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<document id="EB318474F247225D90BD5D4F1246C067" ID-CLB-Dataset="9528" ID-DOI="10.11646/zootaxa.4881.3.3" ID-GBIF-Dataset="56a43dc1-9576-49d5-8650-255e66c48add" ID-ISSN="1175-5326" ID-Zenodo-Dep="4283896" ID-ZooBank="8A41020C-A131-4163-B3C6-D3398647EB4B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1605980153811" checkinUser="plazi" docAuthor="Curini-Galletti, Marco, Carcupino, Marcella, Stocchino, Giacinta A., Leasi, Francesca &amp; Norenburg, Jon L." docDate="2020" docId="03B42573FFC3206FFF56FDB0AF6DFD76" docLanguage="en" docName="zootaxa.4881.3.3.pdf" docOrigin="Zootaxa 4881 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Duplominona hystricina Curini-Galletti &amp; Carcupino &amp; Stocchino &amp; Leasi &amp; Norenburg 2020, n. sp." docType="treatment" docVersion="7" lastPageNumber="487" masterDocId="FF8D5D0BFFC7206AFFC1FF92AD30FFF2" masterDocTitle="New species of Duplominona Karling, 1966 (Platyhelminthes, Proseriata) from the Pacific coast of Panama" masterLastPageNumber="498" masterPageNumber="482" pageNumber="486" updateTime="1698846483825" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="E045AD2DE9E6582A5E7FDF0214FA78F5">New species of Duplominona Karling, 1966 (Platyhelminthes, Proseriata) from the Pacific coast of Panama</mods:title>
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<mods:namePart id="985AA2AE19A0ED40BB837AF01CB4EAEF">Curini-Galletti, Marco</mods:namePart>
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<mods:namePart id="7D898BAFF2D8D8723DA9CE7C9CA4C7CE">Carcupino, Marcella</mods:namePart>
<mods:affiliation id="D97E1E8B4071621910A648C066826874">Dipartimento di Medicina Veterinaria, Università di Sassari, Sassari, Italy &amp; carcupin @ uniss. it; https: // orcid. org / 0000 - 0002 - 0484 - 3043</mods:affiliation>
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<mods:namePart id="320FE149D592DD843E571DE528B3125E">Stocchino, Giacinta A.</mods:namePart>
<mods:affiliation id="73FEF5599BE39F79CE3AB00106D91101">Dipartimento di Medicina Veterinaria, Università di Sassari, Sassari, Italy &amp; stocchin @ uniss. it; https: // orcid. org / 0000 - 0002 - 7005 - 208 X</mods:affiliation>
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<mods:namePart id="2A1C98C31858F2D6B5EEFB7DEA98C7E3">Leasi, Francesca</mods:namePart>
<mods:affiliation id="1E38B1BCA9BB6ECA5EFE6AD26F4EF3B4">Department of Biology, Geology and Environmental Sciences, University of Tennessee at Chattanooga, USA francesca-leasi @ utc. edu; https: // orcid. org / 0000 - 0001 - 6995 - 0934</mods:affiliation>
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<mods:namePart id="6D1B1A3EC7B504E9557E046C75B24321">Norenburg, Jon L.</mods:namePart>
<mods:affiliation id="DC31984B49AC088347B6F570858CA9A8">Department of Invertebrate Zoology, National Museum of Natural History, Smithsonian Institution, Washington, DC, USA NORENBUR @ si. edu; https: // orcid. org / 0000 - 0001 - 7776 - 1527</mods:affiliation>
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<emphasis id="B9694877FFC3206EFF56FDB0AFFCFDCF" bold="true" box="[151,716,546,573]" pageId="4" pageNumber="486">
