treatments-xml/data/03/9B/C5/039BC5481C7CC34DC8F2A1673E1F6E8A.xml
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<document id="634FE5462EB71DED620762E26A9AEC81" ID="10.11646/zootaxa.4175.5.3" ID-DOI="10.11646/zootaxa.4175.5.3" ID-GBIF-Dataset="e1ca0b14-99a0-4176-a8f0-430cba3c4a99" ID-ISSN="1175-5326" ID-Zenodo-Dep="263137" ID-ZooBank="975F0772-9864-49F4-B99A-0826E480E584" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1476771634842" checkinUser="plazi" docAuthor="Sáfián, Szabolcs, Pyrcz, Tomasz &amp; Brattström, Oskar" docDate="2016" docId="039BC5481C7CC34DC8F2A1673E1F6E8A" docLanguage="en" docName="zootaxa.4175.5.3.pdf" docOrigin="Zootaxa 4175 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Bebearia wojtusiaki Safian, Brattstrom &amp; Pyrcz, sp. nov." docType="treatment" docVersion="7" lastPageNumber="459" masterDocId="FFA2BD301C7BC347C865A51D3A546A65" masterDocTitle="Two new species of Bebearia Hemming, 1960, as further evidence of centre of endemism of butterflies in Western Nigeria (Lepidoptera: Nymphalidae: Limenitinae)" masterLastPageNumber="462" masterPageNumber="449" pageNumber="456" updateTime="1698390000371" updateUser="plazi">
<mods:mods id="5E084A3F1C153D359AAC4ECEB5C0FFD3" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="BCDF701747818BDB9F5581C728A49BE5">
<mods:title id="A0DC8E15AD3A456CCAD94171B6457176">Two new species of Bebearia Hemming, 1960, as further evidence of centre of endemism of butterflies in Western Nigeria (Lepidoptera: Nymphalidae: Limenitinae)</mods:title>
</mods:titleInfo>
<mods:name id="6418ACE2EF375C92FE40BF8D42E218D4" type="personal">
<mods:role id="77C123C00CAC70F20A08F8A60C9802FB">
<mods:roleTerm id="61DABBEA6417F90B1679AE2B4A6699C4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="19B0E6A098678ECFFFAAFD7A58C36807">Sáfián, Szabolcs</mods:namePart>
</mods:name>
<mods:name id="273CCE8FB900A3FBD302F5B6C6369CD9" type="personal">
<mods:role id="76ECB1787BA5C16EBD583679B585BF41">
<mods:roleTerm id="C894DB616E9CDB07C20B11B4568AF206">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FD73D0A4C8832FBF84AEE349A9A8B984">Pyrcz, Tomasz</mods:namePart>
</mods:name>
<mods:name id="4E96B0ADE2B9A4154819513B5D7CCF91" type="personal">
<mods:role id="A2079F447A5082FA0706EF81594933F5">
<mods:roleTerm id="CA860E978AD917C8338852CB722951FA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7ACB187EF59BA20B3BFDF5D0DC18C602">Brattström, Oskar</mods:namePart>
</mods:name>
<mods:typeOfResource id="1A7EB1AC2EF1BA166D5974F5234C084C">text</mods:typeOfResource>
<mods:relatedItem id="787265B1DAEA5BA32BD28C5EDD86043A" type="host">
<mods:titleInfo id="8AC7770A9962720FC45FF4BECE436DCB">
<mods:title id="0B02B2B7412FEFE44A94B375EBEF7919">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="C14602F6712231BC0FB6B2CC8F96720D">
<mods:date id="BF2DD12BACEAAABC412B7F6C1BAD94A6">2016</mods:date>
<mods:detail id="A000930D331D9F292592D85291E4706D" type="volume">
<mods:number id="213D73AE58FFFFDC412C56DDE32B1405">4175</mods:number>
</mods:detail>
<mods:detail id="0BAFE4E57FC14F916361D4C6404E37EE" type="issue">
