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<document ID-DOI="10.11646/phytotaxa.524.2.1" ID-GBIF-Dataset="7f01db51-02e7-49aa-adc5-191d90c14275" ID-ISSN="1179-3163" ID-Zenodo-Dep="5642036" checkinTime="1635920993234" checkinUser="plazi" docAuthor="Salgado, Arthur Edward" docDate="2021" docId="03A5792B772F707AFF1FF89DFA10FC9D" docLanguage="en" docName="phytotaxa.524.2.1.pdf" docOrigin="Phytotaxa 524 (2)" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.5:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="5" docTitle="Asplenium pahli Salgado, Wenzel 2021, sp. nov." docType="treatment" docVersion="5" lastPageNumber="75" masterDocId="FF9C0153772D707EFF97FA29FFBAFFE4" masterDocTitle="Taxonomic and nomenclatural notes on Philippine ferns. IV. Asplenium pahli (Polypodiales, Aspleniineae, Aspleniaceae), a new species from Leyte, Philippines" masterLastPageNumber="84" masterPageNumber="71" pageNumber="73" updateTime="1673526947003" updateUser="felipe" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Taxonomic and nomenclatural notes on Philippine ferns. IV. Asplenium pahli (Polypodiales, Aspleniineae, Aspleniaceae), a new species from Leyte, Philippines</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Salgado, Arthur Edward</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-11-03</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>524</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
</mods:detail>
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<mods:start>71</mods:start>
<mods:end>84</mods:end>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/phytotaxa.524.2.1</mods:identifier>
<mods:identifier type="GBIF-Dataset">7f01db51-02e7-49aa-adc5-191d90c14275</mods:identifier>
<mods:identifier type="ISSN">1179-3163</mods:identifier>
<mods:identifier type="Zenodo-Dep">5642036</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.5642042" ID-GBIF-Taxon="190304841" ID-Zenodo-Dep="5642042" LSID="urn:lsid:plazi:treatment:03A5792B772F707AFF1FF89DFA10FC9D" httpUri="http://treatment.plazi.org/id/03A5792B772F707AFF1FF89DFA10FC9D" lastPageId="4" lastPageNumber="75" pageId="2" pageNumber="73">
<subSubSection box="[136,649,692,719]" pageId="2" pageNumber="73" type="nomenclature">
<paragraph blockId="2.[136,649,692,719]" box="[136,649,692,719]" pageId="2" pageNumber="73">
<heading box="[136,649,692,719]" fontSize="11" level="2" pageId="2" pageNumber="73" reason="3">
<taxonomicName authority="Salgado" authorityName="Salgado, Wenzel" authorityYear="2021" box="[136,427,692,719]" class="Polypodiopsida" family="Aspleniaceae" genus="Asplenium" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="73" phylum="Tracheophyta" rank="species" species="pahli" status="sp. nov.">
<emphasis bold="true" box="[136,327,692,719]" italics="true" pageId="2" pageNumber="73">Asplenium pahli</emphasis>
Salgado
</taxonomicName>
,
<emphasis box="[438,523,693,718]" italics="true" pageId="2" pageNumber="73">
<taxonomicNameLabel box="[438,523,693,718]" pageId="2" pageNumber="73" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation box="[537,636,692,718]" captionStart="FIGURE 2" captionStartId="3.[136,229,1926,1948]" captionTargetBox="[163,1421,205,1897]" captionTargetId="figure-18@3.[151,1436,190,1902]" captionTargetPageId="3" captionText="FIGURE 2. Asplenium pahli Salgado, Wenzel 572, holotype (P). Note the plantlets near the apex of two fronds. Muséum national dhistoire naturelle, Paris, France. Collection: Vascular plants P00370339. License CC Attribution 4.0 International.Available at: https:// science.mnhn.fr/institution/mnhn/collection/p/item/p00370339" figureDoi="http://doi.org/10.5281/zenodo.5642044" httpUri="https://zenodo.org/record/5642044/files/figure.png" pageId="2" pageNumber="73">Figure 2</figureCitation>
).
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="73" type="materials_examined">
<paragraph blockId="2.[136,1452,768,970]" pageId="2" pageNumber="73">
<emphasis bold="true" box="[136,218,768,789]" pageId="2" pageNumber="73">Type:—</emphasis>
<materialsCitation ID-GBIF-Occurrence="3400200309" collectingDate="1914-02-05" collectionCode="C, A, F" country="Philippines" elevation="500" location="Leyte" pageId="2" pageNumber="73" specimenCode="P00370339, P00370338, P00370340, C0364253" specimenCount="1" stateProvince="Southern Leyte" typeStatus="holotype">
<collectingCountry box="[218,362,768,789]" name="Philippines" pageId="2" pageNumber="73">PHILIPPINES</collectingCountry>
,
<collectingRegion box="[373,430,768,789]" country="Philippines" name="Southern Leyte" pageId="2" pageNumber="73">Leyte</collectingRegion>
,
<quantity box="[440,504,768,790]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="73" unit="m" value="500.0">
<elevation box="[440,504,768,790]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="73" unit="m" value="500.0">500 m</elevation>
</quantity>
,
<date box="[515,678,768,790]" pageId="2" pageNumber="73" value="1914-02-05">
<collectingDate box="[515,678,768,790]" pageId="2" pageNumber="73" value="1914-02-05">5 February 1914</collectingDate>
</date>
.
<emphasis box="[688,848,768,789]" italics="true" pageId="2" pageNumber="73">
<collectionCode box="[688,706,768,789]" country="Denmark" name="University of Copenhagen" pageId="2" pageNumber="73" type="Herbarium">C</collectionCode>
.
<collectionCode box="[710,728,768,789]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="2" pageNumber="73" type="Herbarium">A</collectionCode>
. Wenzel 572
</emphasis>
(
<typeStatus box="[862,948,768,790]" pageId="2" pageNumber="73">holotype</typeStatus>
<specimenCode box="[954,1069,768,789]" collectionCode="P" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="2" pageNumber="73" type="Herbarium">P00370339</specimenCode>
! designated here, isotypes
<specimenCode box="[1328,1445,768,789]" collectionCode="P" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="2" pageNumber="73" type="Herbarium">P00370338</specimenCode>
!,
<specimenCode box="[189,306,804,825]" collectionCode="P" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="2" pageNumber="73" type="Herbarium">P00370340</specimenCode>
!,
<specimenCode box="[319,419,804,826]" collectionCode="C" country="Denmark" name="University of Copenhagen" pageId="2" pageNumber="73" type="Herbarium">C0364253</specimenCode>
<collectionCode box="[419,432,804,826]" country="USA" lsid="urn:lsid:biocol.org:col:15707" name="Field Museum of Natural History, Botany Department" pageId="2" pageNumber="73" type="Herbarium">F</collectionCode>
(
<collectionCode box="[447,465,804,826]" country="USA" lsid="urn:lsid:biocol.org:col:15707" name="Field Museum of Natural History, Botany Department" pageId="2" pageNumber="73" type="Herbarium">F</collectionCode>
!),
</materialsCitation>
<collectingCountry box="[487,519,804,826]" name="United States of America" pageId="2" pageNumber="73">US</collectingCountry>
01516038!).
</paragraph>
</subSubSection>
<subSubSection lastPageId="4" lastPageNumber="75" pageId="2" pageNumber="73" type="diagnosis">
<paragraph blockId="2.[136,1452,768,970]" pageId="2" pageNumber="73">
<emphasis bold="true" box="[136,243,839,861]" pageId="2" pageNumber="73">Diagnosis:</emphasis>
<taxonomicName authorityName="Salgado, Wenzel" authorityYear="2021" box="[267,424,840,861]" class="Polypodiopsida" family="Aspleniaceae" genus="Asplenium" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="73" phylum="Tracheophyta" rank="species" species="pahli">
<emphasis box="[267,424,840,861]" italics="true" pageId="2" pageNumber="73">Asplenium pahli</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="C. Presl, Cuming" authorityYear="1849" box="[474,626,840,861]" class="Polypodiopsida" family="Aspleniaceae" genus="Asplenium" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="73" phylum="Tracheophyta" rank="species" species="anisodontum">
<emphasis box="[474,626,840,861]" italics="true" pageId="2" pageNumber="73">A. anisodontum</emphasis>
</taxonomicName>
have a proliferous rachis but differ in the pinna shape and serration.
