176 lines
19 KiB
XML
176 lines
19 KiB
XML
<document id="105B106ADA9AC01CB85114758D446873" ID="10.11646/zootaxa.4179.3.10" ID-DOI="10.11646/zootaxa.4179.3.10" ID-GBIF-Dataset="3aab38df-01c7-4489-875f-31c510b24e19" ID-ISSN="1175-5326" ID-Zenodo-Dep="258837" ID-ZooBank="0D339758-0B56-4CEB-A0CC-567E1321F4B8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1477994955485" checkinUser="plazi" docAuthor="Fominykh, Dmitriy D. & Zamotajlov, Alexandr S." docDate="2016" docId="695D0E767C1BFFC624DF5D46797AFE41" docLanguage="en" docName="zootaxa.4179.3.10.pdf" docOrigin="Zootaxa 4179 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Carabus (Neoplectes) martviliensis Retezar & Djavelidze 1992" docType="treatment" docVersion="7" lastPageNumber="567" masterDocId="9564760E7C1EFFC024485B717C16FFEC" masterDocTitle="Further contribution to the knowledge of Georgian species of the Carabus Linnaeus, 1758 subgenus Neoplectes Reitter, 1885, with description of a new subspecies of C. titarenkoi Zamotajlov & Fominykh, 2014 (Coleoptera: Carabidae: Carabini)" masterLastPageNumber="599" masterPageNumber="561" pageNumber="566" updateTime="1698391590155" updateUser="plazi">
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<mods:title id="D35915DB172AB42573CA8E3E3CB43C73">Further contribution to the knowledge of Georgian species of the Carabus Linnaeus, 1758 subgenus Neoplectes Reitter, 1885, with description of a new subspecies of C. titarenkoi Zamotajlov & Fominykh, 2014 (Coleoptera: Carabidae: Carabini)</mods:title>
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<mods:name id="B955660C73224A935D4DD2E748E8F4FA" type="personal">
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<mods:namePart id="B548C123AD12C35E3260351C9EE1947D">Fominykh, Dmitriy D.</mods:namePart>
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<mods:name id="D299546DBCEB0A9C94140E5526043238" type="personal">
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<mods:namePart id="2919833EC21D55C24D00643D7B7D2464">Zamotajlov, Alexandr S.</mods:namePart>
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<mods:typeOfResource id="86EF4E5413EEDA4BAD55FD329EE2AF5D">text</mods:typeOfResource>
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<mods:title id="2553B1234A9609E78802207FE77A7406">Zootaxa</mods:title>
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<mods:date id="A18DE8C41CCE83F1AFC4DFC5C4605AB5">2016</mods:date>
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<mods:number id="01CDB75FF6E289B0090D72FA6D9A66C1">4179</mods:number>
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<mods:identifier id="2EDECB3C8A79496B637D2F31030519F3" type="DOI">10.11646/zootaxa.4179.3.10</mods:identifier>
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<treatment id="695D0E767C1BFFC624DF5D46797AFE41" ID-DOI="http://doi.org/10.5281/zenodo.6071827" ID-GBIF-Taxon="124560921" ID-Zenodo-Dep="6071827" LSID="urn:lsid:plazi:treatment:695D0E767C1BFFC624DF5D46797AFE41" httpUri="http://treatment.plazi.org/id/695D0E767C1BFFC624DF5D46797AFE41" lastPageId="6" lastPageNumber="567" pageId="5" pageNumber="566">
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<subSubSection id="A9EEECEB7C1BFFC524DF5D467DC0F99F" pageId="5" pageNumber="566" type="nomenclature">
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<paragraph id="E14BBF607C1BFFC524DF5D46782EF9BE" blockId="5.[151,1080,1591,1651]" box="[151,1080,1591,1618]" pageId="5" pageNumber="566">
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<heading id="BA03080C7C1BFFC524DF5D46782EF9BE" bold="true" box="[151,1080,1591,1618]" fontSize="11" level="1" pageId="5" pageNumber="566" reason="1">
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<emphasis id="D38063727C1BFFC524DF5D46782EF9BE" bold="true" box="[151,1080,1591,1618]" pageId="5" pageNumber="566">
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<taxonomicName id="26F4C4E37C1BFFC524DF5D467F8CF9BE" ID-CoL="8MJ3X" authority="Retezar & Djavelidze, 1992" authorityName="Retezar & Djavelidze" authorityYear="1992" box="[151,922,1591,1618]" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="566" phylum="Arthropoda" rank="species" species="martviliensis" subGenus="Neoplectes">
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<emphasis id="D38063727C1BFFC524DF5D467E2BF9BD" bold="true" box="[151,573,1591,1617]" italics="true" pageId="5" pageNumber="566">Carabus (Neoplectes) martviliensis</emphasis>
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Retezár & Djavelidze, 1992
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</taxonomicName>
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, stat. resurr.
