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<document id="5D59F89C9713925B828E8038D9DFCF11" ID-DOI="10.11646/zootaxa.5406.3.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="10628032" ID-ZooBank="A1F77DF6-4F1B-4858-85FB-22620BCE0484" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1707284769071" checkinUser="plazi" docAuthor="Zuñiga, Ronald &amp; Hanson, Paul" docDate="2024" docId="052387912127FFBCFF4DFD58FCE54438" docLanguage="en" docName="zootaxa.5406.3.3.pdf" docOrigin="Zootaxa 5406 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5406.3.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Epirhyssa leonoreae Zuniga &amp; Hanson 2024, sp. n." docType="treatment" docVersion="2" lastPageNumber="445" masterDocId="F91AFFE92124FFB8FFDAFFC2FFBF4605" masterDocTitle="Two new species of Epirhyssa (Hymenoptera: Ichneumonidae: Rhyssinae) from Costa Rica" masterLastPageNumber="450" masterPageNumber="441" pageNumber="444" updateTime="1707285214245" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="233B7E46F9E5F925C8F34A229F689B29" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="AFC4166C1BA83EE34CE5DD1B1A60C455">
<mods:title id="8576B300E7AB801D3A70656DA1E95FBB">Two new species of Epirhyssa (Hymenoptera: Ichneumonidae: Rhyssinae) from Costa Rica</mods:title>
</mods:titleInfo>
<mods:name id="96A2D6CB2BC15C9C53C38C1827812B0D" type="personal">
<mods:role id="BD17097D54C2E6E29CAAAE23D9D82CE1">
<mods:roleTerm id="0C5CA2B5FE4393F0A9C68197B0836456">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="43FBB7519265239E60453C07C98E89E0">Zuñiga, Ronald</mods:namePart>
<mods:affiliation id="07BB6F4530D786FC16FD3D93E02B8449">Guanacaste Dry Forest Conservation Fund, Sector Santa Rosa, Area de Conservación Guanacaste, Costa Rica.</mods:affiliation>
</mods:name>
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<mods:role id="5361C092C9308EAB273C1ED95702FA7B">
<mods:roleTerm id="D7D8BEBB5DE1144C624A46FCE54F9BE8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="700159FE48A92459AB6846FF7A3B9C54">Hanson, Paul</mods:namePart>
<mods:affiliation id="55D429B149B335939DB6DE4FB61E9096">Centro de Investigación en Biodiversidad y Ecología Tropical, Escuela de Biologia, Universidad de Costa Rica, San Pedro, 11501 - 2060, San José, Costa Rica.</mods:affiliation>
</mods:name>
<mods:typeOfResource id="BA5FE0FB3354451176D8A40B883D2E36">text</mods:typeOfResource>
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<mods:titleInfo id="4E14F742AB645B5D61A7F4CE23D5C87B">
<mods:title id="CC38BC6B5B6F9575B2AD59E860171329">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="5E0F917A01FE7CA9D91BDCF54700B546">
<mods:date id="BF72CE730BEF6931097781667A58A5C8">2024</mods:date>
<mods:detail id="FBE88F5F5E70073A87065DCFCE68A426" type="pubDate">
<mods:number id="9719D0851CAB02D0AB16B13BBF7DC77B">2024-02-07</mods:number>
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<mods:detail id="86E7DA4CDCC7EDBCD92C48C30E7C508F" type="volume">
<mods:number id="08BA3BE9ED21EB9130C6FD4F116894A7">5406</mods:number>
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<mods:number id="12E763F43D00BBD17408C6F8042DF98A">3</mods:number>
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<mods:start id="ED15E698019C1D6699775F957DA46AA7">441</mods:start>
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<mods:location id="DEB6ACF3BE476114B31AC65DA21ECD80">
<mods:url id="570FFB6A66EE31A8819FFA713D919F29">http://dx.doi.org/10.11646/zootaxa.5406.3.3</mods:url>
</mods:location>
<mods:classification id="569B60C7E0889412E33FB6609F454EB0">journal article</mods:classification>
<mods:identifier id="EB0168D1DD0D26ECEBF89DDA213AEEEF" type="DOI">10.11646/zootaxa.5406.3.3</mods:identifier>
<mods:identifier id="DB957BF0E13D42A228D879BF261B224C" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="8127AF3AA1EE95EFDCBAC2BEA3B755A0" type="Zenodo-Dep">10628032</mods:identifier>
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</mods:mods>
<treatment id="052387912127FFBCFF4DFD58FCE54438" LSID="urn:lsid:plazi:treatment:052387912127FFBCFF4DFD58FCE54438" httpUri="http://treatment.plazi.org/id/052387912127FFBCFF4DFD58FCE54438" lastPageId="4" lastPageNumber="445" pageId="3" pageNumber="444">
<subSubSection id="C590650C2127FFBBFF4DFD58FD7844B0" box="[151,711,666,693]" pageId="3" pageNumber="444" type="nomenclature">
<paragraph id="8D3536872127FFBBFF4DFD58FD7844B0" blockId="3.[151,711,666,729]" box="[151,711,666,693]" pageId="3" pageNumber="444">
