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<mods:title id="AA90806551177C8775CB08A2C81E5FF1">New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores)</mods:title>
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<mods:namePart id="E66C00A263A92C3B59D822DC824B410F">Pinto-da-Rocha, Ricardo</mods:namePart>
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<mods:namePart id="F72BC8D726C637597BFDA6C49CC6B527">Benedetti, Alipio Rezende</mods:namePart>
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<mods:namePart id="491310D1B2B774759766518DFFEEF6CE">de Vasconcelos, Eduardo Gomes</mods:namePart>
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<mods:namePart id="A77058CCF7E1580B9D0838A10E032A1F">Hara, Marcos Ryotaro</mods:namePart>
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<treatment id="3C56F9A55B29EF714C40B1B7F0B9B6FB" ID-GBIF-Taxon="152035932" LSID="urn:lsid:plazi:treatment:3C56F9A55B29EF714C40B1B7F0B9B6FB" httpUri="http://treatment.plazi.org/id/3C56F9A55B29EF714C40B1B7F0B9B6FB" lastPageNumber="37" pageId="12" pageNumber="37" scope_class="Arachnida" scope_family="Gonyleptidae" scope_order="Opiliones" scope_subFamily="Gonyleptinae" scope_subOrder="Laniatores" scope_superFamily="Gonyleptoidea">
<subSubSection id="E598875D191D29423E3DDAA04BB6EBC7" pageId="12" pageNumber="37" type="nomenclature">
<paragraph id="98835CBEF13503E81D398066168C9DF6" pageId="12" pageNumber="37">
<taxonomicName id="0430BEC6A3E32CDA091755997CA11E35" ID-CoL="7WCXD" LSID="http://species-id.net/wiki/Gonyleptes_fragilis" authority="Mello-Leitao, 1923" authorityName="Mello-Leitao" authorityYear="1923" class="Arachnida" family="Gonyleptidae" genus="Gonyleptes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonyleptes fragilis" order="Opiliones" pageId="12" pageNumber="37" phylum="Arthropoda" rank="species" species="fragilis">
Gonyleptes fragilis
<normalizedToken id="70EC44FFF3E3DE7AF846DA76ABCF5FD1" originalValue="Mello-Leitão">Mello-Leitao</normalizedToken>
, 1923
</taxonomicName>
Fig. 11
</paragraph>
</subSubSection>
<subSubSection id="F31F808DBDD08FF5850DFFED990A9FF5" pageId="12" pageNumber="37" type="reference_group">
<paragraph id="B73BF9535C488148EA9244F78220B158" pageId="12" pageNumber="37">
<taxonomicName id="7E3F44B80D3DB65D5B16217BDFCE4397" class="Arachnida" family="Gonyleptidae" genus="Gonyleptes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonyleptes fragilis" order="Opiliones" pageId="12" pageNumber="37" phylum="Arthropoda" rank="species" species="fragilis">Gonyleptes fragilis</taxonomicName>
<normalizedToken id="8A986082C6FA2FCC5E9B4E124BEC7C95" originalValue="Mello-Leitão">Mello-Leitao</normalizedToken>
, 1923: 127;
<bibRefCitation id="D0C1F981B0C101E0F08926BA5F4D592A" author="Kury, AB" journalOrPublisher="Revista Iberica de Aracnologia" pageId="23" pageNumber="48" pagination="5 - 337" title="Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones)" volume="1" year="2003">Kury 2003</bibRefCitation>
: 127; (female holotype, Brazil,
<normalizedToken id="11CA65908628E3345C01DFAA0BB72A19" originalValue="São">Sao</normalizedToken>
Paulo, Santo
<normalizedToken id="9EAB86DBC63FDBBCF34B22AAB831D5E9" originalValue="André">Andre</normalizedToken>
, Alto da Serra, MZSP 56, examined).
