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<document id="D0CC6A2D7E4F44AF8035D807DCEFA14E" ID-CLB-Dataset="32753" ID-DOI="10.11646/zootaxa.4286.2.3" ID-GBIF-Dataset="dfbcd01b-2ea6-4898-af4b-bb8ee425d26b" ID-ISSN="1175-5326" ID-Zenodo-Dep="828403" ID-ZooBank="2B3C978D-C70B-4A31-9F2B-8E082FBBE51B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1500108776027" checkinUser="plazi" docAuthor="Nilsai, Areeruk, Jantarit, Sopark, Satasook, Chutamas &amp; Zhang, Feng" docDate="2017" docId="3E0D8E51FFFAFFCAAAD8FACAFB1B5ACD" docLanguage="en" docName="zootaxa.4286.2.3.pdf" docOrigin="Zootaxa 4286 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Coecobrya" docType="treatment" docVersion="8" lastPageNumber="200" masterDocId="C234F629FFF7FFC4AA4FFFB1FFB05D75" masterDocTitle="Three new species of Coecobrya (Collembola: Entomobryidae) from caves in the Thai Peninsula" masterLastPageNumber="202" masterPageNumber="187" pageNumber="200" updateTime="1698465156621" updateUser="ExternalLinkService">
<mods:mods id="CDCC86F061F1DAFF76004D9D7CE80D3B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="9728D72B6EEE9E55F09D6248E32139B7">
<mods:title id="3E68254B7E1FBD35AE69F6405CA88EED">Three new species of Coecobrya (Collembola: Entomobryidae) from caves in the Thai Peninsula</mods:title>
</mods:titleInfo>
<mods:name id="0B3B5F3ABDC23506EC54B7FD829A48C2" type="personal">
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<mods:namePart id="5BA8E6362BF8A283B0FE290DD6A95B44">Nilsai, Areeruk</mods:namePart>
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<mods:name id="AB54AF33F08C170D4461A08F06E2E936" type="personal">
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<mods:namePart id="B2A63A650A2C55505AD2074253AFA497">Jantarit, Sopark</mods:namePart>
</mods:name>
<mods:name id="14E857A35637CFBE02365EBFBAE9356D" type="personal">
<mods:role id="9DD691424317C8A44E2AA64FE2E0DE9B">
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<mods:namePart id="A2EF16B60617021DC74586B2D7B49802">Satasook, Chutamas</mods:namePart>
</mods:name>
<mods:name id="2B44263CBE18A9972608F6B51419533D" type="personal">
<mods:role id="865E280E7B312A60CA3EF68D18573D46">
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</mods:role>
<mods:namePart id="4C4811DC919B16E7372BE17CA1D0C5E7">Zhang, Feng</mods:namePart>
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<mods:title id="79DF74AA0EA3D5823D1A20F08DD43FE1">Zootaxa</mods:title>
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<mods:part id="009299AA8E6AD2D926ACFC34FB4D74A1">
<mods:date id="62CA8351902A7027198CE8BFDE1DEF50">2017</mods:date>
<mods:detail id="689AF3BE98271C87E8F3BE4D20448CA4" type="volume">
<mods:number id="0E7F375F23A81AF67ABA657FEEF13BC5">4286</mods:number>
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<mods:classification id="5274D358277B7EE99DBC46DE87819839">journal article</mods:classification>
<mods:identifier id="47D2AC341053B3C87DBFA20B45A1ADEB" type="CLB-Dataset">32753</mods:identifier>