<heading id="D67D81EB2127FFBBFF4DFD58FD7844B0" bold="true" box="[151,711,666,693]" fontSize="11" level="1" pageId="3" pageNumber="444" reason="1">
<emphasis id="BFFEEA952127FFBBFF4DFD58FD7844B0" bold="true" box="[151,711,666,693]" pageId="3" pageNumber="444">
<taxonomicName id="4A8A4D042127FFBBFF4DFD58FDC744B0" authority="Zuniga &amp; Hanson" authorityName="Zuniga &amp; Hanson" authorityYear="2024" box="[151,632,666,693]" class="Insecta" family="Ichneumonidae" genus="Epirhyssa" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="444" phylum="Arthropoda" rank="species" species="leonoreae" status="sp. nov.">
<emphasis id="BFFEEA952127FFBBFF4DFD58FE3144B0" bold="true" box="[151,398,666,693]" italics="true" pageId="3" pageNumber="444">Epirhyssa leonoreae</emphasis>
Zúñiga &amp; Hanson
</taxonomicName>
,
<taxonomicNameLabel id="A4CD57EE2127FFBBFD5FFD59FD7844B0" box="[645,711,667,693]" pageId="3" pageNumber="444" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C590650C2127FFBBFF4DFD7CFEAD44DC" box="[151,274,702,729]" pageId="3" pageNumber="444" type="description">
<paragraph id="8D3536872127FFBBFF4DFD7CFEAD44DC" blockId="3.[151,711,666,729]" box="[151,274,702,729]" pageId="3" pageNumber="444">
(
<figureCitation id="15B12A022127FFBBFF45FD7CFF5044DC" box="[159,239,702,729]" captionStart="FIGURES 3–4" captionStartId="4.[152,270,1883,1907]" captionTargetBox="[354,1233,597,1860]" captionTargetId="figure-182@4.[354,1233,597,1860]" captionTargetPageId="4" captionText="FIGURES 3–4. Epirhyssa spp., lateral habitus. 3. E. curtisi Gauld. 4. E. leonoreae Zuñiga &amp; Hanson." figureDoi="http://doi.org/10.5281/zenodo.10628036" httpUri="https://zenodo.org/record/10628036/files/figure.png" pageId="3" pageNumber="444">Figs. 4</figureCitation>
,
<figureCitation id="15B12A022127FFBBFF21FD7DFEB544DC" box="[251,266,703,729]" captionStart="FIGURES 7–8" captionStartId="6.[152,270,1737,1761]" captionTargetBox="[350,1237,489,1713]" captionTargetId="figure-180@6.[350,1237,489,1713]" captionTargetPageId="6" captionText="FIGURES 7–8. Epirhyssa spp., mesoscutum, dorsal view. 7. E. lewisi Zuñiga &amp; Hanson. 8. E. praecincta Porter." figureDoi="http://doi.org/10.5281/zenodo.10628040" httpUri="https://zenodo.org/record/10628040/files/figure.png" pageId="3" pageNumber="444">7</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C590650C2127FFBBFF4DFCC4FC6845B5" pageId="3" pageNumber="444" type="diagnosis">
<paragraph id="8D3536872127FFBBFF4DFCC4FC6845B5" blockId="3.[151,1437,774,2025]" pageId="3" pageNumber="444">
<emphasis id="BFFEEA952127FFBBFF4DFCC4FEB54525" bold="true" box="[151,266,774,800]" pageId="3" pageNumber="444">Diagnosis</emphasis>
.
<taxonomicName id="4A8A4D042127FFBBFEC0FCC4FDBD4525" box="[282,514,774,800]" class="Insecta" family="Ichneumonidae" genus="Epirhyssa" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="444" phylum="Arthropoda" rank="species" species="leonoreae">
<emphasis id="BFFEEA952127FFBBFEC0FCC4FDBD4525" box="[282,514,774,800]" italics="true" pageId="3" pageNumber="444">Epirhyssa leonoreae</emphasis>
</taxonomicName>
is very similar to
<taxonomicName id="4A8A4D042127FFBBFD0CFCC4FC044525" box="[726,955,774,800]" class="Insecta" family="Ichneumonidae" genus="Epirhyssa" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="444" phylum="Arthropoda" rank="species" species="oranensis">
<emphasis id="BFFEEA952127FFBBFD0CFCC4FC044525" box="[726,955,774,800]" italics="true" pageId="3" pageNumber="444">Epirhyssa oranensis</emphasis>
</taxonomicName>
, but the former differs in having different colour patterns on the propodeum and metasoma. In the key to Peruvian species (
<bibRefCitation id="E91B4B762127FFBBFBF3FCE9FAB94540" author="Gomez, I. C. &amp; Saaksjarvi, I. E. &amp; Puhakka, L. &amp; Castillo, C. &amp; Bordera, S." box="[1065,1286,810,837]" pageId="3" pageNumber="444" pagination="311 - 336" refId="ref5396" refString="Gomez, I. C., Saaksjarvi, I. E., Puhakka, L., Castillo, C. &amp; Bordera, S. (2015) The Peruvian Amazonian species of Epirhyssa Cresson (Hymenoptera: Ichneumonidae: Rhyssinae), with notes on tropical species richness. Zootaxa, 3937 (2), 311 - 336. https: // doi. org / 10.11646 / zootaxa. 3937.2.4" type="journal article" year="2015">
Gómez
<emphasis id="BFFEEA952127FFBBFB59FCE9FB0B4541" box="[1155,1204,810,836]" italics="true" pageId="3" pageNumber="444">et al</emphasis>
., 2015
</bibRefCitation>
),
