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<document id="B625670111EC7B978FF680D798CA1C59" ID-CLB-Dataset="54218" ID-DOI="10.11646/zootaxa.5249.2.4" ID-GBIF-Dataset="34ef8754-e9dc-490a-affd-27027f406df2" ID-ISSN="1175-5326" ID-Zenodo-Dep="7687418" ID-ZooBank="81426096-9E0D-4E3D-9B1A-660B4D1BBB65" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1677656213532" checkinUser="plazi" docAuthor="Páll-Gergely, Barna" docDate="2023" docId="8B1E6842FF86733FFF34F99AFB98229B" docLanguage="en" docName="zootaxa.5249.2.4.pdf" docOrigin="Zootaxa 5249 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5249.2.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pincerna viginticostatus Páll-Gergely 2023, n. sp." docType="treatment" docVersion="6" lastPageNumber="274" masterDocId="7727103AFF95732AFFA3FFB9FFB2234B" masterDocTitle="Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) II: The genera Alycaeus and Pincerna" masterLastPageNumber="276" masterPageNumber="253" pageNumber="272" updateTime="1699475959789" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="C7870E61E685C645CA518952645E73EA">Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) II: The genera Alycaeus and Pincerna</mods:title>
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<treatment id="8B1E6842FF86733FFF34F99AFB98229B" ID-DOI="http://doi.org/10.5281/zenodo.7687462" ID-GBIF-Taxon="209384114" ID-Zenodo-Dep="7687462" LSID="urn:lsid:plazi:treatment:8B1E6842FF86733FFF34F99AFB98229B" httpUri="http://treatment.plazi.org/id/8B1E6842FF86733FFF34F99AFB98229B" lastPageId="21" lastPageNumber="274" pageId="19" pageNumber="272">
<subSubSection id="4BAD8ADFFF867339FF34F99AFDB72576" box="[151,517,1571,1597]" pageId="19" pageNumber="272" type="nomenclature">
<paragraph id="0308D954FF867339FF34F99AFDB72576" blockId="19.[151,517,1571,1633]" box="[151,517,1571,1597]" pageId="19" pageNumber="272">
<heading id="58406E38FF867339FF34F99AFDB72576" bold="true" box="[151,517,1571,1597]" fontSize="11" level="1" pageId="19" pageNumber="272" reason="1">
<emphasis id="31C30546FF867339FF34F99AFDB72576" bold="true" box="[151,517,1571,1597]" pageId="19" pageNumber="272">
<taxonomicName id="C4B7A2D7FF867339FF34F99AFE092576" ID-CoL="9c23a448-abdd-4870-b665-63644546074f" authority="Páll-Gergely, 2023" authorityName="Páll-Gergely" authorityYear="2023" box="[151,443,1571,1597]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="19" pageNumber="272" phylum="Mollusca" rank="species" species="viginticostatus" status="sp. nov.">
<emphasis id="31C30546FF867339FF34F99AFE092576" bold="true" box="[151,443,1571,1597]" italics="true" pageId="19" pageNumber="272">Pincerna viginticostatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2AF0B83DFF867339FE60F99AFDB72576" box="[451,517,1571,1597]" pageId="19" pageNumber="272" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="4BAD8ADFFF867339FF34F9FEFE9D252A" box="[151,303,1607,1633]" pageId="19" pageNumber="272" type="description">
<paragraph id="0308D954FF867339FF34F9FEFE9D252A" blockId="19.[151,517,1571,1633]" box="[151,303,1607,1633]" pageId="19" pageNumber="272">
(
<figureCitation id="9B8CC5D1FF867339FF3CF9FEFE95252A" box="[159,295,1607,1633]" captionStart="FIGURE 14" captionStartId="20.[152,253,1979,2003]" captionTargetBox="[168,1419,489,1956]" captionTargetId="figure-160@20.[168,1419,489,1956]" captionTargetPageId="20" captionText="FIGURE 14. Shells of Pincerna Preston, 1907 species.A–F: Pincerna viginticostatus n. sp. (holotype, HNHM 105383); G: R2 of Pincerna vanbuensis (Bavay &amp; Dautzenberg, 1900) (MNHN-IM-2012-27215, same as on Fig. 12E). All photos: B. Páll-Gergely." figureDoi="http://doi.org/10.5281/zenodo.7687452" httpUri="https://zenodo.org/record/7687452/files/figure.png" pageId="19" pageNumber="272">Figs 14A–F</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="4BAD8ADFFF867339FF34F936FA9025BA" pageId="19" pageNumber="272" type="materials_examined">
