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<mods:title id="2617FC2AB3490FEB4394C982F3053926">Revision and phylogenetic analysis of the African genus Ischnopoderona (Scheerpeltz, 1974) new status (Coleoptera, Staphylinidae: Aleocharinae)</mods:title>
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<subSubSection id="C3169BA8FFAA9475FF7872BE634E7FAB" pageId="7" pageNumber="8" type="nomenclature">
<paragraph id="8BB3C823FFAA9475FF7872BE61637F88" blockId="7.[151,779,946,1007]" box="[151,779,946,972]" pageId="7" pageNumber="8">
<heading id="D0FB7F4FFFAA9475FF7872BE61637F88" bold="true" box="[151,779,946,972]" fontSize="11" level="1" pageId="7" pageNumber="8" reason="1">
<emphasis id="B9781431FFAA9475FF7872BE61637F88" bold="true" box="[151,779,946,972]" pageId="7" pageNumber="8">
Genus
<taxonomicName id="4C0CB3A0FFAA9475FF0172BE60C17F88" ID-CoL="8X7H5" authority="Scheerpeltz, 1974" authorityName="Scheerpeltz" authorityYear="1974" box="[238,681,946,972]" class="Insecta" family="Staphylinidae" genus="Ischnopoderona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B9781431FFAA9475FF0172BE63DB7F88" bold="true" box="[238,435,946,972]" italics="true" pageId="7" pageNumber="8">Ischnopoderona</emphasis>
(
<bibRefCitation id="EF9DB5D2FFAA9475FE2D72BE60C97F88" author="Scheerpeltz" box="[450,673,946,972]" pageId="7" pageNumber="54" refString="Scheerpeltz, O. (1974) Coleoptera: Staphylinidae (exclus. Subfam. Paederinae, except. pars min.). In: Hanstrom B., Brinck P. &amp; Rudebeck G. (Ed.) South African Animal Life, 15, 43 - 394." type="journal article" year="1974">Scheerpeltz, 1974</bibRefCitation>
)
</taxonomicName>
, stat. n.
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB3C823FFAA9475FF7872D9634E7FAB" blockId="7.[151,779,946,1007]" box="[151,294,981,1007]" pageId="7" pageNumber="8">
(
<figureCitation id="1337D4A6FFAA9475FF7072D963757FAB" box="[159,285,981,1007]" captionStart-0="FIGURE 3" captionStart-1="FIGURES 4 11" captionStart-2="FIGURES 12 17" captionStartId-0="8.[151,255,1574,1598]" captionStartId-1="9.[151,269,1603,1627]" captionStartId-2="10.[151,269,1532,1556]" captionTargetBox-1="[303,1284,194,1585]" captionTargetBox-2="[403,1195,196,1503]" captionTargetId-1="figure@9.[295,1291,183,1592]" captionTargetId-2="figure@10.[390,1209,182,1516]" captionTargetPageId-1="9" captionTargetPageId-2="10" captionText-0="FIGURE 3. Ischnopoderona gracilicornis (Scheerp.): habitus." captionText-1="FIGURES 4 11. Ischnopoderona gracilicornis (Scheerp.): 4 — maxilla, 5 — labium, 6 — right mandible, 7 — labrum, 8 — mentum, 9 — shape of elytron, 10 — pronotal pubescence, 11 — tergite VIII." captionText-2="FIGURES 12 17. Variability of mesosternum and metasternum of Ischnopoderona sp.: 12 — I. longicollis (Bernh.), 13 — I. lucidula sp. n., 14 — I. nigerrima (Cam.), 15 — I. namibensis (Pace), 16 — I. gracilicornis (Scheerp.), 17 — I. madecassa (Cam.)" httpUri-1="https://zenodo.org/record/175637/files/figure.png" httpUri-2="https://zenodo.org/record/175638/files/figure.png" pageId="7" pageNumber="8">Figs. 317</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3169BA8FFAA9475FF78752960AA7878" box="[151,706,1061,1084]" pageId="7" pageNumber="8" type="reference_group">
<paragraph id="8BB3C823FFAA9475FF78752960AA7878" blockId="7.[151,1168,1061,1116]" box="[151,706,1061,1084]" pageId="7" pageNumber="8">
<treatmentCitationGroup id="AB1CEF0DFFAA9475FF78752960AA7878" box="[151,706,1061,1084]" pageId="7" pageNumber="8">
<taxonomicName id="4C0CB3A0FFAA9475FF78752960D67878" authority="Scheerpeltz, 1974: 229" authorityName="Scheerpeltz" authorityPageNumber="229" authorityYear="1974" box="[151,702,1061,1084]" class="Insecta" family="Staphylinidae" genus="Tachyusa" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="subGenus" subGenus="Ischnopoderona">
<emphasis id="B9781431FFAA9475FF78752962907878" box="[151,248,1061,1084]" italics="true" pageId="7" pageNumber="8">Tachyusa</emphasis>
(
<emphasis id="B9781431FFAA9475FEE5752963DE7878" box="[266,438,1061,1084]" italics="true" pageId="7" pageNumber="8">Ischnopoderona</emphasis>
)
<treatmentCitation id="0AADEE32FFAA9475FE2A752960D67878" author="Scheerpeltz" box="[453,702,1061,1084]" page="229" pageId="7" pageNumber="8" year="1974">
<bibRefCitation id="EF9DB5D2FFAA9475FE2A752960E07878" author="Scheerpeltz" box="[453,648,1061,1084]" pageId="7" pageNumber="54" refString="Scheerpeltz, O. (1974) Coleoptera: Staphylinidae (exclus. Subfam. Paederinae, except. pars min.). In: Hanstrom B., Brinck P. &amp; Rudebeck G. (Ed.) South African Animal Life, 15, 43 - 394." type="journal article" year="1974">Scheerpeltz, 1974</bibRefCitation>
: 229