<taxonomicName id="4A8A4D042127FFBBFAC3FCE9FA224541" box="[1305,1437,810,836]" class="Insecta" family="Ichneumonidae" genus="Epirhyssa" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="444" phylum="Arthropoda" rank="species" species="leonorae">
<emphasis id="BFFEEA952127FFBBFAC3FCE9FA224541" box="[1305,1437,810,836]" italics="true" pageId="3" pageNumber="444">E. leonorae</emphasis>
</taxonomicName>
goes to couplet 16, but differs from the included species in having the occipital carina joining the hypostomal carina and having tergite I about 1.5 × as long as wide. The darkened shallow groove extending from the subalar prominence to the mesopleural fovea is unusual among
<taxonomicName id="4A8A4D042127FFBBFCD2FC54FC6845B5" box="[776,983,918,944]" class="Insecta" family="Ichneumonidae" genus="Epirhyssa" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="444" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="BFFEEA952127FFBBFCD2FC54FCC845B5" box="[776,887,918,944]" italics="true" pageId="3" pageNumber="444">Epirhyssa</emphasis>
species.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C590650C2127FFBBFF1DFC78FC0041A5" pageId="3" pageNumber="444" type="description">
<paragraph id="8D3536872127FFBBFF1DFC78FE2143F5" blockId="3.[151,1437,774,2025]" pageId="3" pageNumber="444">
<emphasis id="BFFEEA952127FFBBFF1DFC78FE0B45D0" bold="true" box="[199,436,954,981]" pageId="3" pageNumber="444">Description. Female</emphasis>
. Fore wing length
<quantity id="4A729B622127FFBBFDA4FC78FD4845D0" box="[638,759,954,981]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" metricValueMax="1.6" metricValueMin="1.0" pageId="3" pageNumber="444" unit="mm" value="13.0" valueMax="16.0" valueMin="10.0">10–16 mm</quantity>
.
<emphasis id="BFFEEA952127FFBBFD25FC79FC8045D0" bold="true" box="[767,831,955,981]" pageId="3" pageNumber="444">Head</emphasis>
with occipital carina more or less complete, interrupted dorsally behind ocelli and often weak ventrally, but joining the hypostomal carina above base of mandible; posterior ocellus separated from eye by about 1.2–1.3 × its diameter; clypeus small, trapezoidal, with ventral margin straight to slightly convex; face as broad as medially high (from supraclypeal suture to base of antenna), smooth; antenna with 31 flagellomeres, apical flagellomeres much shorter and slightly wider than basal flagellomeres; gena smooth with sparse long setae ventrally.
<emphasis id="BFFEEA952127FFBBFDC0FBADFD27428C" bold="true" box="[538,664,1135,1161]" pageId="3" pageNumber="444">Mesosoma</emphasis>
with strong epomia diverging from anterior margin of pronotum; mesoscutum with central lobe not overhanging pronotum; mesopleuron polished; subalar prominence strongly inflated; epicnemial carina quite weak, laterally reaching just above level of the lower corner of pronotum; hind coxa flat dorsally; fore wing with
<emphasis id="BFFEEA952127FFBBFDFDFB19FDE142F1" box="[551,606,1243,1268]" italics="true" pageId="3" pageNumber="444">cu–a</emphasis>
more or less opposite the base of
<emphasis id="BFFEEA952127FFBBFC2EFB19FBAF42F1" box="[1012,1040,1243,1268]" italics="true" pageId="3" pageNumber="444">Rs</emphasis>
&amp;
<emphasis id="BFFEEA952127FFBBFBFFFB19FB8342F1" box="[1061,1084,1243,1268]" italics="true" pageId="3" pageNumber="444">M</emphasis>
; 1st subdiscal cell moderately broadened distally, its width near outer end about 1.8–2.2 × it basal width; abscissa of
<emphasis id="BFFEEA952127FFBBFB8FFB3DFBD3431D" box="[1109,1132,1279,1304]" italics="true" pageId="3" pageNumber="444">M</emphasis>
between 2
<emphasis id="BFFEEA952127FFBBFB3EFB3DFAA3431D" box="[1252,1308,1279,1305]" italics="true" pageId="3" pageNumber="444">rs–m</emphasis>
and
<quantity id="4A729B622127FFBBFA88FB3DFACB431D" box="[1362,1396,1279,1305]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="444" unit="m" value="2.0">
2
<emphasis id="BFFEEA952127FFBBFABAFB3DFACB431D" box="[1376,1396,1279,1304]" italics="true" pageId="3" pageNumber="444">m</emphasis>
</quantity>
–cu 0.2–0.4 × as long as 2
<emphasis id="BFFEEA952127FFBBFE55FAE1FE794339" box="[399,454,1315,1340]" italics="true" pageId="3" pageNumber="444">rs–m</emphasis>
;
<emphasis id="BFFEEA952127FFBBFE0CFAE1FE4D4339" box="[470,498,1315,1340]" italics="true" pageId="3" pageNumber="444">Rs</emphasis>
weakly and evenly bowed; propodeum 2.4 times as long (measured along midline) as width of propodeal orifice, polished with sparse setae laterally, without evident median longitudinal depression.