<paragraph id="0308D954FF867339FF34F936FA9025BA" blockId="19.[151,1437,1678,2029]" pageId="19" pageNumber="272">
<emphasis id="31C30546FF867339FF34F936FEF625E3" bold="true" box="[151,324,1678,1705]" pageId="19" pageNumber="272">Type material.</emphasis>
<materialsCitation id="B3DFD309FF867339FEE8F936FD7025BA" ID-GBIF-Occurrence="4039909347" collectingDate="2020-02-18" collectionCode="HNHM" collectorName="Province, B &amp; Hunyadi, A." country="Vietnam" elevation="241390" latitude="21.945467" location="District" longLatPrecision="1" longitude="106.32412" municipality="Long Dong" pageId="19" pageNumber="272" specimenCode="HNHM 105383" specimenCount="1" typeStatus="holotype">
<emphasis id="31C30546FF867339FEE8F936FE0A25E2" bold="true" box="[331,440,1679,1705]" pageId="19" pageNumber="272">
<typeStatus id="DC0C67F6FF867339FEE8F936FE0A25E2" box="[331,440,1679,1705]" pageId="19" pageNumber="272" type="holotype">Holotype</typeStatus>
</emphasis>
(D=
<quantity id="C44F74B1FF867339FE4AF936FDF125E2" box="[489,579,1679,1705]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="19" pageNumber="272" unit="mm" value="4.2">4.2 mm</quantity>
, H=4.0 mm):
<collectingCountry id="7BA099C4FF867339FD7EF936FCD325E2" box="[733,865,1679,1705]" name="Vietnam" pageId="19" pageNumber="272">VIETNAM</collectingCountry>
, Lạng SƠn
<collectorName id="AE42BC82FF867339FC7CF936FBD325E0" box="[991,1121,1679,1707]" pageId="19" pageNumber="272">Province, B</collectorName>
ắc SƠn
<location id="06688F8FFF867339FB17F936FAB925E2" LSID="urn:lsid:plazi:treatment:8B1E6842FF86733FFF34F99AFB98229B:06688F8FFF867339FB17F936FAB925E2" box="[1204,1291,1679,1705]" country="Vietnam" latitude="21.945467" longLatPrecision="1" longitude="106.32412" municipality="Long Dong" name="District" pageId="19" pageNumber="272">District</location>
,
<collectingMunicipality id="E36C432EFF867339FAB6F936FA2A25E2" box="[1301,1432,1679,1705]" pageId="19" pageNumber="272">Long Dong</collectingMunicipality>
,
<quantity id="C44F74B1FF867339FF34F90AFF542586" box="[151,230,1715,1741]" metricMagnitude="3" metricUnit="m" metricValue="3.8" pageId="19" pageNumber="272" unit="km" value="3.8">3.8 km</quantity>
from junction 1B-
<elevation id="889A3E67FF867339FE15F90AFDC72586" box="[438,629,1715,1741]" metricMagnitude="5" metricUnit="m" metricValue="2.4139" pageId="19" pageNumber="272" unit="m" value="241390.0">
<quantity id="C44F74B1FF867339FE15F90AFD8A2586" box="[438,568,1715,1741]" metricMagnitude="5" metricUnit="m" metricValue="2.4139" pageId="19" pageNumber="272" unit="m" value="241390.0">241, 390 m</quantity>
a.s.l.
</elevation>
,
<geoCoordinate id="6683BF93FF867339FD21F90DFCA02584" box="[642,786,1716,1743]" degrees="21" direction="north" minutes="56.728" orientation="latitude" pageId="19" pageNumber="272" precision="1" value="21.945467">21°56.728′N</geoCoordinate>
,
<geoCoordinate id="6683BF93FF867339FCBCF90DFC0A2584" box="[799,952,1716,1743]" degrees="106" direction="east" minutes="19.447" orientation="longitude" pageId="19" pageNumber="272" precision="1" value="106.32412">106°19.447′E</geoCoordinate>
(locality code: 2020/51), leg.