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3169BA8FFAA9475FF78754966F87818" box="[151,1168,1093,1116]" pageId="7" pageNumber="8" type="materials_examined">
<paragraph id="8BB3C823FFAA9475FF78754966F87818" blockId="7.[151,1168,1061,1116]" box="[151,1168,1093,1116]" pageId="7" pageNumber="8">
<typeStatus id="54B77681FFAA9475FF78754962A47818" box="[151,204,1093,1116]" pageId="7" pageNumber="8">Type</typeStatus>
species:
<taxonomicName id="4C0CB3A0FFAA9475FEC27549618B7818" authority="Scheerpeltz, 1974: 229" authorityName="Scheerpeltz" authorityPageNumber="229" authorityYear="1974" box="[301,995,1093,1116]" class="Insecta" family="Staphylinidae" genus="Tachyusa" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="gracilicornis" subGenus="Ischnopoderona">
<emphasis id="B9781431FFAA9475FEC2754963F87818" box="[301,400,1093,1116]" italics="true" pageId="7" pageNumber="8">Tachyusa</emphasis>
(
<emphasis id="B9781431FFAA9475FE4E754960257818" box="[417,589,1093,1116]" italics="true" pageId="7" pageNumber="8">Ischnopoderona</emphasis>
)
<emphasis id="B9781431FFAA9475FDB37549608E7818" box="[604,742,1093,1116]" italics="true" pageId="7" pageNumber="8">gracilicornis</emphasis>
<bibRefCitation id="EF9DB5D2FFAA9475FD02754961D87818" author="Scheerpeltz" box="[749,944,1093,1116]" pageId="7" pageNumber="54" refString="Scheerpeltz, O. (1974) Coleoptera: Staphylinidae (exclus. Subfam. Paederinae, except. pars min.). In: Hanstrom B., Brinck P. &amp; Rudebeck G. (Ed.) South African Animal Life, 15, 43 - 394." type="journal article" year="1974">Scheerpeltz, 1974</bibRefCitation>
: 229
</taxonomicName>
(by monotypy).
</paragraph>
</subSubSection>
<subSubSection id="C3169BA8FFAA9478FF787581672A7B0A" lastPageId="10" lastPageNumber="11" pageId="7" pageNumber="8" type="diagnosis">
<paragraph id="8BB3C823FFAA9475FF787581632A79FB" blockId="7.[151,1438,1165,2035]" pageId="7" pageNumber="8">
<emphasis id="B9781431FFAA9475FF787581637F78E3" bold="true" box="[151,279,1165,1191]" pageId="7" pageNumber="8">Diagnosis.</emphasis>
<taxonomicName id="4C0CB3A0FFAA9475FEF0758163B278E3" box="[287,474,1165,1191]" class="Insecta" family="Staphylinidae" genus="Ischnopoderona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">Ischnopoderona</taxonomicName>
can be recognized by the combination of the following characters: body slender, parallel­sided; ligula narrow (
<figureCitation id="1337D4A6FFAA9475FE1B75B96053788B" box="[500,571,1205,1231]" captionStart="FIGURES 4 11" captionStartId="9.[151,269,1603,1627]" captionTargetBox="[303,1284,194,1585]" captionTargetId="figure@9.[295,1291,183,1592]" captionTargetPageId="9" captionText="FIGURES 4 11. Ischnopoderona gracilicornis (Scheerp.): 4 — maxilla, 5 — labium, 6 — right mandible, 7 — labrum, 8 — mentum, 9 — shape of elytron, 10 — pronotal pubescence, 11 — tergite VIII." httpUri="https://zenodo.org/record/175637/files/figure.png" pageId="7" pageNumber="8">Fig. 5</figureCitation>
), bifid to the base or nearly so, lobes parallel­sided; maxillary palpi elongate and weakly widened apically (
<figureCitation id="1337D4A6FFAA9475FDDC75D1601378B3" box="[563,635,1245,1271]" captionStart="FIGURES 4 11" captionStartId="9.[151,269,1603,1627]" captionTargetBox="[303,1284,194,1585]" captionTargetId="figure@9.[295,1291,183,1592]" captionTargetPageId="9" captionText="FIGURES 4 11. Ischnopoderona gracilicornis (Scheerp.): 4 — maxilla, 5 — labium, 6 — right mandible, 7 — labrum, 8 — mentum, 9 — shape of elytron, 10 — pronotal pubescence, 11 — tergite VIII." httpUri="https://zenodo.org/record/175637/files/figure.png" pageId="7" pageNumber="8">Fig. 4</figureCitation>
); antennae long, extending from 1/2 to 3/4 of elytral length and very slightly increasing in width apically; head without infraorbital ridge; pronotal hypomera fully visible in lateral view; elytral lateral sides arcuate, widest in the middle (
<figureCitation id="1337D4A6FFAA9475FCCB742161047903" box="[804,876,1325,1351]" captionStart="FIGURES 4 11" captionStartId="9.[151,269,1603,1627]" captionTargetBox="[303,1284,194,1585]" captionTargetId="figure@9.[295,1291,183,1592]" captionTargetPageId="9" captionText="FIGURES 4 11. Ischnopoderona gracilicornis (Scheerp.): 4 — maxilla, 5 — labium, 6 — right mandible, 7 — labrum, 8 — mentum, 9 — shape of elytron, 10 — pronotal pubescence, 11 — tergite VIII." httpUri="https://zenodo.org/record/175637/files/figure.png" pageId="7" pageNumber="8">Fig. 9</figureCitation>
); abdomen distinctly constricted at base, widest at level of tergites V and VI, tergites IIIV distinctly transversely impressed at base, at least first impression coarsely punctate, tergite V distinctly longer than VII; sternites strongly transversely impressed at base; tarsal formula 4­5­5.
</paragraph>
<paragraph id="8BB3C823FFAA9475FF2974DC63217A57" blockId="7.[151,1438,1165,2035]" pageId="7" pageNumber="8">
<emphasis id="B9781431FFAA9475FF2974DC631479AE" bold="true" box="[198,380,1488,1514]" pageId="7" pageNumber="8">Redescription.</emphasis>
Body. Small to medium sized, length 2.0
<quantity id="4CF465C6FFAA9475FC9474DD61B279AF" box="[891,986,1489,1515]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="7" pageNumber="8" unit="mm" value="3.4">3.4 mm</quantity>
. Body slender, parallel­sided, convex, glossy (
<figureCitation id="1337D4A6FFAA9475FF1D74F563517A57" box="[242,313,1529,1555]" captionStart="FIGURE 3" captionStartId="8.[151,255,1574,1598]" captionText="FIGURE 3. Ischnopoderona gracilicornis (Scheerp.): habitus." pageId="7" pageNumber="8">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="8BB3C823FFAA9475FF29772D61287ACF" blockId="7.[151,1438,1165,2035]" pageId="7" pageNumber="8">Head. Circular, temples parallel­sided or arcuately narrowed to hind angles; eyes protruding from lateral contours of head; head without infraorbital ridge. Antennae long to very long, very slightly increasing in width apically, extending from 1/2 to 3/4 of elytral length.</paragraph>
<paragraph id="8BB3C823FFAA9475FF297795619C7BB7" blockId="7.[151,1438,1165,2035]" pageId="7" pageNumber="8">
Mouthparts. Maxillary palpi 4­segmented (
<figureCitation id="1337D4A6FFAA9475FD507795616F7AF7" box="[703,775,1689,1715]" captionStart="FIGURES 4 11" captionStartId="9.[151,269,1603,1627]" captionTargetBox="[303,1284,194,1585]" captionTargetId="figure@9.[295,1291,183,1592]" captionTargetPageId="9" captionText="FIGURES 4 11. Ischnopoderona gracilicornis (Scheerp.): 4 — maxilla, 5 — labium, 6 — right mandible, 7 — labrum, 8 — mentum, 9 — shape of elytron, 10 — pronotal pubescence, 11 — tergite VIII." httpUri="https://zenodo.org/record/175637/files/figure.png" pageId="7" pageNumber="8">Fig. 4</figureCitation>
), palpomere 1 very short, palpomere 2 elongate, slightly widened apically, palpomere 3 distinctly longer than 2, weakly widened towards apex, palpomere 4 short, subuliform. Galea long and slender, slightly shorter than lacinia (
<figureCitation id="1337D4A6FFAA9475FC9D77E561D47B47" box="[882,956,1769,1795]" captionStart="FIGURES 4 11" captionStartId="9.[151,269,1603,1627]" captionTargetBox="[303,1284,194,1585]" captionTargetId="figure@9.[295,1291,183,1592]" captionTargetPageId="9" captionText="FIGURES 4 11. Ischnopoderona gracilicornis (Scheerp.): 4 — maxilla, 5 — labium, 6 — right mandible, 7 — labrum, 8 — mentum, 9 — shape of elytron, 10 — pronotal pubescence, 11 — tergite VIII." httpUri="https://zenodo.org/record/175637/files/figure.png" pageId="7" pageNumber="8">Fig. 4</figureCitation>
). Labial palpi 3­segmented (
<figureCitation id="1337D4A6FFAA9475FAE077E5673F7B47" box="[1295,1367,1769,1795]" captionStart="FIGURES 4 11" captionStartId="9.[151,269,1603,1627]" captionTargetBox="[303,1284,194,1585]" captionTargetId="figure@9.[295,1291,183,1592]" captionTargetPageId="9" captionText="FIGURES 4 11. Ischnopoderona gracilicornis (Scheerp.): 4 — maxilla, 5 — labium, 6 — right mandible, 7 — labrum, 8 — mentum, 9 — shape of elytron, 10 — pronotal pubescence, 11 — tergite VIII." httpUri="https://zenodo.org/record/175637/files/figure.png" pageId="7" pageNumber="8">Fig. 5</figureCitation>
), palpomere 1 elongate, longer than palpomeres 2 and 3 together, palpomere 2 shorter and narrower than 1, palpomere 3 narrower and much longer than 2. Ligula (
<figureCitation id="1337D4A6FFAA9475FCF3763561097B17" box="[796,865,1849,1875]" captionStart="FIGURES 4 11" captionStartId="9.[151,269,1603,1627]" captionTargetBox="[303,1284,194,1585]" captionTargetId="figure@9.[295,1291,183,1592]" captionTargetPageId="9" captionText="FIGURES 4 11. Ischnopoderona gracilicornis (Scheerp.): 4 — maxilla, 5 — labium, 6 — right mandible, 7 — labrum, 8 — mentum, 9 — shape of elytron, 10 — pronotal pubescence, 11 — tergite VIII." httpUri="https://zenodo.org/record/175637/files/figure.png" pageId="7" pageNumber="8">Fig. 5</figureCitation>
) narrow, bifid to the base or nearly so, lobes parallel­sided. Mandibles curved towards apex, right mandible (
<figureCitation id="1337D4A6FFAA9475FC8C766D61C27B3F" box="[867,938,1889,1915]" captionStart="FIGURES 4 11" captionStartId="9.[151,269,1603,1627]" captionTargetBox="[303,1284,194,1585]" captionTargetId="figure@9.[295,1291,183,1592]" captionTargetPageId="9" captionText="FIGURES 4 11. Ischnopoderona gracilicornis (Scheerp.): 4 — maxilla, 5 — labium, 6 — right mandible, 7 — labrum, 8 — mentum, 9 — shape of elytron, 10 — pronotal pubescence, 11 — tergite VIII." httpUri="https://zenodo.org/record/175637/files/figure.png" pageId="7" pageNumber="8">Fig. 6</figureCitation>
) with very small, indistinct molar tooth at middle of inner margin, faintly serrate in dorsal molar region. Labrum (
<figureCitation id="1337D4A6FFAA9475FC0B768566427BE7" box="[996,1066,1929,1955]" captionStart="FIGURES 4 11" captionStartId="9.[151,269,1603,1627]" captionTargetBox="[303,1284,194,1585]" captionTargetId="figure@9.[295,1291,183,1592]" captionTargetPageId="9" captionText="FIGURES 4 11. Ischnopoderona gracilicornis (Scheerp.): 4 — maxilla, 5 — labium, 6 — right mandible, 7 — labrum, 8 — mentum, 9 — shape of elytron, 10 — pronotal pubescence, 11 — tergite VIII." httpUri="https://zenodo.org/record/175637/files/figure.png" pageId="7" pageNumber="8">Fig. 7</figureCitation>
) transverse, moderately deeply arcuately emarginate apically and with characteristic arrangement of setae. Mentum (
<figureCitation id="1337D4A6FFAA9475FB6F76BD66AE7B8F" box="[1152,1222,1969,1995]" captionStart="FIGURES 4 11" captionStartId="9.[151,269,1603,1627]" captionTargetBox="[303,1284,194,1585]" captionTargetId="figure@9.[295,1291,183,1592]" captionTargetPageId="9" captionText="FIGURES 4 11. Ischnopoderona gracilicornis (Scheerp.): 4 — maxilla, 5 — labium, 6 — right mandible, 7 — labrum, 8 — mentum, 9 — shape of elytron, 10 — pronotal pubescence, 11 — tergite VIII." httpUri="https://zenodo.org/record/175637/files/figure.png" pageId="7" pageNumber="8">Fig. 8</figureCitation>
) trapezoidal, with 4 long setae on each side, apical margin straight, anterior angles rounded.
</paragraph>
<caption id="DF7398ABFFA5947AFF78772A615C7A7A" box="[151,820,1574,1598]" pageId="8" pageNumber="9">
<paragraph id="8BB3C823FFA5947AFF78772A615C7A7A" blockId="8.[151,820,1574,1598]" box="[151,820,1574,1598]" pageId="8" pageNumber="9">
<emphasis id="B9781431FFA5947AFF78772A63717A7A" bold="true" box="[151,281,1574,1598]" pageId="8" pageNumber="9">FIGURE 3.</emphasis>
<taxonomicName id="4C0CB3A0FFA5947AFECF772B60BD7A7A" authority="Scheerp." authorityName="Scheerp." box="[288,725,1575,1598]" class="Insecta" family="Staphylinidae" genus="Ischnopoderona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="gracilicornis">
<emphasis id="B9781431FFA5947AFECF772B60347A7A" box="[288,604,1575,1598]" italics="true" pageId="8" pageNumber="9">Ischnopoderona gracilicornis</emphasis>
(Scheerp.)
</taxonomicName>
: habitus.
</paragraph>
</caption>
<paragraph id="8BB3C823FFA5947AFF29777E63637BB0" blockId="8.[151,1436,1650,2036]" pageId="8" pageNumber="9">
Thorax. Pronotum quadrate to longer than wide, moderately to distinctly convex, glossy, widest in apical third, lateral sides sinuately narrowed to hind angles. Pubescence at midline of three
<typeStatus id="54B77681FFA5947AFB63779666A67AF0" box="[1164,1230,1690,1716]" pageId="8" pageNumber="9">types</typeStatus>
: directed entirely anteriorly, directed posteriorly in apical 1/4 and anteriorly in basal ¾ or directed posteriorly in apical half and anteriorly in basal half; pubescence at postero-lateral parts directed obliquely laterally or straight anteriorly (
<figureCitation id="1337D4A6FFA5947AFF70761E62907B68" box="[159,248,1810,1836]" captionStart="FIGURES 4 11" captionStartId="9.[151,269,1603,1627]" captionTargetBox="[303,1284,194,1585]" captionTargetId="figure@9.[295,1291,183,1592]" captionTargetPageId="9" captionText="FIGURES 4 11. Ischnopoderona gracilicornis (Scheerp.): 4 — maxilla, 5 — labium, 6 — right mandible, 7 — labrum, 8 — mentum, 9 — shape of elytron, 10 — pronotal pubescence, 11 — tergite VIII." httpUri="https://zenodo.org/record/175637/files/figure.png" pageId="8" pageNumber="9">Fig. 10</figureCitation>
). Hypomera fully visible in lateral view. Mesocoxae varies from narrowly to widely separated (
<figureCitation id="1337D4A6FFA5947AFA8E761E628C7B10" captionStart="FIGURES 12 17" captionStartId="10.[151,269,1532,1556]" captionTargetBox="[403,1195,196,1503]" captionTargetId="figure@10.[390,1209,182,1516]" captionTargetPageId="10" captionText="FIGURES 12 17. Variability of mesosternum and metasternum of Ischnopoderona sp.: 12 — I. longicollis (Bernh.), 13 — I. lucidula sp. n., 14 — I. nigerrima (Cam.), 15 — I. namibensis (Pace), 16 — I. gracilicornis (Scheerp.), 17 — I. madecassa (Cam.)" httpUri="https://zenodo.org/record/175638/files/figure.png" pageId="8" pageNumber="9">Figs. 1217</figureCitation>
). Mesosternal process varies from narrowly triangular, rounded at apex, extending to 1/2 of mesocoxae and nearly reaching apex of metasternal process (
<figureCitation id="1337D4A6FFA5947AFD30766E61097B38" box="[735,865,1890,1916]" captionStart="FIGURES 12 17" captionStartId="10.[151,269,1532,1556]" captionTargetBox="[403,1195,196,1503]" captionTargetId="figure@10.[390,1209,182,1516]" captionTargetPageId="10" captionText="FIGURES 12 17. Variability of mesosternum and metasternum of Ischnopoderona sp.: 12 — I. longicollis (Bernh.), 13 — I. lucidula sp. n., 14 — I. nigerrima (Cam.), 15 — I. namibensis (Pace), 16 — I. gracilicornis (Scheerp.), 17 — I. madecassa (Cam.)" httpUri="https://zenodo.org/record/175638/files/figure.png" pageId="8" pageNumber="9">Fig. 1213</figureCitation>
) throughout intermediate states (
<figureCitation id="1337D4A6FFA5947AFB08766E671D7B38" box="[1255,1397,1890,1916]" captionStart="FIGURES 12 17" captionStartId="10.[151,269,1532,1556]" captionTargetBox="[403,1195,196,1503]" captionTargetId="figure@10.[390,1209,182,1516]" captionTargetPageId="10" captionText="FIGURES 12 17. Variability of mesosternum and metasternum of Ischnopoderona sp.: 12 — I. longicollis (Bernh.), 13 — I. lucidula sp. n., 14 — I. nigerrima (Cam.), 15 — I. namibensis (Pace), 16 — I. gracilicornis (Scheerp.), 17 — I. madecassa (Cam.)" httpUri="https://zenodo.org/record/175638/files/figure.png" pageId="8" pageNumber="9">Figs. 1416</figureCitation>
) to wide, broadly rounded at apex, extending to about 1/3 of mesocoxae and separated from long, rectangular metasternal process by short, transverse isthmus (
<figureCitation id="1337D4A6FFA5947AFD3176BE615D7B88" box="[734,821,1970,1996]" captionStart="FIGURES 12 17" captionStartId="10.[151,269,1532,1556]" captionTargetBox="[403,1195,196,1503]" captionTargetId="figure@10.[390,1209,182,1516]" captionTargetPageId="10" captionText="FIGURES 12 17. Variability of mesosternum and metasternum of Ischnopoderona sp.: 12 — I. longicollis (Bernh.), 13 — I. lucidula sp. n., 14 — I. nigerrima (Cam.), 15 — I. namibensis (Pace), 16 — I. gracilicornis (Scheerp.), 17 — I. madecassa (Cam.)" httpUri="https://zenodo.org/record/175638/files/figure.png" pageId="8" pageNumber="9">Fig. 17</figureCitation>
). Mesocoxal cavities not or margined posteriorly by fine bead.
</paragraph>
<caption id="DF7398ABFFA4947BFF78774F61C67A3B" httpUri="https://zenodo.org/record/175637/files/figure.png" pageId="9" pageNumber="10" targetBox="[303,1284,194,1585]" targetPageId="9">
<paragraph id="8BB3C823FFA4947BFF78774F61C67A3B" blockId="9.[151,1436,1603,1663]" pageId="9" pageNumber="10">
<emphasis id="B9781431FFA4947BFF78774F63277A1F" bold="true" box="[151,335,1603,1627]" pageId="9" pageNumber="10">FIGURES 411.</emphasis>
<taxonomicName id="4C0CB3A0FFA4947BFEB8774F61667A1E" authority="Scheerp." authorityName="Scheerp." box="[343,782,1603,1626]" class="Insecta" family="Staphylinidae" genus="Ischnopoderona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="gracilicornis">
<emphasis id="B9781431FFA4947BFEB8774F60FC7A1E" box="[343,660,1603,1626]" italics="true" pageId="9" pageNumber="10">Ischnopoderona gracilicornis</emphasis>
(Scheerp.)
</taxonomicName>
: 4—maxilla, 5—labium, 6—right mandible, 7—labrum, 8— mentum, 9—shape of elytron, 10—pronotal pubescence, 11—tergite VIII.
</paragraph>
</caption>
<paragraph id="8BB3C823FFA4947BFF2977BA63177ABC" blockId="9.[151,1437,1718,2024]" pageId="9" pageNumber="10">
Elytra. Quadrate, lateral sides distinctly arcuate, widest in the middle, postero­lateral angles distinctly emarginate (
<figureCitation id="1337D4A6FFA4947BFEC777D263187ABC" box="[296,368,1758,1784]" captionStart="FIGURES 4 11" captionStartId="9.[151,269,1603,1627]" captionTargetBox="[303,1284,194,1585]" captionTargetId="figure@9.[295,1291,183,1592]" captionTargetPageId="9" captionText="FIGURES 4 11. Ischnopoderona gracilicornis (Scheerp.): 4 — maxilla, 5 — labium, 6 — right mandible, 7 — labrum, 8 — mentum, 9 — shape of elytron, 10 — pronotal pubescence, 11 — tergite VIII." httpUri="https://zenodo.org/record/175637/files/figure.png" pageId="9" pageNumber="10">Fig. 9</figureCitation>
).
</paragraph>
<paragraph id="8BB3C823FFA4947BFF29760A66207B0C" blockId="9.[151,1437,1718,2024]" pageId="9" pageNumber="10">Legs. Elongate, metatarsus long, ranging from 3/4 to 4/5 the length of metatibia; basal segment of metatarsus longer than following two tarsal segments combined; tarsal formula 4­5­5.</paragraph>
<paragraph id="8BB3C823FFA4947BFF29765A60287BAC" blockId="9.[151,1437,1718,2024]" pageId="9" pageNumber="10">Abdomen. Distinctly constricted at base, widened posteriorly, widest at level of tergites V and VI, tergites IIIV distinctly transversely impressed at base, at least first impressions coarsely and densely punctate, tergite V distinctly longer than VII, sternites IIIV moderately to strongly transversely impressed at base, impressions coarsely and densely punctate.</paragraph>
<caption id="DF7398ABFFA79478FF7874F0634C7A1A" httpUri="https://zenodo.org/record/175638/files/figure.png" pageId="10" pageNumber="11" targetBox="[403,1195,196,1503]" targetPageId="10">
<paragraph id="8BB3C823FFA79478FF7874F0634C7A1A" blockId="10.[151,1436,1532,1630]" pageId="10" pageNumber="11">
<emphasis id="B9781431FFA79478FF7874F063097A50" bold="true" box="[151,353,1532,1556]" pageId="10" pageNumber="11">FIGURES 1217.</emphasis>
Variability of mesosternum and metasternum of
<taxonomicName id="4C0CB3A0FFA79478FC6674F0665D7A57" box="[905,1077,1532,1555]" class="Insecta" family="Staphylinidae" genus="Ischnopoderona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B9781431FFA79478FC6674F0665D7A57" box="[905,1077,1532,1555]" italics="true" pageId="10" pageNumber="11">Ischnopoderona</emphasis>
</taxonomicName>
sp.: 12—
<taxonomicName id="4C0CB3A0FFA79478FB4974F067F07A57" authority="Bernh." authorityName="Bernh." box="[1190,1432,1532,1555]" class="Insecta" family="Staphylinidae" genus="Ischnopoderona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="longicollis">
<emphasis id="B9781431FFA79478FB4974F067587A57" box="[1190,1328,1532,1555]" italics="true" pageId="10" pageNumber="11">I. longicollis</emphasis>
(Bernh.)
</taxonomicName>
, 13—
<taxonomicName id="4C0CB3A0FFA79478FF23772E63527A7D" box="[204,314,1570,1593]" class="Insecta" family="Staphylinidae" genus="Ischnopoderona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="lucidula" status="sp. nov.">
<emphasis id="B9781431FFA79478FF23772E63527A7D" box="[204,314,1570,1593]" italics="true" pageId="10" pageNumber="11">I. lucidula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24BA94AFFA79478FEAF772D63137A7D" box="[320,379,1569,1593]" pageId="10" pageNumber="11" rank="species">
<emphasis id="B9781431FFA79478FEAF772D63137A7D" bold="true" box="[320,379,1569,1593]" pageId="10" pageNumber="11">sp. n.</emphasis>
</taxonomicNameLabel>
, 14—
<taxonomicName id="4C0CB3A0FFA79478FE51772E60FA7A7C" authority="Cam." authorityName="Cam." box="[446,658,1569,1593]" class="Insecta" family="Staphylinidae" genus="Ischnopoderona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="nigerrima">
<emphasis id="B9781431FFA79478FE51772E60577A7D" box="[446,575,1570,1593]" italics="true" pageId="10" pageNumber="11">I. nigerrima</emphasis>
(Cam.)
</taxonomicName>
, 15—
<taxonomicName id="4C0CB3A0FFA79478FD3D772E61C57A7C" authority="Pace" authorityName="Pace" box="[722,941,1569,1593]" class="Insecta" family="Staphylinidae" genus="Ischnopoderona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="namibensis">
<emphasis id="B9781431FFA79478FD3D772E61087A7D" box="[722,864,1570,1593]" italics="true" pageId="10" pageNumber="11">I. namibensis</emphasis>
(Pace)
</taxonomicName>
, 16—
<taxonomicName id="4C0CB3A0FFA79478FC03772E676C7A7C" authority="Scheerp." authorityName="Scheerp." box="[1004,1284,1569,1593]" class="Insecta" family="Staphylinidae" genus="Ischnopoderona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="gracilicornis">
<emphasis id="B9781431FFA79478FC03772E66E47A7D" box="[1004,1164,1570,1593]" italics="true" pageId="10" pageNumber="11">I. gracilicornis</emphasis>
(Scheerp.)