<mods:identifier id="7E3895456046D596D601CDBA20CFBA89" type="DOI">10.11646/zootaxa.4286.2.3</mods:identifier>
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<treatment id="3E0D8E51FFFAFFCAAAD8FACAFB1B5ACD" ID-DOI="http://doi.org/10.5281/zenodo.6045295" ID-GBIF-Taxon="132548719" ID-Zenodo-Dep="6045295" LSID="urn:lsid:plazi:treatment:3E0D8E51FFFAFFCAAAD8FACAFB1B5ACD" httpUri="http://treatment.plazi.org/id/3E0D8E51FFFAFFCAAAD8FACAFB1B5ACD" lastPageId="14" lastPageNumber="200" pageId="13" pageNumber="200">
<subSubSection id="FEBE6CCCFFFAFFC9AAD8FACAFE1258C3" pageId="13" pageNumber="200" type="nomenclature">
<paragraph id="B61B3F47FFFAFFC9AAD8FACAFDEF58E1" blockId="13.[151,607,1402,1462]" box="[151,607,1402,1429]" pageId="13" pageNumber="200">
<heading id="ED53882BFFFAFFC9AAD8FACAFDEF58E1" bold="true" box="[151,607,1402,1429]" fontSize="11" level="1" pageId="13" pageNumber="200" reason="1">
<emphasis id="84D0E355FFFAFFC9AAD8FACAFDEF58E1" bold="true" box="[151,607,1402,1429]" pageId="13" pageNumber="200">
Key to the Thai species of
<taxonomicName id="71A444C4FFFAFFC9AB91FACBFDEF58E1" box="[478,607,1402,1428]" class="Entognatha" family="Entomobryidae" genus="Coecobrya" kingdom="Animalia" order="Collembola" pageId="13" pageNumber="200" phylum="Arthropoda" rank="genus">
<emphasis id="84D0E355FFFAFFC9AB91FACBFDEF58E1" bold="true" box="[478,607,1402,1428]" italics="true" pageId="13" pageNumber="200">Coecobrya</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="B61B3F47FFFAFFC9AAD8FA2CFE1258C3" blockId="13.[151,607,1402,1462]" box="[151,418,1437,1462]" pageId="13" pageNumber="200">*Cave-dwelling species</paragraph>
</subSubSection>
<keyStep id="0D5027E2FFFAFFC9AAD8FA6CFA2B5B64" pageId="13" pageNumber="200">
<subSubSection id="FEBE6CCCFFFAFFC9AAD8FA6CFA2C5886" box="[151,1436,1501,1523]" pageId="13" pageNumber="200" type="key">
<paragraph id="B61B3F47FFFAFFC9AAD8FA6CFA2C5886" blockId="13.[151,1436,1501,1914]" box="[151,1436,1501,1523]" pageId="13" pageNumber="200">
<keyLead id="0D559C72FFFAFFC9AAD8FA6CFA2C5886" box="[151,1436,1501,1523]" pageId="13" pageNumber="200">1. Antennae more than 2.5 times as long as the cephalic diagonal..................................................2</keyLead>
</paragraph>
</subSubSection>
<paragraph id="B61B3F47FFFAFFC9AAD8FA4DFA2B5B64" blockId="13.[151,1436,1501,1914]" box="[151,1435,1531,1553]" pageId="13" pageNumber="200">
<keyLead id="0D559C72FFFAFFC9AAD8FA4DFA2B5B64" box="[151,1435,1531,1553]" pageId="13" pageNumber="200">- Antennae less than 2.5 times as long as the cephalic diagonal...................................................5</keyLead>
</paragraph>
</keyStep>
<keyStep id="0D5027E2FFFAFFC9AAD8F9A9FA2B5B3E" pageId="13" pageNumber="200">
<paragraph id="B61B3F47FFFAFFC9AAD8F9A9FA2B5B5B" blockId="13.[151,1436,1501,1914]" box="[151,1435,1560,1583]" pageId="13" pageNumber="200">
<keyLead id="0D559C72FFFAFFC9AAD8F9A9FA2B5B5B" box="[151,1435,1560,1583]" pageId="13" pageNumber="200">
