>
<mods:title>Handbook of the Mammals of the World Volume 8 Insectivores, Sloths and Colugos</mods:title>
</mods:titleInfo>
<mods:part>
<mods:extent unit="page">
<mods:start>332</mods:start>
<mods:end>551</mods:end>
</mods:extent>
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<mods:classification>book chapter</mods:classification>
<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.6870843</mods:identifier>
<mods:identifier type="ISBN">978-84-16728-08-4</mods:identifier>
<mods:identifier type="Zenodo-Dep">6870843</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.6869926" ID-Zenodo-Dep="6869926" LSID="urn:lsid:plazi:treatment:3D474A54A010877DFAF7A73C16AEF95E" httpUri="http://treatment.plazi.org/id/3D474A54A010877DFAF7A73C16AEF95E" lastPageId="57" lastPageNumber="452" pageId="56" pageNumber="451">
<heading pageId="56" pageNumber="451">
<subSubSection box="[1402,1481,3195,3241]" pageId="56" pageNumber="451" type="multiple">
<paragraph blockId="56.[1397,2506,3195,3324]" box="[1402,1481,3195,3241]" pageId="56" pageNumber="451">
<figureCitation box="[1402,1481,3195,3241]" captionStart="Plate 18: Soricidae" captionStartId="48.[122,151,3054,3079]" captionTargetBox="[12,2744,15,3636]" captionTargetPageId="47" captionText="136. Elliots Short-tailed Shrew (Blarina hulophaga), 137. Northern Short-tailed Shrew (Blarina brevicaudus), 138. Southern Short-tailed Shrew (Blarina carolinensis), 139. Everglades Short-tailed Shrew (Blarina peninsulae), 140. Shermans Short-tailed Shrew (Blarina shermani), 141. Sichuan Short-tailed Shrew (Blarinella quadraticauda), 142. Burmese Short-tailed Shrew (Blarinella wardi), 143. Indochinese Short-tailed Shrew (Blarinella griselda), 144. Chinese Mole Shrew (Anourosorex squamaipes), 145. Taiwanese Mole Shrew (Anowrosorex yamashinai), 146. Assam Mole Shrew (Anowrosorex assamensis), 147. Giant Mole Shrew (Anourosorex schmid), 148. Desert Gray Shrew (Notiosorex crawfordi), 149. Cockrums Gray Shrew (Notiosorex cockrumi), 150. Large-eared Gray Shrew (Notiosorex evolis), 151. Villas Gray Shrew (Notiosorex villa), 152. Mexican Shrew (Megasorex gigas), 153. Taiwanese Brown-toothed (Shrew Epusoriculusfumidus), 154. Arboreal Brown-toothed Shrew (Episoriculus macrurus), 155. Bailey's Brown-toothed Shrew (Episoriculus baileyi), 156. Long-tailed Brown-toothed Shrew (Episoriculus leucops), 157. Hodgsons Brown-toothed Shrew (Episoriculus caudatus), 158. Sichuan Brown-toothed Shrew (Episoriculus sacratus), 159. Hidden Brown-toothed Shrew (Episoriculus umbrinus), 160. Nepalese Brown-toothed Shrew (Episoriculus soluensis), 161. Himalayan Shrew (Soriculus nigrescens), 162. De Wintons Brown-toothed Shrew (Chodsigoa hypsibia), 163. Pygmy Brown-toothed Shrew (Chodsigoa parva), 164. Smiths Brown-toothed Shrew (Chodsigoa smithii), 165. Salenskis Brown-toothed Shrew (Chodsigoa salenskii), 166. Dusky Brown-toothed Shrew (Chodsigoa furva), 167. Lesser Taiwanese Brown-toothed Shrew (Chodsigoa sodalis), 168. Van Sungs Brown-toothed Shrew (Chodsigoa caovansunga), 169. Hoffmanns Brown-toothed Shrew (Chodsigoa hoffmanni), 170. Lowes Brown-toothed Shrew (Chodsigoa parca), 171. Bornean Water Shrew (Chimarrogale phaeura), 172. Sumatran Water Shrew (Chimarrogale sumatrana), 173. Malayan Water Shrew (Chimarrogale hantu), 174. Chinese Water Shrew (Chimarrogale styani), 175. Himalayan Water Shrew (Chimarrogale himalayica), 176. Leanders Water Shrew (Chimarrogale leander), 177. Japanese Water Shrew (Chimarrogale platycephala), 178. Elegant Water Shrew (Nectogale elegans), 179. Mediterranean Water Shrew (Neomys anomalus), 180. Transcaucasian Water (Shrew Neomysteres), 181. Eurasian Water Shrew (Neomys fodiens)" figureDoi="http://doi.org/10.5281/zenodo.6871913" httpUri="https://zenodo.org/record/6871913/files/figure.png" pageId="56" pageNumber="451">161.</figureCitation>