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="E14BBF607C1BFFC524DF5D2B7DC0F99F" blockId="5.[151,1080,1591,1651]" box="[151,470,1626,1651]" pageId="5" pageNumber="566">
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(
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<figureCitation id="79CFA3E57C1BFFC524D75D2B7D31F99F" box="[159,295,1626,1651]" captionStart="FIGURES 51 – 55" captionStartId="16.[151,264,1967,1990]" captionTargetBox="[278,1316,210,1929]" captionTargetId="figure@16.[258,1328,193,1947]" captionTargetPageId="16" captionText="FIGURES 51 – 55. Dorsal habitus of Carabus martviliensis Retezár & Djavelidze: 51 — Kinchkha-Perdi env.; 52 – 55 — Muchera env. Scale bar 10.0 mm." httpUri="https://zenodo.org/record/258846/files/figure.png" pageId="5" pageNumber="566">Figs. 51–55</figureCitation>
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,
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<figureCitation id="79CFA3E57C1BFFC5257A5D2B7D81F99F" box="[306,407,1626,1651]" captionStart-0="FIGURES 153 – 154" captionStart-1="FIGURES 155 – 158" captionStartId-0="27.[151,264,923,946]" captionStartId-1="27.[151,264,1960,1983]" captionTargetBox-0="[213,1364,201,875]" captionTargetBox-1="[361,1226,1036,1922]" captionTargetId-0="figure@27.[200,1377,193,902]" captionTargetId-1="figure@27.[340,1248,1026,1939]" captionTargetPageId-0="27" captionTargetPageId-1="27" captionText-0="FIGURES 153 – 154. Apical portion of aedeagus and fully inflated endophallus preparation of Carabus martviliensis Retezár & Djavelidze, Kinchkha-Perdi env.: 153 — right lateral view; 154 — dorsolateral view. Scale bar 10.0 mm." captionText-1="FIGURES 155 – 158. Apical portion of aedeagus and fully inflated endophallus preparation of Carabus martviliensis Retezár & Djavelidze, Muchera env.: 155 – 156 — right lateral view; 157 – 158 — dorsolateral view. Scale bar 10.0 mm." httpUri-0="https://zenodo.org/record/258863/files/figure.png" httpUri-1="https://zenodo.org/record/258864/files/figure.png" pageId="5" pageNumber="566">153–158</figureCitation>
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,
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<figureCitation id="79CFA3E57C1BFFC525EA5D2A7DDBF99F" box="[418,461,1627,1651]" captionStart="FIGURE 210" captionStartId="35.[151,250,2016,2038]" captionTargetBox="[150,1435,1140,1993]" captionTargetId="figure@35.[150,1436,1140,1994]" captionTargetPageId="35" captionText="FIGURE 210. Carabus martviliensis Retezár & Djavelidze in nature, Georgia, Muchera env., 1000 m." httpUri="https://zenodo.org/record/258880/files/figure.png" pageId="5" pageNumber="566">210</figureCitation>
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)
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</paragraph>
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</subSubSection>
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||
<subSubSection id="A9EEECEB7C1BFFC524DF5DD37CEAF8A7" pageId="5" pageNumber="566" type="materials_examined">
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<paragraph id="E14BBF607C1BFFC524DF5DD37CEAF8A7" blockId="5.[151,1437,1698,2011]" pageId="5" pageNumber="566">
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<materialsCitation id="519CB53D7C1BFFC524DF5DD37E77F8EF" ID-GBIF-Occurrence="1324406384" collectingDate="2015-06-20" collectionCode="CFDD, CTAY" collectorName="D. Fominykh & A. Zubov" country="Georgia" county="Muchera Mt." elevation="1009" latitude="42.52589" location="Samegrelo" longLatPrecision="1" longitude="42.414223" municipality="Meore-Balda" pageId="5" pageNumber="566" specimenCount="40" specimenCount-female="22" specimenCount-male="18">
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<emphasis id="D38063727C1BFFC524DF5DD37D10F957" bold="true" box="[151,262,1698,1723]" pageId="5" pageNumber="566">Material.</emphasis>
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||
<specimenCount id="F7F274E97C1BFFC5255A5DD27D4CF957" box="[274,346,1699,1723]" pageId="5" pageNumber="566" type="male">18♂♂</specimenCount>
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||
,
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<specimenCount id="F7F274E97C1BFFC525235DD27DA4F957" box="[363,434,1699,1723]" pageId="5" pageNumber="566" type="female">22♀♀</specimenCount>