<collectorName id="AE42BC82FF867339FAAEF90AFA252586" box="[1293,1431,1715,1741]" pageId="19" pageNumber="272">Hunyadi, A.</collectorName>
,
<date id="7709FF94FF867339FF34F96EFEEC25BA" box="[151,350,1751,1777]" pageId="19" pageNumber="272" value="2020-02-18">
<collectingDate id="674D067CFF867339FF34F96EFEEC25BA" box="[151,350,1751,1777]" pageId="19" pageNumber="272" value="2020-02-18">18 February 2020</collectingDate>
</date>
(
<specimenCode id="5311712FFF867339FECEF96EFD9025BA" box="[365,546,1750,1777]" collectionCode="HNHM" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="19" pageNumber="272">HNHM 105383</specimenCode>
,
<figureCitation id="9B8CC5D1FF867339FD8EF96EFD0A25BA" box="[557,696,1751,1777]" captionStart="FIGURE 14" captionStartId="20.[152,253,1979,2003]" captionTargetBox="[168,1419,489,1956]" captionTargetId="figure-160@20.[168,1419,489,1956]" captionTargetPageId="20" captionText="FIGURE 14. Shells of Pincerna Preston, 1907 species.A–F: Pincerna viginticostatus n. sp. (holotype, HNHM 105383); G: R2 of Pincerna vanbuensis (Bavay &amp; Dautzenberg, 1900) (MNHN-IM-2012-27215, same as on Fig. 12E). All photos: B. Páll-Gergely." figureDoi="http://doi.org/10.5281/zenodo.7687452" httpUri="https://zenodo.org/record/7687452/files/figure.png" pageId="19" pageNumber="272">Figs 14A, B</figureCitation>
)
</materialsCitation>
.
<emphasis id="31C30546FF867339FD6EF96EFCFC25BA" bold="true" box="[717,846,1751,1777]" pageId="19" pageNumber="272">
<typeStatus id="DC0C67F6FF867339FD6EF96EFCFB25BA" box="[717,841,1751,1777]" pageId="19" pageNumber="272" type="paratype">Paratypes</typeStatus>
:
</emphasis>
Same data as for holotype, HA (2 shells).
</paragraph>
</subSubSection>
<subSubSection id="4BAD8ADFFF867339FF64F943FEDD2416" pageId="19" pageNumber="272" type="diagnosis">
<paragraph id="0308D954FF867339FF64F943FEDD2416" blockId="19.[151,1437,1678,2029]" pageId="19" pageNumber="272">
<emphasis id="31C30546FF867339FF64F943FEF0245F" bold="true" box="[199,322,1786,1812]" pageId="19" pageNumber="272">Diagnosis.</emphasis>
A medium sized
<taxonomicName id="C4B7A2D7FF867339FDAAF942FD7B245E" box="[521,713,1787,1813]" pageId="19" pageNumber="272">
<emphasis id="31C30546FF867339FDAAF942FDC2245F" box="[521,624,1787,1812]" italics="true" pageId="19" pageNumber="272">Pincerna</emphasis>
species
</taxonomicName>
with regularly ribbed R1, long R2 with ca. 20 slightly elevated ribs, R3 with blunt central swelling, clearly separate inner and outer peristomes, a protruding inner peristome, and a closed umbilicus.
</paragraph>
</subSubSection>
<subSubSection id="4BAD8ADFFF86733EFF64F8DFFE66220A" lastPageId="20" lastPageNumber="273" pageId="19" pageNumber="272" type="description">
<paragraph id="0308D954FF86733EFF64F8DFFA8E2256" blockId="19.[151,1437,1678,2029]" lastBlockId="20.[151,1437,151,465]" lastPageId="20" lastPageNumber="273" pageId="19" pageNumber="272">
<emphasis id="31C30546FF867339FF64F8DFFEEB24CB" bold="true" box="[199,345,1894,1920]" pageId="19" pageNumber="272">Description.</emphasis>
Shell yellowish-corneous or off-white, although all three available shells were somewhat corroded; slightly wider than high; protoconch smooth, yellowish, consisting of ca. 1.5–1.75 whorls; R1 of ca. 1.5 whorls, with widely-spaced, thread-like, regular ribs (strongest near the suture) and extremely fine spiral striation; ribs ca. 26–28 on last half whorl of R1; R2 and R3 approx. 110 degrees combines, R2 ca. half of length of R3; constriction between R2 and R3 relatively shallow; R2 with ca. 20–22 regular, slightly elevated, dense ribs; cross-sectional view not examined; R3 with long, central swelling, with 3–4 thread-like ribs on its anterior part; aperture rounded; outer peristome expanded and reflected at the basal and columellar part; inner peristome protruding, expanded; boundary between inner and outer peristomes clearly visible; umbilicus entirely closed by reflected outer peristome.
