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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1046.57052" ID-GBIF-Dataset="348e46a4-8819-4e3c-b4b9-b5e88bee10d7" ID-PMC="PMC8238928" ID-Pensoft-Pub="1313-2970-1046-1" ID-Pensoft-UUID="DBC42C98E4DA5074B86525CF2FB2FA64" ID-PubMed="34239337" ID-ZooBank="66E61C4EFAFE45DE9145DB38199BDEC3" ModsDocID="1313-2970-1046-1" checkinTime="1624307367557" checkinUser="pensoft" docAuthor="Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott &amp; Meyer, Marc De" docDate="2021" docId="2D27B621B82A5446BA6D65C5ACCF7564" docLanguage="en" docName="ZooKeys 1046: 1-141" docOrigin="ZooKeys 1046" docPubDate="2021-06-21" docSource="http://dx.doi.org/10.3897/zookeys.1046.57052" docTitle="Mesembrius vockerothi Jordaens, Goergen &amp; De Meyer 2021, sp. nov." docType="treatment" docUuid="7EE04C53-0B7E-4D02-AB1D-0C71EABC9F7C" docUuidSource="ZooBank" docVersion="5" id="DBC42C98E4DA5074B86525CF2FB2FA64" lastPageNumber="1" masterDocId="DBC42C98E4DA5074B86525CF2FB2FA64" masterDocTitle="Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data" masterLastPageNumber="141" masterPageNumber="1" pageNumber="1" updateTime="1732487030840" updateUser="ExternalLinkService">
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<mods:title id="0F847E4C97F1A1396C85A9722F314A17">Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data</mods:title>
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<mods:namePart id="1F7D077C21BEE3808705B9323FAED634">Jordaens, Kurt</mods:namePart>
<mods:affiliation id="718324BA6B180638768B7E5AA687DC8C">Royal Museum for Central Africa, Invertebrates Section and JEMU, Leuvensesteenweg 13, B- 3080 Tervuren, Belgium</mods:affiliation>
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<mods:namePart id="3ABDE922E3E39E2D14058D285FE3B066">Goergen, Georg</mods:namePart>
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<mods:affiliation id="4F157E6DFB3CFE0A67479FDA4EDD9200">International Institute for Tropical Agriculture (IITA), Biodiversity Centre, 08 BP 0932 Tri Postal, Cotonou, Benin</mods:affiliation>
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<mods:namePart id="57AA76A1B6873F4FA3F05E2CCD98433B">Skevington, Jeffrey H.</mods:namePart>
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<mods:affiliation id="FC14A7FDE16B0EA0288C89F98B2CAF9B">Canadian National Collection of Insects, Arachnids and Nematodes, Agriculture and Agri-Food Canada, K. W. Neatby Building, 960 Carling Avenue, Ottawa, ON K 1 A 0 C 6, Canada</mods:affiliation>
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<mods:namePart id="10B4013CFB8495D66254EB08DEEDEC48">Kelso, Scott</mods:namePart>
<mods:affiliation id="110CAAE92998849486B1111761D6EE92">Canadian National Collection of Insects, Arachnids and Nematodes, Agriculture and Agri-Food Canada, K. W. Neatby Building, 960 Carling Avenue, Ottawa, ON K 1 A 0 C 6, Canada</mods:affiliation>
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<mods:namePart id="84C6DF82DCF8C7C6B731FDD56A27822E">Meyer, Marc De</mods:namePart>
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<mods:affiliation id="FBDCD0B5AE5C21A6F138F5F8243356A8">Royal Museum for Central Africa, Invertebrates Section and JEMU, Leuvensesteenweg 13, B- 3080 Tervuren, Belgium</mods:affiliation>
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<mods:date id="EE56DBE9DE4FF156E90F9D9643813824">2021</mods:date>
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<mods:classification id="DEBD0729BBD408224EB7644F14622BA6">journal article</mods:classification>
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<subSection id="493430CB9F15F828E3BF4CF42BF6B21D" pageId="0" pageNumber="1" type="results">
<treatment id="2D27B621B82A5446BA6D65C5ACCF7564" ID-GBIF-Taxon="183114814" LSID="urn:lsid:zoobank.org:act:7EE04C53-0B7E-4D02-AB1D-0C71EABC9F7C" httpUri="http://treatment.plazi.org/id/2D27B621B82A5446BA6D65C5ACCF7564" lastPageNumber="1" pageId="0" pageNumber="1" scope_family="Syrphidae" scope_order="Diptera">
<subSubSection id="AB951BBEBE2E374458448235F7509F0D" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="3A8EA14EFF2F9485B9042643A4537BD8" pageId="0" pageNumber="1">
<taxonomicName id="315E933D79FCFA6AEFFCDD676CC95190" ID-CoL="9HYWV" LSID="http://zoobank.org/7EE04C53-0B7E-4D02-AB1D-0C71EABC9F7C" authority="Jordaens, Goergen &amp; De Meyer" authorityName="Jordaens, Goergen &amp; De Meyer" authorityYear="2021" class="Insecta" family="Syrphidae" genus="Mesembrius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesembrius vockerothi" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="vockerothi" status="sp. nov.">Mesembrius vockerothi Jordaens, Goergen &amp; De Meyer</taxonomicName>
<taxonomicNameLabel id="CF054F4A1601A9B0F4CDF71A3CCF1D18" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