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,
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<collectingCountry id="99E3FFF07C1BFFC5258C5DD37E37F957" box="[452,545,1698,1723]" name="Georgia" pageId="5" pageNumber="566">Georgia</collectingCountry>
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,
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<location id="E42BE9BB7C1BFFC526795DD37EBAF957" LSID="urn:lsid:plazi:treatment:695D0E767C1BFFC624DF5D46797AFE41:E42BE9BB7C1BFFC526795DD37EBAF957" box="[561,684,1698,1723]" country="Georgia" county="Muchera Mt." latitude="42.52589" longLatPrecision="1" longitude="42.414223" municipality="Meore-Balda" name="Samegrelo" pageId="5" pageNumber="566">Samegrelo</location>
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,
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||
<quantity id="260C12857C1BFFC526F45DD37F19F957" box="[700,783,1698,1723]" metricMagnitude="3" metricUnit="m" metricValue="3.5" pageId="5" pageNumber="566" unit="km" value="3.5">3.5 km</quantity>
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of
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<collectingMunicipality id="012F251A7C1BFFC527765DD37FC4F957" box="[830,978,1698,1723]" pageId="5" pageNumber="566">Meore-Balda</collectingMunicipality>
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||
vill.,
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||
<locationDeviation id="F0F9A6427C1BFFC520555DD3798AF957" box="[1053,1436,1698,1723]" pageId="5" pageNumber="566">between Abasha and Chachkheri</locationDeviation>
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rivers, SE slopes of
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<collectingCounty id="082AC7EC7C1BFFC525375DB77E00F932" box="[383,534,1734,1759]" pageId="5" pageNumber="566">Muchera Mt.</collectingCounty>
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,
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||
<quantity id="260C12857C1BFFC5266B5DB67E6AF932" box="[547,636,1735,1759]" metricMagnitude="3" metricUnit="m" metricValue="1.009" pageId="5" pageNumber="566" unit="m" value="1009.0">
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||
<elevation id="6AD958537C1BFFC5266B5DB67E6AF932" box="[547,636,1735,1759]" metricMagnitude="3" metricUnit="m" metricValue="1.009" pageId="5" pageNumber="566" unit="m" value="1009.0">1009 m</elevation>
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</quantity>
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,
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||
<geoCoordinate id="84C0D9A77C1BFFC526C15DB67F2FF933" box="[649,825,1735,1759]" direction="north" orientation="latitude" pageId="5" pageNumber="566" precision="1" value="42.52589">N 42°31'33.20"</geoCoordinate>
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/
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<geoCoordinate id="84C0D9A77C1BFFC5271B5DB67814F933" box="[851,1026,1734,1759]" direction="east" orientation="longitude" pageId="5" pageNumber="566" precision="1" value="42.414223">E 42°24'51.20"</geoCoordinate>
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,
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||
<collectingMethod id="38B5C7777C1BFFC520475DB77882F932" box="[1039,1172,1734,1759]" pageId="5" pageNumber="566">pitfall traps</collectingMethod>
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||
, 15.04.-
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||
<collectingDate id="850E60487C1BFFC520A35DB77979F933" box="[1259,1391,1734,1759]" pageId="5" pageNumber="566" value="2015-06-20">
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||
<date id="954A99A07C1BFFC520A35DB77979F933" box="[1259,1391,1734,1759]" pageId="5" pageNumber="566" value="2015-06-20">20.06.2015</date>
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</collectingDate>
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||
(
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<collectorName id="4C01DAB67C1BFFC521C95DB67D1CF8EF" pageId="5" pageNumber="566">D. Fominykh</collectorName>
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||
&
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||
<collectorName id="4C01DAB67C1BFFC525665D9A7D8AF8EF" box="[302,412,1770,1795]" pageId="5" pageNumber="566">A. Zubov</collectorName>