2. Unguis without unpaired inner teeth...........................................
<taxonomicName id="71A444C4FFFAFFC9AE0EF9A8FA8D5B5B" authority="Nilsai &amp; Zhang" authorityName="Nilsai &amp; Zhang" box="[1089,1341,1560,1582]" class="Entognatha" family="Entomobryidae" genus="Coecobrya" kingdom="Animalia" order="Collembola" pageId="13" pageNumber="200" phylum="Arthropoda" rank="species" species="cavicta" status="sp. nov.">
<emphasis id="84D0E355FFFAFFC9AE0EF9A8FB125B5B" box="[1089,1186,1561,1582]" italics="true" pageId="13" pageNumber="200">C. cavicta</emphasis>
Nilsai &amp; Zhang
</taxonomicName>
<taxonomicNameLabel id="9FE35E2EFFFAFFC9AF0CF9A8FA3F5B5A" box="[1347,1423,1561,1583]" pageId="13" pageNumber="200" rank="species">
<emphasis id="84D0E355FFFAFFC9AF0CF9A8FA3F5B5A" bold="true" box="[1347,1423,1561,1583]" pageId="13" pageNumber="200">sp. nov.</emphasis>
</taxonomicNameLabel>
*
</keyLead>
</paragraph>
<paragraph id="B61B3F47FFFAFFC9AAD8F987FA2B5B3E" blockId="13.[151,1436,1501,1914]" box="[151,1435,1589,1611]" pageId="13" pageNumber="200">
<keyLead id="0D559C72FFFAFFC9AAD8F987FA2B5B3E" box="[151,1435,1589,1611]" pageId="13" pageNumber="200">- Unguis with unpaired inner tooth.........................................................................3</keyLead>
</paragraph>
</keyStep>
<keyStep id="0D5027E2FFFAFFC9AAD8F9E2FA2C5BFC" pageId="13" pageNumber="200">
<paragraph id="B61B3F47FFFAFFC9AAD8F9E2FA2B5B1C" blockId="13.[151,1436,1501,1914]" box="[151,1435,1619,1645]" pageId="13" pageNumber="200">
<keyLead id="0D559C72FFFAFFC9AAD8F9E2FA2B5B1C" box="[151,1435,1619,1645]" pageId="13" pageNumber="200">
3. Labial chaetae as mRel1l2; Abd. IV with 7+7 central mac...................
<taxonomicName id="71A444C4FFFAFFC9A9BDF9E5FA8D5B1C" authority="Zhang &amp; Nilsai" authorityName="Zhang &amp; Nilsai" box="[1010,1341,1619,1641]" class="Entognatha" family="Entomobryidae" genus="Coecobrya" kingdom="Animalia" order="Collembola" pageId="13" pageNumber="200" phylum="Arthropoda" rank="species" species="chumphonensis" status="sp. nov.">
<emphasis id="84D0E355FFFAFFC9A9BDF9E5FB125B1C" box="[1010,1186,1620,1641]" italics="true" pageId="13" pageNumber="200">C. chumphonensis</emphasis>
Zhang &amp; Nilsai
</taxonomicName>
<taxonomicNameLabel id="9FE35E2EFFFAFFC9AF0CF9E2FA3F5B1C" box="[1347,1423,1619,1641]" pageId="13" pageNumber="200" rank="species">
<emphasis id="84D0E355FFFAFFC9AF0CF9E2FA3F5B1C" bold="true" box="[1347,1423,1619,1641]" pageId="13" pageNumber="200">sp. nov.</emphasis>
</taxonomicNameLabel>
*
</keyLead>
</paragraph>
<paragraph id="B61B3F47FFFAFFC9AAD8F9C2FA2C5BFC" blockId="13.[151,1436,1501,1914]" box="[151,1436,1651,1676]" pageId="13" pageNumber="200">
<keyLead id="0D559C72FFFAFFC9AAD8F9C2FA2C5BFC" box="[151,1436,1651,1676]" pageId="13" pageNumber="200">
- Labial chaetae as M
<quantity id="715C92A2FFFAFFC9ABEEF931FE0A5BFD" box="[417,442,1651,1676]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="200" unit="m" value="1.0">1m</quantity>
2rel1l2; Abd. IV with 6+6 central mac....................................................4
</keyLead>
</paragraph>
</keyStep>
<keyStep id="0D5027E2FFFAFFC9AAD8F923FA2B5BBD" pageId="13" pageNumber="200">
<paragraph id="B61B3F47FFFAFFC9AAD8F923FA2B5BDD" blockId="13.[151,1436,1501,1914]" box="[151,1435,1682,1708]" pageId="13" pageNumber="200">
<keyLead id="0D559C72FFFAFFC9AAD8F923FA2B5BDD" box="[151,1435,1682,1708]" pageId="13" pageNumber="200">
4. Postlabial chaeta
<date id="C21A1987FFFAFFC9ABC6F922FE115BD9" box="[393,417,1683,1708]" pageId="13" pageNumber="200">X4</date>
ciliate; mac a1 present on Abd. I..........................