</paragraph>
<paragraph id="0308D954FF81733EFF64FE9FFE66220A" blockId="20.[151,1437,151,465]" box="[199,468,294,321]" pageId="20" pageNumber="273">
<emphasis id="31C30546FF81733EFF64FE9FFEE5220B" bold="true" box="[199,343,294,320]" pageId="20" pageNumber="273">Operculum.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="4BAD8ADFFF81733EFF64FEF3FDED229A" pageId="20" pageNumber="273" type="diagnosis">
<paragraph id="0308D954FF81733EFF64FEF3FDED229A" blockId="20.[151,1437,151,465]" pageId="20" pageNumber="273">
<emphasis id="31C30546FF81733EFF64FEF3FE62222F" bold="true" box="[199,464,330,356]" pageId="20" pageNumber="273">Differential diagnosis.</emphasis>
This new species is similar to
<taxonomicName id="C4B7A2D7FF81733EFC95FEF2FC7D222E" baseAuthorityName="Bavay &amp; Dautzenberg" baseAuthorityYear="1900" box="[822,975,331,357]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="20" pageNumber="273" phylum="Mollusca" rank="species" species="vanbuensis">
<emphasis id="31C30546FF81733EFC95FEF2FC7D222E" box="[822,975,331,357]" italics="true" pageId="20" pageNumber="273">P. vanbuensis</emphasis>
</taxonomicName>
in shell shape, but differs from it in the R2 sculpture. While in
<taxonomicName id="C4B7A2D7FF81733EFE3EFED6FD8722C2" baseAuthorityName="Bavay &amp; Dautzenberg" baseAuthorityYear="1900" box="[413,565,367,393]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="20" pageNumber="273" phylum="Mollusca" rank="species" species="vanbuensis">
<emphasis id="31C30546FF81733EFE3EFED6FD8722C2" box="[413,565,367,393]" italics="true" pageId="20" pageNumber="273">P. vanbuensis</emphasis>
</taxonomicName>
the R2 is smooth and consists of 46–60 lighter and darker alternating stripes, this new species has ca. 20–22 slightly elevated ribs.
<taxonomicName id="C4B7A2D7FF81733EFCA2FE2AFC4C22E6" baseAuthorityName="Luo, Zhang &amp; Zhuo" baseAuthorityYear="2009" box="[769,1022,403,429]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="20" pageNumber="273" phylum="Mollusca" rank="species" species="maolanensis">
<emphasis id="31C30546FF81733EFCA2FE2AFC4C22E6" box="[769,1022,403,429]" italics="true" pageId="20" pageNumber="273">Pincerna maolanensis</emphasis>
</taxonomicName>
(which is possibly a synonym of
<taxonomicName id="C4B7A2D7FF81733EFA2BFE2AFEA1229A" baseAuthorityName="Bavay &amp; Dautzenberg" baseAuthorityYear="1900" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="20" pageNumber="273" phylum="Mollusca" rank="species" species="vanbuensis">
<emphasis id="31C30546FF81733EFA2BFE2AFEA1229A" italics="true" pageId="20" pageNumber="273">P. vanbuensis</emphasis>
</taxonomicName>
) also possesses a smooth R2.
