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<document id="B88A390A775D04741757B8C8E63CBC58" ID="10.11646/zootaxa.4136.1.4" ID-DOI="10.11646/zootaxa.4136.1.4" ID-GBIF-Dataset="d620461c-f45a-4ea5-800c-ed6298e631bd" ID-ISSN="1175-5326" ID-Zenodo-Dep="261786" ID-ZooBank="BBC55637-6919-43B3-BB85-83E2AFFE2CED" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1467726852934" checkinUser="plazi" docAuthor="Stonis, Jonas R., Diškus, Arūnas, Remeikis, Andrius, Davis, Donald R., Solis, M. Alma &amp; Torres, Nixon Cumbicus" docDate="2016" docId="03B47A7DFFF0A123D7C143B8A8A4F864" docLanguage="en" docName="zootaxa.4136.1.4.pdf" docOrigin="Zootaxa 4136 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Stigmella tripartita Diskus &amp; Stonis, sp. nov." docType="treatment" docVersion="7" lastPageNumber="110" masterDocId="FF8D0205FFF9A12AD7564160A92DFFBF" masterDocTitle="The first record of Baccharis L. (Asteraceae) as a host-plant genus for Nepticulidae (Lepidoptera), with description of new Stigmella species from South America" masterLastPageNumber="128" masterPageNumber="101" pageNumber="110" updateTime="1698363551601" updateUser="plazi">
<mods:mods id="5E2B0E58E8247147C981F187C2F72310" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="0E488D2F6CC067B8AF2413F78C9D2952">
<mods:title id="C6E320C0A4ACC2473D55AA313C6D7E9B">The first record of Baccharis L. (Asteraceae) as a host-plant genus for Nepticulidae (Lepidoptera), with description of new Stigmella species from South America</mods:title>
</mods:titleInfo>
<mods:name id="087D183974793533C7DF655C651001AD" type="personal">
<mods:role id="0FF3587CC805146C116506722758A505">
<mods:roleTerm id="B0826A058384C9E175C1FC28BA73E86E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2E6492F5F9679A615CB73306F43EE833">Stonis, Jonas R.</mods:namePart>
</mods:name>
<mods:name id="4FD4F97532761D998C5D650A69A0EC79" type="personal">
<mods:role id="17585ED0F4941BECC3C3CE5B121655B8">
<mods:roleTerm id="3FA7E01CBC0D5B31BFC78CE9D4D8BD68">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9E8D4D36CA6441876D221909094A2445">Diškus, Arūnas</mods:namePart>
</mods:name>
<mods:name id="0994136C04F81BD5940A3559BF9A11D2" type="personal">
<mods:role id="346F6D279D0643A6BB0E6C8899B19AFA">
<mods:roleTerm id="4653846361D0567557F1B3CF17F55FED">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="90C479CC2CF62E99A6B7112B806FB3A1">Remeikis, Andrius</mods:namePart>
</mods:name>
<mods:name id="E78DCEC810495461A82634ABF24C4997" type="personal">
<mods:role id="96DF84D0CBA32C1BDA6BCC66CADDF8C5">
<mods:roleTerm id="D9C660DB98903C0737A7FAEF3C8A5CC6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="007B25B988CB837DE794C04CDB5360B4">Davis, Donald R.</mods:namePart>
</mods:name>
<mods:name id="2B50449D8BE402CBDA8A55CFEDD20952" type="personal">
<mods:role id="9C17774BAF8E41793588755D84EF265E">
<mods:roleTerm id="2560F38A5563D714101A44F5B79AED34">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2BEE6BE8E82FB8A6862F278EE350E123">Solis, M. Alma</mods:namePart>
</mods:name>
<mods:name id="042ADDE11201536E894504937D40D817" type="personal">
<mods:role id="328841FB93904B952D7AAF9DD3425C77">
<mods:roleTerm id="3B1B8E6EA4C10BDB685FFA3417112759">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="39D421613559318E58B0A1A63160F075">Torres, Nixon Cumbicus</mods:namePart>
</mods:name>
<mods:typeOfResource id="722093D77C1A463D3739BB5425C65B6A">text</mods:typeOfResource>
<mods:relatedItem id="61E22AD801E026F1029279F743A65C4F" type="host">
<mods:titleInfo id="A906986EBD8DEEF0D08926B9FB720793">
<mods:title id="44238278E4B53A3F714BF131F30163AB">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="2DE95185CC791A7EB9AFC993DEE37D1E">
<mods:date id="784D1E2194F126C05B3BD0A7B9C7AFD8">2016</mods:date>
<mods:detail id="53BC2F4B611AA57EA9C0095D2E7DD2E3" type="volume">
<mods:number id="47C361028C2DEDD30F97E0036C27B5ED">4136</mods:number>
</mods:detail>
<mods:detail id="15E4F626A4C1C3913AF4CB8427852A38" type="issue">
<mods:number id="5CA0CD0EF5379FE23FD3C52D3F95A2D3">1</mods:number>
</mods:detail>
<mods:extent id="15E1AA95083D8E89035E93D456DE524B" unit="page">
<mods:start id="794A5386E77285F446EDEF79B61E8E24">101</mods:start>
<mods:end id="A017789AC979E6AF91F752A3F83D2AD3">128</mods:end>
</mods:extent>
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<mods:classification id="859F0D8083F41CEB0B83E71952B574FB">journal article</mods:classification>
<mods:identifier id="A49B7F9C5D50352F0D085BC5C8D26734" type="DOI">10.11646/zootaxa.4136.1.4</mods:identifier>