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||
) (
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||
<collectionCode id="87E527A57C1BFFC525FA5D9A7E17F8EF" box="[434,513,1771,1795]" pageId="5" pageNumber="566">CFDD</collectionCode>
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||
,
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||
<collectionCode id="87E527A57C1BFFC526455D9A7E4CF8EF" box="[525,602,1771,1795]" pageId="5" pageNumber="566">CTAY</collectionCode>
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||
)
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||
</materialsCitation>
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||
;
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<materialsCitation id="519CB53D7C1BFFC526245D9A7CEEF8A7" ID-GBIF-Occurrence="1324406385" collectingDate="2015-06-20" collectionCode="CFDD" collectorName="D. Fominykh & A. Zubov" country="Georgia" county="Satsikvilo" elevation="816" latitude="42.483334" location="Samegrelo" longLatPrecision="1149" longitude="42.55" municipality="Okatse" pageId="5" pageNumber="566" specimenCount="1" specimenCount-male="1">
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||
<specimenCount id="F7F274E97C1BFFC526245D9A7E87F8EF" box="[620,657,1771,1795]" pageId="5" pageNumber="566" type="male">1♂</specimenCount>
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,
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<collectingCountry id="99E3FFF07C1BFFC526D65D9B7EEDF8EF" box="[670,763,1770,1795]" name="Georgia" pageId="5" pageNumber="566">Georgia</collectingCountry>
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,
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||
<location id="E42BE9BB7C1BFFC5274E5D9B7F97F8EF" LSID="urn:lsid:plazi:treatment:695D0E767C1BFFC624DF5D46797AFE41:E42BE9BB7C1BFFC5274E5D9B7F97F8EF" box="[774,897,1770,1795]" country="Georgia" county="Satsikvilo" latitude="42.483334" longLatPrecision="1149" longitude="42.55" municipality="Okatse" name="Samegrelo" pageId="5" pageNumber="566">Samegrelo</location>
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||
, near
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||
<location id="E42BE9BB7C1BFFC5278B5D9B7862F8EF" LSID="urn:lsid:plazi:treatment:695D0E767C1BFFC624DF5D46797AFE41:E42BE9BB7C1BFFC5278B5D9B7862F8EF" box="[963,1140,1770,1795]" country="Georgia" county="Satsikvilo" latitude="42.483334" longLatPrecision="1149" longitude="42.55" municipality="Okatse" name="Kinchkha-Perdi" pageId="5" pageNumber="566">Kinchkha-Perdi</location>
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||
vill., right bank of
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||
<collectingMunicipality id="012F251A7C1BFFC521065D9B798AF8EF" box="[1358,1436,1770,1795]" pageId="5" pageNumber="566">Okatse</collectingMunicipality>
|
||
(=
|
||
<collectingCounty id="082AC7EC7C1BFFC524E65C7F7D34F8CB" box="[174,290,1806,1831]" pageId="5" pageNumber="566">Satsikvilo</collectingCounty>
|
||
) river,
|
||
<quantity id="260C12857C1BFFC525365C7F7DDBF8CA" box="[382,461,1806,1831]" metricMagnitude="2" metricUnit="m" metricValue="8.16" pageId="5" pageNumber="566" unit="m" value="816.0">
|
||
<elevation id="6AD958537C1BFFC525365C7F7DDBF8CA" box="[382,461,1806,1831]" metricMagnitude="2" metricUnit="m" metricValue="8.16" pageId="5" pageNumber="566" unit="m" value="816.0">816 m</elevation>
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||
</quantity>
|
||
,
|
||
<geoCoordinate id="84C0D9A77C1BFFC525975C7E7E5CF8CB" box="[479,586,1807,1831]" direction="north" orientation="latitude" pageId="5" pageNumber="566" precision="925" value="42.483334">N 42°29'</geoCoordinate>
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/
|
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<geoCoordinate id="84C0D9A77C1BFFC526225C7E7EDEF8CB" box="[618,712,1807,1831]" direction="east" orientation="longitude" pageId="5" pageNumber="566" precision="925" value="42.55">E42°33'</geoCoordinate>
|
||
,
|
||
<collectingMethod id="38B5C7777C1BFFC526905C7F7F77F8CA" box="[728,865,1806,1831]" pageId="5" pageNumber="566">pitfall traps</collectingMethod>
|
||
, 15.04.-
|
||
<date id="954A99A07C1BFFC527F45C7F7856F8CB" box="[956,1088,1806,1831]" pageId="5" pageNumber="566" value="2015-06-20">
|
||