</paragraph>
</subSubSection>
<caption id="57C889DCFF81733EFF3BF802FA2B24B3" ID-DOI="http://doi.org/10.5281/zenodo.7687452" ID-Zenodo-Dep="7687452" httpUri="https://zenodo.org/record/7687452/files/figure.png" pageId="20" pageNumber="273" startId="20.[152,253,1979,2003]" targetBox="[168,1419,489,1956]" targetPageId="20">
<paragraph id="0308D954FF81733EFF3BF802FA2B24B3" blockId="20.[151,1437,1979,2040]" pageId="20" pageNumber="273">
<emphasis id="31C30546FF81733EFF3BF802FE93249F" bold="true" box="[152,289,1979,2004]" pageId="20" pageNumber="273">FIGURE 14.</emphasis>
Shells of
<taxonomicName id="C4B7A2D7FF81733EFE20F805FD67249F" box="[387,725,1980,2004]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="20" pageNumber="273" phylum="Mollusca" rank="species" species="undefined-a">
<emphasis id="31C30546FF81733EFE20F805FE6C2498" box="[387,478,1980,2003]" italics="true" pageId="20" pageNumber="273">Pincerna</emphasis>
<bibRefCitation id="6726A4A5FF81733EFE40F805FDD9249F" author="Preston, H. B." box="[483,619,1980,2004]" pageId="20" pageNumber="273" pagination="206" refId="ref15909" refString="Preston, H. B. (1907) Description of a new subgenus and species of Alycaeus from Ke-Lan-Tan. Proceedings of the Malacological Society of London, 7, 206. https: // doi. org / 10.1093 / oxfordjournals. mollus. a 066171" type="journal article" year="1907">Preston, 1907</bibRefCitation>
species. A
</taxonomicName>
–F:
<taxonomicName id="C4B7A2D7FF81733EFD59F805FC592498" authority="Páll-Gergely, 2023" authorityName="Páll-Gergely" authorityYear="2023" box="[762,1003,1980,2003]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="20" pageNumber="273" phylum="Mollusca" rank="species" species="viginticostatus" status="sp. nov.">
<emphasis id="31C30546FF81733EFD59F805FC592498" box="[762,1003,1980,2003]" italics="true" pageId="20" pageNumber="273">Pincerna viginticostatus</emphasis>
</taxonomicName>
<emphasis id="31C30546FF81733EFC53F805FB95249F" bold="true" box="[1008,1063,1980,2004]" pageId="20" pageNumber="273">
<taxonomicNameLabel id="2AF0B83DFF81733EFC53F805FB95249F" box="[1008,1063,1980,2004]" pageId="20" pageNumber="273" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(holotype, HNHM 105383); G: R2 of
<taxonomicName id="C4B7A2D7FF81733EFF34F859FD2B24B3" authority="(Bavay &amp; Dautzenberg, 1900)" baseAuthorityName="Bavay &amp; Dautzenberg" baseAuthorityYear="1900" box="[151,665,2016,2040]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="20" pageNumber="273" phylum="Mollusca" rank="species" species="vanbuensis">
<emphasis id="31C30546FF81733EFF34F859FED424B3" box="[151,358,2016,2040]" italics="true" pageId="20" pageNumber="273">Pincerna vanbuensis</emphasis>
(Bavay &amp; Dautzenberg, 1900)
</taxonomicName>
(MNHN-IM-2012-27215, same as on Fig. 12E). All photos: B. Páll-Gergely.
</paragraph>
</caption>
<subSubSection id="4BAD8ADFFF80733FFF64FF2FFCED23FB" box="[199,863,150,177]" pageId="21" pageNumber="274" type="description">
<paragraph id="0308D954FF80733FFF64FF2FFCED23FB" blockId="21.[151,1437,150,464]" box="[199,863,150,177]" pageId="21" pageNumber="274">
<emphasis id="31C30546FF80733FFF64FF2FFE5623FB" bold="true" box="[199,484,150,176]" pageId="21" pageNumber="274">Measurements (in mm).</emphasis>
D = 4.1–4.2, H = 3.9–4.0 (n = 3).