<mods:identifier id="7EB7656EC94B935FD9C099CF3B79BD9F" type="GBIF-Dataset">d620461c-f45a-4ea5-800c-ed6298e631bd</mods:identifier>
<mods:identifier id="D126ABDA3293E1AC6EDA3F44126192FD" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="1634FAB63303BB23D21C7DE42EE4451C" type="Zenodo-Dep">261786</mods:identifier>
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<treatment id="03B47A7DFFF0A123D7C143B8A8A4F864" ID-DOI="http://doi.org/10.5281/zenodo.6070507" ID-GBIF-Taxon="121269110" ID-Zenodo-Dep="6070507" LSID="urn:lsid:plazi:treatment:03B47A7DFFF0A123D7C143B8A8A4F864" httpUri="http://treatment.plazi.org/id/03B47A7DFFF0A123D7C143B8A8A4F864" lastPageNumber="110" pageId="9" pageNumber="110">
<subSubSection id="C30798E0FFF0A123D7C143B8ABFFFD4D" box="[151,722,727,754]" pageId="9" pageNumber="110" type="nomenclature">
<paragraph id="8BA2CB6BFFF0A123D7C143B8ABFFFD4D" blockId="9.[151,722,727,787]" box="[151,722,727,754]" pageId="9" pageNumber="110">
<heading id="D0EA7C07FFF0A123D7C143B8ABFFFD4D" bold="true" box="[151,722,727,754]" fontSize="11" level="1" pageId="9" pageNumber="110" reason="1">
<emphasis id="B9691779FFF0A123D7C143B8ABFFFD4D" bold="true" box="[151,722,727,754]" pageId="9" pageNumber="110">
3.
<taxonomicName id="4C1DB0E8FFF0A123D7E243B7AB4AFD4D" ID-CoL="6ZSRH" authority="Diskus &amp; Stonis" authorityName="Diskus &amp; Stonis" box="[180,615,727,754]" class="Insecta" family="Nepticulidae" genus="Stigmella" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="110" phylum="Arthropoda" rank="species" species="tripartita" status="sp. nov.">
<emphasis id="B9691779FFF0A123D7E243B7A8BAFD4E" bold="true" box="[180,407,727,753]" italics="true" pageId="9" pageNumber="110">Stigmella tripartita</emphasis>
Diškus &amp; Stonis
</taxonomicName>
,
<taxonomicNameLabel id="A25AAA02FFF0A123D52243B8ABFFFD4D" box="[628,722,728,754]" pageId="9" pageNumber="110" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30798E0FFF0A123D7C1439AA867FCAC" box="[151,330,762,787]" pageId="9" pageNumber="110" type="description">
<paragraph id="8BA2CB6BFFF0A123D7C1439AA867FCAC" blockId="9.[151,722,727,787]" box="[151,330,762,787]" pageId="9" pageNumber="110">
(
<figureCitation id="1326D7EEFFF0A123D7C9439AA9C0FCAC" box="[159,237,762,787]" captionStart="FIGURE 3" captionStartId="2.[151,250,1980,2002]" captionTargetBox="[151,1436,193,1956]" captionTargetId="figure@2.[151,1436,193,1959]" captionTargetPageId="2" captionText="FIGURE 3. Current records of Stigmella species feeding on plants from the genus Baccharis L." httpUri="https://zenodo.org/record/261788/files/figure.png" pageId="9" pageNumber="110">Figs. 3</figureCitation>
,
<figureCitation id="1326D7EEFFF0A123D7AC439BA86CFCAC" box="[250,321,763,787]" captionStart-0="FIGURES 33 – 38" captionStart-1="FIGURES 39 – 43" captionStartId-0="10.[151,264,1551,1574]" captionStartId-1="11.[151,264,1896,1919]" captionTargetBox-0="[151,1425,195,1520]" captionTargetBox-1="[190,1429,384,1865]" captionTargetId-0="figure@10.[151,1436,195,1530]" captionTargetId-1="figure@11.[151,1436,373,1876]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionText-0="FIGURES 33 – 38. Stigmella tripartita Diškus &amp; Stonis, sp. nov. 33, habitat, SW of Cajabamba, Chimborazo Province, Ecuador, the equatorial Andes, elevation 3870 m; 34, 36, host-plant Baccharis buxifolia (Lam.) Pers.; 35, usual habitat of the species occurrence along roadsides; 37, cocoon; 38, forewing (reconstructed)." captionText-1="FIGURES 39 – 43. Male genitalia of Stigmella tripartita Diškus &amp; Stonis, sp. nov. (ZMUC). 39, capsule (without phallus), ventral view, holotype, genitalia slide no. AD 622; 40, phallus, paratype, genitalia slide no. AD 624; 41, capsule, dorsal view, holotype, genitalia slide no. AD 622; 42, same, phallus; 43, capsule with phallus, paratype, genitalia slide no. AD 624." httpUri-0="https://zenodo.org/record/261794/files/figure.png" httpUri-1="https://zenodo.org/record/261795/files/figure.png" pageId="9" pageNumber="110">33–43</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30798E0FFF0A123D7C14222AA12FC78" pageId="9" pageNumber="110" type="materials_examined">
<paragraph id="8BA2CB6BFFF0A123D7C14222AA12FC78" blockId="9.[151,1437,834,2011]" pageId="9" pageNumber="110">
<emphasis id="B9691779FFF0A123D7C14222A86BFCE4" bold="true" box="[151,326,834,859]" pageId="9" pageNumber="110">
<typeStatus id="54A675C9FFF0A123D7C14222A9FFFCE4" box="[151,210,834,859]" pageId="9" pageNumber="110">Type</typeStatus>
material
</emphasis>
.