<mods:number id="72910A34A648D95940895E1354E2A1E3">5</mods:number>
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<mods:end id="B668C3B4F341A42AC2DFD71C5F168A7E">462</mods:end>
</mods:extent>
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</mods:relatedItem>
<mods:classification id="1D069E95F9C913D2CFCA0B18E2EAD055">journal article</mods:classification>
<mods:identifier id="84C97AF34599453F8366152D4232760A" type="DOI">10.11646/zootaxa.4175.5.3</mods:identifier>
<mods:identifier id="2BF32FD2ED38B569EC0AA2D301E2FE52" type="GBIF-Dataset">e1ca0b14-99a0-4176-a8f0-430cba3c4a99</mods:identifier>
<mods:identifier id="1885B5723224286B0323134A23E4FE26" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="12C50B8C1A4AEC483E8771F0A613B4B9" type="Zenodo-Dep">263137</mods:identifier>
<mods:identifier id="835F38B017A027554A3536709CBA6DB0" type="ZooBank">975F0772-9864-49F4-B99A-0826E480E584</mods:identifier>
</mods:mods>
<treatment id="039BC5481C7CC34DC8F2A1673E1F6E8A" ID-DOI="http://doi.org/10.5281/zenodo.6063295" ID-GBIF-Taxon="124996699" ID-Zenodo-Dep="6063295" LSID="urn:lsid:plazi:treatment:039BC5481C7CC34DC8F2A1673E1F6E8A" httpUri="http://treatment.plazi.org/id/039BC5481C7CC34DC8F2A1673E1F6E8A" lastPageId="10" lastPageNumber="459" pageId="7" pageNumber="456">
<subSubSection id="C32827D51C7CC340C8F2A167385A6ED3" pageId="7" pageNumber="456" type="nomenclature">
<paragraph id="8B8D745E1C7CC340C8F2A167391F6EF0" blockId="7.[151,843,1146,1206]" box="[151,843,1146,1173]" pageId="7" pageNumber="456">
<heading id="D0C5C3321C7CC340C8F2A167391F6EF0" bold="true" box="[151,843,1146,1173]" fontSize="11" level="1" pageId="7" pageNumber="456" reason="1">
<emphasis id="B946A84C1C7CC340C8F2A167391F6EF0" bold="true" box="[151,843,1146,1173]" pageId="7" pageNumber="456">
<taxonomicName id="4C320FDD1C7CC340C8F2A16738B26EF0" authority="Safian, Brattstrom &amp; Pyrcz" authorityName="Safian, Brattstrom &amp; Pyrcz" box="[151,742,1146,1173]" class="Insecta" family="Nymphalidae" genus="Bebearia" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="456" phylum="Arthropoda" rank="species" species="wojtusiaki" status="sp. nov.">
<emphasis id="B946A84C1C7CC340C8F2A1673BD36EF1" bold="true" box="[151,391,1146,1172]" italics="true" pageId="7" pageNumber="456">Bebearia wojtusiaki</emphasis>
Sáfián, Brattström &amp; Pyrcz
</taxonomicName>
<taxonomicNameLabel id="A27515371C7CC340CA8BA166391F6EF0" box="[750,843,1147,1173]" pageId="7" pageNumber="456" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8D745E1C7CC340C8F2A183385A6ED3" blockId="7.[151,843,1146,1206]" box="[151,526,1181,1206]" pageId="7" pageNumber="456">FIGS.: 6A, D; 7A; 8A, D; 9A; 10</paragraph>
</subSubSection>
<subSubSection id="C32827D51C7CC340C8F2A1F83F346D7F" pageId="7" pageNumber="456" type="materials_examined">
<paragraph id="8B8D745E1C7CC340C8F2A1F83F036F47" blockId="7.[151,1437,1253,2037]" pageId="7" pageNumber="456">