</paragraph>
</subSubSection>
<subSubSection id="4BAD8ADFFF80733FFF64FF03FB1B23B3" pageId="21" pageNumber="274" type="etymology">
<paragraph id="0308D954FF80733FFF64FF03FB1B23B3" blockId="21.[151,1437,150,464]" pageId="21" pageNumber="274">
<emphasis id="31C30546FF80733FFF64FF03FEFC239F" bold="true" box="[199,334,186,212]" pageId="21" pageNumber="274">Etymology.</emphasis>
The specific epithet (meaning twenty ribs) refers to the key character of this species (presence of ca. 20 R2 ribs instead more numerous lighter and darker alternating stripes of
<taxonomicName id="C4B7A2D7FF80733FFBA2FF66FB2B23B3" baseAuthorityName="Bavay &amp; Dautzenberg" baseAuthorityYear="1900" box="[1025,1177,222,248]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="21" pageNumber="274" phylum="Mollusca" rank="species" species="vanbuensis">
<emphasis id="31C30546FF80733FFBA2FF66FB2B23B3" box="[1025,1177,222,248]" italics="true" pageId="21" pageNumber="274">P. vanbuensis</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="4BAD8ADFFF80733FFF64FEBBFB872257" box="[199,1077,258,285]" pageId="21" pageNumber="274" type="distribution">
<paragraph id="0308D954FF80733FFF64FEBBFB872257" blockId="21.[151,1437,150,464]" box="[199,1077,258,285]" pageId="21" pageNumber="274">
<emphasis id="31C30546FF80733FFF64FEBBFED32257" bold="true" box="[199,353,258,284]" pageId="21" pageNumber="274">Distribution.</emphasis>
This new species is known only from the
<typeStatus id="DC0C67F6FF80733FFC99FEBAFCD82256" box="[826,874,259,285]" pageId="21" pageNumber="274">type</typeStatus>
locality (
<figureCitation id="9B8CC5D1FF80733FFC76FEBBFB9A2257" box="[981,1064,258,284]" captionStart="FIGURE 13" captionStartId="19.[152,255,1356,1380]" captionTargetBox="[151,1436,237,1333]" captionTargetId="figure-54@19.[151,1436,237,1333]" captionTargetPageId="19" captionText="FIGURE 13. Distribution of Pincerna Preston, 1907 species. Circles: Pincerna vanbuensis (Bavay &amp; Dautzenberg, 1900); White rhomb: Pincerna maolanensis (Luo, Zhang &amp; Zhuo, 2009); Green triangle: Pincerna viginticostatus n. sp. Yellow circle: samples examined in this study; Red circles: Samples mentioned by Do &amp; Nguyen 2022; White circle: type locality of Pincerna clausus Do &amp; Nguyen, 2022 (synonym of P. vanbuensis). The map was made using Google Earth Pro." figureDoi="http://doi.org/10.5281/zenodo.7687448" httpUri="https://zenodo.org/record/7687448/files/figure.png" pageId="21" pageNumber="274">Fig. 13</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="4BAD8ADFFF80733FFF64FE9FFB98229B" pageId="21" pageNumber="274" type="discussion">
<paragraph id="0308D954FF80733FFF64FE9FFB98229B" blockId="21.[151,1437,150,464]" pageId="21" pageNumber="274">
<emphasis id="31C30546FF80733FFF64FE9FFE89220B" bold="true" box="[199,315,294,320]" pageId="21" pageNumber="274">Remarks.</emphasis>
This species differs from
<taxonomicName id="C4B7A2D7FF80733FFDC2FE9EFCF9220B" baseAuthorityName="Bavay &amp; Dautzenberg" baseAuthorityYear="1900" box="[609,843,294,320]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="21" pageNumber="274" phylum="Mollusca" rank="species" species="vanbuensis">
<emphasis id="31C30546FF80733FFDC2FE9EFCF9220B" box="[609,843,294,320]" italics="true" pageId="21" pageNumber="274">Pincerna vanbuensis</emphasis>
</taxonomicName>
only in the fine morphology of R2 sculpture. This is not unusual in the family
<taxonomicName id="C4B7A2D7FF80733FFE18FEF3FD8F222F" box="[443,573,330,356]" class="Gastropoda" family="Alycaeidae" kingdom="Animalia" order="Architaenioglossa" pageId="21" pageNumber="274" phylum="Mollusca" rank="family">Alycaeidae</taxonomicName>
. For example,