<typeStatus id="54A675C9FFF0A123D60A4222A8E4FCE4" box="[348,457,834,859]" pageId="9" pageNumber="110" type="holotype">Holotype</typeStatus>
: ♂ (see Remarks),
<collectingCountry id="F30A8BFBFFF0A123D5964223AA61FCE4" box="[704,844,835,859]" name="Ecuador" pageId="9" pageNumber="110">ECUADOR</collectingCountry>
:
<quantity id="4CE5668EFFF0A123D4094222AA82FCE4" box="[863,943,834,859]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="110" unit="km" value="15.0">15 km</quantity>
SW Cajabamba (Chimborazo Province),
<geoCoordinate id="EE29ADACFFF0A123D7C14206A827FCC0" box="[151,266,870,895]" direction="south" orientation="latitude" pageId="9" pageNumber="110" precision="15" value="-1.7613889">1°45'41&quot;S</geoCoordinate>
,
<geoCoordinate id="EE29ADACFFF0A123D6434206A88DFCC0" box="[277,416,870,895]" direction="west" orientation="longitude" pageId="9" pageNumber="110" precision="15" value="-78.81417">78°48'51&quot;W</geoCoordinate>
, elevation
<quantity id="4CE5668EFFF0A123D5414207AB41FCC1" box="[535,620,871,895]" metricMagnitude="3" metricUnit="m" metricValue="3.87" pageId="9" pageNumber="110" unit="m" value="3870.0">3870 m</quantity>
a.s.l., mining larvae on
<taxonomicName id="4C1DB0E8FFF0A123D4204206ADCEFCC1" authority="(Lam.) Pers." authorityName="(Lam.) Pers." box="[886,1251,870,895]" class="Magnoliopsida" family="Asteraceae" genus="Baccharis" kingdom="Plantae" order="Asterales" pageId="9" pageNumber="110" phylum="Tracheophyta" rank="species" species="buxifolia">
<emphasis id="B9691779FFF0A123D4204206AD7FFCC0" box="[886,1106,870,895]" italics="true" pageId="9" pageNumber="110">Baccharis buxifolia</emphasis>
(Lam.)
<collectingCountry id="F30A8BFBFFF0A123D3FA4207ADCDFCC1" box="[1196,1248,871,894]" name="Iran" pageId="9" pageNumber="110">Pers</collectingCountry>
.
</taxonomicName>
<date id="FFA3EDABFFF0A123D3BF4206AC73FCC0" box="[1257,1374,870,895]" pageId="9" pageNumber="110" value="2007-02-19">19.ii.2007</date>
, field card no. 4874, leg. A. Diškus, J. R. Stonis, genitalia slide nos AD
<specimenCount id="9D1B00E2FFF0A123D42442EAAA9DFC1C" box="[882,944,906,931]" pageId="9" pageNumber="110" type="generic">622♂</specimenCount>
(
<collectionCode id="ED0C53AEFFF0A123D4E842EBAD38FC1C" box="[958,1045,907,931]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="9" pageNumber="110">ZMUC</collectionCode>
).