<emphasis id="BFFEEA952127FFBBFF4DFAA9FEAC4380" bold="true" box="[151,275,1387,1413]" pageId="3" pageNumber="444">Metasoma</emphasis>
with tergite I 1.5–1.8× as long as posteriorly broad, dorsal surface mostly smooth; tergites II–VII with alutaceous sculpture over entire surface and more or less uniformly covered with short hairs; suranal cornus short, covered with dark hairs dorsally, apex circular; ovipositor projecting beyond apex of metasoma by 4.2–5.2× the length of the hind tibia.
</paragraph>
<paragraph id="8D3536872127FFBBFF1DFA38FBF24179" blockId="3.[151,1437,774,2025]" pageId="3" pageNumber="444">
<emphasis id="BFFEEA952127FFBBFF1DFA38FEF74011" bold="true" box="[199,328,1530,1556]" pageId="3" pageNumber="444">Coloration</emphasis>
. Antenna mostly brownish with first and part of second flagellomeres black. Head yellow with black on apical 2/3 of mandibles and with a transverse black band between inner eye margins and passing through ocellar triangle; dark brown arc above occipital carina. Mesosoma yellow except black (or dark brown) on following: hind margin of propleuron; median posterior part of pronotum, extending back near dorsal margin of lateral pronotum but not reaching hind margin; three longitudinal stripes on mesoscutum, scuto-scutellar groove and hind margin of scutellum; tegula; anterior margin of mesopleuron, on a very shallow groove extending from subalar prominence to mesopleural fovea and on mesopleural groove; anterior and posterior margins of propodeum, and with a broad posteromedial mark on dorsal surface. Metasoma with tergite I entirely yellow, tergites II–VII with anterior 0.7 (approximately) brownish and posterior part yellow, except extreme posterior margin dark brown; ovipositor sheath brown. Legs predominantly yellow, with some light brown markings on femora, distal tarsal segments blackish; hind coxa dark brown to black on ventral and dorsal surfaces. Wings light yellowish.
</paragraph>
<paragraph id="8D3536872127FFBBFF1DF845FC0041A5" blockId="3.[151,1437,774,2025]" box="[199,959,1926,1953]" pageId="3" pageNumber="444">
<emphasis id="BFFEEA952127FFBBFF1DF845FEBB41A4" bold="true" box="[199,260,1927,1953]" pageId="3" pageNumber="444">Male</emphasis>
. Structure similar to female, metasoma with extensive yellow.
</paragraph>
</subSubSection>
<subSubSection id="C590650C2127FFBCFF1DF868FBAE47A8" lastPageId="4" lastPageNumber="445" pageId="3" pageNumber="444" type="materials_examined">
<paragraph id="8D3536872127FFBBFF1DF868FA2341EC" blockId="3.[151,1437,774,2025]" pageId="3" pageNumber="444">
<emphasis id="BFFEEA952127FFBBFF1DF868FE9041C1" bold="true" box="[199,303,1962,1988]" pageId="3" pageNumber="444">Material</emphasis>
.