<collectingDate id="850E60487C1BFFC527F45C7F7856F8CB" box="[956,1088,1806,1831]" pageId="5" pageNumber="566" value="2015-06-20">20.06.2015</collectingDate>
|
||
</date>
|
||
(
|
||
<collectorName id="4C01DAB67C1BFFC5201E5C7E78E7F8CB" box="[1110,1265,1806,1831]" pageId="5" pageNumber="566">D. Fominykh</collectorName>
|
||
&
|
||
<collectorName id="4C01DAB67C1BFFC521695C7E7983F8CB" box="[1313,1429,1806,1831]" pageId="5" pageNumber="566">A. Zubov</collectorName>
|
||
) (
|
||
<collectionCode id="87E527A57C1BFFC524D75C427CE6F8A7" box="[159,240,1843,1867]" pageId="5" pageNumber="566">CFDD</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A9EEECEB7C1BFFC6248F5C277930FF39" lastPageId="6" lastPageNumber="567" pageId="5" pageNumber="566" type="distribution">
|
||
<paragraph id="E14BBF607C1BFFC6248F5C277930FF39" blockId="5.[151,1437,1698,2011]" lastBlockId="6.[151,1436,151,429]" lastPageId="6" lastPageNumber="567" pageId="5" pageNumber="566">
|
||
<emphasis id="D38063727C1BFFC5248F5C277DEAF883" bold="true" box="[199,508,1878,1903]" pageId="5" pageNumber="566">Distribution and habitat.</emphasis>
|
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Described from Tekhuri valley in Martvili vicinities. Inhabits the eastern and southern slopes of the Askhi Plateau between the left bank of the Tekhuri river and the right bank of the Tskhenis- Tskhali river. The specimens listed above were collected in a beech forest at
|
||
<quantity id="260C12857C1BFFC527BC5CEF7892F85A" box="[1012,1156,1950,1975]" metricMagnitude="2" metricUnit="m" metricValue="9.25" metricValueMax="10.5" metricValueMin="8.0" pageId="5" pageNumber="566" unit="m" value="925.0" valueMax="1050.0" valueMin="800.0">800–1050 m</quantity>
|
||
. Two sites were sampled in the vicinity of Mt. Muchera, the first a climax forest community, and the second a site with missing leaf litter as a result of a recent fire. The latter site had a population density about five times higher than the climax community, probably demonstrating the population phenomenon earlier discussed by
|
||
<bibRefCitation id="8565C2917C18FFC627845BCD7934FF39" author="Knysh" box="[972,1314,188,213]" pageId="6" pageNumber="567" refString="Knysh, V. G. & Zamotajlov, A. S. (2005) Behavioral strategy of two ground beetle species of the genus Carabus (Coleoptera, Carabidae) colonizing some types of natural and anthropogenic biocenoses. Entomological Review, 85, 835 - 844." type="journal article" year="2005">Knysh and Zamotajlov (2005)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A9EEECEB7C18FFC6248F5BAE797AFE41" pageId="6" pageNumber="567" type="discussion">
|
||
<paragraph id="E14BBF607C18FFC6248F5BAE7E17FE89" blockId="6.[151,1436,151,429]" pageId="6" pageNumber="567">
|
||
<emphasis id="D38063727C18FFC6248F5BAE7D2DFF14" bold="true" box="[199,315,223,248]" pageId="6" pageNumber="567">Remarks.</emphasis>
|
||
A single male specimen with an extremely small body size was collected near Kinchkha-Perdi village. This is probably an aberrant individual, but further specimens of this population would need to be sampled to confirm this. Two specimens were collected at Muchera Mountain Mass, which possessed a rare bright green pronotal and elytral colouration.
|
||
</paragraph>
|
||
<paragraph id="E14BBF607C18FFC6248F5A01797AFE41" blockId="6.[151,1436,151,429]" pageId="6" pageNumber="567">
|
||
We resurrect the specific status of this form based on the consistent differences in external morphology and male genitalia structure between
|
||
<taxonomicName id="26F4C4E37C18FFC626415AE47EADFE41" box="[521,699,404,429]" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="567" phylum="Arthropoda" rank="species" species="martviliensis">
|
||
<emphasis id="D38063727C18FFC626415AE47EADFE41" box="[521,699,404,429]" italics="true" pageId="6" pageNumber="567">C. martviliensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="26F4C4E37C18FFC626B95AE47F40FE41" box="[753,854,404,429]" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="567" phylum="Arthropoda" rank="species" species="mellyi">
|
||
<emphasis id="D38063727C18FFC626B95AE47F40FE41" box="[753,854,404,429]" italics="true" pageId="6" pageNumber="567">C. mellyi</emphasis>
|
||
</taxonomicName>
|
||
(see identification key and illustrations below).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |