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<document id="431666CF916043E5A3C2B546D94B4EE2" ID-CLB-Dataset="36382" ID-DOI="10.11646/zootaxa.3828.1.1" ID-GBIF-Dataset="8c1cda9b-0d82-41f3-a1c1-d8eb574a1ba7" ID-ISSN="1175-5326" ID-Zenodo-Dep="286513" ID-ZooBank="B2FEAE60-7E51-45FA-A38F-930A084A5AAA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460452080076" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2014" docId="03F68791FFCC18158FFE3A8B0760FB85" docLanguage="en" docName="zt03828p076.pdf" docOrigin="Zootaxa 3828 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Scydmaenozila macrosticta Lea, comb. n." docType="treatment" docVersion="7" lastPageNumber="30" masterDocId="FFCFFFE9FFD718088F693A1C034CFC3C" masterDocTitle="' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae)" masterLastPageNumber="76" masterPageNumber="1" pageNumber="28" updateTime="1698338417320" updateUser="ExternalLinkService">
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<mods:title id="FD3ABF798202DC892C0E795537B3DB5A">' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae)</mods:title>
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<mods:namePart id="A9FEE80D922CA04010124CC9A3D9556B">Jałoszyński, Paweł</mods:namePart>
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<treatment id="03F68791FFCC18158FFE3A8B0760FB85" ID-DOI="http://doi.org/10.5281/zenodo.6141064" ID-GBIF-Taxon="127711776" ID-Zenodo-Dep="6141064" LSID="urn:lsid:plazi:treatment:03F68791FFCC18158FFE3A8B0760FB85" httpUri="http://treatment.plazi.org/id/03F68791FFCC18158FFE3A8B0760FB85" lastPageId="29" lastPageNumber="30" pageId="27" pageNumber="28">
<subSubSection id="C345650CFFCC18138FFE3A8B0122FCEF" pageId="27" pageNumber="28" type="nomenclature">
<paragraph id="8BE03687FFCC18138FFE3A8B01D7FC8E" blockId="27.[151,667,151,211]" box="[151,667,151,178]" pageId="27" pageNumber="28">
<heading id="D0A881EBFFCC18138FFE3A8B01D7FC8E" bold="true" box="[151,667,151,178]" fontSize="11" level="1" pageId="27" pageNumber="28" reason="1">
<emphasis id="B92BEA95FFCC18138FFE3A8B01D7FC8E" bold="true" box="[151,667,151,178]" pageId="27" pageNumber="28">
<taxonomicName id="4C5F4D04FFCC18138FFE3A8B0164FC8E" ID-CoL="4W3MN" authority="Lea" authorityName="Lea" box="[151,552,151,178]" class="Insecta" family="Staphylinidae" genus="Scydmaenozila" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="macrosticta" status="comb. nov.">
<emphasis id="B92BEA95FFCC18138FFE3A8B0293FC8D" bold="true" box="[151,479,151,177]" italics="true" pageId="27" pageNumber="28">Scydmaenozila macrosticta</emphasis>
(Lea)
</taxonomicName>
<taxonomicNameLabel id="A21857EEFFCC18138D593A8401D7FC8E" box="[560,667,152,178]" pageId="27" pageNumber="28">comb. n.</taxonomicNameLabel>
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<paragraph id="8BE03687FFCC18138FFE3AA60122FCEF" blockId="27.[151,667,151,211]" box="[151,622,186,211]" pageId="27" pageNumber="28">
(
<figureCitation id="13642A02FFCC18138FF63AA60237FCEF" box="[159,379,186,211]" captionStart-0="FIGURES 57 61" captionStart-1="FIGURES 62 66" captionStart-2="FIGURES 67 71" captionStart-3="FIGURES 72 79" captionStartId-0="28.[151,264,1783,1806]" captionStartId-1="30.[151,264,1734,1757]" captionStartId-2="31.[151,264,1849,1872]" captionStartId-3="33.[151,264,1586,1609]" captionTargetBox-0="[151,1411,193,1762]" captionTargetBox-1="[151,1436,193,1712]" captionTargetBox-2="[162,1426,193,1831]" captionTargetBox-3="[155,1432,200,1548]" captionTargetId-0="figure@28.[151,1436,193,1763]" captionTargetId-1="figure@30.[151,1436,193,1713]" captionTargetId-2="figure@31.[162,1427,193,1833]" captionTargetId-3="figure@33.[151,1436,193,1565]" captionTargetPageId-0="28" captionTargetPageId-1="30" captionTargetPageId-2="31" captionTargetPageId-3="33" captionText-0="FIGURES 57 61. Morphology of Scydmaenozila macrosticta (Lea) comb. n. Head in ventral view (57), distal part of labium in ventral view (58), labrum in dorsal view (59), right mandible in dorsal view (60), and right maxilla in ventral view (61). Abbreviations: bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hr, hypostomal ridge; lac, lacinia; lg, ligula; lp, labial palpus; lp 1 3, labial palpomere 1 3; md, mandible; mn, mentum; mst, mediostipes; mxp, maxillary palpus; mxp 1 4, maxillary palpomere 1 4; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; sat, subapical tooth; smn, submentum." captionText-1="FIGURES 62 66. Morphology of Scydmaenozila macrosticta (Lea) comb. n. Prothorax in ventral and slightly lateral view (62 63), base of pronotum in dorsal view (64), mesonotum and elytral base in dorsal view (65), and mesocoxa in dorsal view (66). Abbreviations: bef, basal elytral foveae; bst, basisternal part of prosternum; cx 2, mesocoxa; eal, elytral articulating lobe; fm 2, mesofemur; hy, hypomeron; ip, internal antebasal pit; lp, lateral antebasal pit; nss, pronosternal suture; pcc. procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; sc 2, mesoscutum; scl 2, mesoscutellum; sss, mesoscutoscutellar suture; tr 2, mesotrochanter." captionText-2="FIGURES 67 71. Morphology of Scydmaenozila macrosticta (Lea) comb. n. Mesothorax in ventral view (67), pterothorax in ventral view (68 69), aedeagus in ventral and slightly lateral view (70), and metaventral intercoxal process in ventral view (71). Abbreviations: aest 3, metanepisternum; aml, apex of median lobe; ar, anterior ridge; cx 3, metacoxa; dlf, dorsoventral fovea; fm 3, metafemur; lmfa, lateral mesofurcal arm; ma, membranous area; mcp, mesocoxal process; mscc, mesocoxal cavity; mscs, mesocoxal socket; msf, mesofurca; msvp, mesoventral process; mtvp, metaventral intercoxal process; pcr, procoxal rest; pm, paramere; pre, prepectus; tr 3, metatrochanter; vlf, ventrolateral fovea." captionText-3="FIGURES 72 79. Scydmaenozila spp. Dorsal habitus (72 75), original set of labels of lectotypes (76, 79), paralectotype (77), and mounting card with three syntypes (with reverse side in bottom) before re-mounting during the present study (78). Scydmaenozila macrosticta (Lea) comb. n. (72, 76 77); Scydmaenozila latipennis (Lea) comb. n. (73, 78 79), Scydmaenozila mallacootana sp. n. (74), and Scydmaenozila inexpectata sp. n. (75)." httpUri-0="https://zenodo.org/record/286527/files/figure.png" httpUri-1="https://zenodo.org/record/286528/files/figure.png" httpUri-2="https://zenodo.org/record/286529/files/figure.png" httpUri-3="https://zenodo.org/record/286530/files/figure.png" pageId="27" pageNumber="28">Figs. 5772, 7677</figureCitation>
,
<figureCitation id="13642A02FFCC18138EEF3AA70282FCEF" box="[390,462,187,211]" captionStart="FIGURES 80 88" captionStartId="34.[151,264,1601,1624]" captionTargetBox="[163,1426,499,1576]" captionTargetId="figure@34.[151,1436,481,1580]" captionTargetPageId="34" captionText="FIGURES 80 88. Aedeagi of Scydmaenozila spp. in ventral (80, 82, 83, 85, 87) and lateral (81, 84, 86, 88) views. Scydmaenozila macrosticta (Lea) comb. n., intact (80 81) and erected (82); Scydmaenozila latipennis (Lea) comb. n. (83 84), Scydmaenozila mallacootana sp. n. (85 86), and Scydmaenozila inexpectata sp. n. (87 88)." httpUri="https://zenodo.org/record/286531/files/figure.png" pageId="27" pageNumber="28">8082</figureCitation>
,
<figureCitation id="13642A02FFCC18138EB33AA7016FFCEF" box="[474,547,187,211]" captionStart="FIGURES 89 96" captionStartId="35.[151,264,1860,1883]" captionTargetBox="[166,1428,430,1831]" captionTargetId="figure@35.[151,1436,411,1839]" captionTargetPageId="35" captionText="FIGURES 89 96. Aedeagi of Scydmaenozila spp. in ventral (89, 91, 93, 95) and lateral (90, 92, 94, 96) views. Scydmaenozila macrosticta (Lea) comb. n. (89 90); Scydmaenozila latipennis (Lea) comb. n. (91 92), Scydmaenozila mallacootana sp. n. (93 94), and Scydmaenozila inexpectata sp. n. (95 96)." httpUri="https://zenodo.org/record/286532/files/figure.png" pageId="27" pageNumber="28">8990</figureCitation>
,
<figureCitation id="13642A02FFCC18138D473AA60115FCEF" box="[558,601,186,211]" captionStart="FIGURES 193 198" captionStartId="72.[151,264,1762,1785]" captionTargetBox="[195,1376,193,1707]" captionTargetId="figure@72.[191,1378,193,1739]" captionTargetPageId="72" captionText="FIGURES 193 198. Distribution of revised Australian species of Horaeomorphus Schaufuss (193), Leascydmus gen. n. (194), Scydmaenozila gen. n. (195), Spinosciacharis gen. n. (196), Sciacharoides gen. n. (197), and Sciacharis Broun. Horaeomorphus anthicoides (Lea) comb. n. (193 a), Horaeomorphus verus sp. n. (193 b), Leascydmus simplicicornis (Lea) comb. n. (194 a), Leascydmus armstrongi sp. n. (194 b), Scydmaenozila macrosticta (Lea) comb. n. (195 a), Scydmaenozila latipennis (Lea) comb. n. (195 b), Scydmaenozila mallacootana sp. n. (195 c), Scydmaenozila inexpectata sp. n. (195 d), Spinosciacharis puncticeps (Franz) comb. n. (196 a), Spinosciacharis sassy sp. n. (196 b), Spinosciacharis clydensis sp. n. (196 c), Spinosciacharis acheron sp. n. (196 d), Sciacharoides thompsoni (Franz) comb. n. (197 a), Sciacharoides eucalypti (Franz) comb. n. (197 b), Sciacharoides maipotonensis (Franz) comb. n. (197 c), Sciacharoides mantistamborinensis (Franz) comb. n. (197 d), and Sciacharis (Maorinus) tasmaniensis (Franz) comb. n. (198)." httpUri="https://zenodo.org/record/286555/files/figure.png" pageId="27" pageNumber="28">195</figureCitation>
a)
</paragraph>
</subSubSection>
<subSubSection id="C345650CFFCC18138FFE3B1E0066FDAE" pageId="27" pageNumber="28" type="reference_group">
<paragraph id="8BE03687FFCC18138FFE3B1E0118FD24" blockId="27.[151,1436,258,403]" box="[151,596,258,280]" pageId="27" pageNumber="28">
<treatmentCitationGroup id="AB4F11A9FFCC18138FFE3B1E0118FD24" box="[151,596,258,280]" pageId="27" pageNumber="28">
<taxonomicName id="4C5F4D04FFCC18138FFE3B1E011CFD24" ID-CoL="777DK" authority="Lea, 1910: 189" authorityName="Lea" authorityPageNumber="189" authorityYear="1910" box="[151,592,258,280]" class="Insecta" family="Staphylinidae" genus="Phagonophana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="macrosticta">
<emphasis id="B92BEA95FFCC18138FFE3B1E02E3FD24" box="[151,431,258,280]" italics="true" pageId="27" pageNumber="28">Phagonophana macrosticta</emphasis>
<treatmentCitation id="0AFE1096FFCC18138EDD3B1F011CFD24" author="Lea" box="[436,592,258,280]" page="189" pageId="27" pageNumber="28" year="1910">
<bibRefCitation id="EFCE4B76FFCC18138EDD3B1F0151FD24" author="Lea" box="[436,541,258,280]" pageId="27" pageNumber="28" refString="Lea, A. M. (1910) Australian and Tasmanian Coleoptera inhabiting or resorting to the nests of ants, bees, and termites. Proceedings of the Royal Society of Victoria, Melbourne (N. S.), 23, 116 - 230." type="journal article" year="1910">Lea, 1910</bibRefCitation>
: 189
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BE03687FFCC18138FFE3B3D0241FD6A" blockId="27.[151,1436,258,403]" pageId="27" pageNumber="28">
<taxonomicName id="4C5F4D04FFCC18138FFE3B3D02F7FD0B" authority="Lea" authorityName="Lea" box="[151,443,289,311]" class="Insecta" family="Staphylinidae" genus="Syndicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="macrostictus">
<emphasis id="B92BEA95FFCC18138FFE3B3D0235FD0B" box="[151,377,289,311]" italics="true" pageId="27" pageNumber="28">Syndicus macrostictus</emphasis>
(Lea)
</taxonomicName>
; implied new combination ex
<taxonomicName id="4C5F4D04FFCC18138D9F3B3D00DDFD0B" ID-CoL="9LG39" box="[758,913,289,311]" class="Insecta" family="Staphylinidae" genus="Phagonophana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEA95FFCC18138D9F3B3D00DDFD0B" box="[758,913,289,311]" italics="true" pageId="27" pageNumber="28">Phagonophana</emphasis>
</taxonomicName>
, which was synonymized with
<taxonomicName id="4C5F4D04FFCC18138BB13B3D067EFD0B" ID-CoL="8XQ4R" box="[1240,1330,289,311]" class="Insecta" family="Staphylinidae" genus="Syndicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEA95FFCC18138BB13B3D067EFD0B" box="[1240,1330,289,311]" italics="true" pageId="27" pageNumber="28">Syndicus</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFCE4B76FFCC18138A353B3E0245FD6A" author="Franz" pageId="27" pageNumber="28" refString="Franz, H. (1971 b) Revision der Gattung Syndicus Motsch. (Coleopt., Scydmaenidae). Koleopterologische Rundschau, 49, 11 - 28." type="journal article" year="1971" yearSuffix="b">Franz, 1971b</bibRefCitation>
.
</paragraph>
<paragraph id="8BE03687FFCC18138FFE3B42006EFD48" blockId="27.[151,1436,258,403]" box="[151,802,350,373]" pageId="27" pageNumber="28">
<treatmentCitationGroup id="AB4F11A9FFCC18138FFE3B42006EFD48" box="[151,802,350,373]" pageId="27" pageNumber="28">
<taxonomicName id="4C5F4D04FFCC18138FFE3B42014FFD48" authority="Lea" authorityName="Lea" box="[151,515,350,373]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="macrostictus">
<emphasis id="B92BEA95FFCC18138FFE3B420288FD49" box="[151,452,350,373]" italics="true" pageId="27" pageNumber="28">Horaeomorphus macrostictus</emphasis>
(Lea)
</taxonomicName>
;
<treatmentCitation id="0AFE1096FFCC18138D673B4301F0FD48" author="Franz" box="[526,700,350,372]" page="148" pageId="27" pageNumber="28" year="1975">
<bibRefCitation id="EFCE4B76FFCC18138D673B4301C6FD48" author="Franz" box="[526,650,350,372]" pageId="27" pageNumber="28" refString="Franz, H. (1975) Revision der Scydmaeniden von Australien, Neuseeland und den benachbarten Inseln. Denkschriften Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 118, 1 - 312." type="journal article" year="1975">Franz, 1975</bibRefCitation>
: 148
</treatmentCitation>
,
<figureCitation id="13642A02FFCC18138DAE3B420052FD48" box="[711,798,350,372]" captionStart="FIGURES 121 128" captionStartId="46.[151,264,1877,1900]" captionTargetBox="[168,1425,220,1825]" captionTargetId="figure@46.[151,1436,193,1856]" captionTargetPageId="46" captionText="FIGURES 121 128. Aedeagi of Spinosciacharis spp. in ventral (121, 123, 125, 127) and lateral (122, 124, 126, 128) views. Spinosciacharis puncticeps (Franz) comb. n. (121 122); Spinosciacharis sassy sp. n. (123 124), Spinosciacharis clydensis sp. n. (125 126), and Spinosciacharis acheron sp. n. (127 128)." httpUri="https://zenodo.org/record/286539/files/figure.png" pageId="27" pageNumber="28">Fig. 125</figureCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BE03687FFCC18138FFE3B610066FDAE" blockId="27.[151,1436,258,403]" box="[151,810,381,403]" pageId="27" pageNumber="28">
<treatmentCitationGroup id="AB4F11A9FFCC18138FFE3B610066FDAE" box="[151,810,381,403]" pageId="27" pageNumber="28">
<taxonomicName id="4C5F4D04FFCC18138FFE3B610135FDAF" ID-CoL="3MNJC" authority="Franz, 1975: 149" authorityName="Franz" authorityPageNumber="149" authorityYear="1975" box="[151,633,381,403]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="australiensis">
<emphasis id="B92BEA95FFCC18138FFE3B610289FDAF" box="[151,453,381,403]" italics="true" pageId="27" pageNumber="28">Horaeomorphus australiensis</emphasis>
<treatmentCitation id="0AFE1096FFCC18138EA23B620135FDAF" author="Franz" box="[459,633,381,403]" page="149" pageId="27" pageNumber="28" year="1975">
<bibRefCitation id="EFCE4B76FFCC18138EA23B62010BFDAF" author="Franz" box="[459,583,381,403]" pageId="27" pageNumber="28" refString="Franz, H. (1975) Revision der Scydmaeniden von Australien, Neuseeland und den benachbarten Inseln. Denkschriften Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 118, 1 - 312." type="journal article" year="1975">Franz, 1975</bibRefCitation>
: 149
</treatmentCitation>
</taxonomicName>
,
<figureCitation id="13642A02FFCC18138DEA3B610196FDAF" box="[643,730,381,403]" captionStart="FIGURES 121 128" captionStartId="46.[151,264,1877,1900]" captionTargetBox="[168,1425,220,1825]" captionTargetId="figure@46.[151,1436,193,1856]" captionTargetPageId="46" captionText="FIGURES 121 128. Aedeagi of Spinosciacharis spp. in ventral (121, 123, 125, 127) and lateral (122, 124, 126, 128) views. Spinosciacharis puncticeps (Franz) comb. n. (121 122); Spinosciacharis sassy sp. n. (123 124), Spinosciacharis clydensis sp. n. (125 126), and Spinosciacharis acheron sp. n. (127 128)." httpUri="https://zenodo.org/record/286539/files/figure.png" pageId="27" pageNumber="28">Fig. 126</figureCitation>
,
<emphasis id="B92BEA95FFCC18138D8D3B610066FDAE" bold="true" box="[740,810,381,402]" pageId="27" pageNumber="28">syn. n.</emphasis>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C345650CFFCC18138FFE3BA3001AF94F" pageId="27" pageNumber="28" type="materials_examined">
<paragraph id="8BE03687FFCC18138FFE3BA302EFFF2B" blockId="27.[151,1437,447,2033]" pageId="27" pageNumber="28">
<emphasis id="B92BEA95FFCC18138FFE3BA302D1FDE4" bold="true" box="[151,413,447,472]" pageId="27" pageNumber="28">
<typeStatus id="54E48825FFCC18138FFE3BA3039EFDE4" box="[151,210,447,472]" pageId="27" pageNumber="28">Type</typeStatus>
material studied
</emphasis>
.