<taxonomicName id="4C1DEFE6FFC3206EFF56FDB0AFB1FDCE" authority="Curini-Galletti" authorityName="Curini-Galletti &amp; Carcupino &amp; Stocchino &amp; Leasi &amp; Norenburg" authorityYear="2020" box="[151,641,546,573]" class="Rhabditophora" family="Monocelididae" genus="Duplominona" kingdom="Animalia" order="Seriata" pageId="4" pageNumber="486" phylum="Platyhelminthes" rank="species" species="hystricina" status="sp. nov.">
<emphasis id="B9694877FFC3206EFF56FDB0AC8FFDCF" bold="true" box="[151,447,546,573]" italics="true" pageId="4" pageNumber="486">Duplominona hystricina</emphasis>
Curini-Galletti
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<taxonomicNameLabel id="A25AF50CFFC3206EFD48FDB1AFFCFDCF" box="[649,716,547,573]" pageId="4" pageNumber="486" rank="species">n. sp.</taxonomicNameLabel>
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<paragraph id="8BA29465FFC3206EFF56FDD5ADDBFD93" blockId="4.[151,716,546,609]" box="[151,235,583,609]" pageId="4" pageNumber="486">
(
<figureCitation id="132688E0FFC3206EFF61FDD5ADD2FD93" box="[160,226,583,609]" captionStart="FIGURE 3" captionStartId="4.[151,250,1770,1794]" captionTargetBox="[162,1437,921,1738]" captionTargetId="figure-258@4.[151,1437,921,1747]" captionTargetPageId="4" captionText="FIGURE 3. Duplominona hystricina n. sp. (A) Habitus of a living animal; see Fig. 2A for interpretation of anatomical features. (B) Reconstruction of the cirrus. (C) Spine morphology. (D, E) Micrographs of cirri. (F) Reconstruction of the genital areas from sagittal sections. (G) Stylet of the accessory organ. (Scale bars: B, C = 5 μm; D, E, G = 10 μm)." figureDoi="http://doi.org/10.5281/zenodo.4283902" httpUri="https://zenodo.org/record/4283902/files/figure.png" pageId="4" pageNumber="486">Fig. 3</figureCitation>
)
</paragraph>
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<subSubSection id="C307C7EEFFC3206EFF56FD1DADC1FCAF" pageId="4" pageNumber="486" type="materials_examined">
<paragraph id="8BA29465FFC3206EFF56FD1DAC3AFD03" blockId="4.[151,1437,655,897]" pageId="4" pageNumber="486">
<materialsCitation id="3B759E38FFC3206EFF56FD1DAC3AFD03" ID-GBIF-Occurrence="2985239301" collectingDate="2011-12" collectionCode="USNM" country="Panama" elevation="20" latitude="8.673019" location="Isla Pacheca" longLatPrecision="1" longitude="-79.0535" pageId="4" pageNumber="486" specimenCode="USNM 1622573" specimenCount="1" typeStatus="holotype">
<emphasis id="B9694877FFC3206EFF56FD1DAC34FD5B" bold="true" box="[151,260,655,681]" pageId="4" pageNumber="486">
<typeStatus id="54A62AC7FFC3206EFF56FD1DAC34FD5B" box="[151,260,655,681]" pageId="4" pageNumber="486" type="holotype">Holotype</typeStatus>
</emphasis>
. Pacific Ocean: Gulf of
<collectingCountry id="F30AD4F5FFC3206EFDEBFD1DAFB6FD5B" box="[554,646,655,681]" name="Panama" pageId="4" pageNumber="486">Panama</collectingCountry>
, off
<location id="8EC2C2BEFFC3206EFD01FD1DAE60FD5B" LSID="urn:lsid:plazi:treatment:03B42573FFC3206FFF56FDB0AF6DFD76:8EC2C2BEFFC3206EFD01FD1DAE60FD5B" box="[704,848,655,681]" country="Panama" latitude="8.673019" longLatPrecision="1" longitude="-79.0535" name="Isla Pacheca" pageId="4" pageNumber="486">Isla Pacheca</location>
(
<locationDeviation id="9A108D47FFC3206EFCA2FD1DA934FD5B" box="[867,1028,655,681]" pageId="4" pageNumber="486">Las Perlas Is.</locationDeviation>
,
<collectingCountry id="F30AD4F5FFC3206EFBD5FD1DA942FD5B" box="[1044,1138,655,681]" name="Panama" pageId="4" pageNumber="486">Panama</collectingCountry>
)(Lat.
<geoCoordinate id="EE29F2A2FFC3206EFB76FD1DA869FD5B" box="[1207,1369,655,681]" degrees="8" direction="north" minutes="40" orientation="latitude" pageId="4" pageNumber="486" precision="1" seconds="22.87" value="8.673019">8°4022.87”N</geoCoordinate>
; Lon.
<geoCoordinate id="EE29F2A2FFC3206EFF56FD21AC78FD3F" box="[151,328,690,717]" degrees="79" direction="west" minutes="3" orientation="longitude" pageId="4" pageNumber="486" precision="1" seconds="12.58" value="-79.0535">79° 312.58”W</geoCoordinate>
), about
<quantity id="4CE53980FFC3206EFE65FD21ACEFFD3F" box="[420,479,691,717]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="486" unit="m" value="20.0">
<elevation id="00307356FFC3206EFE65FD21ACEFFD3F" box="[420,479,691,717]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="486" unit="m" value="20.0">20 m</elevation>
</quantity>
deep, coarse sand with shell fragments,
<date id="FFA3B2A5FFC3206EFC71FD21A957FD3F" box="[944,1127,691,717]" pageId="4" pageNumber="486" value="2011-12">
<collectingDate id="EFE74B4DFFC3206EFC71FD21A957FD3F" box="[944,1127,691,717]" pageId="4" pageNumber="486" value="2011-12">December 2011</collectingDate>
</date>
: one whole mount (
<specimenCode id="DBBB3C1EFFC3206EFA88FD21ADCEFD03" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="486" type="Museum">USNM 1622573</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BA29465FFC3206EFF06FD69A84EFCE7" blockId="4.[151,1437,655,897]" box="[199,1406,762,789]" pageId="4" pageNumber="486">
<materialsCitation id="3B759E38FFC3206EFF06FD69A84AFCE7" ID-GBIF-Occurrence="2985239318" box="[199,1402,762,789]" collectingDate="2011-12" collectionCode="USNM" country="Panama" elevation="20" latitude="8.673019" location="Panama" longLatPrecision="1" longitude="-79.0535" pageId="4" pageNumber="486" specimenCode="USNM 1622574, USNM 1622575" specimenCount="1" typeStatus="paratype">
<emphasis id="B9694877FFC3206EFF06FD69AC0FFCE7" bold="true" box="[199,319,763,789]" pageId="4" pageNumber="486">
<typeStatus id="54A62AC7FFC3206EFF06FD69AC0FFCE7" box="[199,319,763,789]" pageId="4" pageNumber="486" type="paratype">Paratypes</typeStatus>
</emphasis>
. Same data as holotype, two specimens sagittally sectioned (
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;
<specimenCode id="DBBB3C1EFFC3206EFB72FD69A842FCE7" box="[1203,1394,762,789]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="486" type="Museum">USNM 1622575</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BA29465FFC3206EFF06FC8CADC1FCAF" blockId="4.[151,1437,655,897]" pageId="4" pageNumber="486">
<emphasis id="B9694877FFC3206EFF06FC8CAC4FFCCA" bold="true" box="[199,383,798,824]" pageId="4" pageNumber="486">Other material</emphasis>
. Same data as holotype, two specimens observed alive, and prepared as semi-permanent mounts.
</paragraph>
</subSubSection>
<subSubSection id="C307C7EEFFC3206EFF06FCF5A97BFC73" box="[199,1099,871,897]" pageId="4" pageNumber="486" type="etymology">
<paragraph id="8BA29465FFC3206EFF06FCF5A97BFC73" blockId="4.[151,1437,655,897]" box="[199,1099,871,897]" pageId="4" pageNumber="486">
<emphasis id="B9694877FFC3206EFF06FCF5AC78FC73" bold="true" box="[199,328,871,897]" pageId="4" pageNumber="486">Etymology</emphasis>
. The name refers to the spiky,
<taxonomicName id="4C1DEFE6FFC3206EFD63FCF5AFC4FC72" box="[674,756,871,896]" class="Mammalia" family="Hystricidae" genus="Hystrix" kingdom="Animalia" order="Rodentia" pageId="4" pageNumber="486" phylum="Chordata" rank="genus">
<emphasis id="B9694877FFC3206EFD63FCF5AFC4FC72" box="[674,756,871,896]" italics="true" pageId="4" pageNumber="486">Hystrix</emphasis>
</taxonomicName>
-like cirrus of the new species.