</paragraph>
</subSubSection>
<subSubSection box="[1498,1865,3195,3241]" pageId="56" pageNumber="451" type="vernacular_names">
<paragraph blockId="56.[1397,2506,3195,3324]" box="[1498,1865,3195,3241]" pageId="56" pageNumber="451">
<vernacularName box="[1498,1865,3195,3241]" pageId="56" pageNumber="451">Himalayan Shrew</vernacularName>
</paragraph>
</subSubSection>
<subSubSection box="[1934,2286,3195,3241]" pageId="56" pageNumber="451" type="nomenclature">
<paragraph blockId="56.[1397,2506,3195,3324]" box="[1934,2286,3195,3241]" pageId="56" pageNumber="451">
<taxonomicName baseAuthorityName="Gray" baseAuthorityYear="1910" box="[1934,2286,3195,3241]" class="Mammalia" family="Soricidae" genus="Soriculus" kingdom="Animalia" order="Soricomorpha" pageId="56" pageNumber="451" phylum="Chordata" rank="species" species="nigrescens">
<emphasis box="[1934,2286,3195,3241]" italics="true" pageId="56" pageNumber="451">Soriculus nigrescens</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="56" pageNumber="451" type="vernacular_names">
<paragraph blockId="56.[1397,2506,3195,3324]" box="[1398,2505,3259,3280]" pageId="56" pageNumber="451">
<emphasis bold="true" box="[1398,1474,3259,3280]" pageId="56" pageNumber="451">French:</emphasis>
<vernacularName box="[1484,1740,3259,3280]" pageId="56" pageNumber="451">Musaraigne de |'Himalaya</vernacularName>
/
<emphasis bold="true" box="[1761,1852,3259,3280]" pageId="56" pageNumber="451">German:</emphasis>
<vernacularName box="[1861,2151,3259,3280]" pageId="56" pageNumber="451">Sikkim-GrofRRklauenspitzmaus</vernacularName>
/
<emphasis bold="true" box="[2172,2263,3259,3280]" pageId="56" pageNumber="451">Spanish:</emphasis>
<vernacularName box="[2274,2505,3259,3280]" pageId="56" pageNumber="451">Musarana del Himalaya</vernacularName>
</paragraph>
<paragraph blockId="56.[1397,2506,3195,3324]" box="[1398,1932,3298,3319]" pageId="56" pageNumber="451">
<emphasis bold="true" box="[1398,1645,3298,3319]" pageId="56" pageNumber="451">Other common names:</emphasis>
<vernacularName box="[1654,1932,3298,3319]" pageId="56" pageNumber="451">Sikkim Large-clawed Shrew</vernacularName>
</paragraph>
</subSubSection>
</heading>
<subSubSection box="[1397,1952,3365,3398]" pageId="56" pageNumber="451" type="reference_group">
<paragraph blockId="56.[1396,2604,3365,3486]" box="[1397,1952,3365,3398]" pageId="56" pageNumber="451">
<emphasis bold="true" box="[1397,1553,3365,3398]" pageId="56" pageNumber="451">Taxonomy.</emphasis>
<taxonomicName authority="Gray, 1842" authorityName="Gray" authorityYear="1842" box="[1567,1947,3365,3398]" class="Mammalia" family="Soricidae" genus="Corsira" kingdom="Animalia" order="Soricomorpha" pageId="56" pageNumber="451" phylum="Chordata" rank="species" species="nigrescens">Corsira nigrescens Gray, 1842</taxonomicName>
,
</paragraph>
</subSubSection>
<subSubSection box="[1963,2068,3365,3398]" pageId="56" pageNumber="451" type="materials_examined">
<paragraph blockId="56.[1396,2604,3365,3486]" box="[1963,2068,3365,3398]" pageId="56" pageNumber="451">
<materialsCitation box="[1963,2068,3365,3398]" country="India" location="India" pageId="56" pageNumber="451" specimenCount="1">
<collectingCountry box="[1977,2050,3365,3398]" name="India" pageId="56" pageNumber="451">India</collectingCountry>
.”