</taxonomicName>
, 17—
<taxonomicName id="4C0CB3A0FFA79478FAAC772E634C7A1A" authority="Cam." authorityName="Cam." class="Insecta" family="Staphylinidae" genus="Ischnopoderona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="madecassa">
<emphasis id="B9781431FFA79478FAAC772E62BA7A1A" italics="true" pageId="10" pageNumber="11">I. madecassa</emphasis>
(Cam.)
</taxonomicName>
</paragraph>
</caption>
<paragraph id="8BB3C823FFA79478FF287798672A7B0A" blockId="10.[151,1437,1684,2030]" pageId="10" pageNumber="11">
Median lobe of aedeagus short to relatively long and slender, straight or more­or­less bent ventrally in lateral view; gradually narrowed to apex or subparallel­sided and abruptly narrowed apically in ventral view; apex of median lobe rounded, hooked or constricted apically (lateral view); ventral process large and prominent, differs in size and shape between species; internal sac with a few pairs of sclerites differing in size and shape. A combination of these characters is most useful for differentiating species of
<taxonomicName id="4C0CB3A0FFA79478FB92763867527B09" box="[1149,1338,1844,1869]" class="Insecta" family="Staphylinidae" genus="Ischnopoderona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B9781431FFA79478FB92763867527B09" box="[1149,1338,1844,1869]" italics="true" pageId="10" pageNumber="11">Ischnopoderona</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3169BA8FFA79479FF287650629E7C9E" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11" type="discussion">
<paragraph id="8BB3C823FFA79478FF28765067097B82" blockId="10.[151,1437,1684,2030]" pageId="10" pageNumber="11">
<emphasis id="B9781431FFA79478FF28765063287B32" bold="true" box="[199,320,1884,1910]" pageId="10" pageNumber="11">Remarks.</emphasis>
The examination of the
<typeStatus id="54B77681FFA79478FD9B765060CE7B32" box="[628,678,1884,1910]" pageId="10" pageNumber="11">type</typeStatus>
species of
<taxonomicName id="4C0CB3A0FFA79478FCD97650619B7B31" box="[822,1011,1884,1909]" class="Insecta" family="Staphylinidae" genus="Ischnopoderona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B9781431FFA79478FCD97650619B7B31" box="[822,1011,1884,1909]" italics="true" pageId="10" pageNumber="11">Ischnopoderona</emphasis>
</taxonomicName>
as well as other species classified within this genus, clearly show that they form a well­defined monophyletic group characterised by several synapomorphies (see phylogenetic part). Therefore,
<taxonomicName id="4C0CB3A0FFA79478FD1376A061D27B81" box="[764,954,1964,1989]" class="Insecta" family="Staphylinidae" genus="Ischnopoderona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B9781431FFA79478FD1376A061D27B81" box="[764,954,1964,1989]" italics="true" pageId="10" pageNumber="11">Ischnopoderona</emphasis>
</taxonomicName>
is here regarded as a distinct genus.
</paragraph>
<paragraph id="8BB3C823FFA79479FF2976D8629E7C9E" blockId="10.[151,1437,1684,2030]" lastBlockId="11.[151,1434,152,218]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
The genus
<taxonomicName id="4C0CB3A0FFA79478FEB776D8607E7BA9" box="[344,534,2004,2029]" class="Insecta" family="Staphylinidae" genus="Ischnopoderona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B9781431FFA79478FEB776D8607E7BA9" box="[344,534,2004,2029]" italics="true" pageId="10" pageNumber="11">Ischnopoderona</emphasis>
</taxonomicName>
is closely related to the genera
<taxonomicName id="4C0CB3A0FFA79478FC2B76D866967BAA" authority="Erichson, 1837" authorityName="Erichson" authorityYear="1837" box="[964,1278,2004,2030]" class="Insecta" family="Staphylinidae" genus="Tachyusa" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B9781431FFA79478FC2B76D8665A7BA9" box="[964,1074,2004,2029]" italics="true" pageId="10" pageNumber="11">Tachyusa</emphasis>
<bibRefCitation id="EF9DB5D2FFA79478FBAD76D866967BAA" author="Erichson" box="[1090,1278,2004,2030]" pageId="10" pageNumber="54" refString="Erichson, W. F. (1837) Die Kafer der Mark Brandenburg. Bd. 1, Abt. 1. F. H. Morion, Berlin, 384 pp." type="book" year="1837">Erichson, 1837</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C0CB3A0FFA79479FAFC76D863237CF6" authority="Stephens, 1835" authorityName="Stephens" authorityYear="1835" class="Insecta" family="Staphylinidae" genus="Ischnopoda" higherTaxonomySource="CoL" kingdom="Animalia" lastPageId="11" lastPageNumber="12" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B9781431FFA79478FAFC76D867F37BA9" box="[1299,1435,2004,2029]" italics="true" pageId="10" pageNumber="11">Ischnopoda</emphasis>
Stephens, 1835
</taxonomicName>
and
<taxonomicName id="4C0CB3A0FFA69479FE6A719460C47CF6" authority="Thomson, 1858" authorityName="Thomson" authorityYear="1858" box="[389,684,152,178]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="B9781431FFA69479FE6A719463807CF5" box="[389,488,152,177]" italics="true" pageId="11" pageNumber="12">Gnypeta</emphasis>
Thomson, 1858
</taxonomicName>
. It can be easily distinguished from these genera by the following key:
</paragraph>
</subSubSection>
<subSubSection id="C3169BA8FFA69479FF78701F67F27868" pageId="11" pageNumber="12" type="key">
<key id="FF9D9F12FFA69479FF78701F67F27868" pageId="11" pageNumber="12">