<typeStatus id="54A675C9FFF0A123D37E42EBADA2FC1C" box="[1064,1167,907,931]" pageId="9" pageNumber="110" type="paratype">Paratype</typeStatus>
: ♂ (see Remarks), same label data as
<typeStatus id="54A675C9FFF0A123D67E42CEA8A0FC78" box="[296,397,942,967]" pageId="9" pageNumber="110" type="holotype">holotype</typeStatus>
, genitalia slide no. AD
<specimenCount id="9D1B00E2FFF0A123D5D942CEABE0FC78" box="[655,717,942,967]" pageId="9" pageNumber="110" type="generic">624♂</specimenCount>
(
<collectionCode id="ED0C53AEFFF0A123D58A42CFAA1EFC78" box="[732,819,943,967]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="9" pageNumber="110">ZMUC</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C30798E0FFF0A123D79142B2ADEEFBB0" pageId="9" pageNumber="110" type="diagnosis">
<paragraph id="8BA2CB6BFFF0A123D79142B2ADEEFBB0" blockId="9.[151,1437,834,2011]" pageId="9" pageNumber="110">
<emphasis id="B9691779FFF0A123D79142B2A86CFC54" bold="true" box="[199,321,978,1003]" pageId="9" pageNumber="110">Diagnosis.</emphasis>
In the male genitalia, the set of three very large cornuti distinguishes
<taxonomicName id="4C1DB0E8FFF0A123D30342B3ADF4FC54" box="[1109,1241,979,1003]" class="Insecta" family="Nepticulidae" genus="Stigmella" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="110" phylum="Arthropoda" rank="species" species="tripartita" status="sp. nov.">
<emphasis id="B9691779FFF0A123D30342B3ADF4FC54" box="[1109,1241,979,1003]" italics="true" pageId="9" pageNumber="110">S. tripartita</emphasis>
</taxonomicName>
<emphasis id="B9691779FFF0A123D3B642B2AC16FC54" bold="true" box="[1248,1339,978,1003]" pageId="9" pageNumber="110">
<taxonomicNameLabel id="A25AAA02FFF0A123D3B642B2AC16FC54" box="[1248,1339,978,1003]" pageId="9" pageNumber="110" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from all other
<taxonomicName id="4C1DB0E8FFF0A123D7814296A86DFBB0" box="[215,320,1014,1039]" class="Insecta" family="Nepticulidae" genus="Stigmella" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="110" phylum="Arthropoda" rank="genus">
<emphasis id="B9691779FFF0A123D7814296A86DFBB0" box="[215,320,1014,1039]" italics="true" pageId="9" pageNumber="110">Stigmella</emphasis>
</taxonomicName>
species; the host-plant (
<taxonomicName id="4C1DB0E8FFF0A123D5064296AA01FBB0" box="[592,812,1014,1039]" class="Magnoliopsida" family="Asteraceae" genus="Baccharis" kingdom="Plantae" order="Asterales" pageId="9" pageNumber="110" phylum="Tracheophyta" rank="species" species="buxifolia">
<emphasis id="B9691779FFF0A123D5064296AA01FBB0" box="[592,812,1014,1039]" italics="true" pageId="9" pageNumber="110">Baccharis buxifolia</emphasis>
</taxonomicName>
) also makes this species distinctive.
</paragraph>
</subSubSection>
<subSubSection id="C30798E0FFF0A123D791457AA846F960" pageId="9" pageNumber="110" type="description">
<paragraph id="8BA2CB6BFFF0A123D791457AAAFCFB7C" blockId="9.[151,1437,834,2011]" pageId="9" pageNumber="110">
<emphasis id="B9691779FFF0A123D791457AA829FB8C" bold="true" box="[199,260,1050,1075]" pageId="9" pageNumber="110">Male</emphasis>
(
<figureCitation id="1326D7EEFFF0A123D645457AA84AFB8C" box="[275,359,1050,1075]" captionStart="FIGURES 33 – 38" captionStartId="10.[151,264,1551,1574]" captionTargetBox="[151,1425,195,1520]" captionTargetId="figure@10.[151,1436,195,1530]" captionTargetPageId="10" captionText="FIGURES 33 – 38. Stigmella tripartita Diškus &amp; Stonis, sp. nov. 33, habitat, SW of Cajabamba, Chimborazo Province, Ecuador, the equatorial Andes, elevation 3870 m; 34, 36, host-plant Baccharis buxifolia (Lam.) Pers.; 35, usual habitat of the species occurrence along roadsides; 37, cocoon; 38, forewing (reconstructed)." httpUri="https://zenodo.org/record/261794/files/figure.png" pageId="9" pageNumber="110">Fig. 38</figureCitation>
). Forewing length about 3.0 mm; wingspan about 6.5 mm (n=2). Head: palpi cream; frontal tuft brown to brown cream; scape cream to yellowish cream; antenna longer than half the length of forewing; flagellum with approx. 26 segments, grey.Thorax and tegula dark grey-brown to fuscous. Forewing dark brown-grey to fuscous, with slightly postmedian fascia; fascia broad to narrow, whitish grey or grey cream; terminal and tornal cilia pale grey-brown. Abdomen dark grey to fuscous (also see Remarks).
</paragraph>
<paragraph id="8BA2CB6BFFF0A123D79145AEA88FFB58" blockId="9.[151,1437,834,2011]" box="[199,418,1230,1255]" pageId="9" pageNumber="110">
<emphasis id="B9691779FFF0A123D79145AEA808FB58" bold="true" box="[199,293,1230,1255]" pageId="9" pageNumber="110">Female.</emphasis>
Unknown.
</paragraph>
<paragraph id="8BA2CB6BFFF0A123D7914592AABDF994" blockId="9.[151,1437,834,2011]" pageId="9" pageNumber="110">
<emphasis id="B9691779FFF0A123D7914592A85DFAB4" bold="true" box="[199,368,1266,1291]" pageId="9" pageNumber="110">Male genitalia</emphasis>
(
<figureCitation id="1326D7EEFFF0A123D6294592AB2FFAB4" box="[383,514,1266,1291]" captionStart="FIGURES 39 – 43" captionStartId="11.[151,264,1896,1919]" captionTargetBox="[190,1429,384,1865]" captionTargetId="figure@11.[151,1436,373,1876]" captionTargetPageId="11" captionText="FIGURES 39 – 43. Male genitalia of Stigmella tripartita Diškus &amp; Stonis, sp. nov. (ZMUC). 39, capsule (without phallus), ventral view, holotype, genitalia slide no. AD 622; 40, phallus, paratype, genitalia slide no. AD 624; 41, capsule, dorsal view, holotype, genitalia slide no. AD 622; 42, same, phallus; 43, capsule with phallus, paratype, genitalia slide no. AD 624." httpUri="https://zenodo.org/record/261795/files/figure.png" pageId="9" pageNumber="110">Figs 39–43</figureCitation>
). Capsule short (230–295 µm) but broad (235–240 µm). Vinculum with very small (25–30 µm), triangularly shaped lateral (anterior) lobes; ventral plate of vinculum short. Uncus 85–90 µm broad, with two small lobes, each bearing one (
<figureCitation id="1326D7EEFFF0A123D5D7445AABF5FAEC" box="[641,728,1338,1363]" captionStart="FIGURES 39 – 43" captionStartId="11.[151,264,1896,1919]" captionTargetBox="[190,1429,384,1865]" captionTargetId="figure@11.[151,1436,373,1876]" captionTargetPageId="11" captionText="FIGURES 39 – 43. Male genitalia of Stigmella tripartita Diškus &amp; Stonis, sp. nov. (ZMUC). 39, capsule (without phallus), ventral view, holotype, genitalia slide no. AD 622; 40, phallus, paratype, genitalia slide no. AD 624; 41, capsule, dorsal view, holotype, genitalia slide no. AD 622; 42, same, phallus; 43, capsule with phallus, paratype, genitalia slide no. AD 624." httpUri="https://zenodo.org/record/261795/files/figure.png" pageId="9" pageNumber="110">Fig. 41</figureCitation>
) or two papillae. Gnathos stout, with two caudal processes broadening basally (
<figureCitation id="1326D7EEFFF0A123D62A443EA8FDFAC8" box="[380,464,1374,1399]" captionStart="FIGURES 39 – 43" captionStartId="11.[151,264,1896,1919]" captionTargetBox="[190,1429,384,1865]" captionTargetId="figure@11.[151,1436,373,1876]" captionTargetPageId="11" captionText="FIGURES 39 – 43. Male genitalia of Stigmella tripartita Diškus &amp; Stonis, sp. nov. (ZMUC). 39, capsule (without phallus), ventral view, holotype, genitalia slide no. AD 622; 40, phallus, paratype, genitalia slide no. AD 624; 41, capsule, dorsal view, holotype, genitalia slide no. AD 622; 42, same, phallus; 43, capsule with phallus, paratype, genitalia slide no. AD 624." httpUri="https://zenodo.org/record/261795/files/figure.png" pageId="9" pageNumber="110">Fig. 41</figureCitation>
); central plate of gnathos large to very large, 85 µm broad, in the form of a trapezium (
<figureCitation id="1326D7EEFFF0A123D7C944E2A9DBFA24" box="[159,246,1410,1435]" captionStart="FIGURES 39 – 43" captionStartId="11.[151,264,1896,1919]" captionTargetBox="[190,1429,384,1865]" captionTargetId="figure@11.[151,1436,373,1876]" captionTargetPageId="11" captionText="FIGURES 39 – 43. Male genitalia of Stigmella tripartita Diškus &amp; Stonis, sp. nov. (ZMUC). 39, capsule (without phallus), ventral view, holotype, genitalia slide no. AD 622; 40, phallus, paratype, genitalia slide no. AD 624; 41, capsule, dorsal view, holotype, genitalia slide no. AD 622; 42, same, phallus; 43, capsule with phallus, paratype, genitalia slide no. AD 624." httpUri="https://zenodo.org/record/261795/files/figure.png" pageId="9" pageNumber="110">Fig. 41</figureCitation>
). Valva (
<figureCitation id="1326D7EEFFF0A123D63644E2A899FA24" box="[352,436,1410,1435]" captionStart="FIGURES 39 – 43" captionStartId="11.[151,264,1896,1919]" captionTargetBox="[190,1429,384,1865]" captionTargetId="figure@11.[151,1436,373,1876]" captionTargetPageId="11" captionText="FIGURES 39 – 43. Male genitalia of Stigmella tripartita Diškus &amp; Stonis, sp. nov. (ZMUC). 39, capsule (without phallus), ventral view, holotype, genitalia slide no. AD 622; 40, phallus, paratype, genitalia slide no. AD 624; 41, capsule, dorsal view, holotype, genitalia slide no. AD 622; 42, same, phallus; 43, capsule with phallus, paratype, genitalia slide no. AD 624." httpUri="https://zenodo.org/record/261795/files/figure.png" pageId="9" pageNumber="110">Fig. 39</figureCitation>
) 180 µm long, 80–85 µm broad, with two apical processes (ventral process short and pointed); inner lobe almost straight or slightly bulged; transtilla with rounded sublateral processes or without processes, rounded at corners (
<figureCitation id="1326D7EEFFF0A123D6BB44AAAB42FA5C" box="[493,623,1482,1507]" captionStart="FIGURES 39 – 43" captionStartId="11.[151,264,1896,1919]" captionTargetBox="[190,1429,384,1865]" captionTargetId="figure@11.[151,1436,373,1876]" captionTargetPageId="11" captionText="FIGURES 39 – 43. Male genitalia of Stigmella tripartita Diškus &amp; Stonis, sp. nov. (ZMUC). 39, capsule (without phallus), ventral view, holotype, genitalia slide no. AD 622; 40, phallus, paratype, genitalia slide no. AD 624; 41, capsule, dorsal view, holotype, genitalia slide no. AD 622; 42, same, phallus; 43, capsule with phallus, paratype, genitalia slide no. AD 624." httpUri="https://zenodo.org/record/261795/files/figure.png" pageId="9" pageNumber="110">Figs 41, 43</figureCitation>
). Juxta membranous, distinctly triangular, broadening caudally (
<figureCitation id="1326D7EEFFF0A123D26A44AAACBDFA5C" box="[1340,1424,1482,1507]" captionStart="FIGURES 39 – 43" captionStartId="11.[151,264,1896,1919]" captionTargetBox="[190,1429,384,1865]" captionTargetId="figure@11.[151,1436,373,1876]" captionTargetPageId="11" captionText="FIGURES 39 – 43. Male genitalia of Stigmella tripartita Diškus &amp; Stonis, sp. nov. (ZMUC). 39, capsule (without phallus), ventral view, holotype, genitalia slide no. AD 622; 40, phallus, paratype, genitalia slide no. AD 624; 41, capsule, dorsal view, holotype, genitalia slide no. AD 622; 42, same, phallus; 43, capsule with phallus, paratype, genitalia slide no. AD 624." httpUri="https://zenodo.org/record/261795/files/figure.png" pageId="9" pageNumber="110">Fig. 39</figureCitation>
). Phallus (
<figureCitation id="1326D7EEFFF0A123D7AF448EA889F9B8" box="[249,420,1518,1543]" captionStart="FIGURES 39 – 43" captionStartId="11.[151,264,1896,1919]" captionTargetBox="[190,1429,384,1865]" captionTargetId="figure@11.[151,1436,373,1876]" captionTargetPageId="11" captionText="FIGURES 39 – 43. Male genitalia of Stigmella tripartita Diškus &amp; Stonis, sp. nov. (ZMUC). 39, capsule (without phallus), ventral view, holotype, genitalia slide no. AD 622; 40, phallus, paratype, genitalia slide no. AD 624; 41, capsule, dorsal view, holotype, genitalia slide no. AD 622; 42, same, phallus; 43, capsule with phallus, paratype, genitalia slide no. AD 624." httpUri="https://zenodo.org/record/261795/files/figure.png" pageId="9" pageNumber="110">Figs 40, 42, 43</figureCitation>
) very short (145–200 µm) but broad (120 µm); vesica with a set of three very large (up to 100 µm long) cornuti (
<figureCitation id="1326D7EEFFF0A123D6C24772A8C4F994" box="[404,489,1554,1579]" captionStart="FIGURES 39 – 43" captionStartId="11.[151,264,1896,1919]" captionTargetBox="[190,1429,384,1865]" captionTargetId="figure@11.[151,1436,373,1876]" captionTargetPageId="11" captionText="FIGURES 39 – 43. Male genitalia of Stigmella tripartita Diškus &amp; Stonis, sp. nov. (ZMUC). 39, capsule (without phallus), ventral view, holotype, genitalia slide no. AD 622; 40, phallus, paratype, genitalia slide no. AD 624; 41, capsule, dorsal view, holotype, genitalia slide no. AD 622; 42, same, phallus; 43, capsule with phallus, paratype, genitalia slide no. AD 624." httpUri="https://zenodo.org/record/261795/files/figure.png" pageId="9" pageNumber="110">Fig. 40</figureCitation>
); no small spinose cornuti developed.
</paragraph>
<paragraph id="8BA2CB6BFFF0A123D7914756A846F960" blockId="9.[151,1437,834,2011]" pageId="9" pageNumber="110">
<emphasis id="B9691779FFF0A123D7914756A867F9F0" bold="true" box="[199,330,1590,1615]" pageId="9" pageNumber="110">Bionomics.</emphasis>
Larvae mine in leaves. Host-plant:
<taxonomicName id="4C1DB0E8FFF0A123D5BC4756ADDBF9F0" authority="(Lam.) Pers." authorityName="(Lam.) Pers." box="[746,1270,1590,1615]" class="Magnoliopsida" family="Asteraceae" genus="Baccharis" kingdom="Plantae" order="Asterales" pageId="9" pageNumber="110" phylum="Tracheophyta" rank="species" species="buxifolia">
<emphasis id="B9691779FFF0A123D5BC4756AAE4F9F0" box="[746,969,1590,1615]" italics="true" pageId="9" pageNumber="110">Baccharis buxifolia</emphasis>
(Lam.)
<collectingCountry id="F30A8BFBFFF0A123D37E4757AD71F9F1" box="[1064,1116,1591,1614]" name="Iran" pageId="9" pageNumber="110">Pers</collectingCountry>
. (Asteraceae)
</taxonomicName>
(
<figureCitation id="1326D7EEFFF0A123D25E4756ACBDF9F0" box="[1288,1424,1590,1615]" captionStart="FIGURES 33 – 38" captionStartId="10.[151,264,1551,1574]" captionTargetBox="[151,1425,195,1520]" captionTargetId="figure@10.[151,1436,195,1530]" captionTargetPageId="10" captionText="FIGURES 33 – 38. Stigmella tripartita Diškus &amp; Stonis, sp. nov. 33, habitat, SW of Cajabamba, Chimborazo Province, Ecuador, the equatorial Andes, elevation 3870 m; 34, 36, host-plant Baccharis buxifolia (Lam.) Pers.; 35, usual habitat of the species occurrence along roadsides; 37, cocoon; 38, forewing (reconstructed)." httpUri="https://zenodo.org/record/261794/files/figure.png" pageId="9" pageNumber="110">Figs 34, 36</figureCitation>
). Larvae yellow to pale yellow, and mine in February. Contorted gallery of mine with central line of dark brown to black frass (approximately resembling the mines of the European
<taxonomicName id="4C1DB0E8FFF0A123D42C471FAD62F928" authority="Stt." authorityName="Stt." box="[890,1103,1662,1687]" class="Insecta" family="Nepticulidae" genus="Stigmella" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="110" phylum="Arthropoda" rank="species" species="myrtilella">
<emphasis id="B9691779FFF0A123D42C471FAD2EF928" box="[890,1027,1662,1687]" italics="true" pageId="9" pageNumber="110">S. myrtilella</emphasis>
(Stt.))
</taxonomicName>
. Larval exit slit on upper side of the leaf. Cocoon grey-beige to grey-ochre; length 2.6–3.2 mm, maximal width 1.3–1.8 mm (
<figureCitation id="1326D7EEFFF0A123D3B347C2AC11F904" box="[1253,1340,1698,1723]" captionStart="FIGURES 33 – 38" captionStartId="10.[151,264,1551,1574]" captionTargetBox="[151,1425,195,1520]" captionTargetId="figure@10.[151,1436,195,1530]" captionTargetPageId="10" captionText="FIGURES 33 – 38. Stigmella tripartita Diškus &amp; Stonis, sp. nov. 33, habitat, SW of Cajabamba, Chimborazo Province, Ecuador, the equatorial Andes, elevation 3870 m; 34, 36, host-plant Baccharis buxifolia (Lam.) Pers.; 35, usual habitat of the species occurrence along roadsides; 37, cocoon; 38, forewing (reconstructed)." httpUri="https://zenodo.org/record/261794/files/figure.png" pageId="9" pageNumber="110">Fig. 37</figureCitation>
). Adults emerged in March.
</paragraph>
</subSubSection>
<subSubSection id="C30798E0FFF0A123D791478AABF8F898" pageId="9" pageNumber="110" type="distribution">
<paragraph id="8BA2CB6BFFF0A123D791478AABF8F898" blockId="9.[151,1437,834,2011]" pageId="9" pageNumber="110">
<emphasis id="B9691779FFF0A123D791478AA874F8BC" bold="true" box="[199,345,1770,1795]" pageId="9" pageNumber="110">Distribution</emphasis>
(
<figureCitation id="1326D7EEFFF0A123D63C478AA882F8BC" box="[362,431,1770,1795]" captionStart="FIGURE 3" captionStartId="2.[151,250,1980,2002]" captionTargetBox="[151,1436,193,1956]" captionTargetId="figure@2.[151,1436,193,1959]" captionTargetPageId="2" captionText="FIGURE 3. Current records of Stigmella species feeding on plants from the genus Baccharis L." httpUri="https://zenodo.org/record/261788/files/figure.png" pageId="9" pageNumber="110">Fig. 3</figureCitation>
). This species occurs high in the equatorial Andes (
<collectingCountry id="F30A8BFBFFF0A123D4AA478AAD70F8BC" box="[1020,1117,1770,1795]" name="Ecuador" pageId="9" pageNumber="110">Ecuador</collectingCountry>
) at altitudes between
<quantity id="4CE5668EFFF0A123D200478BA9C1F899" metricMagnitude="3" metricUnit="m" metricValue="3.835" metricValueMax="3.87" metricValueMin="3.8" pageId="9" pageNumber="110" unit="m" value="3835.0" valueMax="3870.0" valueMin="3800.0">3800– 3870 m</quantity>
(
<figureCitation id="1326D7EEFFF0A123D7AD466EA863F898" box="[251,334,1806,1831]" captionStart="FIGURES 33 – 38" captionStartId="10.[151,264,1551,1574]" captionTargetBox="[151,1425,195,1520]" captionTargetId="figure@10.[151,1436,195,1530]" captionTargetPageId="10" captionText="FIGURES 33 – 38. Stigmella tripartita Diškus &amp; Stonis, sp. nov. 33, habitat, SW of Cajabamba, Chimborazo Province, Ecuador, the equatorial Andes, elevation 3870 m; 34, 36, host-plant Baccharis buxifolia (Lam.) Pers.; 35, usual habitat of the species occurrence along roadsides; 37, cocoon; 38, forewing (reconstructed)." httpUri="https://zenodo.org/record/261794/files/figure.png" pageId="9" pageNumber="110">Fig. 33</figureCitation>
), mostly along roadsides (
<figureCitation id="1326D7EEFFF0A123D522466EABE5F898" box="[628,712,1806,1831]" captionStart="FIGURES 33 – 38" captionStartId="10.[151,264,1551,1574]" captionTargetBox="[151,1425,195,1520]" captionTargetId="figure@10.[151,1436,195,1530]" captionTargetPageId="10" captionText="FIGURES 33 – 38. Stigmella tripartita Diškus &amp; Stonis, sp. nov. 33, habitat, SW of Cajabamba, Chimborazo Province, Ecuador, the equatorial Andes, elevation 3870 m; 34, 36, host-plant Baccharis buxifolia (Lam.) Pers.; 35, usual habitat of the species occurrence along roadsides; 37, cocoon; 38, forewing (reconstructed)." httpUri="https://zenodo.org/record/261794/files/figure.png" pageId="9" pageNumber="110">Fig. 35</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30798E0FFF0A123D7914652AA54F8D0" pageId="9" pageNumber="110" type="etymology">
<paragraph id="8BA2CB6BFFF0A123D7914652AA54F8D0" blockId="9.[151,1437,834,2011]" pageId="9" pageNumber="110">
<emphasis id="B9691779FFF0A123D7914652A863F8F4" bold="true" box="[199,334,1842,1867]" pageId="9" pageNumber="110">Etymology.</emphasis>
The species name is derived from the Latin
<emphasis id="B9691779FFF0A123D41E4654AA97F8F4" box="[840,954,1844,1867]" italics="true" pageId="9" pageNumber="110">tripartitus</emphasis>
(divided in three parts; triple) in reference to the unusual set of three very large cornuti of the male genitalia.
</paragraph>
</subSubSection>
<subSubSection id="C30798E0FFF0A123D791461AA8A4F864" pageId="9" pageNumber="110" type="discussion">
<paragraph id="8BA2CB6BFFF0A123D791461AA8A4F864" blockId="9.[151,1437,834,2011]" pageId="9" pageNumber="110">
<emphasis id="B9691779FFF0A123D791461AA819F82C" bold="true" box="[199,308,1914,1939]" pageId="9" pageNumber="110">Remarks</emphasis>
. This distinctive new species, associated with
<taxonomicName id="4C1DB0E8FFF0A123D403461AAD18F82C" box="[853,1077,1914,1939]" class="Magnoliopsida" family="Asteraceae" genus="Baccharis" kingdom="Plantae" order="Asterales" pageId="9" pageNumber="110" phylum="Tracheophyta" rank="species" species="buxifolia">
<emphasis id="B9691779FFF0A123D403461AAD18F82C" box="[853,1077,1914,1939]" italics="true" pageId="9" pageNumber="110">Baccharis buxifolia</emphasis>
</taxonomicName>
as a host-plant and possessing three large cornuti in the male genitalia, is described from the genitalia slides dissected from adults still enclosed within the pupa shell.
</paragraph>
</subSubSection>
</treatment>
</document>
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