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection lastPageId="57" lastPageNumber="452" pageId="56" pageNumber="451" type="discussion">
<paragraph blockId="56.[1396,2604,3365,3486]" pageId="56" pageNumber="451">
<taxonomicName authorityName="Blyth" authorityYear="1854" box="[1399,1515,3408,3437]" class="Mammalia" family="Soricidae" genus="Soriculus" kingdom="Animalia" order="Soricomorpha" pageId="56" pageNumber="451" phylum="Chordata" rank="genus">Soriculus</taxonomicName>
is currently monotypic.
<taxonomicName authorityName="Ellermann &amp; Morrison-Scott" authorityYear="1966" box="[1864,2019,3408,3437]" class="Mammalia" family="Soricidae" genus="Episoriculus" kingdom="Animalia" order="Soricomorpha" pageId="56" pageNumber="451" phylum="Chordata" rank="genus">Episoriculus</taxonomicName>
and
<taxonomicName authorityName="Kastchenko" authorityYear="1907" box="[2096,2224,3408,3437]" class="Mammalia" family="Soricidae" genus="Chodsigoa" kingdom="Animalia" order="Soricomorpha" pageId="56" pageNumber="451" phylum="Chordata" rank="genus">Chodsigoa</taxonomicName>
were included in
<taxonomicName authorityName="Blyth" authorityYear="1854" box="[2487,2603,3408,3437]" class="Mammalia" family="Soricidae" genus="Soriculus" kingdom="Animalia" order="Soricomorpha" pageId="56" pageNumber="451" phylum="Chordata" rank="genus">Soriculus</taxonomicName>
as subgenera but are generally accepted as full genera. Notably, although
<taxonomicName authorityName="Blyth" authorityYear="1854" box="[2487,2603,3443,3476]" class="Mammalia" family="Soricidae" genus="Soriculus" kingdom="Animalia" order="Soricomorpha" pageId="56" pageNumber="451" phylum="Chordata" rank="genus">Soriculus</taxonomicName>
</paragraph>
<paragraph blockId="57.[775,1367,290,717]" pageId="57" pageNumber="452">is monotypic, it might still contain undescribed species based on molecular studies. Two subspecies recognized.</paragraph>
</subSubSection>
<subSubSection pageId="57" pageNumber="452" type="synonymic_list">
<caption ID-DOI="http://doi.org/10.5281/zenodo.6871219" ID-Zenodo-Dep="6871219" httpUri="https://zenodo.org/record/6871219/files/figure.png" inLine="true" pageId="57" pageNumber="452" targetBox="[159,750,297,712]" targetPageId="57">
<paragraph blockId="57.[775,1367,290,717]" box="[775,1175,409,442]" pageId="57" pageNumber="452">
<emphasis bold="true" box="[775,1175,409,442]" pageId="57" pageNumber="452">Subspecies and Distribution.</emphasis>
</paragraph>
<paragraph blockId="57.[775,1367,290,717]" pageId="57" pageNumber="452">
<taxonomicName authority="Gray, 1842" authorityName="Gray" authorityYear="1842" baseAuthorityName="Gray" baseAuthorityYear="1842" box="[776,1117,448,481]" class="Mammalia" family="Soricidae" genus="Soriculus" kingdom="Animalia" order="Soricomorpha" pageId="57" pageNumber="452" phylum="Chordata" rank="subSpecies" species="nigrescens" subSpecies="nigrescens">S.n.nigrescensGray,1842—NW&amp;NEIndia(Uttarakhand,WestBengal,andSikkim)andNepal.</taxonomicName>
</paragraph>
<paragraph blockId="57.[775,1367,290,717]" pageId="57" pageNumber="452">
<taxonomicName authority="Dobson, 1890" authorityName="Dobson" authorityYear="1890" box="[776,1146,571,600]" class="Mammalia" family="Soricidae" genus="Soriculus" kingdom="Animalia" order="Soricomorpha" pageId="57" pageNumber="452" phylum="Chordata" rank="subSpecies" species="nigrescens" subSpecies="minor">S. n. minor Dobson, 1890</taxonomicName>
— Bhutan, NE India, SW China (S Tibet [= Xizang] and NW Yunnan, W of the Salween River), and adjacent N Myanmar.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="57" pageNumber="452" type="description">
<paragraph blockId="57.[159,1366,728,1703]" pageId="57" pageNumber="452">
<emphasis bold="true" box="[161,415,728,757]" pageId="57" pageNumber="452">Descriptive notes.</emphasis>
Head-body 70-94 mm, tail 32-50 mm, hindfoot 12-17 mm. No specific data are available for body weight. Condylo-incisive lengths are 19-7-23-8 mm, and tooth rows are 8:8-10-7 mm. Dorsum of the Himalayan Shrew is dust in color. Forefeet and claws are enlarged and larger than hindfeet, indicating digging ability. Tail is ¢.50% of head—body length. Ears are small and hidden under fur. Skull is bony, and braincase is low. Low talon (posterior cusp) of I' and first upper unicuspid are similar in height. Second upper unicuspid is the largest. M, is tiny. Pigmentation on teeth is very light and,if present, only occurs on tips of teeth. Dental formula is I 3/2, C1/0,P2/1,M 3/3 (x2) = 30. Chromosomal complement has 2n = 64 and FNa = 86 + XY for specimens from Gandaki, Nepal.