<emphasis id="B92BEA95FFCC18138EC53BA300F4FDE4" bold="true" box="[428,952,447,472]" pageId="27" pageNumber="28">
<typeStatus id="54E48825FFCC18138EC53BA3016EFDE4" box="[428,546,447,472]" pageId="27" pageNumber="28" type="lectotype">Lectotype</typeStatus>
(here designated):
<collectingCountry id="F3487617FFCC18138C633BA300FFFDE4" box="[778,947,447,472]" name="Australia" pageId="27" pageNumber="28">AUSTRALIA</collectingCountry>
:
</emphasis>
♂: five labels (
<figureCitation id="13642A02FFCC18138B033BA3078CFDE4" box="[1130,1216,447,472]" captionStart="FIGURES 72 79" captionStartId="33.[151,264,1586,1609]" captionTargetBox="[155,1432,200,1548]" captionTargetId="figure@33.[151,1436,193,1565]" captionTargetPageId="33" captionText="FIGURES 72 79. Scydmaenozila spp. Dorsal habitus (72 75), original set of labels of lectotypes (76, 79), paralectotype (77), and mounting card with three syntypes (with reverse side in bottom) before re-mounting during the present study (78). Scydmaenozila macrosticta (Lea) comb. n. (72, 76 77); Scydmaenozila latipennis (Lea) comb. n. (73, 78 79), Scydmaenozila mallacootana sp. n. (74), and Scydmaenozila inexpectata sp. n. (75)." httpUri="https://zenodo.org/record/286530/files/figure.png" pageId="27" pageNumber="28">Fig. 76</figureCitation>
): &quot;Co-type&quot; [white, printed], [
<taxonomicName id="4C5F4D04FFCC18138E6A3BFF02F7FDC0" box="[259,443,483,508]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="76" phylum="Arthropoda" rank="genus">Horaeomorphus</taxonomicName>
/
<taxonomicName id="4C5F4D04FFCC18138EB93BFF0111FDC0" box="[464,605,483,508]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="macrostictus">macrostictus</taxonomicName>
/ Lea / det. H. Franz&quot; [white, handwritten and printed], &quot;S. Aust. Museum / specimen&quot; [orange, printed], &quot;
<emphasis id="B92BEA95FFCC18138D44381B0065FE22" box="[557,809,519,542]" italics="true" pageId="27" pageNumber="28">
<taxonomicName id="4C5F4D04FFCC18138D44381B0041FE22" box="[557,781,519,542]" class="Insecta" family="Staphylinidae" genus="Phagonophana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">PHAGONOPHANA</taxonomicName>
/
</emphasis>
<taxonomicName id="4C5F4D04FFCC18138C573814071FFE23" authority="Lea, 1910" authorityName="Lea" authorityYear="1910" box="[830,1107,519,543]" class="Insecta" family="Staphylinidae" genus="Phagonophana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="macrosticta">
<emphasis id="B92BEA95FFCC18138C573814008DFE23" box="[830,961,520,543]" italics="true" pageId="27" pageNumber="28">macrosticta</emphasis>
<bibRefCitation id="EFCE4B76FFCC18138CBC3814071FFE23" author="Lea" box="[981,1107,519,543]" pageId="27" pageNumber="28" refString="Lea, A. M. (1910) Australian and Tasmanian Coleoptera inhabiting or resorting to the nests of ants, bees, and termites. Proceedings of the Royal Society of Victoria, Melbourne (N. S.), 23, 116 - 230." type="journal article" year="1910">Lea, 1910</bibRefCitation>
</taxonomicName>
/ P. Jałoszyński, 2014 /
<typeStatus id="54E48825FFCC18138FFE38370200FE7F" box="[151,332,555,579]" pageId="27" pageNumber="28" type="lectotype">LECTOTYPUS</typeStatus>
&quot; [white, printed], &quot;
<taxonomicName id="4C5F4D04FFCC18138D5D3837005EFE7E" box="[564,786,555,578]" class="Insecta" family="Staphylinidae" genus="Scydmaenozila" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="73" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B92BEA95FFCC18138D5D3837005EFE7E" box="[564,786,555,578]" italics="true" pageId="27" pageNumber="28">SCYDMAENOZILA</emphasis>
</taxonomicName>
/
<taxonomicName id="4C5F4D04FFCC18138C5D38370700FE7F" authority="Lea, 1910" authorityName="Lea" authorityYear="1910" box="[820,1100,555,579]" class="Insecta" family="Staphylinidae" genus="Phagonophana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="macrosticta">
<emphasis id="B92BEA95FFCC18138C5D383700FBFE7E" box="[820,951,555,578]" italics="true" pageId="27" pageNumber="28">macrosticta</emphasis>
(
<bibRefCitation id="EFCE4B76FFCC18138CA538370708FE7F" author="Lea" box="[972,1092,555,579]" pageId="27" pageNumber="28" refString="Lea, A. M. (1910) Australian and Tasmanian Coleoptera inhabiting or resorting to the nests of ants, bees, and termites. Proceedings of the Royal Society of Victoria, Melbourne (N. S.), 23, 116 - 230." type="journal article" year="1910">Lea, 1910</bibRefCitation>
)
</taxonomicName>
/ det. P. Jałoszyński, 2014&quot; [white, printed] (
<collectionCode id="ED4EAE42FFCC18138E3C385202DBFE5A" box="[341,407,590,614]" httpUri="http://grbio.org/cool/mhew-9czz" name="South African Museum" pageId="27" pageNumber="28">SAM</collectionCode>
).
<typeStatus id="54E48825FFCC18138EC33851010BFE5A" box="[426,583,589,614]" pageId="27" pageNumber="28" type="paralectotype">Paralectotype</typeStatus>
: ♀: seven labels (
<figureCitation id="13642A02FFCC18138C673851002EFE5A" box="[782,866,589,614]" captionStart="FIGURES 72 79" captionStartId="33.[151,264,1586,1609]" captionTargetBox="[155,1432,200,1548]" captionTargetId="figure@33.[151,1436,193,1565]" captionTargetPageId="33" captionText="FIGURES 72 79. Scydmaenozila spp. Dorsal habitus (72 75), original set of labels of lectotypes (76, 79), paralectotype (77), and mounting card with three syntypes (with reverse side in bottom) before re-mounting during the present study (78). Scydmaenozila macrosticta (Lea) comb. n. (72, 76 77); Scydmaenozila latipennis (Lea) comb. n. (73, 78 79), Scydmaenozila mallacootana sp. n. (74), and Scydmaenozila inexpectata sp. n. (75)." httpUri="https://zenodo.org/record/286530/files/figure.png" pageId="27" pageNumber="28">Fig. 77</figureCitation>
): &quot;14300 /
<taxonomicName id="4C5F4D04FFCC18138CB7385107CFFE5A" box="[990,1155,589,614]" class="Insecta" family="Staphylinidae" genus="Phagonophana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="65" phylum="Arthropoda" rank="genus">Phagonophana</taxonomicName>
/
<taxonomicName id="4C5F4D04FFCC18138BF338510657FE5A" box="[1178,1307,589,614]" class="Insecta" family="Staphylinidae" genus="Phagonophana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="macrosticta">macrosticta</taxonomicName>
/ Tasmania /
<typeStatus id="54E48825FFCC18138FC0386E03BEFEB6" box="[169,242,626,650]" pageId="27" pageNumber="28">
<collectionCode id="ED4EAE42FFCC18138FC0386E03BEFEB6" box="[169,242,626,650]" pageId="27" pageNumber="28">TYPE</collectionCode>
</typeStatus>
&quot; [white, handwritten, &quot;
<typeStatus id="54E48825FFCC18138E95386E010BFEB6" box="[508,583,626,650]" pageId="27" pageNumber="28">
<collectionCode id="ED4EAE42FFCC18138E95386E010BFEB6" box="[508,583,626,650]" pageId="27" pageNumber="28">TYPE</collectionCode>
</typeStatus>
&quot; in red ink], &quot;
<taxonomicName id="4C5F4D04FFCC18138D86386D003FFEB6" box="[751,883,625,650]" class="Insecta" family="Staphylinidae" genus="Phagonophana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="macrosticta">macrosticta</taxonomicName>
/ Lea
<typeStatus id="54E48825FFCC18138CAA386E0745FEB5" box="[963,1033,626,649]" pageId="27" pageNumber="28">
<collectionCode id="ED4EAE42FFCC18138CAA386E0745FEB5" box="[963,1033,626,649]" pageId="27" pageNumber="28">TYPE</collectionCode>
</typeStatus>
/ Marrawah&quot; [white, handwritten], &quot;
<taxonomicName id="4C5F4D04FFCC18138FF63888021BFE91" box="[159,343,660,685]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="76" phylum="Arthropoda" rank="genus">Horaeomorphus</taxonomicName>
/
<taxonomicName id="4C5F4D04FFCC18138E05388802B5FE91" box="[364,505,660,685]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="macrostictus">macrostictus</taxonomicName>
/ Lea / det. H. Franz&quot; [white, handwritten and printed], &quot;
<collectionCode id="ED4EAE42FFCC18138B1838890789FE91" LSID="urn:lsid:biocol.org:col:34982" box="[1137,1221,661,685]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34982" name="South Australia Museum" pageId="27" pageNumber="28">SAMA</collectionCode>
Database / No. 25- 037062&quot; [white, printed], &quot;
<emphasis id="B92BEA95FFCC18138E9438A501B3FEEC" box="[509,767,697,720]" italics="true" pageId="27" pageNumber="28">
<taxonomicName id="4C5F4D04FFCC18138E9438A50191FEEC" box="[509,733,697,720]" class="Insecta" family="Staphylinidae" genus="Phagonophana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">PHAGONOPHANA</taxonomicName>
/
</emphasis>
<taxonomicName id="4C5F4D04FFCC18138C7138A50774FEEC" authority="Lea, 1910" authorityName="Lea" authorityYear="1910" box="[792,1080,696,720]" class="Insecta" family="Staphylinidae" genus="Phagonophana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="macrosticta">
<emphasis id="B92BEA95FFCC18138C7138A500D7FEEC" box="[792,923,697,720]" italics="true" pageId="27" pageNumber="28">macrosticta</emphasis>
<bibRefCitation id="EFCE4B76FFCC18138CDC38A50774FEEC" author="Lea" box="[949,1080,696,720]" pageId="27" pageNumber="28" refString="Lea, A. M. (1910) Australian and Tasmanian Coleoptera inhabiting or resorting to the nests of ants, bees, and termites. Proceedings of the Royal Society of Victoria, Melbourne (N. S.), 23, 116 - 230." type="journal article" year="1910">Lea, 1910</bibRefCitation>
</taxonomicName>
/ P. Jałoszyński, 2014 /
<typeStatus id="54E48825FFCC18138FFE38C002D9FEC8" box="[151,405,732,756]" pageId="27" pageNumber="28" type="paralectotype">PARALECTOTYPUS</typeStatus>
&quot; [white, printed], &quot;
<taxonomicName id="4C5F4D04FFCC18138D1E38C0001AFECF" box="[631,854,732,755]" class="Insecta" family="Staphylinidae" genus="Scydmaenozila" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="73" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B92BEA95FFCC18138D1E38C0001AFECF" box="[631,854,732,755]" italics="true" pageId="27" pageNumber="28">SCYDMAENOZILA</emphasis>
</taxonomicName>
/
<taxonomicName id="4C5F4D04FFCC18138C1D38C107C5FEC8" authority="Lea, 1910" authorityName="Lea" authorityYear="1910" box="[884,1161,732,756]" class="Insecta" family="Staphylinidae" genus="Phagonophana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="macrosticta">
<emphasis id="B92BEA95FFCC18138C1D38C100BBFEC8" box="[884,1015,733,756]" italics="true" pageId="27" pageNumber="28">macrosticta</emphasis>
(
<bibRefCitation id="EFCE4B76FFCC18138B6338C007CDFEC8" author="Lea" box="[1034,1153,732,756]" pageId="27" pageNumber="28" refString="Lea, A. M. (1910) Australian and Tasmanian Coleoptera inhabiting or resorting to the nests of ants, bees, and termites. Proceedings of the Royal Society of Victoria, Melbourne (N. S.), 23, 116 - 230." type="journal article" year="1910">Lea, 1910</bibRefCitation>
)
</taxonomicName>
/ P. Jałoszyński, 2014&quot; [white, printed] (
<collectionCode id="ED4EAE42FFCC18138E3D38E302DAFF2B" box="[340,406,767,791]" httpUri="http://grbio.org/cool/mhew-9czz" name="South African Museum" pageId="27" pageNumber="28">SAM</collectionCode>
).
</paragraph>
<paragraph id="8BE03687FFCC18138FAE393E0076F934" blockId="27.[151,1437,447,2033]" pageId="27" pageNumber="28">
<emphasis id="B92BEA95FFCC18138FAE393E02F6FF07" bold="true" box="[199,442,802,827]" pageId="27" pageNumber="28">Additional material</emphasis>
(38 exx.).
<specimenCount id="9D59FD0EFFCC18138D22393F0132FF06" box="[587,638,803,827]" pageId="27" pageNumber="28" type="generic">1 ♂</specimenCount>
, 1 ♀ (
<typeStatus id="54E48825FFCC18138DB0393E0071FF07" box="[729,829,802,827]" pageId="27" pageNumber="28" type="holotype">holotype</typeStatus>
and
<typeStatus id="54E48825FFCC18138CEA393F00A8FF07" box="[899,996,803,827]" pageId="27" pageNumber="28" type="paratype">paratype</typeStatus>
of
<taxonomicName id="4C5F4D04FFCC18138B7E393E0621FF07" box="[1047,1389,802,827]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="australiensis">
<emphasis id="B92BEA95FFCC18138B7E393E0621FF07" box="[1047,1389,802,827]" italics="true" pageId="27" pageNumber="28">Horaeomorphus australiensis</emphasis>
</taxonomicName>
): S
<collectingCountry id="F3487617FFCC18138FFE395A024DFF63" box="[151,257,838,863]" name="Australia" pageId="27" pageNumber="28">Australia</collectingCountry>
, Bakewell (NHM); 1 ex., Australian Capital Territory, Blundells Creek,
<quantity id="4CA79B62FFCC18138B45395B0729FF63" box="[1068,1125,838,863]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="27" pageNumber="28" unit="km" value="3.0">3 km</quantity>
E of Piccadilly Circus,
<quantity id="4CA79B62FFCC18138A1B395A03E3FFBD" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="27" pageNumber="28" unit="m" value="850.0">850 m</quantity>
,
<date id="FFE11047FFCC18138FD239750263FFBE" box="[187,303,873,898]" pageId="27" pageNumber="28" value="1984-02">Feb. 1984</date>
, &quot;flight intercept window / trough trap&quot;, leg. Lawrence &amp; Johnson; 8 exx., same data except for
<date id="FFE11047FFCC18138A193976039EFF9A" pageId="27" pageNumber="28" value="1985-01">Jan. 1985</date>
, leg. Weir, Lawrence, Johnson; 3 exx., same data, except for
<date id="FFE11047FFCC18138CFE39920743FF9A" box="[919,1039,909,934]" pageId="27" pageNumber="28" value="1985-03">Mar. 1985</date>
; 5 exx., same data except for
<date id="FFE11047FFCC18138A023991039EFFF5" pageId="27" pageNumber="28" value="1985-04">Apr. 1985</date>
; 1 ex., same data, except for
<date id="FFE11047FFCC18138D7239AD01DDFFF5" box="[539,657,944,969]" pageId="27" pageNumber="28" value="1985-05">May 1985</date>
; 2 exx., same data, except for
<date id="FFE11047FFCC18138C8E39AD0717FFF5" box="[999,1115,944,969]" pageId="27" pageNumber="28" value="1985-09">Sep. 1985</date>
; 1 ex., same data, except for
<date id="FFE11047FFCC18138FFE39C80244FFD1" box="[151,264,980,1005]" pageId="27" pageNumber="28" value="1985-10">Oct. 1985</date>
; 1 ex., same data except for
<quantity id="4CA79B62FFCC18138D2039C801CEFFD1" box="[585,642,980,1005]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="27" pageNumber="28" unit="km" value="6.0">6 km</quantity>
NE of Piccadilly Circus,
<quantity id="4CA79B62FFCC18138CCA39C800A0FFD0" box="[931,1004,980,1005]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="27" pageNumber="28" unit="m" value="750.0">750 m</quantity>
,
<date id="FFE11047FFCC18138C9139C80727FFD1" box="[1016,1131,980,1005]" pageId="27" pageNumber="28" value="1985-02">Feb. 1985</date>
; 2 exx., Blundells Creek,
<quantity id="4CA79B62FFCC18138AE739C803F7F82C" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="27" pageNumber="28" unit="km" value="3.0">3 km</quantity>
E of Piccadilly Circus,
<quantity id="4CA79B62FFCC18138EBA39EB0153F833" box="[467,543,1015,1040]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="27" pageNumber="28" unit="m" value="850.0">850 m</quantity>
,
<date id="FFE11047FFCC18138D4439E4018DF82C" box="[557,705,1015,1040]" pageId="27" pageNumber="28" value="1986-09-07">7 Sept. 1986</date>
, &quot;Berlesate,
<collectionCode id="ED4EAE42FFCC18138C3D39E400D4F82C" LSID="urn:lsid:biocol.org:col:32981" box="[852,920,1016,1040]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="27" pageNumber="28">ANIC</collectionCode>
1066, leaf litter&quot;, leg. J. F. Lawrence; Black Mountain near Canberra, &quot;near light trap&quot;, flight intercept trap,
<date id="FFE11047FFCC18138C0B3E070754F808" box="[866,1048,1051,1076]" pageId="27" pageNumber="28" value="1996-09-09" valueMax="1996-09-30" valueMin="1996-09-09">9-30 Sept. 1996</date>
, leg. T. Weir &amp; W. Dressler; 1 ex., Tasmania, Cradle Mountain, leg. J. Armstrong; 1 ex., Tasmania,
<quantity id="4CA79B62FFCC18138C103E2300FFF86B" box="[889,947,1086,1111]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="27" pageNumber="28" unit="km" value="7.0">7 km</quantity>
SWW Derwent Bridge, pantrap,
<date id="FFE11047FFCC18138A583E220245F846" pageId="27" pageNumber="28" value="1983-01-06" valueMax="1983-02-02" valueMin="1983-01-06">6 Jan. - 2 Feb. 1983</date>
, leg. I. Naumann &amp; J. Cardale; 1 ex., Tasmania, Hobart, Mt. Nelson, dry sclerophyll, under fallen branch,
<date id="FFE11047FFCC18138FFE3E990274F8A2" box="[151,312,1157,1182]" pageId="27" pageNumber="28" value="1992-02-02" valueMax="1992-02-05" valueMin="1992-02-02">2-5 Feb. 1992</date>
, leg. C. Reid; 2 exx., Tasmania, Anson's Bay Road, pitfall trap, &quot;Leptospermum, in wet heath&quot;,
<date id="FFE11047FFCC18138A1E3E9A0247F8FD" pageId="27" pageNumber="28" value="1993-02-20" valueMax="1993-02-25" valueMin="1993-02-20">20- 25.ii.1993</date>
, leg. J. Trueman et al.; 2 exx., same data except for Peters Link Road, dry sclerophyll; 1 ex., same data, except for
<date id="FFE11047FFCC18138E7A3ED102E6F8D9" box="[275,426,1229,1253]" pageId="27" pageNumber="28" value="1993-05-22" valueMax="1993-05-27" valueMin="1993-05-22">22-27.v.1993</date>
; 2 exx., same data, except for Eddystone Point,
<date id="FFE11047FFCC18138CBF3ED00723F8D9" box="[982,1135,1228,1253]" pageId="27" pageNumber="28" value="1993-02-20" valueMax="1993-02-25" valueMin="1993-02-20">20-25.ii.1993</date>
, coastal heath; 1 ex. same data, except for
<date id="FFE11047FFCC18138E223EEC0293F934" box="[331,479,1264,1288]" pageId="27" pageNumber="28" value="1993-05-22" valueMax="1993-05-27" valueMin="1993-05-22">22-27.v.1993</date>
(specimens in
<collectionCode id="ED4EAE42FFCC18138DEE3EEC0187F934" LSID="urn:lsid:biocol.org:col:32981" box="[647,715,1264,1288]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="27" pageNumber="28">ANIC</collectionCode>
and cPJ).
</paragraph>
<paragraph id="8BE03687FFCC18138FAE3F0F001AF94F" blockId="27.[151,1437,447,2033]" pageId="27" pageNumber="28">
<emphasis id="B92BEA95FFCC18138FAE3F0F02E8F910" bold="true" box="[199,420,1299,1324]" pageId="27" pageNumber="28">Revised diagnosis.</emphasis>
Male and female: elytra with variously distinct pattern: base of elytra and subtriangular subapical spots dark brown, remaining surface light brown. Male: aedeagus slender, in ventral view with broadly rounded apex; apical part of median lobe abruptly demarcated.
</paragraph>
</subSubSection>
<subSubSection id="C345650CFFCC18158FAE3F610394FF28" lastPageId="29" lastPageNumber="30" pageId="27" pageNumber="28" type="description">
<paragraph id="8BE03687FFCC18138FAE3F61067DFA3C" blockId="27.[151,1437,447,2033]" pageId="27" pageNumber="28">
<emphasis id="B92BEA95FFCC18138FAE3F610238F9AA" bold="true" box="[199,372,1405,1430]" pageId="27" pageNumber="28">Redescription.</emphasis>
Body of male (
<figureCitation id="13642A02FFCC18138D5C3F6101C6F9AA" box="[565,650,1405,1430]" captionStart="FIGURES 72 79" captionStartId="33.[151,264,1586,1609]" captionTargetBox="[155,1432,200,1548]" captionTargetId="figure@33.[151,1436,193,1565]" captionTargetPageId="33" captionText="FIGURES 72 79. Scydmaenozila spp. Dorsal habitus (72 75), original set of labels of lectotypes (76, 79), paralectotype (77), and mounting card with three syntypes (with reverse side in bottom) before re-mounting during the present study (78). Scydmaenozila macrosticta (Lea) comb. n. (72, 76 77); Scydmaenozila latipennis (Lea) comb. n. (73, 78 79), Scydmaenozila mallacootana sp. n. (74), and Scydmaenozila inexpectata sp. n. (75)." httpUri="https://zenodo.org/record/286530/files/figure.png" pageId="27" pageNumber="28">Fig. 72</figureCitation>
) strongly convex, elongate and slender, with long appendages, BL
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(mean
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); cuticle glossy, head, pronotum and appendages light to dark brown, each elytron with dark suture, base and subtriangular subapical spot, remaining surface lighter (in some cases indistinctly but the pattern still discernible), vestiture dark brown, on elytra slightly darker than on remaining body parts.