<taxonomicName id="C4B7A2D7FF80733FFD46FEF3FAF9222F" authority="(Kobelt &amp; Mollendorff, 1886)" baseAuthorityName="Kobelt &amp; Mollendorff" baseAuthorityYear="1886" box="[741,1355,330,356]" class="Gastropoda" family="Alycaeidae" genus="Dicharax" kingdom="Animalia" order="Architaenioglossa" pageId="21" pageNumber="274" phylum="Mollusca" rank="species" species="moellendorffi">
<emphasis id="31C30546FF80733FFD46FEF3FC5C222F" box="[741,1006,330,356]" italics="true" pageId="21" pageNumber="274">Dicharax moellendorffi</emphasis>
(Kobelt &amp;
<bibRefCitation id="6726A4A5FF80733FFBD1FEF3FAF1222F" author="Mollendorff, O. F. von" box="[1138,1347,330,356]" pageId="21" pageNumber="274" pagination="156 - 210" refId="ref15225" refString="Mollendorff, O. F. von (1886) Materialen zur Fauna for China. Jahrbucher der Deutschen Malakozoologischen Gesellschaft, 13, 156 - 210." type="journal article" year="1886">Möllendorff, 1886</bibRefCitation>
)
</taxonomicName>
differs from the otherwise very similar
<taxonomicName id="C4B7A2D7FF80733FFDB1FED7FC6522C3" authority="(Mollendorff, 1886)" baseAuthorityName="Mollendorff" baseAuthorityYear="1886" box="[530,983,366,392]" class="Gastropoda" family="Alycaeidae" genus="Dicharax" kingdom="Animalia" order="Architaenioglossa" pageId="21" pageNumber="274" phylum="Mollusca" rank="species" species="cristatus">
<emphasis id="31C30546FF80733FFDB1FED7FD5522C3" box="[530,743,366,392]" italics="true" pageId="21" pageNumber="274">Dicharax cristatus</emphasis>
(
<bibRefCitation id="6726A4A5FF80733FFD5AFED7FC7C22C3" author="Mollendorff, O. F. von" box="[761,974,366,392]" pageId="21" pageNumber="274" pagination="156 - 210" refId="ref15225" refString="Mollendorff, O. F. von (1886) Materialen zur Fauna for China. Jahrbucher der Deutschen Malakozoologischen Gesellschaft, 13, 156 - 210." type="journal article" year="1886">Möllendorff, 1886</bibRefCitation>
)
</taxonomicName>
by the smooth R2 (Páll-Gergely
<emphasis id="31C30546FF80733FFAC3FED6FA2F22C3" box="[1376,1437,366,392]" italics="true" pageId="21" pageNumber="274">et al.</emphasis>
2017). Also,
<taxonomicName id="C4B7A2D7FF80733FFE8DFE2BFD4922E7" authority="Pall-Gergely, 2017" authorityName="Pall-Gergely" authorityYear="2017" box="[302,763,402,428]" class="Gastropoda" family="Alycaeidae" genus="Metalycaeus" kingdom="Animalia" order="Architaenioglossa" pageId="21" pageNumber="274" phylum="Mollusca" rank="species" species="minatoi">
<emphasis id="31C30546FF80733FFE8DFE2BFDA922E7" box="[302,539,402,428]" italics="true" pageId="21" pageNumber="274">Metalycaeus minatoi</emphasis>
Páll-Gergely, 2017
</taxonomicName>
possesses a smooth R2, while that of
<taxonomicName id="C4B7A2D7FF80733FFB1EFE2AFF69229B" authority="(Pilsbry, 1902)" baseAuthorityName="Pilsbry" baseAuthorityYear="1902" class="Gastropoda" family="Alycaeidae" genus="Metalycaeus" kingdom="Animalia" order="Architaenioglossa" pageId="21" pageNumber="274" phylum="Mollusca" rank="species" species="vinctus">
<emphasis id="31C30546FF80733FFB1EFE2AFA8622E7" box="[1213,1332,403,428]" italics="true" pageId="21" pageNumber="274">M. vinctus</emphasis>
(
<bibRefCitation id="6726A4A5FF80733FFAE5FE2BFF61229B" author="Pilsbry, H. A." pageId="21" pageNumber="274" pagination="53 - 57" refId="ref15883" refString="Pilsbry, H. A. (1902) New land molluscs from the Japanese Empire. The Nautilus, 16, 53 - 57." type="journal article" year="1902">Pilsbry, 1902</bibRefCitation>
)
</taxonomicName>
, which is its putative sister species, is ribbed (
<bibRefCitation id="6726A4A5FF80733FFD7CFE0FFBAD229B" author="Pall-Gergely, B. &amp; Asami, T." box="[735,1055,438,464]" pageId="21" pageNumber="274" pagination="1 - 16" refId="ref15401" refString="Pall-Gergely, B. &amp; Asami, T. (2017) A new species of Metalycaeus hitherto undistinguished from M. vinctus (Pilsbry, 1902). Venus, 75 (1 - 4), 1 - 16." type="journal article" year="2017">Páll-Gergely &amp; Asami 2017</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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