<taxonomicName id="71A444C4FFFAFFC9AE52F922FA8D5BDD" authority="Zhang &amp; Nilsai" authorityName="Zhang &amp; Nilsai" box="[1053,1341,1682,1704]" class="Entognatha" family="Entomobryidae" genus="Coecobrya" kingdom="Animalia" order="Collembola" pageId="13" pageNumber="200" phylum="Arthropoda" rank="species" species="polychaeta" status="sp. nov.">
<emphasis id="84D0E355FFFAFFC9AE52F922FB125BDD" box="[1053,1186,1683,1704]" italics="true" pageId="13" pageNumber="200">C. polychaeta</emphasis>
Zhang &amp; Nilsai
</taxonomicName>
<taxonomicNameLabel id="9FE35E2EFFFAFFC9AF0CF923FA3F5BDD" box="[1347,1423,1682,1704]" pageId="13" pageNumber="200" rank="species">
<emphasis id="84D0E355FFFAFFC9AF0CF923FA3F5BDD" bold="true" box="[1347,1423,1682,1704]" pageId="13" pageNumber="200">sp. nov.</emphasis>
</taxonomicNameLabel>
*
</keyLead>
</paragraph>
<paragraph id="B61B3F47FFFAFFC9AAD8F902FA2B5BBD" blockId="13.[151,1436,1501,1914]" box="[151,1435,1714,1740]" pageId="13" pageNumber="200">
<keyLead id="0D559C72FFFAFFC9AAD8F902FA2B5BBD" box="[151,1435,1714,1740]" pageId="13" pageNumber="200">
- Postlabial chaeta
<date id="C21A1987FFFAFFC9ABC6F902FE115BB9" box="[393,417,1715,1740]" pageId="13" pageNumber="200">X4</date>
smooth; mac a1 absent on Abd. I.......................
<taxonomicName id="71A444C4FFFAFFC9A9B1F902FA8D5BBD" authority="Zhang &amp; Nilsai" authorityName="Zhang &amp; Nilsai" box="[1022,1341,1714,1736]" class="Entognatha" family="Entomobryidae" genus="Coecobrya" kingdom="Animalia" order="Collembola" pageId="13" pageNumber="200" phylum="Arthropoda" rank="species" species="polychaeta" status="sp. nov.">
<emphasis id="84D0E355FFFAFFC9A9B1F902FBBE5BBD" box="[1022,1038,1715,1736]" italics="true" pageId="13" pageNumber="200">C</emphasis>
. cf.
<emphasis id="84D0E355FFFAFFC9AE76F902FB125BBD" box="[1081,1186,1715,1736]" italics="true" pageId="13" pageNumber="200">polychaeta</emphasis>
Zhang &amp; Nilsai
</taxonomicName>
<taxonomicNameLabel id="9FE35E2EFFFAFFC9AF0CF903FA3F5BBD" box="[1347,1423,1714,1736]" pageId="13" pageNumber="200" rank="species">
<emphasis id="84D0E355FFFAFFC9AF0CF903FA3F5BBD" bold="true" box="[1347,1423,1714,1736]" pageId="13" pageNumber="200">sp. nov.</emphasis>
</taxonomicNameLabel>
*
</keyLead>
</paragraph>
</keyStep>
<keyStep id="0D5027E2FFFAFFC9AAD8F963FA2B5A70" pageId="13" pageNumber="200">
<paragraph id="B61B3F47FFFAFFC9AAD8F963FA2C5B9D" blockId="13.[151,1436,1501,1914]" box="[151,1436,1746,1768]" pageId="13" pageNumber="200">
<keyLead id="0D559C72FFFAFFC9AAD8F963FA2C5B9D" box="[151,1436,1746,1768]" pageId="13" pageNumber="200">5. Abd. IV with more than 4+4 central mac...................................................................6</keyLead>
</paragraph>
<paragraph id="B61B3F47FFFAFFC9AAD8F941FA2B5A70" blockId="13.[151,1436,1501,1914]" box="[151,1435,1775,1797]" pageId="13" pageNumber="200">
<keyLead id="0D559C72FFFAFFC9AAD8F941FA2B5A70" box="[151,1435,1775,1797]" pageId="13" pageNumber="200">- Abd. IV with 3+3 or 4+4 central mac...................................................................... 7</keyLead>
</paragraph>
</keyStep>
<keyStep id="0D5027E2FFFAFFC9AAD8F8BDFA2C5A35" pageId="13" pageNumber="200">
<paragraph id="B61B3F47FFFAFFC9AAD8F8BDFA2C5A57" blockId="13.[151,1436,1501,1914]" box="[151,1436,1804,1826]" pageId="13" pageNumber="200">
<keyLead id="0D559C72FFFAFFC9AAD8F8BDFA2C5A57" box="[151,1436,1804,1826]" pageId="13" pageNumber="200">