<figureCitation id="CF589A3C5C75440423074B39D58864B2" captionStart="Figures 25, 26" captionStartId="F13" captionText="Figures 25, 26. Mesembrius spp., habitus, lateral view 25 M. tibialis sp. nov. (♂) 26 M. vockerothi sp. nov. (♂)." figureDoi="10.3897/zookeys.1046.57052.figures25-26" httpUri="https://binary.pensoft.net/fig/557474" pageId="0" pageNumber="1">Figs 26</figureCitation>
<figureCitation id="62FDE502552BA17AE552FB35E6A608A1" captionStart="Figure 45" captionStartId="F23" captionText="Figure 45. Mesembrius spp., habitus, lateral view. M. vockerothi sp. nov. (♀)." figureDoi="10.3897/zookeys.1046.57052.figure45" httpUri="https://binary.pensoft.net/fig/557484" pageId="0" pageNumber="1">, 45</figureCitation>
<figureCitation id="C9C36ABB8D91F729B7C241A6CF1F4708" captionStart="Figures 6469" captionStartId="F27" captionText="Figures 64 - 69. Mesembrius spp., head, frontal view 64 M. simplicipes Curran (♂) 65 M. strigilatus (Bezzi) (♂) 66 M. sulcus sp. nov. (♂) 67 M. tarsatus (Bigot) (♂) 68 M. tibialis sp. nov. (♂) 69 M. vockerothi sp. nov. (♂)." figureDoi="10.3897/zookeys.1046.57052.figures64-69" httpUri="https://binary.pensoft.net/fig/557488" pageId="0" pageNumber="1">, 69</figureCitation>
<figureCitation id="73ADC6EF57FE4EF6460083654A07913A" captionStart="Figures 7982" captionStartId="F30" captionText="Figures 79 - 82. Mesembrius spp., head, frontal view 79 M. simplicipes Curran (♂) 80 M. sulcus sp. nov. (♀) 81 M. tarsatus (Bigot) (♀) 82 M. vockerothi sp. nov. (♀)." figureDoi="10.3897/zookeys.1046.57052.figures79-82" httpUri="https://binary.pensoft.net/fig/557491" pageId="0" pageNumber="1">, 82</figureCitation>
<figureCitation id="A8BCEEB0D998BAE1A95442792A86DCA1" captionStart="Figures 101106" captionStartId="F34" captionText="Figures 101 - 106. Mesembrius spp., abdomen, dorsal view 101 M. simplicipes Curran (♂) 102 M. strigilatus (Bezzi) (♂) 103 M. sulcus sp. nov. (♂) 104 M. tarsatus (Bigot) (♂) 105 M. tibialis sp. nov. (♂) 106 M. vockerothi sp. nov. (♂)." figureDoi="10.3897/zookeys.1046.57052.figures101-106" httpUri="https://binary.pensoft.net/fig/557495" pageId="0" pageNumber="1">, 106</figureCitation>
<figureCitation id="CE747C2FC0105DCC22F0A09006664F53" captionStart="Figures 124126" captionStartId="F38" captionText="Figures 124 - 126. Mesembrius spp., abdomen, dorsal view 124 M. sulcus sp. nov. (♀) 125 M. tarsatus (Bigot) (♀) 126 M. vockerothi sp. nov. (♀)." figureDoi="10.3897/zookeys.1046.57052.figures124-126" httpUri="https://binary.pensoft.net/fig/557499" pageId="0" pageNumber="1">, 126</figureCitation>
<figureCitation id="B8CF544A37D46BF59A6A183F1EF42327" captionStart="Figures 147150" captionStartId="F44" captionText="Figures 147 - 150. Mesembrius spp., right wing 147 M. sulcus sp. nov. (♂) 148 M. tarsatus (Bigot) (♂) 149 M. tibialis sp. nov. (♂) 150 M. vockerothi sp. nov. (♂)." figureDoi="10.3897/zookeys.1046.57052.figures147-150" httpUri="https://binary.pensoft.net/fig/557505" pageId="0" pageNumber="1">, 150</figureCitation>
<figureCitation id="AAA746F97B7FA07C828B017F5C266940" captionStart="Figures 217228" captionStartId="F59" captionText="Figures 217 - 228. Mesembrius spp., male genitalia, lateral view 217 M. nigriceps Curran 218 M. perforatus (Speiser) 219 M. platytarsis Curran syn. nov. 220 M. regulus (Hull) 221 M. rex Curran 222 M. senegalensis (Macquart) 223 M. simplicipes Curran 224 M. strigilatus (Bezzi) 225 M. sulcus sp. nov. 226 M. tarsatus (Bigot) 227 M. tibialis sp. nov. 228 M. vockerothi sp. nov." figureDoi="10.3897/zookeys.1046.57052.figures217-228" httpUri="https://binary.pensoft.net/fig/557519" pageId="0" pageNumber="1">, 228</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="67DCCCAEDD9B28AD28CBCB2BFA33CF67" pageId="0" pageNumber="1" type="differential diagnosis">
<paragraph id="01B26C087E09B0F0DE2290699E0AA7BA" pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
<paragraph id="D42EA46DF4A01286DA0ED64CE1703A7B" pageId="0" pageNumber="1">
<taxonomicName id="963EABD5C68315CDC92155D597874036" authorityName="Jordaens, Goergen &amp; De Meyer" authorityYear="2021" class="Insecta" family="Syrphidae" genus="Mesembrius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesembrius vockerothi" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="vockerothi">
<emphasis id="816C73651AD61C8932EFCB130B99D905" italics="true" pageId="0" pageNumber="1">Mesembrius vockerothi</emphasis>
</taxonomicName>
sp. nov. is the smallest of the
<taxonomicName id="FD36E612A37A4F6D56CED5EDE480CF4B" authorityName="Rondani" authorityYear="1857" class="Insecta" family="Syrphidae" genus="Mesembrius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesembrius" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis id="E50D176DD54D3C3C292FEF458D5CCAD9" italics="true" pageId="0" pageNumber="1">Mesembrius</emphasis>
</taxonomicName>