<keyStep id="30F8D086FFA69479FF78701F67F27E45" pageId="11" pageNumber="12">
<paragraph id="8BB3C823FFA69479FF78701F67F47DC5" blockId="11.[151,1437,275,1068]" pageId="11" pageNumber="12">
<keyLead id="30FD6B16FFA69479FF78701F67F47DC5" pageId="11" pageNumber="12">1. Abdomen distinctly constricted at base; tergite V moderately transverse, 1.21.8 times wider than long; elytral postero­lateral angles distinctly emarginated; elytral lateral sides strongly arcuate and widest in the middle; sternites IIIV strongly transversely impressed at base...................................... 2</keyLead>
</paragraph>
<paragraph id="8BB3C823FFA69479FF78709F67F27E45" blockId="11.[151,1437,275,1068]" pageId="11" pageNumber="12">
<keyLead id="30FD6B16FFA69479FF78709F67F27E45" pageId="11" pageNumber="12"> Abdomen weakly constricted at base; tergite V strongly transverse, at least 2.0 times wider than long; elytral postero­lateral angles slightly emarginated; elytral lateral sides slightly arcuate and widest in posterior half; sternites IIIV slightly transversely impressed at base .................................................. 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F8D086FFA69479FF78731F67F17F39" pageId="11" pageNumber="12">
<paragraph id="8BB3C823FFA69479FF78731F67F27EED" blockId="11.[151,1437,275,1068]" pageId="11" pageNumber="12">
<keyLead id="30FD6B16FFA69479FF78731F67F27EED" pageId="11" pageNumber="12">
2(1). Ligula bifid in apical half, lobes divergent apically; maxillary palpus 3 clearly widened apically; apical margin of mentum distinctly concave; mandible with large molar tooth on right mandible; antennae clearly increasing in width towards apex; abdomen more or less parallel sided; abdominal transverse impressions with longitudinal ridges between punctures
<emphasis id="B9781431FFA69479FBEE738367437EEC" box="[1025,1323,655,680]" italics="true" pageId="11" pageNumber="12">
.........................
<taxonomicName id="4C0CB3A0FFA69479FB52738367437EEC" ID-CoL="63TGL" authority="Erichson" authorityName="Erichson" box="[1213,1323,655,680]" class="Insecta" family="Staphylinidae" genus="Tachyusa" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">Tachyusa</taxonomicName>
</emphasis>
Erichson
</keyLead>
</paragraph>
<paragraph id="8BB3C823FFA69479FF7873B767F17F39" blockId="11.[151,1437,275,1068]" pageId="11" pageNumber="12">
<keyLead id="30FD6B16FFA69479FF7873B767F17F39" pageId="11" pageNumber="12">
Ligula divided to base, or nearly so, lobes parallel­sided; maxillary palpus 3 weakly widened apically; apical margin of mentum not concave, straight; mandible with small, indistinct molar tooth on right mandible; antennae weakly increasing in width towards apex; abdomen distinctly widened posteriorly; abdominal transverse impressions without longitudinal ridges between punctures .......................
<emphasis id="B9781431FFA69479FF16726F66327F38" box="[249,1114,867,892]" italics="true" pageId="11" pageNumber="12">......................................................................................................................</emphasis>
<taxonomicName id="4C0CB3A0FFA69479FBB2726F67F17F39" ID-CoL="8X7H5" authority="(Scheerp.)" authorityName="Scheerp." baseAuthorityName="Scheerp." box="[1117,1433,867,893]" class="Insecta" family="Staphylinidae" genus="Ischnopoderona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="B9781431FFA69479FBB2726F67727F38" box="[1117,1306,867,892]" italics="true" pageId="11" pageNumber="12">Ischnopoderona</emphasis>
(Scheerp.)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F8D086FFA69479FF78728367F27868" pageId="11" pageNumber="12">
<paragraph id="8BB3C823FFA69479FF78728367F17F90" blockId="11.[151,1437,275,1068]" pageId="11" pageNumber="12">
<keyLead id="30FD6B16FFA69479FF78728367F17F90" pageId="11" pageNumber="12">
3(1). Ligula bifid in apical half, lobes divergent apically; galea length slightly shorter than lacinia; basal segment of metatarsus longer than the following two tarsal segments combined.
<taxonomicName id="4C0CB3A0FFA69479FB4F72B767F17F90" authority="Stephens" authorityName="Stephens" box="[1184,1433,954,980]" class="Insecta" family="Staphylinidae" genus="Ischnopoda" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="B9781431FFA69479FB4F72B767417F90" box="[1184,1321,955,980]" italics="true" pageId="11" pageNumber="12">Ischnopoda</emphasis>
Stephens
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BB3C823FFA69479FF7872EA67F27868" blockId="11.[151,1437,275,1068]" pageId="11" pageNumber="12">
<keyLead id="30FD6B16FFA69479FF7872EA67F27868" pageId="11" pageNumber="12">
Ligula divided to base, or near base, lobes parallel­sided; galea subequal in length to lacinia; basal segment of metatarsus shorter than the following two tarsal segments combined
<emphasis id="B9781431FFA69479FB74751F674C7868" box="[1179,1316,1043,1068]" italics="true" pageId="11" pageNumber="12">
.....
<taxonomicName id="4C0CB3A0FFA69479FB51751F674C7868" ID-CoL="8VRPT" authority="Thomson" authorityName="Thomson" box="[1214,1316,1043,1068]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">Gnypeta</taxonomicName>
</emphasis>
Thomson
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>