</paragraph>
<paragraph id="8BE03687FFCC18138FAE3C17011AFB20" blockId="27.[151,1437,447,2033]" pageId="27" pageNumber="28">
Head (
<figureCitation id="13642A02FFCC18138E783C170223FA18" box="[273,367,1547,1572]" captionStart="FIGURES 57 61" captionStartId="28.[151,264,1783,1806]" captionTargetBox="[151,1411,193,1762]" captionTargetId="figure@28.[151,1436,193,1763]" captionTargetPageId="28" captionText="FIGURES 57 61. Morphology of Scydmaenozila macrosticta (Lea) comb. n. Head in ventral view (57), distal part of labium in ventral view (58), labrum in dorsal view (59), right mandible in dorsal view (60), and right maxilla in ventral view (61). Abbreviations: bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hr, hypostomal ridge; lac, lacinia; lg, ligula; lp, labial palpus; lp 1 3, labial palpomere 1 3; md, mandible; mn, mentum; mst, mediostipes; mxp, maxillary palpus; mxp 1 4, maxillary palpomere 1 4; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; sat, subapical tooth; smn, submentum." httpUri="https://zenodo.org/record/286527/files/figure.png" pageId="27" pageNumber="28">Figs. 57</figureCitation>
,
<figureCitation id="13642A02FFCC18138E133C1002DBFA18" box="[378,407,1548,1572]" captionStart="FIGURES 72 79" captionStartId="33.[151,264,1586,1609]" captionTargetBox="[155,1432,200,1548]" captionTargetId="figure@33.[151,1436,193,1565]" captionTargetPageId="33" captionText="FIGURES 72 79. Scydmaenozila spp. Dorsal habitus (72 75), original set of labels of lectotypes (76, 79), paralectotype (77), and mounting card with three syntypes (with reverse side in bottom) before re-mounting during the present study (78). Scydmaenozila macrosticta (Lea) comb. n. (72, 76 77); Scydmaenozila latipennis (Lea) comb. n. (73, 78 79), Scydmaenozila mallacootana sp. n. (74), and Scydmaenozila inexpectata sp. n. (75)." httpUri="https://zenodo.org/record/286530/files/figure.png" pageId="27" pageNumber="28">72</figureCitation>
) broadest at eyes, HL
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(mean
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), HW
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(mean
<quantity id="4CA79B62FFCC18138A413C1706DCFA18" box="[1320,1424,1547,1572]" metricMagnitude="-4" metricUnit="m" metricValue="5.6" pageId="27" pageNumber="28" unit="mm" value="0.56">0.56 mm</quantity>
); tempora longer than eyes, strongly curved posteromesally; vertex weakly convex; frons subtrapezoidal. Eyes large and strongly projecting laterally from the head silhouette, finely faceted. Punctures on head dorsum fine, separated by spaces 1.5
<date id="FFE11047FFCC18138E553C690206FAB2" box="[316,330,1653,1678]" pageId="27" pageNumber="28">2</date>
x as wide as diameters of punctures; setae on frons and anterior portion of vertex short, sparse and suberect, tempora and posterior portion of vertex densely covered with long bristles directed posteriorly and posterolaterally. Antennae (
<figureCitation id="13642A02FFCC18138EA63CA0016EFAE9" box="[463,546,1724,1749]" captionStart="FIGURES 72 79" captionStartId="33.[151,264,1586,1609]" captionTargetBox="[155,1432,200,1548]" captionTargetId="figure@33.[151,1436,193,1565]" captionTargetPageId="33" captionText="FIGURES 72 79. Scydmaenozila spp. Dorsal habitus (72 75), original set of labels of lectotypes (76, 79), paralectotype (77), and mounting card with three syntypes (with reverse side in bottom) before re-mounting during the present study (78). Scydmaenozila macrosticta (Lea) comb. n. (72, 76 77); Scydmaenozila latipennis (Lea) comb. n. (73, 78 79), Scydmaenozila mallacootana sp. n. (74), and Scydmaenozila inexpectata sp. n. (75)." httpUri="https://zenodo.org/record/286530/files/figure.png" pageId="27" pageNumber="28">Fig. 72</figureCitation>
) slender, AnL
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(mean
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); antennomeres IVII distinctly elongate, VIII slightly longer than wide, IXX each about as long as broad, XI as long or nearly as long as IXX together, 2.5x as long as broad, pointed.
</paragraph>
<paragraph id="8BE03687FFCC18138FAE3D3401D5FBCC" blockId="27.[151,1437,447,2033]" pageId="27" pageNumber="28">
Pronotum (
<figureCitation id="13642A02FFCC18138E203D3B02E7FB7C" box="[329,427,1831,1856]" captionStart="FIGURES 62 66" captionStartId="30.[151,264,1734,1757]" captionTargetBox="[151,1436,193,1712]" captionTargetId="figure@30.[151,1436,193,1713]" captionTargetPageId="30" captionText="FIGURES 62 66. Morphology of Scydmaenozila macrosticta (Lea) comb. n. Prothorax in ventral and slightly lateral view (62 63), base of pronotum in dorsal view (64), mesonotum and elytral base in dorsal view (65), and mesocoxa in dorsal view (66). Abbreviations: bef, basal elytral foveae; bst, basisternal part of prosternum; cx 2, mesocoxa; eal, elytral articulating lobe; fm 2, mesofemur; hy, hypomeron; ip, internal antebasal pit; lp, lateral antebasal pit; nss, pronosternal suture; pcc. procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; sc 2, mesoscutum; scl 2, mesoscutellum; sss, mesoscutoscutellar suture; tr 2, mesotrochanter." httpUri="https://zenodo.org/record/286528/files/figure.png" pageId="27" pageNumber="28">Figs. 64</figureCitation>
,
<figureCitation id="13642A02FFCC18138ED33D340294FB7C" box="[442,472,1832,1856]" captionStart="FIGURES 72 79" captionStartId="33.[151,264,1586,1609]" captionTargetBox="[155,1432,200,1548]" captionTargetId="figure@33.[151,1436,193,1565]" captionTargetPageId="33" captionText="FIGURES 72 79. Scydmaenozila spp. Dorsal habitus (72 75), original set of labels of lectotypes (76, 79), paralectotype (77), and mounting card with three syntypes (with reverse side in bottom) before re-mounting during the present study (78). Scydmaenozila macrosticta (Lea) comb. n. (72, 76 77); Scydmaenozila latipennis (Lea) comb. n. (73, 78 79), Scydmaenozila mallacootana sp. n. (74), and Scydmaenozila inexpectata sp. n. (75)." httpUri="https://zenodo.org/record/286530/files/figure.png" pageId="27" pageNumber="28">72</figureCitation>
) in dorsal view strongly elongate, broadest near anterior third, PL
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, (mean
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), PW
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(mean
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); anterior and lateral margins in anterior half rounded; sides slightly concave in posterior third; posterior margin distinctly bisinuate; base of pronotum with two pairs of large and deep pits, additionally with a pair of shallow but distinct impressions located dorsolaterally. Punctures on pronotal disc fine and inconspicuous; entire dorsal and lateral surface of pronotum covered with moderately dense bristles, dorsally with admixture of thin setae.
</paragraph>
<caption id="DF20660FFFCB18148FFE3CEB02BFFB9B" httpUri="https://zenodo.org/record/286527/files/figure.png" pageId="28" pageNumber="29" targetBox="[151,1411,193,1762]" targetPageId="28">
<paragraph id="8BE03687FFCB18148FFE3CEB02BFFB9B" blockId="28.[151,1436,1783,1959]" pageId="28" pageNumber="29">
<emphasis id="B92BEA95FFCB18148FFE3CEB0218FB31" bold="true" box="[151,340,1783,1806]" pageId="28" pageNumber="29">FIGURES 5761.</emphasis>
Morphology of
<taxonomicName id="4C5F4D04FFCB18148E953CE40003FB32" authority="Lea" authorityName="Lea" box="[508,847,1784,1806]" class="Insecta" family="Staphylinidae" genus="Scydmaenozila" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="macrosticta" status="comb. nov.">
<emphasis id="B92BEA95FFCB18148E953CE4005EFB32" box="[508,786,1784,1806]" italics="true" pageId="28" pageNumber="29">Scydmaenozila macrosticta</emphasis>
(Lea)
</taxonomicName>
<emphasis id="B92BEA95FFCB18148C3F3CE400FFFB31" bold="true" box="[854,947,1784,1805]" pageId="28" pageNumber="29">
<taxonomicNameLabel id="A21857EEFFCB18148C3F3CE400FFFB31" box="[854,947,1784,1805]" pageId="28" pageNumber="29">comb. n.</taxonomicNameLabel>
</emphasis>
Head in ventral view (57), distal part of labium in ventral view (58), labrum in dorsal view (59), right mandible in dorsal view (60), and right maxilla in ventral view (61). Abbreviations: bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hr, hypostomal ridge; lac, lacinia; lg, ligula; lp, labial palpus; lp13, labial palpomere 13; md, mandible; mn, mentum; mst, mediostipes; mxp, maxillary palpus; mxp14, maxillary palpomere 14; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; sat, subapical tooth; smn, submentum.
</paragraph>
</caption>
<paragraph id="8BE03687FFCA18158FAE3A8B078DFD7C" blockId="29.[151,1437,151,1977]" pageId="29" pageNumber="30">
Elytra (
<figureCitation id="13642A02FFCA18158E723A8B0235FC8C" box="[283,377,151,176]" captionStart="FIGURES 62 66" captionStartId="30.[151,264,1734,1757]" captionTargetBox="[151,1436,193,1712]" captionTargetId="figure@30.[151,1436,193,1713]" captionTargetPageId="30" captionText="FIGURES 62 66. Morphology of Scydmaenozila macrosticta (Lea) comb. n. Prothorax in ventral and slightly lateral view (62 63), base of pronotum in dorsal view (64), mesonotum and elytral base in dorsal view (65), and mesocoxa in dorsal view (66). Abbreviations: bef, basal elytral foveae; bst, basisternal part of prosternum; cx 2, mesocoxa; eal, elytral articulating lobe; fm 2, mesofemur; hy, hypomeron; ip, internal antebasal pit; lp, lateral antebasal pit; nss, pronosternal suture; pcc. procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; sc 2, mesoscutum; scl 2, mesoscutellum; sss, mesoscutoscutellar suture; tr 2, mesotrochanter." httpUri="https://zenodo.org/record/286528/files/figure.png" pageId="29" pageNumber="30">Figs. 65</figureCitation>
,
<figureCitation id="13642A02FFCA18158EEF3A8402EFFC8C" box="[390,419,152,176]" captionStart="FIGURES 72 79" captionStartId="33.[151,264,1586,1609]" captionTargetBox="[155,1432,200,1548]" captionTargetId="figure@33.[151,1436,193,1565]" captionTargetPageId="33" captionText="FIGURES 72 79. Scydmaenozila spp. Dorsal habitus (72 75), original set of labels of lectotypes (76, 79), paralectotype (77), and mounting card with three syntypes (with reverse side in bottom) before re-mounting during the present study (78). Scydmaenozila macrosticta (Lea) comb. n. (72, 76 77); Scydmaenozila latipennis (Lea) comb. n. (73, 78 79), Scydmaenozila mallacootana sp. n. (74), and Scydmaenozila inexpectata sp. n. (75)." httpUri="https://zenodo.org/record/286530/files/figure.png" pageId="29" pageNumber="30">72</figureCitation>
) regularly oval, broadest near middle, EL
<quantity id="4CA79B62FFCA18158CED3A8B076BFC93" box="[900,1063,151,176]" metricMagnitude="-3" metricUnit="m" metricValue="1.49" metricValueMax="1.6" metricValueMin="1.38" pageId="29" pageNumber="30" unit="mm" value="1.49" valueMax="1.6" valueMin="1.38">1.381.60 mm</quantity>
(mean
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), EW
<quantity id="4CA79B62FFCA18158A453A84038FFCE8" metricMagnitude="-3" metricUnit="m" metricValue="1.015" metricValueMax="1.13" metricValueMin="0.9" pageId="29" pageNumber="30" unit="mm" value="1.015" valueMax="1.13" valueMin="0.9">0.901.13 mm</quantity>
(mean
<quantity id="4CA79B62FFCA18158E773AA002C5FCE8" box="[286,393,188,212]" metricMagnitude="-3" metricUnit="m" metricValue="1.04" pageId="29" pageNumber="30" unit="mm" value="1.04">1.04 mm</quantity>
), EI 1.331.53 (mean 1.44); basal impressions short but distinct; elytral apices separately rounded. Punctures on elytral disc fine but slightly more distinct than those on pronotum, separated by spaces 2
<date id="FFE11047FFCA18158AEB3AFC06C2FCC4" box="[1410,1422,224,248]" pageId="29" pageNumber="30">3</date>
x as wide as diameters of punctures; setae long, curved sparse and erect, in lightly pigmented specimens setae on light elytral spots contrastingly dark.