species. Males lack an apical pile brush on the profemur, have an unmodified metatibia and are dichoptic and the face is markedly conical. It can be distinguished from the male of other species by its smaller size and the conical face. The yellow pile on the mesotarsomeres is inconspicuous (very prominent on all tarsomeres in
<taxonomicName id="226D0DA47E3BAA34832C426E812996C9" lsidName="M. capensis" pageId="0" pageNumber="1" rank="species" species="capensis">
<emphasis id="168A5872877550829087B84D4F71844D" italics="true" pageId="0" pageNumber="1">M. capensis</emphasis>
</taxonomicName>
) and the scutellum is yellow pilose with short black pile interspersed on its entire surface (yellow pilose only in both morphotypes of
<taxonomicName id="AF8C3D0B3D161638860ED2D5C0AE26EE" lsidName="M. caffer" pageId="0" pageNumber="1" rank="species" species="caffer">
<emphasis id="F6AF8F0D3ED8FD63CA756D81C0B41D5D" italics="true" pageId="0" pageNumber="1">M. caffer</emphasis>
</taxonomicName>
,
<taxonomicName id="BDC22B2F1E8025E1D29F426BAFBC08B1" lsidName="M. capensis" pageId="0" pageNumber="1" rank="species" species="capensis">
<emphasis id="7479FE638C1B64CB52D6F09ED6FFA5D9" italics="true" pageId="0" pageNumber="1">M. capensis</emphasis>
</taxonomicName>
,
<taxonomicName id="1506A3459660024772B085808D79EB17" lsidName="M. minor" pageId="0" pageNumber="1" rank="species" species="minor">
<emphasis id="C7DD43DB4FA32705534A356893AA1183" italics="true" pageId="0" pageNumber="1">M. minor</emphasis>
</taxonomicName>
and
<taxonomicName id="B82CF23D72ED02A93F7D868246940A0C" lsidName="M. senegalensis" pageId="0" pageNumber="1" rank="species" species="senegalensis">
<emphasis id="E8C3469F45CC2A1F53ECFC55EDEEE4F8" italics="true" pageId="0" pageNumber="1">M. senegalensis</emphasis>
</taxonomicName>
; yellow pilose with black pile in posterior half in
<taxonomicName id="D22B37E2C0C7F417210E2EB7487406AE" lsidName="M. strigilatus" pageId="0" pageNumber="1" rank="species" species="strigilatus">
<emphasis id="5E0D277402BC7E56B870A60D79D9B6BE" italics="true" pageId="0" pageNumber="1">M. strigilatus</emphasis>
</taxonomicName>
). Females have a frons which is pale pilose on the ventral half. It can be distinguished from the female of other species by its smaller size and the conical face. The pro- and metafemur are dark brown to black (yellow-brown in
<taxonomicName id="6C5B4CF220037E1CA69453F3D56EAC93" lsidName="M. senegalensis" pageId="0" pageNumber="1" rank="species" species="senegalensis">
<emphasis id="B59625933F33BF8176464BE07043FD46" italics="true" pageId="0" pageNumber="1">M. senegalensis</emphasis>
</taxonomicName>
). Tergite II has a pair of yellow maculae (fascia in
<taxonomicName id="6DB9BE370A41246777D30B5659F32418" lsidName="M. capensis" pageId="0" pageNumber="1" rank="species" species="capensis">
<emphasis id="2537F881F65E737A83C09D4597BAA9FF" italics="true" pageId="0" pageNumber="1">M. capensis</emphasis>
</taxonomicName>
and spined morph of
<taxonomicName id="632B5CA589BC65791F7EE10242C511A6" lsidName="M. caffer" pageId="0" pageNumber="1" rank="species" species="caffer">
<emphasis id="D4C61EF0D4ABE585802C024F8E7A115B" italics="true" pageId="0" pageNumber="1">M. caffer</emphasis>
</taxonomicName>
) and the black posterior marking extends to the lateral margins (not so in
<taxonomicName id="90BEE26EE15C764BAE9D74DAD4272200" lsidName="M. minor" pageId="0" pageNumber="1" rank="species" species="minor">
<emphasis id="511BA2927795FEC014FE8327F55B9FD8" italics="true" pageId="0" pageNumber="1">M. minor</emphasis>
</taxonomicName>
). The metafemur has no ventral swelling in the middle (present in
<taxonomicName id="CBEA2D20A5C91AF8B8EBFC378210838A" lsidName="M. minor" pageId="0" pageNumber="1" rank="species" species="minor">
<emphasis id="A80ED0F5FE7E9B878FEEF48BFBC3DC0E" italics="true" pageId="0" pageNumber="1">M. minor</emphasis>
</taxonomicName>
). The pro- and mesotarsi are uniformly dark brown (brown with a darker medial part in
<taxonomicName id="9ED58CBEC77D1C692A3C28A7495CCC97" lsidName="M. minor" pageId="0" pageNumber="1" rank="species" species="minor">
<emphasis id="AD7E695F6C7BDDE3361AE943CC9C35B7" italics="true" pageId="0" pageNumber="1">M. minor</emphasis>
</taxonomicName>
). The posteroventral side of the metafemur has short black setae at distal 1/2 to 1/3 (only at distal 1/6 in the nominal morph of
<taxonomicName id="3492A7DC0DD4DAFE74DDF3A289FFCEAE" lsidName="M. caffer" pageId="0" pageNumber="1" rank="species" species="caffer">
<emphasis id="E94FBFA65B29FCEFD028E89D5C4021B7" italics="true" pageId="0" pageNumber="1">M. caffer</emphasis>
</taxonomicName>
and in
<taxonomicName id="8E6774B383227E3A98F4CF47E23AD6D7" lsidName="M. strigilatus" pageId="0" pageNumber="1" rank="species" species="strigilatus">
<emphasis id="AE6E540C7E7D1147EF6A69E81773032B" italics="true" pageId="0" pageNumber="1">M. strigilatus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="DAD1601479975E706BFF14EBF34A616E" pageId="0" pageNumber="1" type="materials_examined">