6. Abd. III with 1+1 central mac......................................................
<taxonomicName id="71A444C4FFFAFFC9AEC6F8BCFA2C5A57" authority="Schaffer, 1896" authorityName="Schaffer" authorityYear="1896" box="[1161,1436,1804,1826]" class="Entognatha" family="Entomobryidae" genus="Coecobrya" kingdom="Animalia" order="Collembola" pageId="13" pageNumber="200" phylum="Arthropoda" rank="species" species="hoefti">
<emphasis id="84D0E355FFFAFFC9AEC6F8BCFB2F5A57" box="[1161,1183,1805,1826]" italics="true" pageId="13" pageNumber="200">C.</emphasis>
cf.
<emphasis id="84D0E355FFFAFFC9AE8BF8BCFB4A5A57" box="[1220,1274,1805,1826]" italics="true" pageId="13" pageNumber="200">hoefti</emphasis>
(
<bibRefCitation id="D23542B6FFFAFFC9AF49F8BDFA265A57" author="Schaffer" box="[1286,1430,1804,1826]" pageId="13" pageNumber="200" refString="Schaffer, C. (1896) Die Collembola der Umgebung von Hamburg und benachbarter Gebiete. Mitteilungen aus dem Naturhistorischen Museum in Hamburg, 13, 149 - 216." type="journal article" year="1896">Schäffer, 1896</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="B61B3F47FFFAFFC9AAD8F89AFA2C5A35" blockId="13.[151,1436,1501,1914]" box="[151,1436,1834,1856]" pageId="13" pageNumber="200">
<keyLead id="0D559C72FFFAFFC9AAD8F89AFA2C5A35" box="[151,1436,1834,1856]" pageId="13" pageNumber="200">
- Abd. III with 2+2 central mac...........................................
<taxonomicName id="71A444C4FFFAFFC9AE4BF89AFA2C5A35" authority="Zhang, Deharveng &amp; Chen, 2009" authorityName="Zhang, Deharveng &amp; Chen" authorityYear="2009" box="[1028,1436,1834,1856]" class="Entognatha" family="Entomobryidae" genus="Coecobrya" kingdom="Animalia" order="Collembola" pageId="13" pageNumber="200" phylum="Arthropoda" rank="species" species="lanna">
<emphasis id="84D0E355FFFAFFC9AE4BF89AFBE75A35" box="[1028,1111,1835,1856]" italics="true" pageId="13" pageNumber="200">C. lanna</emphasis>
<bibRefCitation id="D23542B6FFFAFFC9AE13F89BFA2C5A35" author="Zhang" box="[1116,1436,1834,1856]" pageId="13" pageNumber="200" refString="Zhang, F., Deharveng, L. &amp; Chen, J. X. (2009) New species and rediagnosis of Coecobrya (Collembola: Entomobryidae), with a key to the species of the genus. Journal of Natural History, 43, 2597 - 2615. https: // doi. org / 10.1080 / 00222930903243970" type="journal article" year="2009">Zhang, Deharveng &amp; Chen, 2009</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<paragraph id="B61B3F47FFFAFFC9AAD8F8F9FA2B5A28" blockId="13.[151,1436,1501,1914]" box="[151,1435,1863,1885]" pageId="13" pageNumber="200">
<keyLead id="0D559C72FFFAFFC9AAD8F8F9FA2B5A28" box="[151,1435,1863,1885]" pageId="13" pageNumber="200">
<keyStep id="0D5027E2FFFAFFC9AAD8F8F9FA2B5A28" box="[151,1435,1863,1885]" pageId="13" pageNumber="200">