<taxonomicName authorityName="Salgado, Wenzel" authorityYear="2021" box="[1304,1381,840,861]" class="Polypodiopsida" family="Aspleniaceae" genus="Asplenium" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="73" phylum="Tracheophyta" rank="species" species="pahli">
<emphasis box="[1304,1381,840,861]" italics="true" pageId="2" pageNumber="73">A. pahli</emphasis>
</taxonomicName>
differs from
<taxonomicName authorityName="G. Forster" authorityYear="1786" box="[243,366,876,897]" class="Polypodiopsida" family="Aspleniaceae" genus="Asplenium" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="73" phylum="Tracheophyta" rank="species" species="caudatum">
<emphasis box="[243,366,876,897]" italics="true" pageId="2" pageNumber="73">A. caudatum</emphasis>
</taxonomicName>
,
<emphasis box="[379,797,876,897]" italics="true" pageId="2" pageNumber="73">
<taxonomicName authorityName="Lamarck" authorityYear="1786" box="[379,494,876,897]" class="Polypodiopsida" family="Aspleniaceae" genus="Asplenium" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="73" phylum="Tracheophyta" rank="species" species="falcatum">A. falcatum</taxonomicName>
,
<taxonomicName authorityName="Mett. ex Kuhn" authorityYear="1868" box="[504,657,876,897]" class="Polypodiopsida" family="Aspleniaceae" genus="Asplenium" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="73" phylum="Tracheophyta" rank="species" species="gueinzianum">A. gueinzianum</taxonomicName>
,
<taxonomicName authorityName="Wallich" authorityYear="1828" box="[668,797,876,897]" class="Polypodiopsida" family="Aspleniaceae" genus="Asplenium" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="73" phylum="Tracheophyta" rank="species" species="planicaule">A. planicaule</taxonomicName>
</emphasis>
and
<taxonomicName box="[846,963,876,897]" class="Polypodiopsida" family="Aspleniaceae" genus="Asplenium" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="73" phylum="Tracheophyta" rank="species" species="polyodon">
<emphasis box="[846,963,876,897]" italics="true" pageId="2" pageNumber="73">A. polyodon</emphasis>
</taxonomicName>
in the proliferous rachis, the pinna margins being irregularly serrate, sinuses of different depth and width, fewer sori distant from the costa, not contiguous, and straight or slightly falcate.
</paragraph>
<paragraph blockId="2.[136,1452,1016,2087]" pageId="2" pageNumber="73">
Terrestrial. Rhizome short, erect,
<quantity box="[518,645,1016,1043]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="2" pageNumber="73" unit="cm" value="0.45" valueMax="0.5" valueMin="0.4">0.40.5 cm</quantity>
thick, paleae and palea bases few, scattered, clathrate, black, lanceate, margin entire, &lt;
<quantity box="[328,386,1052,1078]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="73" unit="mm" value="1.0">1mm</quantity>
long, upper part of the paleae often broken off, cells small, polygonal. Roots many, crowded at the base of fronds, brown, thin, fragile when dried. Fronds erect, pinnate, crowded at the apex of the rhizome, spirally arranged,
<quantity box="[343,456,1124,1151]" metricMagnitude="-1" metricUnit="m" metricValue="3.5" metricValueMax="4.5" metricValueMin="2.5" pageId="2" pageNumber="73" unit="cm" value="35.0" valueMax="45.0" valueMin="25.0">2545 cm</quantity>
long. Stipes 10.5(13.017.0)21.0 cm long, adaxial side grooved, abaxial side rounded, greenish, glabrous or glabrescent, with sparse paleae, paleae bicolorous, base and margin light brown, central area black, lanceate, ≤
<quantity box="[334,394,1196,1223]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="73" unit="mm" value="1.0">1 mm</quantity>
long, adpressed against the surface, margin entire, irregular with an occasional acute projection formed by a clathrate cell wall, palea apex acute. Lamina oblong,
<quantity box="[879,991,1232,1259]" metricMagnitude="-1" metricUnit="m" metricValue="2.5" metricValueMax="3.5" metricValueMin="1.5" pageId="2" pageNumber="73" unit="cm" value="25.0" valueMax="35.0" valueMin="15.0">1535 cm</quantity>
long, (5.5)