<paragraph id="820BC74351C0416884E6BC158AC4DAE6" pageId="0" pageNumber="1">Examined material.</paragraph>
<paragraph id="EB8E1B179CE429FBACA4930178F2F00E" pageId="0" pageNumber="1">
<taxonomicName id="C07CD8648495993F9BF3C7635E2AEB24" authorityName="Jordaens, Goergen &amp; De Meyer" authorityYear="2021" class="Insecta" family="Syrphidae" genus="Mesembrius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesembrius vockerothi" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="vockerothi">
<emphasis id="6B65A83969018C24D151B0143EF982C9" italics="true" pageId="0" pageNumber="1">Mesembrius vockerothi</emphasis>
</taxonomicName>
<materialsCitation id="725E6CE766B9C2DEBD01AD5E6CC03F33" collectingDate="1934-12-12" collectorName="F. W. Edwards." country="UGANDA" location="Jordaens" specimenCount="1" stateProvince="Kampala" typeStatus="Holotype">
<location id="7BEF40EF952FCE455638616D14A90639" LSID="urn:lsid:plazi:treatment:2D27B621B82A5446BA6D65C5ACCF7564:7BEF40EF952FCE455638616D14A90639" country="UGANDA" name="Jordaens" stateProvince="Kampala">Jordaens</location>
, Goergen &amp; De Meyer:
<typeStatus id="76FC2F6B1F8600A459378E08393BF063">Holotype</typeStatus>
, male,&quot; &quot;
<collectingCountry id="072D69F6B4D89B3FA50B11F5B13B292D" name="Uganda">UGANDA</collectingCountry>
: //
<collectingRegion id="1345A842D1B9FEB5C3E8A0F959E3590D" country="Uganda" name="Kampala">Kampala</collectingRegion>
, //
<collectingDate id="112320E307555CD6F798F1969D884289" value="1934-12-12">12.xii.1934</collectingDate>
, //
<collectorName id="DAD8022E6FB114D99E4304FA18BABC4A">F.W. Edwards.</collectorName>
//B.M. 1935-203.&quot;
<normalizedToken id="B168E28078C22EBCCAE832FCE79A6168" originalValue="“HOLOTYPUS”">
&quot;
<typeStatus id="548F0320254354E4026FB2A904FCF47D">HOLOTYPUS</typeStatus>
&quot;
</normalizedToken>
&quot;
<taxonomicName id="D168C0E31460F6C613AD9A43BFBF93BB" authorityName="Jordaens, Goergen &amp; De Meyer" authorityYear="2021" class="Insecta" family="Syrphidae" genus="Mesembrius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesembrius vockerothi" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="vockerothi">Mesembrius vockerothi</taxonomicName>
// Jordaens &amp; De Meyer 2019&quot; &quot; NHMUK 010369964&quot; [NHMUK].
</materialsCitation>
</paragraph>
<paragraph id="BA8FF856F2418E0685118C9D65DC1881" pageId="0" pageNumber="1">
<materialsCitation id="16B43CD3809E079F5B821EC74A7278F7" collectingDate="1918-07" collectorName="R. Mayne" country="Democratic Republic of the Congo" location="Baraka" municipality="Kalembelembe" specimenCount="1" specimenCount-female="1" typeStatus="Paratypes">
<emphasis id="36C3C0007CFD1E7D9344E6334F1252DB" bold="true" pageId="0" pageNumber="1">
<emphasis id="D900C61E2D9073463DA24B3CC59FAD2E" italics="true" pageId="0" pageNumber="1">
<typeStatus id="B457610C3054DE9CA825358AC6AD9215">Paratypes</typeStatus>
</emphasis>
:
</emphasis>
<collectingCountry id="0829846822C2F1D41361C7DFB21487A4" name="Democratic Republic of the Congo">Democratic Republic of the Congo</collectingCountry>
<specimenCount id="B96BDF2D9BCF784F26CC6227CACAB0D6" type="female">1♀</specimenCount>
;
<collectingMunicipality id="C0ADBAA9D9AEF56C39F9A92024E8F3AB">Kalembelembe</collectingMunicipality>
,
<location id="8F5D38927A110E047D51817540C681B1" LSID="urn:lsid:plazi:treatment:2D27B621B82A5446BA6D65C5ACCF7564:8F5D38927A110E047D51817540C681B1" country="Democratic Republic of the Congo" municipality="Kalembelembe" name="Baraka">Baraka</location>
;
<collectingDate id="B482D5E5DB32D7F6B6897F0EB951B2C5" value="1918-07">Jul 1918</collectingDate>
;
<collectorName id="E4CADD5B3C2C80D1F5BEC5F07532E228">
R.
<normalizedToken id="C48D3D682EA3E7DB46C7126F6544A03F" originalValue="Mayné">Mayne</normalizedToken>
</collectorName>
leg.; RMNH
</materialsCitation>
<materialsCitation id="6A0C29CB7589617AB6D490D90783B840" collectingDate="1911-07" collectorName="Murtula" country="Democratic Republic of the Congo" county="North-Kivu" location="Lesse" municipality="Beni" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="B3E6B53481D72726D77F7CB4B10F8784" type="female">1♀</specimenCount>
;
<collectingCounty id="4F3706C08BEF27EA473FBD3D83669499">North-Kivu</collectingCounty>
,
<collectingMunicipality id="00CF67F72775464D756187B04C7AFDA0">Beni</collectingMunicipality>
<normalizedToken id="4BF3F743EB617125B9C8754EC2290355" originalValue="à">a</normalizedToken>
<location id="FCDB4635329F3E14F1EE5D644B9C85BE" LSID="urn:lsid:plazi:treatment:2D27B621B82A5446BA6D65C5ACCF7564:FCDB4635329F3E14F1EE5D644B9C85BE" country="Democratic Republic of the Congo" county="North-Kivu" municipality="Beni" name="Lesse">Lesse</location>
;
<collectingDate id="E9E9CE33EF3666313280504F42CA2178" value="1911-07">Jul 1911</collectingDate>
;
<collectorName id="4CFDE42E3567FC61A6F37EC740B6A7D6">Murtula</collectorName>
leg.; KMMA
</materialsCitation>
.