<materialsCitation id="3B5A7E031C7CC340C8F2A1F83F036F47" ID-GBIF-Occurrence="1323867395" collectingDate="2010-11-22" collectionCode="ABRI" country="Nigeria" county="Ologbo" elevation="22" latitude="6.0174837" location="Oskar Brattstrom Coordinates" longLatPrecision="1" longitude="5.5407667" municipality="Benin City" pageId="7" pageNumber="456" specimenCount="1" specimenCount-male="1" stateProvince="Edo" typeStatus="holotype">
<emphasis id="B946A84C1C7CC340C8F2A1F83B596E9B" bold="true" box="[151,269,1253,1278]" pageId="7" pageNumber="456">
<typeStatus id="5489CAFC1C7CC340C8F2A1F83B5C6E9B" box="[151,264,1253,1278]" pageId="7" pageNumber="456" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCount id="9D34BFD71C7CC340C971A1FB3B7D6E98" box="[276,297,1254,1277]" pageId="7" pageNumber="456" type="male"></specimenCount>
<collectingCountry id="F32534CE1C7CC340C954A1FB3BF36E9B" box="[305,423,1254,1278]" name="Nigeria" pageId="7" pageNumber="456">NIGERIA</collectingCountry>
,
<collectingCounty id="62EC0CD21C7CC340C9D6A1F8385E6E9B" box="[435,522,1253,1278]" pageId="7" pageNumber="456">Ologbo</collectingCounty>
, South of
<collectingMunicipality id="6BE9EE241C7CC340CA1BA1F838AE6E9B" box="[638,762,1253,1278]" pageId="7" pageNumber="456">Benin City</collectingMunicipality>
,
<collectingRegion id="49F6BABC1C7CC340CB62A1F839606E9B" box="[775,820,1253,1278]" country="Nigeria" name="Edo" pageId="7" pageNumber="456">Edo</collectingRegion>
State
<date id="FF8C529E1C7CC340CB1EA1FB39AA6E9B" box="[891,1022,1254,1278]" pageId="7" pageNumber="456" value="2010-11-22">
<collectingDate id="EFC8AB761C7CC340CB1EA1FB39AA6E9B" box="[891,1022,1254,1278]" pageId="7" pageNumber="456" value="2010-11-22">22.XI.2010</collectingDate>
</date>
. leg.:
<location id="8EED22851C7CC340CC27A1F83FCD6E9B" LSID="urn:lsid:plazi:treatment:039BC5481C7CC34DC8F2A1673E1F6E8A:8EED22851C7CC340CC27A1F83FCD6E9B" box="[1090,1433,1253,1278]" country="Nigeria" county="Ologbo" latitude="6.0174837" longLatPrecision="1" longitude="5.5407667" municipality="Benin City" name="Oskar Brattstrom Coordinates" pageId="7" pageNumber="456" stateProvince="Edo">Oskar Brattström Coordinates</location>
:
<geoCoordinate id="EE0612991C7CC340C8F2A0143B416F47" box="[151,277,1289,1314]" direction="north" orientation="latitude" pageId="7" pageNumber="456" precision="1" value="6.0174837">6°1'2.94&quot;N</geoCoordinate>
,
<geoCoordinate id="EE0612991C7CC340C947A0143BEF6F47" box="[290,443,1289,1314]" direction="east" orientation="longitude" pageId="7" pageNumber="456" precision="1" value="5.5407667">5°32'26.76&quot;E</geoCoordinate>
. Elevation:
<quantity id="4CCAD9BB1C7CC340CA5AA017382D6F44" box="[575,633,1290,1314]" metricMagnitude="1" metricUnit="m" metricValue="2.2" pageId="7" pageNumber="456" unit="m" value="22.0">
<elevation id="001F936D1C7CC340CA5AA017382D6F44" box="[575,633,1290,1314]" metricMagnitude="1" metricUnit="m" metricValue="2.2" pageId="7" pageNumber="456" unit="m" value="22.0">22 m</elevation>
</quantity>
asl. Gen. prep.: OB-BEB-003. Deposited in the
<collectionCode id="ED23EC9B1C7CC340CCF2A0173E8E6F47" box="[1175,1242,1290,1314]" pageId="7" pageNumber="456">ABRI</collectionCode>
collection.
</materialsCitation>
</paragraph>
<paragraph id="8B8D745E1C7CC340C8A2A03039956CEF" blockId="7.[151,1437,1253,2037]" pageId="7" pageNumber="456">
<materialsCitation id="3B5A7E031C7CC340C8A2A03038AC6F20" ID-GBIF-Occurrence="1323867393" box="[199,760,1325,1350]" country="Nigeria" location="Okomu Forest" pageId="7" pageNumber="456" specimenCount="15" specimenCount-female="7" specimenCount-male="8" typeStatus="paratype">
<emphasis id="B946A84C1C7CC340C8A2A0303B1C6F23" bold="true" box="[199,328,1325,1350]" pageId="7" pageNumber="456">
<typeStatus id="5489CAFC1C7CC340C8A2A0303B176F23" box="[199,323,1325,1350]" pageId="7" pageNumber="456" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="9D34BFD71C7CC340C935A0333BDC6F23" box="[336,392,1326,1350]" pageId="7" pageNumber="456" type="male">8♂♂</specimenCount>
<specimenCount id="9D34BFD71C7CC340C9F5A0333B9C6F20" box="[400,456,1326,1349]" pageId="7" pageNumber="456" type="female">7♀♀</specimenCount>
<collectingCountry id="F32534CE1C7CC340C9B5A03338126F23" box="[464,582,1326,1350]" name="Nigeria" pageId="7" pageNumber="456">NIGERIA</collectingCountry>
,
<location id="8EED22851C7CC340CA37A03038AC6F20" LSID="urn:lsid:plazi:treatment:039BC5481C7CC34DC8F2A1673E1F6E8A:8EED22851C7CC340CA37A03038AC6F20" box="[594,760,1325,1350]" country="Nigeria" name="Okomu Forest" pageId="7" pageNumber="456">Okomu Forest</location>
</materialsCitation>
,
<materialsCitation id="3B5A7E031C7CC340CB61A030399D6F23" ID-GBIF-Occurrence="1323867396" box="[772,969,1325,1350]" country="Nigeria" location="Bendel State" pageId="7" pageNumber="456" specimenCount="1" specimenCount-male="1" stateProvince="Bendel State" typeStatus="paratype">
<collectingRegion id="49F6BABC1C7CC340CB61A03039C36F23" box="[772,919,1325,1350]" inferredBySuffix="true" pageId="7" pageNumber="456">Bendel State</collectingRegion>
:
<specimenCount id="9D34BFD71C7CC340CBC6A033399D6F23" box="[931,969,1326,1350]" pageId="7" pageNumber="456" type="male">1♂</specimenCount>
</materialsCitation>
:
<date id="FF8C529E1C7CC340CBB3A0333E626F23" box="[982,1078,1326,1350]" pageId="7" pageNumber="456" value="1983-09">IX.1983</date>
; 1♂: no date; 1♂:
<date id="FF8C529E1C7CC340CD6EA0303FC36F23" box="[1291,1431,1325,1350]" pageId="7" pageNumber="456" value="1985-12-14">14.XII.1985</date>
; 1♂:
<date id="FF8C529E1C7CC340C8B1A04F3B186F0F" box="[212,332,1362,1386]" pageId="7" pageNumber="456" value="1984-10-18">18.X.1984</date>
; 1♂:
<date id="FF8C529E1C7CC340C9C5A04F38776F0F" box="[416,547,1362,1386]" pageId="7" pageNumber="456" value="1984-11-20">20.XI.1984</date>
; 1♂:
<date id="FF8C529E1C7CC340CA13A04C38B96F0F" box="[630,749,1361,1386]" pageId="7" pageNumber="456" value="1986-02-14">14.II.1986</date>
; 2♂:
<date id="FF8C529E1C7CC340CB24A04F39EE6F0F" box="[833,954,1362,1386]" pageId="7" pageNumber="456" value="1984-10-19">19.X.1984</date>
; 1♂:
<date id="FF8C529E1C7CC340CC68A04F3ED36F0F" box="[1037,1159,1362,1386]" pageId="7" pageNumber="456" value="1984-10-20">20.X.1984</date>
;; 1♂:
<date id="FF8C529E1C7CC340CC87A04F3F0F6F0F" box="[1250,1371,1362,1386]" pageId="7" pageNumber="456" value="1984-10-18">18.X.1984</date>
; 2♂:
<date id="FF8C529E1C7CC340C8F2A0683B776FEB" box="[151,291,1397,1422]" pageId="7" pageNumber="456" value="1985-12-14">14.XII.1985</date>
; 3♀: no date; 2♀:
<date id="FF8C529E1C7CC340CA68A06B38D36FEB" box="[525,647,1398,1422]" pageId="7" pageNumber="456" value="1984-10-20">20.X.1984</date>
; 1♀:
<date id="FF8C529E1C7CC340CAAAA06839126FEB" box="[719,838,1397,1422]" pageId="7" pageNumber="456" value="1986-02-13">13.II.1986</date>
; 1♀:
<date id="FF8C529E1C7CC340CBEBA06B3E536FEB" box="[910,1031,1398,1422]" pageId="7" pageNumber="456" value="1984-10-19">19.X.1984</date>
; 1♀:
<date id="FF8C529E1C7CC340CC35A0683E8F6FEB" box="[1104,1243,1397,1422]" pageId="7" pageNumber="456" value="1985-12-14">14.XII.1985</date>
, all leg.: Janusz Wojtusiak 1♀:
<date id="FF8C529E1C7CC340C921A0873BEF6FD7" box="[324,443,1434,1458]" pageId="7" pageNumber="456" value="2000-01-08" valueMax="2000-01-09" valueMin="2000-01-08">8-9.I.2000</date>
, leg.: Tomasz Pyrcz;
<materialsCitation id="3B5A7E031C7CC340CAD5A0873F046FD7" ID-GBIF-Occurrence="1323867402" box="[688,1360,1433,1458]" collectingDate="1984-05-25" country="Nigeria" location="Okomu Forest" pageId="7" pageNumber="456" specimenCount="1" specimenCount-male="1" stateProvince="Bendel State" typeStatus="paratype">
<specimenCount id="9D34BFD71C7CC340CAD5A08738876FD7" box="[688,723,1434,1458]" pageId="7" pageNumber="456" type="male">1♂</specimenCount>
<collectingCountry id="F32534CE1C7CC340CAB9A08739066FD7" box="[732,850,1434,1458]" name="Nigeria" pageId="7" pageNumber="456">NIGERIA</collectingCountry>
,
<location id="8EED22851C7CC340CB3AA0843E506FD4" LSID="urn:lsid:plazi:treatment:039BC5481C7CC34DC8F2A1673E1F6E8A:8EED22851C7CC340CB3AA0843E506FD4" box="[863,1028,1433,1458]" country="Nigeria" name="Okomu Forest" pageId="7" pageNumber="456" stateProvince="Bendel State">Okomu Forest</location>
,
<collectingRegion id="49F6BABC1C7CC340CC74A0843EF46FD7" box="[1041,1184,1433,1458]" inferredBySuffix="true" pageId="7" pageNumber="456">Bendel State</collectingRegion>
<date id="FF8C529E1C7CC340CCCCA0843F4F6FD7" box="[1193,1307,1433,1458]" pageId="7" pageNumber="456" value="1984-05-25">
<collectingDate id="EFC8AB761C7CC340CCCCA0843F4F6FD7" box="[1193,1307,1433,1458]" pageId="7" pageNumber="456" value="1984-05-25">25.V.1984</collectingDate>
</date>
gen
</materialsCitation>
.