<materialsCitation id="3DE23CDA2127FFBBFEE5F868FC8D41EC" collectingDate="1995-04" collectionCode="USNM" collectorName="A. Azofeifa" country="Costa Rica" county="Est. La Casona" elevation="1520" location="Sendero Orquideas" municipality="Monteverde" pageId="3" pageNumber="444" specimenCount="1" stateProvince="Puntarenas" typeStatus="holotype">
<typeStatus id="523188252127FFBBFEE5F868FE1841C1" box="[319,423,1962,1988]" pageId="3" pageNumber="444" type="holotype">Holotype</typeStatus>
F:
<collectingCountry id="F59D76172127FFBBFE0AF869FDF041C1" box="[464,591,1962,1989]" name="Costa Rica" pageId="3" pageNumber="444">Costa Rica</collectingCountry>
,
<collectingRegion id="4F4EF8652127FFBBFD86F869FCA141C0" box="[604,798,1963,1989]" country="Costa Rica" name="Puntarenas" pageId="3" pageNumber="444">Puntarenas Prov.</collectingRegion>
:
<collectingCounty id="64544E0B2127FFBBFCF0F869FC6641C0" box="[810,985,1963,1989]" pageId="3" pageNumber="444">Est. La Casona</collectingCounty>
,
<collectingMunicipality id="6D51ACFD2127FFBBFC3CF868FBCF41C1" box="[998,1136,1962,1988]" pageId="3" pageNumber="444">Monteverde</collectingMunicipality>
,
<location id="8855605C2127FFBBFBA7F868FAE841C1" LSID="urn:lsid:plazi:treatment:052387912127FFBCFF4DFD58FCE54438:8855605C2127FFBBFBA7F868FAE841C1" box="[1149,1367,1962,1988]" country="Costa Rica" county="Est. La Casona" municipality="Monteverde" name="Sendero Orquideas" pageId="3" pageNumber="444" stateProvince="Puntarenas">Sendero Orquideas</location>
,
<quantity id="4A729B622127FFBBFABEF868FF1041EC" metricMagnitude="3" metricUnit="m" metricValue="1.52" pageId="3" pageNumber="444" unit="m" value="1520.0">
<elevation id="06A7D1B42127FFBBFABEF868FF1041EC" metricMagnitude="3" metricUnit="m" metricValue="1.52" pageId="3" pageNumber="444" unit="m" value="1520.0">1520 m</elevation>
</quantity>
,
<date id="F93410472127FFBBFF1BF80CFEA841EC" box="[193,279,1998,2025]" pageId="3" pageNumber="444" value="1995-04">
<collectingDate id="E970E9AF2127FFBBFF1BF80CFEA841EC" box="[193,279,1998,2025]" pageId="3" pageNumber="444" value="1995-04">iv.1995</collectingDate>
</date>
,
<collectorName id="207F53512127FFBBFEFCF80DFE0D41ED" box="[294,434,1998,2025]" pageId="3" pageNumber="444">A. Azofeifa</collectorName>
, INBIOCRI002543759 (
<collectionCode id="EB9BAE422127FFBBFD08F80DFC9641EC" box="[722,809,1999,2025]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="3" pageNumber="444" type="Museum">USNM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3DE23CDA2127FFBBFC98F80DFA2341EC" box="[834,1436,1998,2025]" country="Costa Rica" location="San" municipality="Buen Amigo" pageId="3" pageNumber="444" specimenCount="1" stateProvince="Puntarenas" typeStatus="paratype">
<typeStatus id="523188252127FFBBFC98F80DFC0C41EC" box="[834,947,1999,2025]" pageId="3" pageNumber="444" type="paratype">Paratypes</typeStatus>
: 1F,
<collectingRegion id="4F4EF8652127FFBBFC29F80DFB0541EC" box="[1011,1210,1999,2025]" country="Costa Rica" name="Puntarenas" pageId="3" pageNumber="444">Puntarenas Prov.</collectingRegion>
:
<collectingMunicipality id="6D51ACFD2127FFBBFB10F80DFADE41ED" box="[1226,1377,1998,2025]" pageId="3" pageNumber="444">Buen Amigo</collectingMunicipality>
,
<location id="8855605C2127FFBBFAA8F80DFA2341EC" LSID="urn:lsid:plazi:treatment:052387912127FFBCFF4DFD58FCE54438:8855605C2127FFBBFAA8F80DFA2341EC" box="[1394,1436,1999,2025]" country="Costa Rica" municipality="Buen Amigo" name="San" pageId="3" pageNumber="444" stateProvince="Puntarenas">San</location>
</materialsCitation>
</paragraph>
<paragraph id="8D3536872120FFBCFF4DFF55FBAE47A8" blockId="4.[151,1437,150,573]" pageId="4" pageNumber="445">
Luis Monteverde,
<quantity id="4A729B622120FFBCFEB0FF54FDB546B4" box="[362,522,150,177]" metricMagnitude="3" metricUnit="m" metricValue="1.175" metricValueMax="1.35" metricValueMin="1.0" pageId="4" pageNumber="445" unit="m" value="1175.0" valueMax="1350.0" valueMin="1000.0">1000–1350 m</quantity>
,
<date id="F93410472120FFBCFDC2FF55FDD046B4" box="[536,623,151,177]" pageId="4" pageNumber="445" value="1994-11">xi.1994</date>
, Z. Fuentes, INBIOCRI002111842 (
<collectionCode id="EB9BAE422120FFBCFBCAFF55FBC546B4" box="[1040,1146,151,177]" pageId="4" pageNumber="445">MZUCR</collectionCode>
); 1F,
