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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.275.4571" ID-GBIF-Dataset="4900d4c5-2f4b-42be-90c6-0db97d1f15a6" ID-PMC="PMC3677326" ID-Pensoft-Pub="1313-2970-275-23" ID-PubMed="23794810" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-275-23" ModsDocOrigin="ZooKeys 275" ModsDocTitle="Further studies on the Pselaphodes complex of genera from China (Coleoptera, Staphylinidae, Pselaphinae)" checkinTime="1451247589296" checkinUser="pensoft" docAuthor="Yin, Zi-Wei, Hlavac, Peter &amp; Li, Li-Zhen" docDate="2013" docId="3709BA19E0DC80E24782FFEC3C00B72E" docLanguage="en" docName="ZooKeys 275: 23-65" docOrigin="ZooKeys 275" docSource="http://dx.doi.org/10.3897/zookeys.275.4571" docTitle="Pselaphodes monoceros Yin &amp; Hlavac, sp. n." docType="treatment" docVersion="3" lastPageNumber="60" masterDocId="C11BA9533614BB4BD87BFF85D136156E" masterDocTitle="Further studies on the Pselaphodes complex of genera from China (Coleoptera, Staphylinidae, Pselaphinae)" masterLastPageNumber="65" masterPageNumber="23" pageNumber="56" updateTime="1668155449358" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Further studies on the Pselaphodes complex of genera from China (Coleoptera, Staphylinidae, Pselaphinae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Yin, Zi-Wei</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hlavac, Peter</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Li, Li-Zhen</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2013</mods:date>
<mods:detail type="volume">
<mods:number>275</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>23</mods:start>
<mods:end>65</mods:end>
</mods:extent>
</mods:part>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.275.4571</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.275.4571</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-275-23</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152041390" LSID="urn:lsid:plazi:treatment:3709BA19E0DC80E24782FFEC3C00B72E" httpUri="http://treatment.plazi.org/id/3709BA19E0DC80E24782FFEC3C00B72E" lastPageId="37" lastPageNumber="60" pageId="33" pageNumber="56">
<subSubSection pageId="33" pageNumber="56" type="nomenclature">
<paragraph pageId="33" pageNumber="56">
<taxonomicName LSID="urn:lsid:zoobank.org:act:8A403224-2E7F-4422-802C-957017D73558" authority="Yin &amp; Hlavac" class="Insecta" family="Staphylinidae" genus="Pselaphodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pselaphodes monoceros" order="Coleoptera" pageId="33" pageNumber="56" phylum="Arthropoda" rank="species" species="monoceros">
Pselaphodes monoceros Yin &amp;
<normalizedToken originalValue="Hlaváč">Hlavac</normalizedToken>
</taxonomicName>
<taxonomicNameLabel pageId="33" pageNumber="56">sp. n.</taxonomicNameLabel>
Figs 22B24
</paragraph>
</subSubSection>
<subSubSection pageId="33" pageNumber="56" type="type material">
<paragraph pageId="33" pageNumber="56">Type material</paragraph>
<paragraph pageId="33" pageNumber="56">(5 ♂♂, 1 ♀). Holotype: ♂, labeled 'China: Xizang Prov. / Cuona County / Lexiang, alt. 2500 m / 16.vii.2012, Ye Liu leg.' (SNUC); Paratypes: 4 ♂♂, 1 ♀, same label data as holotype type (SNUC).</paragraph>
</subSubSection>
<subSubSection pageId="34" pageNumber="57" type="diagnosis">
<paragraph pageId="34" pageNumber="57">
<pageBreakToken pageId="34" pageNumber="57" start="start">Diagnosis</pageBreakToken>
.
</paragraph>
<paragraph pageId="34" pageNumber="57">
Reddish brown; length 2.91-3.03; clypeus projected anteriorly, forming a horn-like process in male; postgenae elongate, rounded laterally; antennomeres
<normalizedToken originalValue="IXXI">IX-XI</normalizedToken>
enlarged; pronotum rounded at anterolateral margins; male with greatly elongate metaventral processes; metacoxae simple; aedeagus with symmetric median lobe.
</paragraph>
</subSubSection>
<subSubSection lastPageId="35" lastPageNumber="58" pageId="34" pageNumber="57" type="description">
<paragraph pageId="34" pageNumber="57">Description.</paragraph>
<paragraph lastPageId="35" lastPageNumber="58" pageId="34" pageNumber="57">
Male (Fig. 22B). Length 2.91-3.00. Head slightly longer than wide, HL 0.58-0.59, HW 0.56-0.58; clypeus projecting anteriorly (Fig. 24B); maxillary palpi (Fig. 24C) with segments III indistinctly projected laterally; eyes each composed of about 40 facets. Antennal clubs as in Fig. 24A. Pronotum (Fig. 24D) slightly longer than wide, PL 0.58-0.61, PW 0.55-0.59, rounded at anterolateral margins. Elytra wider than long, EL 0.89-0.90, EW 1.16-1.17. Metaventral processes greatly elongate, apically narrowed (Fig. 24E). Protrochantersand profemora spinose ventrally (Fig. 24F), protibiae with small apical spur (Fig. 24G); mesotrochanters spinose ventrally, mesofemora simple (Fig. 24H), mesotibiae with
<pageBreakToken pageId="35" pageNumber="58" start="start">small</pageBreakToken>
apical spine (Fig. 24I); metatrochanters and metafemora simple (Fig. 24J). Abdomen broad at base and narrowed apically, AL 0.86-0.90, AW 1.16-1.19. Sternite IX as in Fig. 24K. Aedeagus length 0.56, with symmetric median lobe (Figs 24
<normalizedToken originalValue="LN">L-N</normalizedToken>
).