<collectingCountry id="F3487617FFCA18158D503B3B013DFD7C" box="[569,625,295,320]" name="India" pageId="29" pageNumber="30">Hind</collectingCountry>
wings well developed, about twice as long as elytra.
</paragraph>
<paragraph id="8BE03687FFCA18158FAE3B51002AFD59" blockId="29.[151,1437,151,1977]" box="[199,870,332,357]" pageId="29" pageNumber="30">
Legs (
<figureCitation id="13642A02FFCA18158E653B500226FD59" box="[268,362,332,357]" captionStart="FIGURES 62 66" captionStartId="30.[151,264,1734,1757]" captionTargetBox="[151,1436,193,1712]" captionTargetId="figure@30.[151,1436,193,1713]" captionTargetPageId="30" captionText="FIGURES 62 66. Morphology of Scydmaenozila macrosticta (Lea) comb. n. Prothorax in ventral and slightly lateral view (62 63), base of pronotum in dorsal view (64), mesonotum and elytral base in dorsal view (65), and mesocoxa in dorsal view (66). Abbreviations: bef, basal elytral foveae; bst, basisternal part of prosternum; cx 2, mesocoxa; eal, elytral articulating lobe; fm 2, mesofemur; hy, hypomeron; ip, internal antebasal pit; lp, lateral antebasal pit; nss, pronosternal suture; pcc. procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; sc 2, mesoscutum; scl 2, mesoscutellum; sss, mesoscutoscutellar suture; tr 2, mesotrochanter." httpUri="https://zenodo.org/record/286528/files/figure.png" pageId="29" pageNumber="30">Figs. 66</figureCitation>
,
<figureCitation id="13642A02FFCA18158E1F3B5002D8FD58" box="[374,404,332,356]" captionStart="FIGURES 72 79" captionStartId="33.[151,264,1586,1609]" captionTargetBox="[155,1432,200,1548]" captionTargetId="figure@33.[151,1436,193,1565]" captionTargetPageId="33" captionText="FIGURES 72 79. Scydmaenozila spp. Dorsal habitus (72 75), original set of labels of lectotypes (76, 79), paralectotype (77), and mounting card with three syntypes (with reverse side in bottom) before re-mounting during the present study (78). Scydmaenozila macrosticta (Lea) comb. n. (72, 76 77); Scydmaenozila latipennis (Lea) comb. n. (73, 78 79), Scydmaenozila mallacootana sp. n. (74), and Scydmaenozila inexpectata sp. n. (75)." httpUri="https://zenodo.org/record/286530/files/figure.png" pageId="29" pageNumber="30">72</figureCitation>
) long and slender, without modifications.
</paragraph>
<paragraph id="8BE03687FFCA18158FAE3B7307A8FE01" blockId="29.[151,1437,151,1977]" pageId="29" pageNumber="30">
Aedeagus (
<figureCitation id="13642A02FFCA18158E2C3B7302EEFDB4" box="[325,418,367,392]" captionStart="FIGURES 67 71" captionStartId="31.[151,264,1849,1872]" captionTargetBox="[162,1426,193,1831]" captionTargetId="figure@31.[162,1427,193,1833]" captionTargetPageId="31" captionText="FIGURES 67 71. Morphology of Scydmaenozila macrosticta (Lea) comb. n. Mesothorax in ventral view (67), pterothorax in ventral view (68 69), aedeagus in ventral and slightly lateral view (70), and metaventral intercoxal process in ventral view (71). Abbreviations: aest 3, metanepisternum; aml, apex of median lobe; ar, anterior ridge; cx 3, metacoxa; dlf, dorsoventral fovea; fm 3, metafemur; lmfa, lateral mesofurcal arm; ma, membranous area; mcp, mesocoxal process; mscc, mesocoxal cavity; mscs, mesocoxal socket; msf, mesofurca; msvp, mesoventral process; mtvp, metaventral intercoxal process; pcr, procoxal rest; pm, paramere; pre, prepectus; tr 3, metatrochanter; vlf, ventrolateral fovea." httpUri="https://zenodo.org/record/286529/files/figure.png" pageId="29" pageNumber="30">Figs. 70</figureCitation>
,
<figureCitation id="13642A02FFCA18158EC73B6C02BBFDB4" box="[430,503,368,392]" captionStart="FIGURES 80 88" captionStartId="34.[151,264,1601,1624]" captionTargetBox="[163,1426,499,1576]" captionTargetId="figure@34.[151,1436,481,1580]" captionTargetPageId="34" captionText="FIGURES 80 88. Aedeagi of Scydmaenozila spp. in ventral (80, 82, 83, 85, 87) and lateral (81, 84, 86, 88) views. Scydmaenozila macrosticta (Lea) comb. n., intact (80 81) and erected (82); Scydmaenozila latipennis (Lea) comb. n. (83 84), Scydmaenozila mallacootana sp. n. (85 86), and Scydmaenozila inexpectata sp. n. (87 88)." httpUri="https://zenodo.org/record/286531/files/figure.png" pageId="29" pageNumber="30">8082</figureCitation>
,
<figureCitation id="13642A02FFCA18158D6B3B6C0106FDB4" box="[514,586,368,392]" captionStart="FIGURES 89 96" captionStartId="35.[151,264,1860,1883]" captionTargetBox="[166,1428,430,1831]" captionTargetId="figure@35.[151,1436,411,1839]" captionTargetPageId="35" captionText="FIGURES 89 96. Aedeagi of Scydmaenozila spp. in ventral (89, 91, 93, 95) and lateral (90, 92, 94, 96) views. Scydmaenozila macrosticta (Lea) comb. n. (89 90); Scydmaenozila latipennis (Lea) comb. n. (91 92), Scydmaenozila mallacootana sp. n. (93 94), and Scydmaenozila inexpectata sp. n. (95 96)." httpUri="https://zenodo.org/record/286532/files/figure.png" pageId="29" pageNumber="30">8990</figureCitation>
) elongate and relatively slender; AeL
<quantity id="4CA79B62FFCA18158C913B730713FDBB" box="[1016,1119,367,392]" metricMagnitude="-4" metricUnit="m" metricValue="5.8" pageId="29" pageNumber="30" unit="mm" value="0.58">0.58 mm</quantity>
; in ventral view short apical part abruptly demarcated, with broadly and weakly rounded apical margin; in lateral view apical part subtriangular and sharply demarcated from ventral wall; internal armature complicated, with submedian complex of elongate and darkly sclerotized structures and subapical finely denticulate lateral lobes of internal sac; parameres long and slender, with indistinctly broadened apices. Aedeagus erected (
<figureCitation id="13642A02FFCA18158CEC3BE30091FE24" box="[901,989,511,536]" captionStart="FIGURES 80 88" captionStartId="34.[151,264,1601,1624]" captionTargetBox="[163,1426,499,1576]" captionTargetId="figure@34.[151,1436,481,1580]" captionTargetPageId="34" captionText="FIGURES 80 88. Aedeagi of Scydmaenozila spp. in ventral (80, 82, 83, 85, 87) and lateral (81, 84, 86, 88) views. Scydmaenozila macrosticta (Lea) comb. n., intact (80 81) and erected (82); Scydmaenozila latipennis (Lea) comb. n. (83 84), Scydmaenozila mallacootana sp. n. (85 86), and Scydmaenozila inexpectata sp. n. (87 88)." httpUri="https://zenodo.org/record/286531/files/figure.png" pageId="29" pageNumber="30">Fig. 82</figureCitation>
) with all components of internal sac extruded and still easily recognizable submedian darkly sclerotized subtriangular central complex.