</paragraph>
<paragraph id="64F9E93E2CA6D0B6C0BC4F73D2CFBD34" pageId="12" pageNumber="37">
<taxonomicName id="625F2A9586E6FD1355E23B96B46874A3" class="Insecta" family="Tettigoniidae" genus="Mendesius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mendesius guttatus" order="Orthoptera" pageId="12" pageNumber="37" phylum="Arthropoda" rank="species" species="guttatus">Mendesius guttatus</taxonomicName>
Roewer, 1952: 57; (female holotype, Brazil, SMF RII 10311/72, examined).
</paragraph>
<paragraph id="1046899CA63ED4E77C7B008A619BF926" pageId="12" pageNumber="37">
<taxonomicName id="94600B52ADF6D09EABB6DB0EA4A65AF4" class="Arachnida" family="Gonyleptidae" genus="Gonyleptes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonyleptes banana" order="Opiliones" pageId="12" pageNumber="37" phylum="Arthropoda" rank="species" species="banana">Gonyleptes banana</taxonomicName>
Kury, 2003: 127 (replacement name for
<taxonomicName id="A63D711C29EDB7308D4E4FA6607EA66C" class="Insecta" family="Tettigoniidae" genus="Mendesius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mendesius guttatus" order="Orthoptera" pageId="12" pageNumber="37" phylum="Arthropoda" rank="species" species="guttatus">Mendesius guttatus</taxonomicName>
Roewer, 1952, preoccupied by
<taxonomicName id="5CBEE9677FC62C8869D0869B747E2A82" class="Arachnida" family="Gonyleptidae" genus="Gonyleptes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonyleptes guttatus" order="Opiliones" pageId="12" pageNumber="37" phylum="Arthropoda" rank="species" species="guttatus">Gonyleptes guttatus</taxonomicName>
Roewer, 1917). Syn. n.
</paragraph>
</subSubSection>
<subSubSection id="7A7A5A505A59E7427E5CF09B2CCE7D05" pageId="12" pageNumber="37" type="diagnosis">
<paragraph id="DE13F368044E0AB480D880DED47659BC" pageId="12" pageNumber="37">Diagnosis.</paragraph>
<paragraph id="FB8AF38CE6F06257D79B09E83E31B94A" pageId="12" pageNumber="37">
<taxonomicName id="837CCFC9A2347B7B498189DE58E8BCF9" class="Arachnida" family="Gonyleptidae" genus="Gonyleptes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonyleptes fragilis" order="Opiliones" pageId="12" pageNumber="37" phylum="Arthropoda" rank="species" species="fragilis">Gonyleptes fragilis</taxonomicName>
resembles
<taxonomicName id="9D57286072F2DA512B82736C24EB4C47" class="Arachnida" family="Gonyleptidae" genus="Gonyleptes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonyleptes horridus" order="Opiliones" pageId="12" pageNumber="37" phylum="Arthropoda" rank="species" species="horridus">Gonyleptes horridus</taxonomicName>
and
<taxonomicName id="598638D4DC55C74B50AF40439ED48151" class="Arachnida" family="Gonyleptidae" genus="Gonyleptes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonyleptes curvicornis" order="Opiliones" pageId="12" pageNumber="37" phylum="Arthropoda" rank="species" species="curvicornis">Gonyleptes curvicornis</taxonomicName>
because of the overall similarities of the features of the dorsal scutum.
<taxonomicName id="408DB011C97B71D041ADDA2475284114" class="Arachnida" family="Gonyleptidae" genus="Gonyleptes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonyleptes fragilis" order="Opiliones" pageId="12" pageNumber="37" phylum="Arthropoda" rank="species" species="fragilis">Gonyleptes fragilis</taxonomicName>
can be distinguished from the other two species by the ocularium with a pair of spines (tubercles in
<taxonomicName id="24191DA79C4D653E80A0A78C9DA39C77" class="Arachnida" family="Gonyleptidae" genus="Gonyleptes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonyleptes horridus" order="Opiliones" pageId="12" pageNumber="37" phylum="Arthropoda" rank="species" species="horridus">Gonyleptes horridus</taxonomicName>
and
<taxonomicName id="97076116DE8B8D4BDE8A48E21F40D734" class="Arachnida" family="Gonyleptidae" genus="Gonyleptes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonyleptes curvicornis" order="Opiliones" pageId="12" pageNumber="37" phylum="Arthropoda" rank="species" species="curvicornis">Gonyleptes curvicornis</taxonomicName>
) and the pale yellowish spots around the tubercles on abdominal scutum.