<materialsCitation id="00C36072F5B2D9B48C6E7D44DF8A2336" collectingDate="1930-10" collectorName="van Someren" country="Kenya" location="Jinja" specimenCount="1" specimenCount-male="1" stateProvince="Jinja" typeStatus="paratype">
<collectingCountry id="194EC6D1F32AF00F1CEF2E2759CFA7A6" name="Kenya">Kenya</collectingCountry>
<specimenCount id="6AB5ED5B0D36A64565AAC4FF6E9FF935" type="male">1♂</specimenCount>
;
<collectingRegion id="6201CDA31214E7AD34AA757495E17FE4" country="Uganda" name="Jinja">Jinja</collectingRegion>
;
<collectingDate id="964E298AB9BCE4F6EB7331619B073CA9" value="1930-10">Oct 1930</collectingDate>
;
<collectorName id="13C2784AAE6D749B664DE4DBD2A2D4E0">van Someren</collectorName>
leg.; NHMUK
</materialsCitation>
<materialsCitation id="14A81F65D624627AA0CA99D739FB2917" collectingDate="1948-10" collectorName="van Someren" country="Kenya" location="Nyeri" specimenCount="1" specimenCount-female="1" stateProvince="Nyeri" typeStatus="paratype">
<specimenCount id="96E63FEA0E9CFAA636C38FE2B4E01663" type="female">1♀</specimenCount>
;
<collectingRegion id="553BBA35075AC6CE70955121DB746223" country="Kenya" name="Nyeri">Nyeri</collectingRegion>
;
<collectingDate id="672294EC9909BA7484B9A2546D831B08" value="1948-10">Oct 1948</collectingDate>
;
<collectorName id="C49F1ADF6CB833C35C5D508163AB6F4F">van Someren</collectorName>
leg.; NHMUK
</materialsCitation>
.
<materialsCitation id="C5D073DE3C034DF30A69FCEB97A5BEC7" collectingDate="1911-08-17" collectorName="C. C. Gowdey" country="Uganda" location="Entebbe" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<collectingCountry id="59DD44B370282EFD7DDAAC8D0E325315" name="Uganda">Uganda</collectingCountry>
<specimenCount id="50E29E52C7FCFEF41E4FBFE39FF07EA6" type="male">1♂</specimenCount>
<specimenCount id="E955AEE85042FF1E6A7A305A04488B0B" type="female">1♀</specimenCount>
;
<location id="043F9B4BBA66C6B144CCA17756030A72" LSID="urn:lsid:plazi:treatment:2D27B621B82A5446BA6D65C5ACCF7564:043F9B4BBA66C6B144CCA17756030A72" country="Uganda" name="Entebbe">Entebbe</location>
;
<collectingDate id="4F8A2C756A03A836F5E1ACB021529760" value="1911-08-17">17 Aug 1911</collectingDate>
;
<collectorName id="909665FA5526C8FFB3E473E4C12EB539">C.C. Gowdey</collectorName>
leg.; NHMUK
</materialsCitation>
<materialsCitation id="753E95C4B0E13F5A9AD5BF23B0A0C950" collectingDate="1971-11-09" collectorName="H. Falke" country="Uganda" location="Entebbe" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
<specimenCount id="9B31560EA4821ADB40395597BB62A6EB" type="male">2♂♂</specimenCount>
;
<location id="AB841EBFF32AD5A4725BCB6E482E769B" LSID="urn:lsid:plazi:treatment:2D27B621B82A5446BA6D65C5ACCF7564:AB841EBFF32AD5A4725BCB6E482E769B" country="Uganda" name="Entebbe">Entebbe</location>
;
<collectingDate id="8B299739672A5F708F1126A0CDB77656" value="1971-11-09">9 Nov 1971</collectingDate>
;
<collectorName id="DD818D15D16380B208DF1FF65C32413B">H. Falke</collectorName>
leg.; CNC
</materialsCitation>
<materialsCitation id="933680C051AE19E6015A28DA66F939E6" collectingDate="1972-01-05" collectorName="H. Falke" country="Uganda" location="Entebbe" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="52376F29E2DC0A177899BFE99EE3F6CD" type="male">1♂</specimenCount>
;
<location id="127F89D6FF034BC4096D4FFC57ACF0EA" LSID="urn:lsid:plazi:treatment:2D27B621B82A5446BA6D65C5ACCF7564:127F89D6FF034BC4096D4FFC57ACF0EA" country="Uganda" name="Entebbe">Entebbe</location>
;
<collectingDate id="AD83BD9BFD26A2BBF3E8DED989050EBD" value="1972-01-05">5 Jan 1972</collectingDate>
;
<collectorName id="A49768029307E144F4F864F64E6386AC">H. Falke</collectorName>
leg.; CNC
</materialsCitation>
<materialsCitation id="16058CE24FC3C2BC26707D626D00993A" collectingDate="1918-03-12" collectingDateMax="1918-03-20" collectingDateMin="1918-03-12" collectorName="C. C. Gowdey" country="Uganda" location="Kampala" specimenCount="1" specimenCount-female="1" stateProvince="Kampala" typeStatus="paratype">
<specimenCount id="EFE108F3A3B5915D031834D69DF4141C" type="female">1♀</specimenCount>
;
<collectingRegion id="1A10582D51A8E97A7A68E6DD547EEABD" country="Uganda" name="Kampala">Kampala</collectingRegion>
;
<collectingDate id="46C2A69D7179D014E4E5D54EFCFE935B" value="1918-03-12" valueMax="1918-03-20" valueMin="1918-03-12">12-20 Mar 1918</collectingDate>
;
<collectorName id="B10A4523AD873161DA5C1B84DE17FA7F">C.C. Gowdey</collectorName>
leg.; NHMUK
</materialsCitation>
<materialsCitation id="E433C2C99BF6464878E8C3A7E1F7EF25" collectingDate="1934-12-12" collectorName="F. W. Edwards" country="Uganda" location="Kampala" specimenCount="3" specimenCount-male="3" stateProvince="Kampala" typeStatus="paratype">