</paragraph>
</subSubSection>
<caption id="DF62C4EDFFC3206EFF56F978A950F8B8" ID-DOI="http://doi.org/10.5281/zenodo.4283902" ID-Zenodo-Dep="4283902" httpUri="https://zenodo.org/record/4283902/files/figure.png" pageId="4" pageNumber="486" startId="4.[151,250,1770,1794]" targetBox="[162,1437,921,1738]" targetPageId="4">
<paragraph id="8BA29465FFC3206EFF56F978A950F8B8" blockId="4.[151,1437,1770,1867]" pageId="4" pageNumber="486">
<emphasis id="B9694877FFC3206EFF56F978AC22F8F0" bold="true" box="[151,274,1770,1794]" pageId="4" pageNumber="486">FIGURE 3.</emphasis>
<taxonomicName id="4C1DEFE6FFC3206EFED9F978AF3DF8F0" authority="Curini-Galletti &amp; Carcupino &amp; Stocchino &amp; Leasi &amp; Norenburg, 2020" authorityName="Curini-Galletti &amp; Carcupino &amp; Stocchino &amp; Leasi &amp; Norenburg" authorityYear="2020" box="[280,525,1770,1794]" class="Rhabditophora" family="Monocelididae" genus="Duplominona" kingdom="Animalia" order="Seriata" pageId="4" pageNumber="486" phylum="Platyhelminthes" rank="species" species="hystricina" status="sp. nov.">
<emphasis id="B9694877FFC3206EFED9F978AF3DF8F0" box="[280,525,1770,1794]" italics="true" pageId="4" pageNumber="486">Duplominona hystricina</emphasis>
</taxonomicName>
<emphasis id="B9694877FFC3206EFDD2F978AF7CF8F0" bold="true" box="[531,588,1770,1794]" pageId="4" pageNumber="486">
<taxonomicNameLabel id="A25AF50CFFC3206EFDD2F978AF7CF8F0" box="[531,588,1770,1794]" pageId="4" pageNumber="486" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(A) Habitus of a living animal; see Fig. 2 A for interpretation of anatomical features. (B) Reconstruction of the cirrus. (C) Spine morphology. (D, E) Micrographs of cirri. (F) Reconstruction of the genital areas from sagittal sections. (G) Stylet of the accessory organ. (Scale bars: B, C = 5 μm; D, E, G = 10 μm).
</paragraph>
</caption>
<subSubSection id="C307C7EEFFC3206FFF06F8EAAE55FE06" lastPageId="5" lastPageNumber="487" pageId="4" pageNumber="486" type="description">
<paragraph id="8BA29465FFC3206EFF06F8EAAC5AF828" blockId="4.[151,1437,1912,2011]" pageId="4" pageNumber="486">
<emphasis id="B9694877FFC3206EFF06F8EAAC69F861" bold="true" box="[199,345,1912,1939]" pageId="4" pageNumber="486">Description.</emphasis>
A minute species, less than
<quantity id="4CE53980FFC3206EFD54F8EAAFDBF861" box="[661,747,1912,1939]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="486" unit="mm" value="1.5">1.5 mm</quantity>
in length, very similar in habitus to the previous species. Anterior tip with oily dots and tiny rod-shaped rhabdoids, about ¼ the diameter of the statocyst. With a short pharynx at about mid-body.
</paragraph>
<paragraph id="8BA29465FFC2206FFF06FF04A841FEEF" blockId="5.[151,1437,150,645]" pageId="5" pageNumber="487">
<emphasis id="B9694877FFC2206FFF06FF04AC89FF42" bold="true" box="[199,441,150,176]" pageId="5" pageNumber="487">Male genital system</emphasis>
. With few (35) testes in one medio-ventral row (
<figureCitation id="132688E0FFC2206FFBC2FF04A950FF42" box="[1027,1120,150,176]" captionStart="FIGURE 3" captionStartId="4.[151,250,1770,1794]" captionTargetBox="[162,1437,921,1738]" captionTargetId="figure-258@4.[151,1437,921,1747]" captionTargetPageId="4" captionText="FIGURE 3. Duplominona hystricina n. sp. (A) Habitus of a living animal; see Fig. 2A for interpretation of anatomical features. (B) Reconstruction of the cirrus. (C) Spine morphology. (D, E) Micrographs of cirri. (F) Reconstruction of the genital areas from sagittal sections. (G) Stylet of the accessory organ. (Scale bars: B, C = 5 μm; D, E, G = 10 μm)." figureDoi="http://doi.org/10.5281/zenodo.4283902" httpUri="https://zenodo.org/record/4283902/files/figure.png" pageId="5" pageNumber="487">Fig. 3A</figureCitation>
). With an ovoid copulatory organ, about 35 μm long. Small, spiny cirrus about 15 μm long, provided with 1012 rows of spines (
<figureCitation id="132688E0FFC2206FFAD1FF28A863FF26" box="[1296,1363,186,212]" captionStart="FIGURE 3" captionStartId="4.[151,250,1770,1794]" captionTargetBox="[162,1437,921,1738]" captionTargetId="figure-258@4.[151,1437,921,1747]" captionTargetPageId="4" captionText="FIGURE 3. Duplominona hystricina n. sp. (A) Habitus of a living animal; see Fig. 2A for interpretation of anatomical features. (B) Reconstruction of the cirrus. (C) Spine morphology. (D, E) Micrographs of cirri. (F) Reconstruction of the genital areas from sagittal sections. (G) Stylet of the accessory organ. (Scale bars: B, C = 5 μm; D, E, G = 10 μm)." figureDoi="http://doi.org/10.5281/zenodo.4283902" httpUri="https://zenodo.org/record/4283902/files/figure.png" pageId="5" pageNumber="487">Fig. 3</figureCitation>
BE). Spines are 35 μm long, slender, slightly recurved, and acutely pointed, with a small, swollen basis, 11.5 μm across (
<figureCitation id="132688E0FFC2206FFF5EFE90ADC8FEEE" box="[159,248,258,284]" captionStart="FIGURE 3" captionStartId="4.[151,250,1770,1794]" captionTargetBox="[162,1437,921,1738]" captionTargetId="figure-258@4.[151,1437,921,1747]" captionTargetPageId="4" captionText="FIGURE 3. Duplominona hystricina n. sp. (A) Habitus of a living animal; see Fig. 2A for interpretation of anatomical features. (B) Reconstruction of the cirrus. (C) Spine morphology. (D, E) Micrographs of cirri. (F) Reconstruction of the genital areas from sagittal sections. (G) Stylet of the accessory organ. (Scale bars: B, C = 5 μm; D, E, G = 10 μm)." figureDoi="http://doi.org/10.5281/zenodo.4283902" httpUri="https://zenodo.org/record/4283902/files/figure.png" pageId="5" pageNumber="487">Fig 3 C</figureCitation>
). The minute spines of the species are poorly sclerified, and quickly deteriorated in the whole mounts.
</paragraph>
<paragraph id="8BA29465FFC2206FFF06FEB4AF30FE97" blockId="5.[151,1437,150,645]" pageId="5" pageNumber="487">
Accessory organ about 20 μm across, provided with a pen-nib shaped stylet 1213 μm long (
<figureCitation id="132688E0FFC2206FFB14FEB4A800FEB2" box="[1237,1328,294,320]" captionStart="FIGURE 3" captionStartId="4.[151,250,1770,1794]" captionTargetBox="[162,1437,921,1738]" captionTargetId="figure-258@4.[151,1437,921,1747]" captionTargetPageId="4" captionText="FIGURE 3. Duplominona hystricina n. sp. (A) Habitus of a living animal; see Fig. 2A for interpretation of anatomical features. (B) Reconstruction of the cirrus. (C) Spine morphology. (D, E) Micrographs of cirri. (F) Reconstruction of the genital areas from sagittal sections. (G) Stylet of the accessory organ. (Scale bars: B, C = 5 μm; D, E, G = 10 μm)." figureDoi="http://doi.org/10.5281/zenodo.4283902" httpUri="https://zenodo.org/record/4283902/files/figure.png" pageId="5" pageNumber="487">Fig 3 G</figureCitation>
). It opens to the outside with its own pore.