</paragraph>
</subSubSection>
<subSubSection id="B15EC82F8A0646B1271C534DA5F23190" pageId="12" pageNumber="37" type="description">
<paragraph id="053AC98C1D15392FA8029609B50BD1E8" pageId="12" pageNumber="37">Description.</paragraph>
<paragraph id="E9AF881CF03CF7E4589AB7668D616B11" pageId="12" pageNumber="37">
Penis (Fig. 11
<normalizedToken id="309CB6CFFDD61B8658ABA7CBF94057FF" originalValue="AC">A-C</normalizedToken>
): ventral plate inflated in the middle region, with a deep U-cleft on anterior margin (its distal tips convergent), 3 distal pairs of long, curved setae, 1 median pair of short setae, 4 basal pairs of spatulate setae. Glans sac slightly projected distally, with similar projection to a dorsal process. Stylus sigmoid, smooth. Ventral process robust, apex flabelliform with several small projections on distal margin.
</paragraph>
</subSubSection>
<subSubSection id="809D8CB98205A4F6A72436461C217C8F" pageId="12" pageNumber="37" type="taxonomical note">
<paragraph id="1C7C62E1AE0D159130691D408756499A" pageId="12" pageNumber="37">Taxonomical note.</paragraph>
<paragraph id="EB14057B0449333EE6CDF44B2D2AE3FE" pageId="12" pageNumber="37">
<bibRefCitation id="0A3648ADB57C97856DDEBE042ACEE7CA" author="Soares, HEM" journalOrPublisher="Revista Brasileira de Entomologia" pageId="23" pageNumber="48" pagination="227 - 232" title="Opera Opiliologica Varia. XX. Notas sinonimicas (Opiliones, Gonyleptidae)." volume="32" year="1988">Soares and Soares (1988)</bibRefCitation>
proposed the new combination
<taxonomicName id="6B59ADCDC9E09EE9AF530D87C027C4D6" class="Arachnida" family="Gonyleptidae" genus="Gonyleptes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonyleptes guttatus" order="Opiliones" pageId="12" pageNumber="37" phylum="Arthropoda" rank="species" species="guttatus">Gonyleptes guttatus</taxonomicName>
(Roewer, 1952) for
<taxonomicName id="2B3B4E629703703AEE7D81D72520B31D" class="Arachnida" family="Gonyleptidae" genus="Melloleitaniella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Melloleitaniella guttatus" order="Opiliones" pageId="12" pageNumber="37" phylum="Arthropoda" rank="species" species="guttatus">Melloleitaniella guttatus</taxonomicName>
, making it a homonym of
<taxonomicName id="4268CB39A9DEB534DF3133E2A5BBAEE7" class="Arachnida" family="Gonyleptidae" genus="Gonyleptes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonyleptes guttatus" order="Opiliones" pageId="12" pageNumber="37" phylum="Arthropoda" rank="species" species="guttatus">Gonyleptes guttatus</taxonomicName>
Roewer, 1917 [=
<taxonomicName id="B3BE6C919A1701768521E41219CD19C9" class="Arachnida" family="Gonyleptidae" genus="Melloleitaniella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Melloleitaniella guttata" order="Opiliones" pageId="12" pageNumber="37" phylum="Arthropoda" rank="species" species="guttata">Melloleitaniella guttata</taxonomicName>
(Roewer, 1917)].