<specimenCount id="2E99920DA60ACFBFAB327EE65CB9F3C6" type="male">3♂♂</specimenCount>
;
<collectingRegion id="FB154A38C2661720A2F7F0DE300F7E3D" country="Uganda" name="Kampala">Kampala</collectingRegion>
;
<collectingDate id="0F1D90C3E38A1978A2220F2231732AD6" value="1934-12-12">12 Dec 1934</collectingDate>
;
<collectorName id="0FD3FFF83FE34CAF4DF3234C4A587DDE">F.W. Edwards</collectorName>
leg.; NHMUK
</materialsCitation>
<materialsCitation id="1B5D8847FBABD4AA2843A0E0D482CAFA" collectingDate="1934-12-13" collectorName="J. Ford" country="Uganda" location="Namanue" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="818A63DD0BFE92D9017AE9D71A2404FE" type="male">1♂</specimenCount>
<specimenCount id="1594032934E708D812EB9E192C8534B6" type="female">1♀</specimenCount>
;
<location id="57EA28249E076C434EEACFA12B7B4A64" LSID="urn:lsid:plazi:treatment:2D27B621B82A5446BA6D65C5ACCF7564:57EA28249E076C434EEACFA12B7B4A64" country="Uganda" name="Namanue">Namanue</location>
;
<collectingDate id="BCE6181484FE047A96EBF03E3AF93B8C" value="1934-12-13">13 Dec 1934</collectingDate>
;
<collectorName id="419D190627154BF4C1FCE44CE7EBDF73">J. Ford</collectorName>
leg.; NHMUK
</materialsCitation>
<materialsCitation id="43520BFAF84A142D91D377468E29AFFE" collectingDate="1911-09-26" collectingDateMax="1911-09-30" collectingDateMin="1911-09-26" collectorName="S. A. Neave" country="Uganda" location="Tero Forest" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="66113EF0DA727BBA8B50B3E24199C999" type="female">1♀</specimenCount>
;
<location id="46DD53F86479593B2A4D0C50FF80D5E8" LSID="urn:lsid:plazi:treatment:2D27B621B82A5446BA6D65C5ACCF7564:46DD53F86479593B2A4D0C50FF80D5E8" country="Uganda" name="Tero Forest">Tero Forest</location>
;
<collectingDate id="428BFA5E38E4370D063F21A8621F2BFE" value="1911-09-26" valueMax="1911-09-30" valueMin="1911-09-26">26-30 Sep 1911</collectingDate>
;
<collectorName id="FFD3CC087249C5AF5213DFA2D862886B">S.A. Neave</collectorName>
leg.; NHMUK
</materialsCitation>
<materialsCitation id="2B18875F11A27BD9788B801FC7AB50DA" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="C. H. Marshall" country="Uganda" location="Unyoro District" specimenCount="1" specimenCount-female="1" stateProvince="Unyoro District" typeStatus="paratype">
<specimenCount id="AE800F6A76B93C21D9DCE340B2FFFB8B" type="female">1♀</specimenCount>
;
<collectingRegion id="F57295EBE08CCDA9D327ED27DC3827E4" inferredBySuffix="true">Unyoro District</collectingRegion>
;
<collectorName id="41AB12AFEEEE4695C6EEB78ED9C1123A">C.H. Marshall</collectorName>
leg.; NMSA
</materialsCitation>
<materialsCitation id="63EB083F5C0EC3A6F9A1857C6A18D412" collectingDate="2018-12-16" collectorName="G. Stahls" country="Uganda" location="Mabamba Swamp" municipality="Central Region" specimenCount="1" specimenCount-female="1" stateProvince="Wakiso District" typeStatus="paratype">
<specimenCount id="E04CB84921458576F1642BF8AC4DC0FD" type="female">1♀</specimenCount>
;
<collectingMunicipality id="E6EBDB4AC2FB6CB25A9C2718807E0A88">Central Region</collectingMunicipality>
,
<collectingRegion id="1A0ED2A8E0B04A1B6BE6E42F68658A27" country="Uganda" name="Wakiso">Wakiso District</collectingRegion>
,
<location id="62B568CB875E2372F0E7989DD97DAFC7" LSID="urn:lsid:plazi:treatment:2D27B621B82A5446BA6D65C5ACCF7564:62B568CB875E2372F0E7989DD97DAFC7" country="Uganda" municipality="Central Region" name="Mabamba Swamp" stateProvince="Wakiso District">Mabamba Swamp</location>
;
<collectingDate id="8F1B5F97E4A4A89EEC84A43662CD692C" value="2018-12-16">16 Dec 2018</collectingDate>
;
<collectorName id="6441B320A631D7FF61CAFCD5A97EE6F3">
G.
<normalizedToken id="D148DC9315E2C1C911CA2BA5CF52E1E8" originalValue="Ståhls">Stahls</normalizedToken>
</collectorName>
leg.; MZH
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="9B271BCADBEB3A7CE16E1A5C56E2EF9E" pageId="0" pageNumber="1" type="materials_examined">
<paragraph id="A0296D29A006A76007D9588FBE8C024C" pageId="0" pageNumber="1">Other material</paragraph>
<paragraph id="9BFAB6F77CABB94827451AFCC1D62E16" pageId="0" pageNumber="1">
<specimenCount id="DC3E79C6BBED60BF2AAF1C12F571695D" type="female">1♀</specimenCount>
with locality and date unknown, D. Bruce leg. (NHMUK).
</paragraph>
</subSubSection>
<subSubSection id="63304312671B0283DF51DD23C3E95898" pageId="0" pageNumber="1" type="description">
<paragraph id="A33825F9295B2D927EE880C05C72B66D" pageId="0" pageNumber="1">Description male</paragraph>
<paragraph id="226A183BC53D2273A5A62A45A5960611" pageId="0" pageNumber="1">
<emphasis id="CD881044EE7478CEDC3509BEAAAEA4D3" bold="true" pageId="0" pageNumber="1">
(Fig.