</paragraph>
<paragraph id="8BE03687FFCA18158FAE385B0394FF28" blockId="29.[151,1437,151,1977]" pageId="29" pageNumber="30">
<emphasis id="B92BEA95FFCA18158FAE385B0252FE5C" bold="true" box="[199,286,583,608]" pageId="29" pageNumber="30">Female</emphasis>
. Similar to male, on average slightly larger, but body lengths and proportions of body parts overlap with those measured in males; distinguished by antennomere VIII about as long as broad and antennomeres IXX slightly transverse. BL
<quantity id="4CA79B62FFCA18158EC8388C0109FE9B" box="[417,581,656,680]" metricMagnitude="-3" metricUnit="m" metricValue="2.855" metricValueMax="2.98" metricValueMin="2.73" pageId="29" pageNumber="30" unit="mm" value="2.855" valueMax="2.98" valueMin="2.73">2.732.98 mm</quantity>
(mean
<quantity id="4CA79B62FFCA18158DF538930049FE94" box="[668,773,655,680]" metricMagnitude="-3" metricUnit="m" metricValue="2.86" pageId="29" pageNumber="30" unit="mm" value="2.86">2.86 mm</quantity>
); HL
<quantity id="4CA79B62FFCA18158C21389300A1FE9B" box="[840,1005,655,680]" metricMagnitude="-4" metricUnit="m" metricValue="5.4" metricValueMax="5.8" metricValueMin="5.0" pageId="29" pageNumber="30" unit="mm" value="0.54" valueMax="0.58" valueMin="0.5">0.500.58 mm</quantity>
(mean
<quantity id="4CA79B62FFCA18158B2A389307E1FE94" box="[1091,1197,655,680]" metricMagnitude="-4" metricUnit="m" metricValue="5.2" pageId="29" pageNumber="30" unit="mm" value="0.52">0.52 mm</quantity>
), HW
<quantity id="4CA79B62FFCA18158B91389306D1FE9B" box="[1272,1437,655,680]" metricMagnitude="-4" metricUnit="m" metricValue="5.9" metricValueMax="6.3" metricValueMin="5.5" pageId="29" pageNumber="30" unit="mm" value="0.59" valueMax="0.63" valueMin="0.55">0.550.63 mm</quantity>
(mean
<quantity id="4CA79B62FFCA18158F8F38A8021CFEF0" box="[230,336,692,717]" metricMagnitude="-4" metricUnit="m" metricValue="5.8" pageId="29" pageNumber="30" unit="mm" value="0.58">0.58 mm</quantity>
), AnL
<quantity id="4CA79B62FFCA18158ECB38A8010BFEF0" box="[418,583,692,717]" metricMagnitude="-3" metricUnit="m" metricValue="1.575" metricValueMax="1.75" metricValueMin="1.4" pageId="29" pageNumber="30" unit="mm" value="1.575" valueMax="1.75" valueMin="1.4">1.401.75 mm</quantity>
(mean
<quantity id="4CA79B62FFCA18158DF638A80045FEF0" box="[671,777,692,717]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="29" pageNumber="30" unit="mm" value="1.5">1.50 mm</quantity>
); PL
<quantity id="4CA79B62FFCA18158C2138A800A2FEF0" box="[840,1006,692,717]" metricMagnitude="-4" metricUnit="m" metricValue="7.9" metricValueMax="8.299999999999999" metricValueMin="7.5" pageId="29" pageNumber="30" unit="mm" value="0.79" valueMax="0.83" valueMin="0.75">0.750.83 mm</quantity>
(mean
<quantity id="4CA79B62FFCA18158B2C38A807E3FEF0" box="[1093,1199,692,716]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="29" pageNumber="30" unit="mm" value="0.8">0.80 mm</quantity>
), PW
<quantity id="4CA79B62FFCA18158B9138A806D1FEF0" box="[1272,1437,692,717]" metricMagnitude="-4" metricUnit="m" metricValue="5.9" metricValueMax="6.3" metricValueMin="5.5" pageId="29" pageNumber="30" unit="mm" value="0.59" valueMax="0.63" valueMin="0.55">0.550.63 mm</quantity>
(mean
<quantity id="4CA79B62FFCA18158F8F38CB0203FECC" box="[230,335,727,752]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="29" pageNumber="30" unit="mm" value="0.6">0.60 mm</quantity>
); EL
<quantity id="4CA79B62FFCA18158EF938CB0179FED3" box="[400,565,727,752]" metricMagnitude="-3" metricUnit="m" metricValue="1.54" metricValueMax="1.65" metricValueMin="1.43" pageId="29" pageNumber="30" unit="mm" value="1.54" valueMax="1.65" valueMin="1.43">1.431.65 mm</quantity>
(mean
<quantity id="4CA79B62FFCA18158DE538CB01BAFECC" box="[652,758,727,752]" metricMagnitude="-3" metricUnit="m" metricValue="1.54" pageId="29" pageNumber="30" unit="mm" value="1.54">1.54 mm</quantity>
), EW 1.00
<quantity id="4CA79B62FFCA18158C1638CB00A8FED3" box="[895,996,727,752]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" pageId="29" pageNumber="30" unit="mm" value="1.15">1.15 mm</quantity>
(mean
<quantity id="4CA79B62FFCA18158B5238C407E9FECC" box="[1083,1189,728,752]" metricMagnitude="-3" metricUnit="m" metricValue="1.07" pageId="29" pageNumber="30" unit="mm" value="1.07">1.07 mm</quantity>
), EI 1.331.53 (mean 1.44).
</paragraph>
</subSubSection>
<subSubSection id="C345650CFFCA18158FAE390306D1FF04" box="[199,1437,799,824]" pageId="29" pageNumber="30" type="distribution">
<paragraph id="8BE03687FFCA18158FAE390306D1FF04" blockId="29.[151,1437,151,1977]" box="[199,1437,799,824]" pageId="29" pageNumber="30">
<emphasis id="B92BEA95FFCA18158FAE39030215FF04" bold="true" box="[199,345,799,824]" pageId="29" pageNumber="30">Distribution</emphasis>
(
<figureCitation id="13642A02FFCA18158E0139030285FF04" box="[360,457,799,824]" captionStart="FIGURES 193 198" captionStartId="72.[151,264,1762,1785]" captionTargetBox="[195,1376,193,1707]" captionTargetId="figure@72.[191,1378,193,1739]" captionTargetPageId="72" captionText="FIGURES 193 198. Distribution of revised Australian species of Horaeomorphus Schaufuss (193), Leascydmus gen. n. (194), Scydmaenozila gen. n. (195), Spinosciacharis gen. n. (196), Sciacharoides gen. n. (197), and Sciacharis Broun. Horaeomorphus anthicoides (Lea) comb. n. (193 a), Horaeomorphus verus sp. n. (193 b), Leascydmus simplicicornis (Lea) comb. n. (194 a), Leascydmus armstrongi sp. n. (194 b), Scydmaenozila macrosticta (Lea) comb. n. (195 a), Scydmaenozila latipennis (Lea) comb. n. (195 b), Scydmaenozila mallacootana sp. n. (195 c), Scydmaenozila inexpectata sp. n. (195 d), Spinosciacharis puncticeps (Franz) comb. n. (196 a), Spinosciacharis sassy sp. n. (196 b), Spinosciacharis clydensis sp. n. (196 c), Spinosciacharis acheron sp. n. (196 d), Sciacharoides thompsoni (Franz) comb. n. (197 a), Sciacharoides eucalypti (Franz) comb. n. (197 b), Sciacharoides maipotonensis (Franz) comb. n. (197 c), Sciacharoides mantistamborinensis (Franz) comb. n. (197 d), and Sciacharis (Maorinus) tasmaniensis (Franz) comb. n. (198)." httpUri="https://zenodo.org/record/286555/files/figure.png" pageId="29" pageNumber="30">Fig. 195</figureCitation>
a). South-eastern
<collectingCountry id="F3487617FFCA18158DE2390301BAFF04" box="[651,758,799,824]" name="Australia" pageId="29" pageNumber="30">Australia</collectingCountry>
: South
<collectingCountry id="F3487617FFCA18158C20390300FFFF04" box="[841,947,799,824]" name="Australia" pageId="29" pageNumber="30">Australia</collectingCountry>
, Australian Capital Territory and Tasmania.
</paragraph>
</subSubSection>
<subSubSection id="C345650CFFCA18158FAE39580760FB85" pageId="29" pageNumber="30" type="discussion">
<paragraph id="8BE03687FFCA18158FAE39580247FA34" blockId="29.[151,1437,151,1977]" pageId="29" pageNumber="30">
<emphasis id="B92BEA95FFCA18158FAE39580277FF61" bold="true" box="[199,315,836,861]" pageId="29" pageNumber="30">Remarks.</emphasis>
<bibRefCitation id="EFCE4B76FFCA18158E2B39590298FF61" author="Franz" box="[322,468,836,861]" pageId="29" pageNumber="30" refString="Franz, H. (1975) Revision der Scydmaeniden von Australien, Neuseeland und den benachbarten Inseln. Denkschriften Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 118, 1 - 312." type="journal article" year="1975">Franz (1975)</bibRefCitation>
studied two specimens from SAM, and he treated one of them as a
<typeStatus id="54E48825FFCA18158BBD3958067AFF61" box="[1236,1334,836,861]" pageId="29" pageNumber="30" type="holotype">holotype</typeStatus>
(without specifying the sex), the other one as a male co-type.