<bibRefCitation id="9DE5E0557B7C24CFE0ADF4770D75DD7F" author="Kury, AB" journalOrPublisher="Revista Iberica de Aracnologia" pageId="23" pageNumber="48" pagination="5 - 337" title="Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones)" volume="1" year="2003">Kury (2003)</bibRefCitation>
renamed the species as
<taxonomicName id="D4B34374E2B3E5DCAE277C802F452D90" class="Arachnida" family="Gonyleptidae" genus="Gonyleptes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonyleptes banana" order="Opiliones" pageId="12" pageNumber="37" phylum="Arthropoda" rank="species" species="banana">Gonyleptes banana</taxonomicName>
, based on the
<normalizedToken id="0597CB9CAF758CA4B5E5CA891C4E519C" originalValue="Roewers">Roewer's</normalizedToken>
citation about the peculiar collection site of the type specimen, a banana shipment for export. The female holotype of
<taxonomicName id="E78C633EB21CE1FABC86034DD91C44CD" class="Insecta" family="Tettigoniidae" genus="Mendesius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mendesius guttatus" order="Orthoptera" pageId="12" pageNumber="37" phylum="Arthropoda" rank="species" species="guttatus">Mendesius guttatus</taxonomicName>
matches exactly the female holotype of
<taxonomicName id="B9B563E823E58F8FA098AD93C8B7BBA1" class="Arachnida" family="Gonyleptidae" genus="Gonyleptes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonyleptes fragilis" order="Opiliones" pageId="12" pageNumber="37" phylum="Arthropoda" rank="species" species="fragilis">Gonyleptes fragilis</taxonomicName>
. Both males and females of
<taxonomicName id="11963910C6579FA0C99CEC115CF55437" class="Arachnida" family="Gonyleptidae" genus="Gonyleptes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonyleptes fragilis" order="Opiliones" pageId="12" pageNumber="37" phylum="Arthropoda" rank="species" species="fragilis">Gonyleptes fragilis</taxonomicName>
present a brown general coloration with some pale yellowish spots around the tubercles of the mesotergum, a diagnostic characteristic for this species. In addition, the occurrence area of
<taxonomicName id="C3F69D665F553BC63C56F7A0ECD35971" class="Arachnida" family="Gonyleptidae" genus="Gonyleptes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonyleptes fragilis" order="Opiliones" pageId="12" pageNumber="37" phylum="Arthropoda" rank="species" species="fragilis">Gonyleptes fragilis</taxonomicName>
includes the Vale do Ribeira and southern coast of
<normalizedToken id="4C85F953A9E8ED41B4040325079C4602" originalValue="São">Sao</normalizedToken>
Paulo (
<bibRefCitation id="60267660EC08DD6D7999A7D00D12EDB5" author="Kury, AB" journalOrPublisher="Revista Iberica de Aracnologia" pageId="23" pageNumber="48" pagination="5 - 337" title="Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones)" volume="1" year="2003">Kury 2003</bibRefCitation>
), an important area known for producing banana for exportation in the 1940's and 1950's.
</paragraph>
<caption id="8223D10031CFBB6A553F711CD99CE690" pageId="12" pageNumber="37">
<paragraph id="2FAEE4B895520AEAC29C97B297981228" pageId="12" pageNumber="37">
Figure 11.
<taxonomicName id="E552C9572CD190ACDB5FE3D1452E0389" class="Arachnida" family="Gonyleptidae" genus="Gonyleptes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonyleptes fragilis" order="Opiliones" pageId="12" pageNumber="37" phylum="Arthropoda" rank="species" species="fragilis">Gonyleptes fragilis</taxonomicName>
<normalizedToken id="C7C6A38A43A598105B46E1540BF61076" originalValue="Mello-Leitão">Mello-Leitao</normalizedToken>
. Distal part of penis (MZSP 30764) A dorsal view B ditto, ventral view C ditto, left lateral view. Scale bar: 0.02 mm.
</paragraph>
</caption>
</subSubSection>
</treatment>
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