<figureCitation id="46597216EA4AD744118D657B5D85B6B5" captionStart="Figures 25, 26" captionStartId="F13" captionText="Figures 25, 26. Mesembrius spp., habitus, lateral view 25 M. tibialis sp. nov. (♂) 26 M. vockerothi sp. nov. (♂)." figureDoi="10.3897/zookeys.1046.57052.figures25-26" httpUri="https://binary.pensoft.net/fig/557474" pageId="0" pageNumber="1">26</figureCitation>
).
</emphasis>
Body length: 11.0-13.2 mm. Wing length: 9.6-10.5 mm.
</paragraph>
<paragraph id="9D7003D6A3C4FB410530E8DEBC0B293A" pageId="0" pageNumber="1">
<emphasis id="5F5717340B4C67FBC8CA4F1A5CA9906F" bold="true" italics="true" pageId="0" pageNumber="1">Head</emphasis>
(Fig.
<figureCitation id="5ADB0A2E227A9433260A10EA40726F34" captionStart="Figures 6469" captionStartId="F27" captionText="Figures 64 - 69. Mesembrius spp., head, frontal view 64 M. simplicipes Curran (♂) 65 M. strigilatus (Bezzi) (♂) 66 M. sulcus sp. nov. (♂) 67 M. tarsatus (Bigot) (♂) 68 M. tibialis sp. nov. (♂) 69 M. vockerothi sp. nov. (♂)." figureDoi="10.3897/zookeys.1046.57052.figures64-69" httpUri="https://binary.pensoft.net/fig/557488" pageId="0" pageNumber="1">69</figureCitation>
). Eyes dichoptic; distance between eyes approx. the width of anterior ocellus. Face conical; white with dark medial vitta; white pilose. Vertical triangle black; black pilose; yellow pollinose on ventral half. Distance between lateral ocellus and eye margin 1/2 width of ocellus. Occiput yellow; yellow pilose; yellow and white pollinose. Frontal triangle short; yellow-white; with long, black pile medially, yellow pilose on gena; yellow pollinose. Frontal prominence shiny dark brown to black. Antenna dark brown, antennal arista reddish-brown.
</paragraph>
<paragraph id="2DA4FC131028F78A582489B91567C1A9" pageId="0" pageNumber="1">
<emphasis id="DF6C5067F07A77AB6BA341DD19CDAE64" bold="true" italics="true" pageId="0" pageNumber="1">Thorax.</emphasis>
Scutum black with, dorsally, a pair of well-demarcated white pollinose vittae which are connected posteriorly; lateral white pollinose vitta clear; yellow pilose. Scutellum yellow-brown; yellow pilose with shorter black pile interspersed on its entire surface.
</paragraph>
<paragraph id="C02E219617F0B4CB64368BE4AEA33447" pageId="0" pageNumber="1">
<emphasis id="BBE861DDBB0CCEE0A88BCCCB441BCB2E" bold="true" italics="true" pageId="0" pageNumber="1">Legs.</emphasis>
All femora dark brown to black, except for extreme distal ends which are orange-brown; femora yellow to orange. Pro- and mesoleg: Femur with black pile on anterior and dorsal side and with longer yellow pile on posterior and posterodorsal sides. Tarsi yellow to orange. Metaleg: Femur with long and thin yellow pile; with black pile ventrally on distal half. Tibia with yellow and black pile, of which the yellow pile is longer on posterodorsal side. Metatibia unmodified. Metatarsi dark brown.
</paragraph>
<paragraph id="96DAF02FB6269D914C5D401A3CA5FD9D" pageId="0" pageNumber="1">
<emphasis id="E4E262A1CCACBEBB4F5D6B2795F160B1" bold="true" italics="true" pageId="0" pageNumber="1">Wing</emphasis>
(Fig.
<figureCitation id="9D58A950249C75B4FCE3A87BF1F0752E" captionStart="Figures 147150" captionStartId="F44" captionText="Figures 147 - 150. Mesembrius spp., right wing 147 M. sulcus sp. nov. (♂) 148 M. tarsatus (Bigot) (♂) 149 M. tibialis sp. nov. (♂) 150 M. vockerothi sp. nov. (♂)." figureDoi="10.3897/zookeys.1046.57052.figures147-150" httpUri="https://binary.pensoft.net/fig/557505" pageId="0" pageNumber="1">150</figureCitation>
). Entire wing uniformly very dense microtrichose.
</paragraph>
<paragraph id="5AE7F2B69E268468A3D54E484B30624B" pageId="0" pageNumber="1">
<emphasis id="B7A8B58E6A3E7059A5C95937891262CF" bold="true" italics="true" pageId="0" pageNumber="1">Abdomen</emphasis>
(Fig.
<figureCitation id="4D0E473EB45F418FF3E7B3F9F6D27EC0" captionStart="Figures 101106" captionStartId="F34" captionText="Figures 101 - 106. Mesembrius spp., abdomen, dorsal view 101 M. simplicipes Curran (♂) 102 M. strigilatus (Bezzi) (♂) 103 M. sulcus sp. nov. (♂) 104 M. tarsatus (Bigot) (♂) 105 M. tibialis sp. nov. (♂) 106 M. vockerothi sp. nov. (♂)." figureDoi="10.3897/zookeys.1046.57052.figures101-106" httpUri="https://binary.pensoft.net/fig/557495" pageId="0" pageNumber="1">106</figureCitation>
). Tergite II with a pair of very large yellow-orange, rounded maculae; black marking hourglass-shaped; posterior black marking equal in size to anterior black marking and with a medial white pollinose area; yellow pilose, but black pilose on posterior half of black marking. Tergite III with a pair of large yellow-orange maculae; with large black marking on posterior 2/3; yellow pilose on maculae, black pilose on black marking. Tergite IV black, with a pair of small yellow maculae in anterolateral corners; white pilose and strongly white pollinose on anterior and lateral parts; predominantly black pilose on black marking.
</paragraph>
<paragraph id="D0084B8106BBD7D54FC3CC46195E8414" pageId="0" pageNumber="1">
<emphasis id="814BDF67C0906C0598E588D3198474A1" bold="true" italics="true" pageId="0" pageNumber="1">Genitalia</emphasis>
(Fig.