7. Abd. I with 4+4 central mac; Abd. II with 2+2 central mac...............................
<taxonomicName id="71A444C4FFFAFFC9AEC5F8F9FA225A28" authority="Deharveng, 1990" authorityName="Deharveng" authorityYear="1990" box="[1162,1426,1863,1885]" class="Entognatha" family="Entomobryidae" genus="Coecobrya" kingdom="Animalia" order="Collembola" pageId="13" pageNumber="200" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="84D0E355FFFAFFC9AEC5F8F9FB545A28" box="[1162,1252,1864,1885]" italics="true" pageId="13" pageNumber="200">C. similis</emphasis>
<bibRefCitation id="D23542B6FFFAFFC9AEA4F8F6FA225A28" author="Deharveng" box="[1259,1426,1863,1885]" pageId="13" pageNumber="200" refString="Deharveng, L. (1990) Fauna of Thai caves. II. New Entomobryiodea Collembola from Chiang Dao cave, Thailand. Occasional Papers of the Bernice P. Bishop Museum, 30, 279 - 287." type="journal article" year="1990">Deharveng, 1990</bibRefCitation>
</taxonomicName>
*
</keyStep>
</keyLead>
</paragraph>
<paragraph id="B61B3F47FFFAFFCAAAD8F8D4FB1B5ACD" blockId="13.[151,1436,1501,1914]" lastBlockId="14.[151,1436,1700,1976]" lastPageId="14" lastPageNumber="201" pageId="13" pageNumber="200">
<keyLead id="0D559C72FFFAFFC9AAD8F8D4FA2B5A0F" box="[151,1435,1892,1914]" pageId="13" pageNumber="200">
- Abd. I with 5+5 central mac; Abd. II with 3+3 central mac...........................
<taxonomicName id="71A444C4FFFAFFC9AE13F8D4FA225A0F" authority="Deharveng, 1990" authorityName="Deharveng" authorityYear="1990" box="[1116,1426,1892,1914]" class="Entognatha" family="Entomobryidae" genus="Coecobrya" kingdom="Animalia" order="Collembola" pageId="13" pageNumber="200" phylum="Arthropoda" rank="species" species="guanophila">
<emphasis id="84D0E355FFFAFFC9AE13F8D4FB545A0F" box="[1116,1252,1893,1914]" italics="true" pageId="13" pageNumber="200">C. guanophila</emphasis>
<bibRefCitation id="D23542B6FFFAFFC9AEA4F8D5FA225A0F" author="Deharveng" box="[1259,1426,1892,1914]" pageId="13" pageNumber="200" refString="Deharveng, L. (1990) Fauna of Thai caves. II. New Entomobryiodea Collembola from Chiang Dao cave, Thailand. Occasional Papers of the Bernice P. Bishop Museum, 30, 279 - 287." type="journal article" year="1990">Deharveng, 1990</bibRefCitation>
</taxonomicName>
*
</keyLead>
<subSection id="DF2B26ACFFF9FFCAAAD8F914FB1B5ACD" pageId="14" pageNumber="201" type="multiple">
We thank Louis Deharveng for providing partial molecular data and suggestions for this work. Thanks should be given to Department of National Parks, Wildlife and Plant Conservation
<collectingCountry id="CEB37FD7FFF9FFCAA99DF976FB865B95" box="[978,1078,1735,1760]" name="Thailand" pageId="14" pageNumber="201">Thailand</collectingCountry>
for permission to conduct field research, Thale Ban National Park chief Jarun Khunnarong, and Nong Plak Phraya and Khao Raya Bangsa Non Hunting Area chief Yutthana Satjakul for their permission and support, Rueangrit Promdam for his generous fieldwork assistance. This work was supported by the Science Achievement Scholarship of
<collectingCountry id="CEB37FD7FFF9FFCAAEE7F885FABC5A38" box="[1192,1292,1844,1869]" name="Thailand" pageId="14" pageNumber="201">Thailand</collectingCountry>
(SAST), the Faculty of Science Research Fund Year 2015 and Graduate school granted to A. Nilsai, Prince of Songkla University (PSU-SCI580885S) and
<collectingCountry id="CEB37FD7FFF9FFCAA87FF8CDFD245AE0" box="[560,660,1916,1941]" name="Thailand" pageId="14" pageNumber="201">Thailand</collectingCountry>
Research Fund (TRG5880189) granted to S. Jantarit as well as the Fundamental Research Funds for the Central Universities (KYZ201617) granted to F. Zhang.
</subSection>
</paragraph>
</treatment>
</document>
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