<materialsCitation id="3DE23CDA2120FFBCFB61FF55FDBE46D0" collectingDate="1995-02" collectionCode="USNM" pageId="4" pageNumber="445" specimenCount="1">
same data,
<date id="F93410472120FFBCFAE7FF55FA2746B4" box="[1341,1432,150,177]" pageId="4" pageNumber="445" value="1995-02">
<collectingDate id="E970E9AF2120FFBCFAE7FF55FA2746B4" box="[1341,1432,150,177]" pageId="4" pageNumber="445" value="1995-02">ii. 1995</collectingDate>
</date>
, INBIOCRI002543222 (
<collectionCode id="EB9BAE422120FFBCFE78FF79FE4646D0" box="[418,505,187,213]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="445" type="Museum">USNM</collectionCode>
)
</materialsCitation>
; 1F,
<materialsCitation id="3DE23CDA2120FFBCFDE2FF79FB3E46D0" box="[568,1153,187,213]" collectingDate="1993-03" collectionCode="USUC" pageId="4" pageNumber="445" specimenCount="1">
same data,
<date id="F93410472120FFBCFD62FF79FCAE46D0" box="[696,785,187,213]" pageId="4" pageNumber="445" value="1993-03">
<collectingDate id="E970E9AF2120FFBCFD62FF79FCAE46D0" box="[696,785,187,213]" pageId="4" pageNumber="445" value="1993-03">iii.1993</collectingDate>
</date>
, INBIOCRI001123780 (
<collectionCode id="EB9BAE422120FFBCFBF2FF79FBC646D0" box="[1064,1145,187,213]" pageId="4" pageNumber="445">USUC</collectionCode>
)
</materialsCitation>
; 1F,
<materialsCitation id="3DE23CDA2120FFBCFB62FF79FDBC46FC" collectingDate="1995-11" collectionCode="MNCR" pageId="4" pageNumber="445" specimenCount="1">
same data,
<date id="F93410472120FFBCFAE2FF79FA2746D0" box="[1336,1432,186,213]" pageId="4" pageNumber="445" value="1995-11">
<collectingDate id="E970E9AF2120FFBCFAE2FF79FA2746D0" box="[1336,1432,186,213]" pageId="4" pageNumber="445" value="1995-11">xi. 1995</collectingDate>
</date>
, INBIOCRI002358054 (
<collectionCode id="EB9BAE422120FFBCFE78FF1DFE4446FC" box="[418,507,223,249]" country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" pageId="4" pageNumber="445">MNCR</collectionCode>
)
</materialsCitation>
; 1F,
<materialsCitation id="3DE23CDA2120FFBCFDE3FF1DFB2446FC" box="[569,1179,223,249]" collectingDate="1993-02" collectionCode="MZUCR" pageId="4" pageNumber="445" specimenCount="1">
same data,
<date id="F93410472120FFBCFD62FF1DFCAD46FC" box="[696,786,223,249]" pageId="4" pageNumber="445" value="1993-02">
<collectingDate id="E970E9AF2120FFBCFD62FF1DFCAD46FC" box="[696,786,223,249]" pageId="4" pageNumber="445" value="1993-02">ii. 1993</collectingDate>
</date>
, INBIOCRI002130120 (
<collectionCode id="EB9BAE422120FFBCFBF2FF1DFB2D46FC" box="[1064,1170,223,249]" pageId="4" pageNumber="445">MZUCR</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3DE23CDA2120FFBCFB7DFF1DFA274718" collectingDate="1993-07" collectionCode="USUC" collectorName="Z. P. Tenorio &amp; G. Rodriguez" elevation="1050" location="Tierras Morenas" municipality="Rio San Lorenzo" pageId="4" pageNumber="445" specimenCount="1" stateProvince="Guanacaste">
1F,
<collectingRegion id="4F4EF8652120FFBCFB0BFF1DFA2646FC" box="[1233,1433,223,249]" country="Costa Rica" name="Guanacaste" pageId="4" pageNumber="445">Guanacaste Prov.</collectingRegion>
:
<collectingMunicipality id="6D51ACFD2120FFBCFF4DFEC1FEE74718" box="[151,344,259,285]" pageId="4" pageNumber="445">Río San Lorenzo</collectingMunicipality>
,
<location id="8855605C2120FFBCFEBEFEC1FDA14718" LSID="urn:lsid:plazi:treatment:052387912127FFBCFF4DFD58FCE54438:8855605C2120FFBCFEBEFEC1FDA14718" box="[356,542,259,285]" municipality="Rio San Lorenzo" name="Tierras Morenas" pageId="4" pageNumber="445" stateProvince="Guanacaste">Tierras Morenas</location>
,
<collectorName id="207F53512120FFBCFDF0FEC1FD094718" box="[554,694,259,285]" pageId="4" pageNumber="445">Z.P. Tenorio</collectorName>
,
<quantity id="4A729B622120FFBCFD1BFEC0FCA64718" box="[705,793,258,285]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="4" pageNumber="445" unit="m" value="1050.0">
<elevation id="06A7D1B42120FFBCFD1BFEC0FCA64718" box="[705,793,258,285]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="4" pageNumber="445" unit="m" value="1050.0">1050 m</elevation>
</quantity>
,
<date id="F93410472120FFBCFCFFFEC1FC3B4718" box="[805,900,259,285]" pageId="4" pageNumber="445" value="1993-07">
<collectingDate id="E970E9AF2120FFBCFCFFFEC1FC3B4718" box="[805,900,259,285]" pageId="4" pageNumber="445" value="1993-07">vii.1993</collectingDate>
</date>
,