<figureCitation id="F7AAF98A8285896C0C1D419FA973ED76" captionStart="Figures 217228" captionStartId="F59" captionText="Figures 217 - 228. Mesembrius spp., male genitalia, lateral view 217 M. nigriceps Curran 218 M. perforatus (Speiser) 219 M. platytarsis Curran syn. nov. 220 M. regulus (Hull) 221 M. rex Curran 222 M. senegalensis (Macquart) 223 M. simplicipes Curran 224 M. strigilatus (Bezzi) 225 M. sulcus sp. nov. 226 M. tarsatus (Bigot) 227 M. tibialis sp. nov. 228 M. vockerothi sp. nov." figureDoi="10.3897/zookeys.1046.57052.figures217-228" httpUri="https://binary.pensoft.net/fig/557519" pageId="0" pageNumber="1">228</figureCitation>
). Epandrium: Dorsal lobe of surstylus distally broadly rounded, with characteristic large tooth-like projection; entirely pilose, except on tooth and at basis (stalk). Ventral lobe of surstylus with one large black setula in middle section and a row of 4-5 long black setulae.
</paragraph>
</subSubSection>
<subSubSection id="008DEC42533FB5E2473CE01402D486BC" pageId="0" pageNumber="1" type="description">
<paragraph id="E7B995D895C1E098BE5DEC73128320D6" pageId="0" pageNumber="1">Description female</paragraph>
<paragraph id="3274962DD15FFC67B5EEDF99B1A384D9" pageId="0" pageNumber="1">
<emphasis id="029AF132AA830F98048FFD1E39584548" bold="true" pageId="0" pageNumber="1">
(Fig.
<figureCitation id="3E0EB3F7873E1E68D8D7DB4FF1D80206" captionStart="Figure 45" captionStartId="F23" captionText="Figure 45. Mesembrius spp., habitus, lateral view. M. vockerothi sp. nov. (♀)." figureDoi="10.3897/zookeys.1046.57052.figure45" httpUri="https://binary.pensoft.net/fig/557484" pageId="0" pageNumber="1">45</figureCitation>
).
</emphasis>
Body length: 14.0-15.1 mm. Wing length: 9.7-10.3 mm.
</paragraph>
<paragraph id="9A02880846F2AEC83A43B2C114FB14D5" pageId="0" pageNumber="1">
As male, except for the following character states: Eyes dichoptic (Fig.
<figureCitation id="D31E16F312DA0BB5DC4B5AEAB5308952" captionStart="Figures 7982" captionStartId="F30" captionText="Figures 79 - 82. Mesembrius spp., head, frontal view 79 M. simplicipes Curran (♂) 80 M. sulcus sp. nov. (♀) 81 M. tarsatus (Bigot) (♀) 82 M. vockerothi sp. nov. (♀)." figureDoi="10.3897/zookeys.1046.57052.figures79-82" httpUri="https://binary.pensoft.net/fig/557491" pageId="0" pageNumber="1">82</figureCitation>
). Frons white pilose, brown pilose on ocellar triangle and surrounding area; strongly white pollinose to just before ocellar triangle. Pile on legs shorter. Abdomen as in Fig.
<figureCitation id="B9D1C0FF759911D1DD1F4E77460FDACE" captionStart="Figures 124126" captionStartId="F38" captionText="Figures 124 - 126. Mesembrius spp., abdomen, dorsal view 124 M. sulcus sp. nov. (♀) 125 M. tarsatus (Bigot) (♀) 126 M. vockerothi sp. nov. (♀)." figureDoi="10.3897/zookeys.1046.57052.figures124-126" httpUri="https://binary.pensoft.net/fig/557499" pageId="0" pageNumber="1">126</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A032B386D8101A371392EF6737F8317B" pageId="0" pageNumber="1" type="distribution">
<paragraph id="2559DC773A188B9BA23088141479BFC6" pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph id="501491EC7DE1B53C9F8A33B1CD5C6FE2" pageId="0" pageNumber="1">Democratic Republic of the Congo, Kenya and Uganda.</paragraph>
</subSubSection>
<subSubSection id="694405DDE526DA39C625EDF47CA048C4" pageId="0" pageNumber="1" type="comments">
<paragraph id="0C69C7596F66BF4148B286DDA7C3D134" pageId="0" pageNumber="1">Comments.</paragraph>
<paragraph id="731AA3D4A0CF9FC719661CE6351D3E9F" pageId="0" pageNumber="1">
This is a new species and the smallest in size of all Afrotropical
<taxonomicName id="5F734435CDFA0D398914FD20D5FF7299" authorityName="Rondani" authorityYear="1857" class="Insecta" family="Syrphidae" genus="Mesembrius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesembrius" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis id="1667346CD99B2B212D990CFCA6286CC7" italics="true" pageId="0" pageNumber="1">Mesembrius</emphasis>
</taxonomicName>
hitherto known. It is the only Afrotropical
<taxonomicName id="857B350FF941C9470061B62164FD1C63" authorityName="Rondani" authorityYear="1857" class="Insecta" family="Syrphidae" genus="Mesembrius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesembrius" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis id="6FC7C0D90C2E242325EF71528A8B1529" italics="true" pageId="0" pageNumber="1">Mesembrius</emphasis>
</taxonomicName>
species with a conical face.
</paragraph>
</subSubSection>
<subSubSection id="8B718393C493C27AECEFA19034BBBC75" pageId="0" pageNumber="1" type="etymology">
<paragraph id="5B0DC52551967B0EBEC4658F37E23410" pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph id="7D46B1D5A93631FECE2A907214E3807F" pageId="0" pageNumber="1">Named in honour of the Dipterist Dick Vockeroth (1928-2012), who already indicated on the labels that some specimens from Uganda probably belonged to a new species. The specific epithet should be treated as a noun in the genitive case.</paragraph>
</subSubSection>
</treatment>
</subSection>
</document>