</paragraph>
</subSubSection>
<subSubSection pageId="57" pageNumber="452" type="biology_ecology">
<paragraph blockId="57.[159,1366,728,1703]" pageId="57" pageNumber="452">
<emphasis bold="true" box="[161,272,1126,1151]" pageId="57" pageNumber="452">Habitat.</emphasis>
Mixed deciduous-coniferous forest, conifer-rhododendron forest, above timberline in alpine zone, and open rocky areas at elevations of 1000-4300 m.
</paragraph>
</subSubSection>
<subSubSection pageId="57" pageNumber="452" type="food_feeding">
<paragraph blockId="57.[159,1366,728,1703]" pageId="57" pageNumber="452">
<emphasis bold="true" box="[161,428,1197,1230]" pageId="57" pageNumber="452">Food and Feeding.</emphasis>
Diet of the Himalayan Shrew primarily contains insects (beetles, flies, and bees) and earthworms.
</paragraph>
</subSubSection>
<subSubSection pageId="57" pageNumber="452" type="breeding">
<paragraph blockId="57.[159,1366,728,1703]" pageId="57" pageNumber="452">
<emphasis bold="true" box="[161,295,1275,1308]" pageId="57" pageNumber="452">Breeding.</emphasis>
The Himalayan Shrew reportedly has two breeding seasons, a primary one in June-July and a secondary one in August-October. Litters have 3-9 young (average six in spring and 4-5 in autumn).
</paragraph>
</subSubSection>
<subSubSection pageId="57" pageNumber="452" type="activity">
<paragraph blockId="57.[159,1366,728,1703]" pageId="57" pageNumber="452">
<emphasis bold="true" box="[159,393,1393,1426]" pageId="57" pageNumber="452">Activity patterns.</emphasis>
The Himalayan Shrew occurs aboveground and underground. Based on its morphology, it is semi-fossorial.
</paragraph>
</subSubSection>
<subSubSection box="[161,1100,1472,1505]" pageId="57" pageNumber="452" type="biology_ecology">
<paragraph blockId="57.[159,1366,728,1703]" box="[161,1100,1472,1505]" pageId="57" pageNumber="452">
<emphasis bold="true" box="[161,861,1472,1505]" pageId="57" pageNumber="452">Movements, Home range and Social organization.</emphasis>
No information.
</paragraph>
</subSubSection>
<subSubSection pageId="57" pageNumber="452" type="conservation">
<paragraph blockId="57.[159,1366,728,1703]" pageId="57" pageNumber="452">
<emphasis bold="true" box="[161,510,1512,1545]" pageId="57" pageNumber="452">Status and Conservation.</emphasis>
Classified as Least Concern on The IUCN Red List. In South Asia, major threats faced by the Himalayan Shrew are habitat loss due to agricultural expansion and human settlements. It is known from a few specimens in south-western China, but population size is likely large in Nepal, based on large collections in museums.
</paragraph>
</subSubSection>
<subSubSection box="[161,1264,1678,1703]" pageId="57" pageNumber="452" type="bibRefCitation_list">
<paragraph blockId="57.[159,1366,728,1703]" box="[161,1264,1678,1703]" pageId="57" pageNumber="452">
<emphasis bold="true" box="[161,314,1678,1703]" pageId="57" pageNumber="452">Bibliography.</emphasis>
Hoffmann (1985), Motokawa (2003a), Motokawa et al. (2008), Smith &amp; Yan Xie (2008).
</paragraph>
</subSubSection>
</treatment>
</document>