<collectorName id="207F53512120FFBCFC4AFEC1FB954718" box="[912,1066,259,285]" pageId="4" pageNumber="445">G. Rodriguez</collectorName>
, INBIOCRI001954443 (
<collectionCode id="EB9BAE422120FFBCFAE5FEC1FA2F4718" box="[1343,1424,259,285]" pageId="4" pageNumber="445">USUC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3DE23CDA2120FFBCFF4DFEE5FAD14744" box="[151,1390,294,321]" collectingDate="1990-06" collectionCode="MZUCR" collectorName="Malaise" elevation="1100" location="Estacion Cacao" municipality="Arenales" pageId="4" pageNumber="445" specimenCount="1" stateProvince="Guanacaste">
1F,
<collectingRegion id="4F4EF8652120FFBCFF1BFEE5FE364744" box="[193,393,295,321]" country="Costa Rica" name="Guanacaste" pageId="4" pageNumber="445">Guanacaste Prov.</collectingRegion>
:
<collectingMunicipality id="6D51ACFD2120FFBCFE49FEE5FE464744" box="[403,505,295,321]" pageId="4" pageNumber="445">Arenales</collectingMunicipality>
,
<location id="8855605C2120FFBCFDDFFEE5FD084744" LSID="urn:lsid:plazi:treatment:052387912127FFBCFF4DFD58FCE54438:8855605C2120FFBCFDDFFEE5FD084744" box="[517,695,295,321]" municipality="Arenales" name="Estacion Cacao" pageId="4" pageNumber="445" stateProvince="Guanacaste">Estación Cacao</location>
,
<quantity id="4A729B622120FFBCFD19FEE5FCA64744" box="[707,793,295,321]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="4" pageNumber="445" unit="m" value="1100.0">
<elevation id="06A7D1B42120FFBCFD19FEE5FCA64744" box="[707,793,295,321]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="4" pageNumber="445" unit="m" value="1100.0">1100 m</elevation>
</quantity>
,
<date id="F93410472120FFBCFCFCFEE5FCC24744" box="[806,893,295,321]" pageId="4" pageNumber="445" value="1990-06">
<collectingDate id="E970E9AF2120FFBCFCFCFEE5FCC24744" box="[806,893,295,321]" pageId="4" pageNumber="445" value="1990-06">vi.1990</collectingDate>
</date>
,
<collectorName id="207F53512120FFBCFC50FEE5FC5A4744" box="[906,997,295,321]" pageId="4" pageNumber="445">Malaise</collectorName>
, INBIOCRI001692526 (
<collectionCode id="EB9BAE422120FFBCFB26FEE5FAD94744" box="[1276,1382,295,321]" pageId="4" pageNumber="445">MZUCR</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3DE23CDA2120FFBCFAA0FEE5FAD04760" collectingDate="1996-11" collectingDateMax="1996-12-31" collectingDateMin="1996-11" collectionCode="MZUCR" collectorName="A. Masis" elevation="1075" location="Estacion Cacao" municipality="Cruz" pageId="4" pageNumber="445" specimenCount="1" stateProvince="Guanacaste">
1F,
<collectingRegion id="4F4EF8652120FFBCFF4DFE89FEE14760" box="[151,350,331,357]" country="Costa Rica" name="Guanacaste" pageId="4" pageNumber="445">Guanacaste Prov.</collectingRegion>
: la
<collectingMunicipality id="6D51ACFD2120FFBCFE59FE89FE034760" box="[387,444,331,357]" pageId="4" pageNumber="445">Cruz</collectingMunicipality>
,
<location id="8855605C2120FFBCFE12FE89FDC74760" LSID="urn:lsid:plazi:treatment:052387912127FFBCFF4DFD58FCE54438:8855605C2120FFBCFE12FE89FDC74760" box="[456,632,331,357]" municipality="Cruz" name="Estacion Cacao" pageId="4" pageNumber="445" stateProvince="Guanacaste">Estación Cacao</location>
,
<quantity id="4A729B622120FFBCFD59FE88FCA04760" box="[643,799,330,357]" metricMagnitude="3" metricUnit="m" metricValue="1.075" metricValueMax="1.15" metricValueMin="1.0" pageId="4" pageNumber="445" unit="m" value="1075.0" valueMax="1150.0" valueMin="1000.0">
<elevation id="06A7D1B42120FFBCFD59FE88FCA04760" box="[643,799,330,357]" metricMagnitude="3" metricUnit="m" metricValue="1.075" metricValueMax="1.15" metricValueMin="1.0" pageId="4" pageNumber="445" unit="m" value="1075.0" valueMax="1150.0" valueMin="1000.0">1000–1150 m</elevation>
</quantity>
,
<date id="F93410472120FFBCFCF1FE89FC0A4760" box="[811,949,331,357]" pageId="4" pageNumber="445" value="1996-11" valueMax="1996-12-31" valueMin="1996-11-00">
<collectingDate id="E970E9AF2120FFBCFCF1FE89FC0A4760" box="[811,949,331,357]" pageId="4" pageNumber="445" value="1996-11" valueMax="1996-12-31" valueMin="1996-11-00">xi–xii. 1996</collectingDate>
</date>
,
<collectorName id="207F53512120FFBCFC65FE89FB984760" box="[959,1063,331,357]" pageId="4" pageNumber="445">A. Masis</collectorName>
, INB0003138652 (
<collectionCode id="EB9BAE422120FFBCFB27FE89FAD84760" box="[1277,1383,331,357]" pageId="4" pageNumber="445">MZUCR</collectionCode>