</paragraph>
<paragraph id="8BA29465FFC2206FFF06FEFCAE55FE06" blockId="5.[151,1437,150,645]" pageId="5" pageNumber="487">
<emphasis id="B9694877FFC2206FFF06FEFCACE4FE7A" bold="true" box="[199,468,366,392]" pageId="5" pageNumber="487">Female genital system</emphasis>
. Ovaries just in front of the pharynx. Vitellaria extending from of anterior testes to bursa. With a small, spherical pre-penial bursa, provided dorsally with a few large resorbiens vacuoles (
<figureCitation id="132688E0FFC2206FFAD1FE00A85AFE5F" box="[1296,1386,402,429]" captionStart="FIGURE 3" captionStartId="4.[151,250,1770,1794]" captionTargetBox="[162,1437,921,1738]" captionTargetId="figure-258@4.[151,1437,921,1747]" captionTargetPageId="4" captionText="FIGURE 3. Duplominona hystricina n. sp. (A) Habitus of a living animal; see Fig. 2A for interpretation of anatomical features. (B) Reconstruction of the cirrus. (C) Spine morphology. (D, E) Micrographs of cirri. (F) Reconstruction of the genital areas from sagittal sections. (G) Stylet of the accessory organ. (Scale bars: B, C = 5 μm; D, E, G = 10 μm)." figureDoi="http://doi.org/10.5281/zenodo.4283902" httpUri="https://zenodo.org/record/4283902/files/figure.png" pageId="5" pageNumber="487">Fig. 3 F</figureCitation>
:
<emphasis id="B9694877FFC2206FFAB4FE01A8BCFE5E" box="[1397,1420,403,428]" italics="true" pageId="5" pageNumber="487">rv</emphasis>
), abutting the gut lining. With a short and wide vaginal duct. The female duct continues posteriorly to the bursa and opens behind the prostatoid organ pore through the female pore.
</paragraph>
</subSubSection>
<subSubSection id="C307C7EEFFC2206FFF06FE6CAF6DFD76" pageId="5" pageNumber="487" type="diagnosis">
<paragraph id="8BA29465FFC2206FFF06FE6CAFBBFD92" blockId="5.[151,1437,150,645]" pageId="5" pageNumber="487">
<emphasis id="B9694877FFC2206FFF06FE6CAC0AFDEA" bold="true" box="[199,314,510,536]" pageId="5" pageNumber="487">Diagnosis</emphasis>
. Species of
<taxonomicName id="4C1DEFE6FFC2206FFE0AFE6CAF55FDEA" authorityName="Karling" authorityYear="1966" box="[459,613,510,536]" class="Rhabditophora" family="Monocelididae" genus="Duplominona" kingdom="Animalia" order="Seriata" pageId="5" pageNumber="487" phylum="Platyhelminthes" rank="genus">
<emphasis id="B9694877FFC2206FFE0AFE6CAF55FDEA" box="[459,613,510,536]" italics="true" pageId="5" pageNumber="487">Duplominona</emphasis>
</taxonomicName>
with few (35) testes. Small cirrus provided with 1012 rows of acute spines, 35 μm high, with swollen bases. With a spherical bursa in front of the copulatory organ, and external va-gina. Accessory organ stylet 1213 μm long.
</paragraph>
<paragraph id="8BA29465FFC2206FFF06FDF9AF6DFD76" blockId="5.[151,1437,150,645]" box="[199,605,618,645]" pageId="5" pageNumber="487">Pore index: a: b: c: d = =5: 1.5: 2: 1.</paragraph>
</subSubSection>
</treatment>
</document>