<quantity box="[1115,1221,1232,1259]" metricMagnitude="-2" metricUnit="m" metricValue="7.25" metricValueMax="8.0" metricValueMin="6.5" pageId="2" pageNumber="73" unit="cm" value="7.25" valueMax="8.0" valueMin="6.5">6.58 cm</quantity>
wide, base truncate, pinnate, chartaceous, 1215 alternating pinna pairs, pinnae slightly reduced at near apex. Rachis proliferous, abaxial side rounded, adaxial side with a wide U-shaped groove with prominent ridges, greenish, glabrescent, with adpressed small paleae, ≤
<quantity box="[307,367,1340,1367]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="73" unit="mm" value="1.0">1 mm</quantity>
long, lanceate, black, margin entire, apex acute, paleae on both sides of the rachis. Pinnae 3.24.0 cm long, 1.0
<quantity box="[294,370,1376,1403]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="2" pageNumber="73" unit="cm" value="1.3">1.3 cm</quantity>
wide in the middle, noncontiguous, alternating, generally trapezoid, stalked to sessile, stalk ≤
<quantity metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="73" unit="mm" value="1.0">1 mm</quantity>
long, naked or with few paleae, lamina adaxial side glabrous, abaxial side glabrous or with a few reduced paleae on the costa, costa prominent of both sides, adaxial side grooved with prominent ridges gradually disappearing toward the pinna apex, abaxial side rounded, basal pinna pair sub-opposite or alternating, generally oblong, of the same length and shape as median pinnae or slightly smaller; pinna base basiscopic side cuneate, acroscopic side cuneate to broadly cuneate, pinna apex acute, pinna margins irregularly serrate, sinuses of different depth and width, acroscopic margin with 56 oblique widely separated teeth, larger teeth with 23 apical teeth, teeth blunt or acute, basiscopic margin serration similar to acroscopic margin, 35 oblique widely separated teeth, lamina apical segment usually missing, when present deformed with a proliferous bud at the insertion with the rachis. Venation free, costa visible with prominent ridges, 57 pairs of veins oblique to the costa, abaxial side veins not visible, adaxial side veins hardly visible, once forking, ending near the margin; hydathodes absent. Sori in a single row distal from pinna base, on both sides of the costa, originating about
<quantity box="[558,626,1772,1799]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="73" unit="mm" value="1.0">1 mm</quantity>
from costa, oblique, widely separated, 36 sori on the acroscopic side, 25 on the basiscopic side,
<quantity box="[454,549,1808,1835]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="2" pageNumber="73" unit="mm" value="6.0" valueMax="7.0" valueMin="5.0">57 mm</quantity>
long, occupying middle portion of lamina, ending before reaching the margin. Indusium thick, brown, entire, rolls back to expose mature sporangia, margin entire or slightly jagged.
</paragraph>
<paragraph blockId="2.[136,1452,1016,2087]" pageId="2" pageNumber="73">
<emphasis box="[189,477,1880,1906]" italics="true" pageId="2" pageNumber="73">Distribution and ecology</emphasis>
:—Island of
<collectingRegion box="[626,693,1881,1907]" country="Philippines" name="Southern Leyte" pageId="2" pageNumber="73">Leyte</collectingRegion>
, Central-Eastern
<collectingCountry box="[893,1023,1880,1906]" name="Philippines" pageId="2" pageNumber="73">Philippines</collectingCountry>
. A terrestrial species growing on the eastern slopes of the
<collectingRegion box="[370,434,1917,1943]" country="Philippines" name="Southern Leyte" pageId="2" pageNumber="73">Leyte</collectingRegion>
Cordillera, at
<quantity box="[595,666,1916,1943]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="73" unit="m" value="500.0">500 m</quantity>
elevation, in lowland tropical evergreen rainforest. Climate warm and humid; rainfall evenly distributed throughout the year (
<bibRefCitation author="Pagasa" box="[753,900,1952,1978]" pageId="2" pageNumber="73" refId="ref6002" refString="Pagasa. (2020) Philippine Atmospheric, Geophysical and Astronomical Services Administration (PAGASA). Available from: http: // bagong. pagasa. dost. gov. ph / climate / climatological-normals. (accessed 7 September 2020)" type="url" year="2020">Pagasa 2020</bibRefCitation>
).
</paragraph>
<paragraph blockId="2.[136,1452,1016,2087]" pageId="2" pageNumber="73">
<emphasis box="[189,309,1988,2014]" italics="true" pageId="2" pageNumber="73">Etymology</emphasis>
:—The species epithet honors Brother George Pahl, FSC, Ph.D., an American biologist and university professor who taught, mentored, and inspired thousands of students in the
<collectingCountry box="[963,1113,2024,2050]" name="United States of America" pageId="2" pageNumber="73">United States</collectingCountry>
and the
<collectingCountry box="[1207,1333,2024,2050]" name="Philippines" pageId="2" pageNumber="73">Philippines</collectingCountry>
during his teaching career spanning more than 50 years.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.5642044" ID-Zenodo-Dep="5642044" httpUri="https://zenodo.org/record/5642044/files/figure.png" pageId="3" pageNumber="74" startId="3.[136,229,1926,1948]" targetBox="[163,1421,205,1897]" targetPageId="3">
<paragraph blockId="3.[136,1451,1926,2020]" pageId="3" pageNumber="74">
<emphasis bold="true" box="[136,251,1926,1948]" pageId="3" pageNumber="74">FIGURE 2</emphasis>
.
<taxonomicName authority="Salgado, Wenzel" authorityName="Salgado, Wenzel" authorityYear="2021" box="[267,597,1926,1948]" class="Polypodiopsida" family="Aspleniaceae" genus="Asplenium" kingdom="Plantae" order="Polypodiales" pageId="3" pageNumber="74" phylum="Tracheophyta" rank="species" species="pahli">
<emphasis box="[267,427,1926,1947]" italics="true" pageId="3" pageNumber="74">Asplenium pahli</emphasis>
Salgado,
<emphasis box="[530,597,1926,1947]" italics="true" pageId="3" pageNumber="74">Wenzel</emphasis>
</taxonomicName>
<emphasis box="[606,642,1927,1948]" italics="true" pageId="3" pageNumber="74">572</emphasis>
, holotype (P). Note the plantlets near the apex of two fronds. Muséum national dhistoire naturelle, Paris, France. Collection: Vascular plants P00370339. License CC Attribution 4.0 International. Available at: https:// science.mnhn.fr/institution/mnhn/collection/p/item/p00370339
</paragraph>
</caption>
<paragraph blockId="4.[136,1452,158,293]" pageId="4" pageNumber="75">
<emphasis box="[189,416,159,185]" italics="true" pageId="4" pageNumber="75">Conservation status</emphasis>
:—
<taxonomicName authorityName="Salgado, Wenzel" authorityYear="2021" box="[452,544,158,185]" class="Polypodiopsida" family="Aspleniaceae" genus="Asplenium" kingdom="Plantae" order="Polypodiales" pageId="4" pageNumber="75" phylum="Tracheophyta" rank="species" species="pahli">
<emphasis box="[452,544,158,185]" italics="true" pageId="4" pageNumber="75">A. pahli</emphasis>
</taxonomicName>
was collected in an area near the towns of Dagami and Jaro that is presently denuded of its original primary growth forest (
<bibRefCitation author="Lasco, R. D. &amp; Visco, R. G. &amp; Pulhin, J. M." box="[658,856,194,220]" pageId="4" pageNumber="75" pagination="652 - 670" refId="ref5497" refString="Lasco, R. D., Visco, R. G. &amp; Pulhin, J. M. (2001) Secondary forests in the Philippines: Formation and transformation in the 20 th century. Journal of Tropical Forest Science 13: 652 - 670. Available from: https: // www. researchgate. net / publication / 239592580 _ Secondary _ forests _ in _ the _ Philippines _ Formation _ and _ transformation _ in _ the _ 20 th _ century (accessed 3 November 2020)" type="journal article" year="2001">
Lasco
<emphasis box="[734,790,194,220]" italics="true" pageId="4" pageNumber="75">et al.</emphasis>
2001
</bibRefCitation>
;
<bibRefCitation author="Mongabay" box="[868,1056,194,220]" pageId="4" pageNumber="75" refId="ref5831" refString="Mongabay (2020) Deforestation statistics for the Philippines. Available from: https: // rainforests. mongabay. com / deforestation / archive / Philippines. htm (accessed 8 September 2020)" type="url" year="2020">Mongabay 2020</bibRefCitation>
). The only collection known is that of Wenzel in 1914. This is a possible indication that
<taxonomicName authorityName="Salgado, Wenzel" authorityYear="2021" box="[713,801,230,257]" class="Polypodiopsida" family="Aspleniaceae" genus="Asplenium" kingdom="Plantae" order="Polypodiales" pageId="4" pageNumber="75" phylum="Tracheophyta" rank="species" species="pahli">
<emphasis box="[713,801,230,257]" italics="true" pageId="4" pageNumber="75">A. pahli</emphasis>
</taxonomicName>
is endangered or possibly extinct in its
<typeStatus box="[1237,1285,231,257]" pageId="4" pageNumber="75">type</typeStatus>
locality. There might be more plants scattered in the remaining stands of old growth deep in the
<collectingRegion box="[1044,1108,267,293]" country="Philippines" name="Southern Leyte" pageId="4" pageNumber="75">Leyte</collectingRegion>
Cordillera.
</paragraph>
<paragraph blockId="4.[136,1451,331,393]" pageId="4" pageNumber="75">
<emphasis box="[136,703,331,358]" italics="true" pageId="4" pageNumber="75">
<taxonomicName authorityName="G. Forster" authorityYear="1786" box="[136,374,331,357]" class="Polypodiopsida" family="Aspleniaceae" genus="Asplenium" kingdom="Plantae" order="Polypodiales" pageId="4" pageNumber="75" phylum="Tracheophyta" rank="species" species="caudatum">Asplenium caudatum</taxonomicName>
,
<taxonomicName authorityName="Lamarck" authorityYear="1786" box="[385,517,331,358]" class="Polypodiopsida" family="Aspleniaceae" genus="Asplenium" kingdom="Plantae" order="Polypodiales" pageId="4" pageNumber="75" phylum="Tracheophyta" rank="species" species="falcatum">A. falcatum</taxonomicName>
,
<taxonomicName authorityName="C. Presl, Cuming" authorityYear="1849" box="[527,703,331,358]" class="Polypodiopsida" family="Aspleniaceae" genus="Asplenium" kingdom="Plantae" order="Polypodiales" pageId="4" pageNumber="75" phylum="Tracheophyta" rank="species" species="anisodontum">A. anisodontum</taxonomicName>
</emphasis>
and
<taxonomicName authorityName="Salgado, Wenzel" authorityYear="2021" box="[757,846,331,358]" class="Polypodiopsida" family="Aspleniaceae" genus="Asplenium" kingdom="Plantae" order="Polypodiales" pageId="4" pageNumber="75" phylum="Tracheophyta" rank="species" species="pahli">
<emphasis box="[757,846,331,358]" italics="true" pageId="4" pageNumber="75">A. pahli</emphasis>
</taxonomicName>
represent different species as shown by the characters presented in this key.
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="75" type="key">
<key pageId="4" pageNumber="75">
<keyStep pageId="4" pageNumber="75">
<paragraph blockId="4.[136,1452,436,889]" box="[136,1451,436,458]" pageId="4" pageNumber="75">
<keyLead box="[136,1451,436,458]" pageId="4" pageNumber="75">1. Fronds non proliferous .......................................................................................................................................................................2</keyLead>
</paragraph>
<paragraph blockId="4.[136,1452,436,889]" box="[136,1451,465,487]" pageId="4" pageNumber="75">
<keyLead box="[136,1451,465,487]" pageId="4" pageNumber="75">1. Fronds proliferous ..............................................................................................................................................................................3</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="4" pageNumber="75">
<paragraph blockId="4.[136,1452,436,889]" pageId="4" pageNumber="75">
<keyLead pageId="4" pageNumber="75">
2. Basal pinnae not decreasing in size, lamina base truncate; acroscopic basal pinna lobe with a prominent sharp tooth, sori single row, oblique to costa..........................................................................................................................................................
<taxonomicName authorityName="Lamarck" authorityYear="1786" box="[1340,1451,522,544]" class="Polypodiopsida" family="Aspleniaceae" genus="Asplenium" kingdom="Plantae" order="Polypodiales" pageId="4" pageNumber="75" phylum="Tracheophyta" rank="species" species="falcatum">
<emphasis box="[1340,1451,522,544]" italics="true" pageId="4" pageNumber="75">A. falcatum</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="4.[136,1452,436,889]" pageId="4" pageNumber="75">
<keyLead pageId="4" pageNumber="75">
2. Basal pinnae gradually decreasing in size, lamina base attenuate; acroscopic basal pinna lobe without a prominent sharp tooth, sori in two rows, one parallel to costa and one oblique to costa ............................................................................................
<taxonomicName authorityName="G. Forster" authorityYear="1786" box="[1329,1451,580,601]" class="Polypodiopsida" family="Aspleniaceae" genus="Asplenium" kingdom="Plantae" order="Polypodiales" pageId="4" pageNumber="75" phylum="Tracheophyta" rank="species" species="caudatum">
<emphasis box="[1329,1451,580,601]" italics="true" pageId="4" pageNumber="75">A. caudatum</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="4" pageNumber="75">
<paragraph blockId="4.[136,1452,436,889]" pageId="4" pageNumber="75">
<keyLead pageId="4" pageNumber="75">
3. Fronds up to
<quantity box="[340,412,609,630]" metricMagnitude="0" metricUnit="m" metricValue="1.2" pageId="4" pageNumber="75" unit="cm" value="120.0">120 cm</quantity>
long. Stipe and rachis dark brown; rachis hirsute, paleae projecting away from the stipe and rachis, oblong to lanceate or reduced to long twisted biserrate filaments, margin with prominent biserrate projections up to five cells long, palea base and margin concolorous with the rest of the palea, apex a long twisted biseriate filament; median pinnae falcate, 10.0
<quantity metricMagnitude="-1" metricUnit="m" metricValue="1.56" pageId="4" pageNumber="75" unit="cm" value="15.6">15.6 cm</quantity>
long,
<quantity box="[301,408,695,717]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" metricValueMax="1.2" metricValueMin="0.9" pageId="4" pageNumber="75" unit="cm" value="1.05" valueMax="1.2" valueMin="0.9">0.91.2 cm</quantity>
wide; sori close and parallel to costa then curving towards the margin,
<quantity box="[1093,1177,695,716]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="11.0" metricValueMin="7.0" pageId="4" pageNumber="75" unit="mm" value="9.0" valueMax="11.0" valueMin="7.0">711mm</quantity>
long, contiguous ................. .....................................................................................................................................................................................
<taxonomicName authorityName="C. Presl, Cuming" authorityYear="1849" box="[1300,1451,724,745]" class="Polypodiopsida" family="Aspleniaceae" genus="Asplenium" kingdom="Plantae" order="Polypodiales" pageId="4" pageNumber="75" phylum="Tracheophyta" rank="species" species="anisodontum">
<emphasis box="[1300,1451,724,745]" italics="true" pageId="4" pageNumber="75">A. anisodontum</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="4.[136,1452,436,889]" pageId="4" pageNumber="75">
<keyLead pageId="4" pageNumber="75">
3. Fronds up to
<quantity box="[335,393,753,775]" metricMagnitude="-1" metricUnit="m" metricValue="4.5" pageId="4" pageNumber="75" unit="cm" value="45.0">45 cm</quantity>
long. Stipe and rachis greenish; rachis with a few scattered paleae, paleae adpressed against the stipe and rachis surface, deltoid to lanceate in shape, margin entire and irregular, with an occasional short, acute projection caused by a cell wall, palea base and margin bicolorous with light brown margins and black central area, apex acute; median pinna straight, 3.54.0 cm long and 1.0
<quantity box="[343,407,839,861]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="4" pageNumber="75" unit="cm" value="1.3">1.3 cm</quantity>
wide; sori distant from the costa, straight or slightly falcate, oblique,
<quantity box="[1076,1155,839,861]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="4" pageNumber="75" unit="mm" value="6.0" valueMax="7.0" valueMin="5.0">57 mm</quantity>
long, not contiguous............... .................................................................................................................................................................................................
<taxonomicName authorityName="Salgado, Wenzel" authorityYear="2021" box="[1375,1450,868,889]" class="Polypodiopsida" family="Aspleniaceae" genus="Asplenium" kingdom="Plantae" order="Polypodiales" pageId="4" pageNumber="75" phylum="Tracheophyta" rank="species" species="pahli">
<emphasis box="[1375,1450,868,889]" italics="true" pageId="4" pageNumber="75">A. pahli</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>