)
</materialsCitation>
; 1F,
<materialsCitation id="3DE23CDA2120FFBCFF4DFEADFC3E478C" box="[151,897,366,393]" collectingDate="1994-11" collectingDateMax="1996-12-31" collectingDateMin="1996-11" collectionCode="USNM" collectorName="J. F. Corrales" elevation="1075" location="Estacion Cacao" municipality="Cruz" pageId="4" pageNumber="445" specimenCount="1" stateProvince="Guanacaste">
same data,
<date id="F93410472120FFBCFECEFEADFECC478C" box="[276,371,367,393]" pageId="4" pageNumber="445" value="1994-11">
<collectingDate id="E970E9AF2120FFBCFECEFEADFECC478C" box="[276,371,367,393]" pageId="4" pageNumber="445" value="1994-11">xi. 1994</collectingDate>
</date>
,
<collectorName id="207F53512120FFBCFEA4FEADFDB1478C" box="[382,526,367,393]" pageId="4" pageNumber="445">J.F. Corrales</collectorName>
, INBIOCRI002023075 (
<collectionCode id="EB9BAE422120FFBCFCF8FEADFCC6478C" box="[802,889,367,393]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="445" type="Museum">USNM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3DE23CDA2120FFBCFC56FEADFBB247A8" collectingDate="1998-01" collectionCode="USUC" collectorName="R. Villalobos" elevation="1450" location="Estacion Altamira" pageId="4" pageNumber="445" specimenCount="1" stateProvince="Puntarenas">
1M,
<collectingRegion id="4F4EF8652120FFBCFC1BFEADFB3D478C" box="[961,1154,367,393]" country="Costa Rica" name="Puntarenas" pageId="4" pageNumber="445">Puntarenas Prov.</collectingRegion>
:
<location id="8855605C2120FFBCFB56FEADFAE7478C" LSID="urn:lsid:plazi:treatment:052387912127FFBCFF4DFD58FCE54438:8855605C2120FFBCFB56FEADFAE7478C" box="[1164,1368,367,393]" name="Estacion Altamira" pageId="4" pageNumber="445" stateProvince="Puntarenas">Estación Altamira</location>
,
<locationDeviation id="9C872FA52120FFBCFAB9FEADFEFE47A8" location="Cerro Bioley" pageId="4" pageNumber="445">
<quantity id="4A729B622120FFBCFAB9FEADFA23478C" box="[1379,1436,367,393]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="445" unit="km" value="1.0">1 km</quantity>
S Cerro Bioley
</locationDeviation>
,
<quantity id="4A729B622120FFBCFE91FE50FE1D47A8" box="[331,418,402,429]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="4" pageNumber="445" unit="m" value="1450.0">
<elevation id="06A7D1B42120FFBCFE91FE50FE1D47A8" box="[331,418,402,429]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="4" pageNumber="445" unit="m" value="1450.0">1450 m</elevation>
</quantity>
,
<date id="F93410472120FFBCFE74FE51FE4047A8" box="[430,511,403,429]" pageId="4" pageNumber="445" value="1998-01">
<collectingDate id="E970E9AF2120FFBCFE74FE51FE4047A8" box="[430,511,403,429]" pageId="4" pageNumber="445" value="1998-01">i. 1998</collectingDate>
</date>
,
<collectorName id="207F53512120FFBCFDD0FE51FD1F47A8" box="[522,672,403,429]" pageId="4" pageNumber="445">R. Villalobos</collectorName>
, INBIOCRI002413553 (
<collectionCode id="EB9BAE422120FFBCFC6EFE51FBBA47A8" box="[948,1029,403,429]" pageId="4" pageNumber="445">USUC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C590650C2120FFBCFF1DFE74FDB047F0" pageId="4" pageNumber="445" type="discussion">
<paragraph id="8D3536872120FFBCFF1DFE74FDB047F0" blockId="4.[151,1437,150,573]" pageId="4" pageNumber="445">
<emphasis id="BFFEEA952120FFBCFF1DFE74FEF047D5" bold="true" box="[199,335,438,464]" pageId="4" pageNumber="445">Comments.</emphasis>
This species has been collected on the western side of the Tilaran and
<collectingRegion id="4F4EF8652120FFBCFB8CFE75FB6847D4" box="[1110,1239,439,465]" country="Costa Rica" name="Guanacaste" pageId="4" pageNumber="445">Guanacaste</collectingRegion>
mountain ranges. Nothing is known about its hosts.
</paragraph>
</subSubSection>
<subSubSection id="C590650C2120FFBCFF1DFE3DFCE54438" pageId="4" pageNumber="445" type="etymology">
<paragraph id="8D3536872120FFBCFF1DFE3DFCE54438" blockId="4.[151,1437,150,573]" pageId="4" pageNumber="445">
<emphasis id="BFFEEA952120FFBCFF1DFE3DFEF7441C" bold="true" box="[199,328,511,537]" pageId="4" pageNumber="445">Etymology</emphasis>
. This species is named in honor of Leonore Ortega Fonseca, a brilliant and tenacious woman, full of life, for her love and patience, companion of the first author.
</paragraph>
</subSubSection>
</treatment>
</document>
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