<bibRefCitation id="EFCE4B76FFCA18158D8B39740011FFBC" author="Lea" box="[738,861,872,896]" pageId="29" pageNumber="30" refString="Lea, A. M. (1910) Australian and Tasmanian Coleoptera inhabiting or resorting to the nests of ants, bees, and termites. Proceedings of the Royal Society of Victoria, Melbourne (N. S.), 23, 116 - 230." type="journal article" year="1910">Lea (1910)</bibRefCitation>
clearly stated that two specimens were available to him: &quot;One specimen was taken from an ants' nest under a stone, a second under a near by stone but not associated with ants&quot;. These two specimens have a status of
<typeStatus id="54E48825FFCA18158DA239AC007CFFF4" box="[715,816,944,968]" pageId="29" pageNumber="30" type="syntype">syntypes</typeStatus>
, and Franz's (1975) redescription does not contain any data that fulfill the requirements of the formal
<typeStatus id="54E48825FFCA18158DBE39C80072FFD1" box="[727,830,980,1005]" pageId="29" pageNumber="30" type="lectotype">lectotype</typeStatus>
designation. The two specimens presumably were originally mounted on one card (a frequent case in the Lea Collection) and separated in a later time. The &quot;
<typeStatus id="54E48825FFCA18158A4539EB06DFF82C" box="[1324,1427,1015,1040]" pageId="29" pageNumber="30" type="holotype">holotype</typeStatus>
&quot; mentioned by Franz as a specimen bearing the original set of labels is a female, and the &quot;co-type&quot; without original labels is a male, distinctly smaller than the female, and with the aedeagus mounted by Franz in euparal (remounted in
<collectingCountry id="F3487617FFCA18158FDF3E780246F841" box="[182,266,1124,1149]" name="Canada" pageId="29" pageNumber="30">Canada</collectingCountry>
balsam during the present study) and illustrated (
<bibRefCitation id="EFCE4B76FFCA18158C203E78070DF841" author="Franz" box="[841,1089,1124,1149]" pageId="29" pageNumber="30" refString="Franz, H. (1975) Revision der Scydmaeniden von Australien, Neuseeland und den benachbarten Inseln. Denkschriften Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 118, 1 - 312." type="journal article" year="1975">
Franz 1975;
<figureCitation id="13642A02FFCA18158CB73E78070DF841" box="[990,1089,1124,1149]" captionStart="FIGURES 121 128" captionStartId="46.[151,264,1877,1900]" captionTargetBox="[168,1425,220,1825]" captionTargetId="figure@46.[151,1436,193,1856]" captionTargetPageId="46" captionText="FIGURES 121 128. Aedeagi of Spinosciacharis spp. in ventral (121, 123, 125, 127) and lateral (122, 124, 126, 128) views. Spinosciacharis puncticeps (Franz) comb. n. (121 122); Spinosciacharis sassy sp. n. (123 124), Spinosciacharis clydensis sp. n. (125 126), and Spinosciacharis acheron sp. n. (127 128)." httpUri="https://zenodo.org/record/286539/files/figure.png" pageId="29" pageNumber="30">Fig. 125</figureCitation>
</bibRefCitation>
). The aedeagus is erected and distorted (
<figureCitation id="13642A02FFCA18158E673E9B022BF89C" box="[270,359,1159,1184]" captionStart="FIGURES 80 88" captionStartId="34.[151,264,1601,1624]" captionTargetBox="[163,1426,499,1576]" captionTargetId="figure@34.[151,1436,481,1580]" captionTargetPageId="34" captionText="FIGURES 80 88. Aedeagi of Scydmaenozila spp. in ventral (80, 82, 83, 85, 87) and lateral (81, 84, 86, 88) views. Scydmaenozila macrosticta (Lea) comb. n., intact (80 81) and erected (82); Scydmaenozila latipennis (Lea) comb. n. (83 84), Scydmaenozila mallacootana sp. n. (85 86), and Scydmaenozila inexpectata sp. n. (87 88)." httpUri="https://zenodo.org/record/286531/files/figure.png" pageId="29" pageNumber="30">Fig. 82</figureCitation>
), but internal structures are highly similar to those found in all additional males from the Australian Capital Territory and Tasmania, and also in the
<typeStatus id="54E48825FFCA18158C3B3EB000F8F8F9" box="[850,948,1196,1221]" pageId="29" pageNumber="30" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C5F4D04FFCA18158C8A3EB00679F8F9" box="[995,1333,1196,1221]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="australiensis">
<emphasis id="B92BEA95FFCA18158C8A3EB00679F8F9" box="[995,1333,1196,1221]" italics="true" pageId="29" pageNumber="30">Horaeomorphus australiensis</emphasis>
</taxonomicName>
. The
<specimenCount id="9D59FD0EFFCA18158AE93EB00240F8D4" pageId="29" pageNumber="30" type="generic">25 specimens</specimenCount>
from Blundells Creek and 11 from various localities in Tasmania show a considerable variation in the body length and pigmentation, ranging from small (BL
<quantity id="4CA79B62FFCA18158C763EE800CBF930" box="[799,903,1268,1293]" metricMagnitude="-3" metricUnit="m" metricValue="2.63" pageId="29" pageNumber="30" unit="mm" value="2.63">2.63 mm</quantity>
) to large (BL
<quantity id="4CA79B62FFCA18158B5B3EE807D7F930" box="[1074,1179,1268,1292]" metricMagnitude="-3" metricUnit="m" metricValue="2.98" pageId="29" pageNumber="30" unit="mm" value="2.98">2.98 mm</quantity>
) individuals and from sharply contrasting light and dark elytral pattern to only indistinct (but still discernible) spots with diffused margins. The aedeagus of the
<typeStatus id="54E48825FFCA18158E933F200110F969" box="[506,604,1340,1365]" pageId="29" pageNumber="30" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C5F4D04FFCA18158DE03F210072F969" box="[649,830,1340,1365]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="australiensis">
<emphasis id="B92BEA95FFCA18158DE03F210072F969" box="[649,830,1340,1365]" italics="true" pageId="29" pageNumber="30">H. australiensis</emphasis>
</taxonomicName>
does not differ from those found in all males from Blundells Creek and Tasmania, and the specimen's external morphology falls within the range of interindividual variability of
<taxonomicName id="4C5F4D04FFCA18158E2A3F9902B1F9A0" box="[323,509,1413,1436]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="macrostictus">
<emphasis id="B92BEA95FFCA18158E2A3F9902B1F9A0" box="[323,509,1413,1436]" italics="true" pageId="29" pageNumber="30">H. macrostictus</emphasis>
</taxonomicName>
.
<taxonomicName id="4C5F4D04FFCA18158D7F3F980021F9A1" box="[534,877,1412,1437]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="australiensis">
<emphasis id="B92BEA95FFCA18158D7F3F980021F9A1" box="[534,877,1412,1437]" italics="true" pageId="29" pageNumber="30">Horaeomorphus australiensis</emphasis>
</taxonomicName>
is therefore placed as a junior synonym of
<taxonomicName id="4C5F4D04FFCA18158FFE3FBB0281F9FC" box="[151,461,1447,1472]" class="Insecta" family="Staphylinidae" genus="Scydmaenozila" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="macrosticta">
<emphasis id="B92BEA95FFCA18158FFE3FBB0281F9FC" box="[151,461,1447,1472]" italics="true" pageId="29" pageNumber="30">Scydmaenozila macrosticta</emphasis>
</taxonomicName>
(=
<taxonomicName id="4C5F4D04FFCA18158E953FBB0000F9FC" box="[508,844,1447,1472]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="macrostictus">
<emphasis id="B92BEA95FFCA18158E953FBB0000F9FC" box="[508,844,1447,1472]" italics="true" pageId="29" pageNumber="30">Horaeomorphus macrostictus</emphasis>
</taxonomicName>
). The male
<typeStatus id="54E48825FFCA18158CB63FB5077AF9FD" box="[991,1078,1449,1473]" pageId="29" pageNumber="30" type="syntype">syntype</typeStatus>
of
<taxonomicName id="4C5F4D04FFCA18158B0A3FBB06D0F9FC" box="[1123,1436,1447,1472]" class="Insecta" family="Staphylinidae" genus="Phagonophana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="macrosticta">
<emphasis id="B92BEA95FFCA18158B0A3FBB06D0F9FC" box="[1123,1436,1447,1472]" italics="true" pageId="29" pageNumber="30">Phagonophana macrosticta</emphasis>
</taxonomicName>
that does not bear the original labels (
<figureCitation id="13642A02FFCA18158D293FD001DEF9D9" box="[576,658,1484,1509]" captionStart="FIGURES 72 79" captionStartId="33.[151,264,1586,1609]" captionTargetBox="[155,1432,200,1548]" captionTargetId="figure@33.[151,1436,193,1565]" captionTargetPageId="33" captionText="FIGURES 72 79. Scydmaenozila spp. Dorsal habitus (72 75), original set of labels of lectotypes (76, 79), paralectotype (77), and mounting card with three syntypes (with reverse side in bottom) before re-mounting during the present study (78). Scydmaenozila macrosticta (Lea) comb. n. (72, 76 77); Scydmaenozila latipennis (Lea) comb. n. (73, 78 79), Scydmaenozila mallacootana sp. n. (74), and Scydmaenozila inexpectata sp. n. (75)." httpUri="https://zenodo.org/record/286530/files/figure.png" pageId="29" pageNumber="30">Fig. 76</figureCitation>
) is here designated as a
<typeStatus id="54E48825FFCA18158CCF3FD00741F9D9" box="[934,1037,1484,1509]" pageId="29" pageNumber="30" type="lectotype">lectotype</typeStatus>
to provide a name-bearing
<typeStatus id="54E48825FFCA18158A2D3FD10638F9D9" box="[1348,1396,1485,1509]" pageId="29" pageNumber="30">type</typeStatus>
for this name.
</paragraph>
<paragraph id="8BE03687FFCA18158FAE3C08001DFADC" blockId="29.[151,1437,151,1977]" pageId="29" pageNumber="30">
As mentioned above, this species is relatively variable in the body length, proportions of body parts and the elytral pattern. Specimens from Tasmania and
<collectingCountry id="F3487617FFCA18158DDA3C2B0055FA6C" box="[691,793,1591,1616]" name="Australia" pageId="29" pageNumber="30">Australia</collectingCountry>
are similarly variable and do not differ in the range of variability. However, since these populations are isolated, an analysis of larger samples may reveal some subtle differences. The elytral pattern, although variably distinct, is observable even in the most darkened specimens, in which the contrast between lighter and darker spots is low. In all remaining species of
<taxonomicName id="4C5F4D04FFCA18158B013CB8065CFA81" box="[1128,1296,1700,1725]" class="Insecta" family="Staphylinidae" genus="Scydmaenozila" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="73" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B92BEA95FFCA18158B013CB8065CFA81" box="[1128,1296,1700,1725]" italics="true" pageId="29" pageNumber="30">Scydmaenozila</emphasis>
</taxonomicName>
no traces of elytral pattern can be seen and the elytra are uniformly brown.
</paragraph>
<paragraph id="8BE03687FFCA18158FAE3CF10760FB85" blockId="29.[151,1437,151,1977]" pageId="29" pageNumber="30">
Two specimens from NHM (the
<typeStatus id="54E48825FFCA18158D593CF1012CFB39" box="[560,608,1773,1797]" pageId="29" pageNumber="30">type</typeStatus>
series of
<taxonomicName id="4C5F4D04FFCA18158DA23CF10030FB39" box="[715,892,1772,1797]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="australiensis">
<emphasis id="B92BEA95FFCA18158DA23CF10030FB39" box="[715,892,1772,1797]" italics="true" pageId="29" pageNumber="30">H. australiensis</emphasis>
</taxonomicName>
) bear labels with &quot;S
<collectingCountry id="F3487617FFCA18158B0D3CF00782FB39" box="[1124,1230,1772,1797]" name="Australia" pageId="29" pageNumber="30">Australia</collectingCountry>
, Bakewell&quot;. There is Bakewell Bore in South
<collectingCountry id="F3487617FFCA18158EA53D13017FFB14" box="[460,563,1807,1832]" name="Australia" pageId="29" pageNumber="30">Australia</collectingCountry>
(over
<quantity id="4CA79B62FFCA18158D163D13019AFB14" box="[639,726,1807,1832]" metricMagnitude="5" metricUnit="m" metricValue="6.3" pageId="29" pageNumber="30" unit="km" value="630.0">630 km</quantity>
NW of Adelaide), but Bakewell on the labels is most likely a person name (Adam Ślipiński, in an email to the author dated
<date id="FFE11047FFCA18158CF63D28076FFB71" box="[927,1059,1844,1869]" pageId="29" pageNumber="30" value="2014-06-03">06.03.2014</date>
). The exact collecting place of
<taxonomicName id="4C5F4D04FFCA18158FFE3D4B029DFB4C" box="[151,465,1879,1904]" class="Insecta" family="Staphylinidae" genus="Scydmaenozila" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="macrosticta">
<emphasis id="B92BEA95FFCA18158FFE3D4B029DFB4C" box="[151,465,1879,1904]" italics="true" pageId="29" pageNumber="30">Scydmaenozila macrosticta</emphasis>
</taxonomicName>
in South
<collectingCountry id="F3487617FFCA18158D3E3D4B01F1FB4C" box="[599,701,1879,1904]" name="Australia" pageId="29" pageNumber="30">Australia</collectingCountry>
remains unknown, but judging from the distribution in the Australian Capital Territory and in Tasmania (where it is known from scattered localities from nearly entire island), this species may occur in SE part of South
<collectingCountry id="F3487617FFCA18158D113DBC0193FB85" box="[632,735,1952,1977]" name="Australia" pageId="29" pageNumber="30">Australia</collectingCountry>
and possibly also in Victoria.
</paragraph>
</subSubSection>
</treatment>
</document>