</paragraph>
<paragraph pageId="35" pageNumber="58">Female. Similar to male in general; BL 3.03, HL 0.62, HW 0.57, PL 0.62, PW 0.60, EL 0.7, EW 1.19, AL 1.06, AW 1.28. Eyes each composed of about 20 facets. Antennae unmodified; metaventral processes absent.</paragraph>
</subSubSection>
<subSubSection pageId="36" pageNumber="59" type="comparative notes">
<paragraph pageId="36" pageNumber="59">
<pageBreakToken pageId="36" pageNumber="59" start="start">Comparative</pageBreakToken>
notes.
</paragraph>
<paragraph pageId="36" pageNumber="59">
This unusual
<taxonomicName class="Insecta" family="Staphylinidae" genus="Pselaphodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pselaphodes" order="Coleoptera" pageId="36" pageNumber="59" phylum="Arthropoda" rank="genus">Pselaphodes</taxonomicName>
species has simple maxillary palpomeres II and IV, with palpomeres III only slightly projecting laterally on the anterolateral margins. This form of maxillary palpi together with the modified clypeus in the male is not known in any other species of the
<taxonomicName class="Insecta" family="Staphylinidae" genus="Pselaphodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pselaphodes" order="Coleoptera" pageId="36" pageNumber="59" phylum="Arthropoda" rank="genus">Pselaphodes</taxonomicName>
complex of genera. These two characters, in combination with the form of the antennal clubs, and the greatly elongate metaventral processes readily separate the new species from all other congeners of the genus. The generic limit of
<taxonomicName class="Insecta" family="Staphylinidae" genus="Pselaphodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pselaphodes" order="Coleoptera" pageId="36" pageNumber="59" phylum="Arthropoda" rank="genus">Pselaphodes</taxonomicName>
is expanded based on this species. The form of maxillary palpi seems to be occasionally variable within genus (also see comments on
<taxonomicName class="Insecta" family="Staphylinidae" genus="Labomimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Labomimus simplicipalpus" order="Coleoptera" pageId="36" pageNumber="59" phylum="Arthropoda" rank="species" species="simplicipalpus">Labomimus simplicipalpus</taxonomicName>
above). An extensive species-level phylogenetic analysis is needed for the determination of the taxonomic placements of these atypical species.
</paragraph>
</subSubSection>
<subSubSection pageId="37" pageNumber="60" type="distribution">
<paragraph pageId="37" pageNumber="60">
<pageBreakToken pageId="37" pageNumber="60" start="start">Distribution</pageBreakToken>
.
</paragraph>
<paragraph pageId="37" pageNumber="60">Southwest China: Xizang (= Tibet).</paragraph>
</subSubSection>
<subSubSection pageId="37" pageNumber="60" type="biology">
<paragraph pageId="37" pageNumber="60">Biology.</paragraph>
<paragraph pageId="37" pageNumber="60">Adults were collected by beating a pile of mixed live and dead branches in a forest.</paragraph>
</subSubSection>
<subSubSection pageId="37" pageNumber="60" type="etymology">
<paragraph pageId="37" pageNumber="60">Etymology.</paragraph>
<paragraph pageId="37" pageNumber="60">
The Latin word
<normalizedToken originalValue="monoceros">'monoceros'</normalizedToken>
means 'a
<normalizedToken originalValue="unicorn">unicorn'</normalizedToken>
, referring to the unique protuberance on the clypeus in the male.
</paragraph>
<caption pageId="37" pageNumber="60">
<paragraph pageId="37" pageNumber="60">
Figure 24. Diagnostic features of
<taxonomicName class="Insecta" family="Staphylinidae" genus="Pselaphodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pselaphodes monoceros" order="Coleoptera" pageId="37" pageNumber="60" phylum="Arthropoda" rank="species" species="monoceros">Pselaphodes monoceros</taxonomicName>
in male. A antenna B head, in lateral view C maxillary palpus D pronotum E median metaventral process, in lateral view F protrochanter and profemur G apical portion of protibia H mesotrochanter and mesofemur I apical portion of mesotibia J metatrochanter and metafemur K sternite IX L aedeagus, in dorsal view M same, in lateral view N same, in ventral view. Scales (mm): A, B, D, F, H, J = 0.3; E, L, M, N = 0.2; C = 0.1; G, I, K = 0.05.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>