<materialsCitation id="3B5A7E031C7CC340CD39A0873E0F6FB3" ID-GBIF-Occurrence="1323867413" collectingDate="1985-12-15" country="Nigeria" location="Okomu Forest" pageId="7" pageNumber="456" specimenCount="1" specimenCount-male="1" stateProvince="Bendel State" typeStatus="paratype">
prep.: MZUJ-JL-155;
<specimenCount id="9D34BFD71C7CC340C935A0A33B276FB3" box="[336,371,1470,1494]" pageId="7" pageNumber="456" type="male">1♂</specimenCount>
<collectingCountry id="F32534CE1C7CC340C9E1A0A33BAE6FB3" box="[388,506,1470,1494]" name="Nigeria" pageId="7" pageNumber="456">NIGERIA</collectingCountry>
,
<location id="8EED22851C7CC340CA6BA0A038EF6FB0" LSID="urn:lsid:plazi:treatment:039BC5481C7CC34DC8F2A1673E1F6E8A:8EED22851C7CC340CA6BA0A038EF6FB0" box="[526,699,1469,1494]" country="Nigeria" name="Okomu Forest" pageId="7" pageNumber="456" stateProvince="Bendel State">Okomu Forest</location>
,
<collectingRegion id="49F6BABC1C7CC340CAAAA0A039326FB3" box="[719,870,1469,1494]" inferredBySuffix="true" pageId="7" pageNumber="456">Bendel State</collectingRegion>
<date id="FF8C529E1C7CC340CB13A0A03E4B6FB3" box="[886,1055,1469,1494]" pageId="7" pageNumber="456" value="1985-12-15">
<collectingDate id="EFC8AB761C7CC340CB13A0A03E4B6FB3" box="[886,1055,1469,1494]" pageId="7" pageNumber="456" value="1985-12-15">15. XII. 1985</collectingDate>
</date>
gen
</materialsCitation>
.
<materialsCitation id="3B5A7E031C7CC340CC0AA0A33F0D6F9F" ID-GBIF-Occurrence="1323867401" collectingDate="2014-09-05" country="Nigeria" location="Okomu Forest" municipality="Janusz Wojtusiak" pageId="7" pageNumber="456" specimenCount="1" specimenCount-female="1" stateProvince="Bendel State" typeStatus="paratype">
prep.: MZUJ-JL-151;
<specimenCount id="9D34BFD71C7CC340CD1CA0A33FC86FB3" box="[1401,1436,1470,1494]" pageId="7" pageNumber="456" type="female">1♀</specimenCount>
<collectingCountry id="F32534CE1C7CC340C8F2A0FF3B596F9F" box="[151,269,1506,1530]" name="Nigeria" pageId="7" pageNumber="456">NIGERIA</collectingCountry>
,
<location id="8EED22851C7CC340C972A0FC3BEF6F9C" LSID="urn:lsid:plazi:treatment:039BC5481C7CC34DC8F2A1673E1F6E8A:8EED22851C7CC340C972A0FC3BEF6F9C" box="[279,443,1505,1530]" country="Nigeria" municipality="Janusz Wojtusiak" name="Okomu Forest" pageId="7" pageNumber="456" stateProvince="Bendel State">Okomu Forest</location>
,
<collectingRegion id="49F6BABC1C7CC340C9A3A0FC38016F9F" box="[454,597,1505,1530]" inferredBySuffix="true" pageId="7" pageNumber="456">Bendel State</collectingRegion>
, no date, leg.:
<collectingMunicipality id="6BE9EE241C7CC340CA92A0FF39ED6F9F" box="[759,953,1505,1530]" pageId="7" pageNumber="456">Janusz Wojtusiak</collectingMunicipality>
gen. prep.: MZUJ-JL-01/
<date id="FF8C529E1C7CC340CCBDA0FC3F0D6F9F" box="[1240,1369,1505,1530]" pageId="7" pageNumber="456" value="2014-09-05">
<collectingDate id="EFC8AB761C7CC340CCBDA0FC3F0D6F9F" box="[1240,1369,1505,1530]" pageId="7" pageNumber="456" value="2014-09-05">09.05.2014</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B5A7E031C7CC340CD01A0FC38446C78" ID-GBIF-Occurrence="1323867411" country="Nigeria" location="Ologbo Forest" pageId="7" pageNumber="456" specimenCount="8" specimenCount-female="3" specimenCount-male="5" typeStatus="paratype">
<specimenCount id="9D34BFD71C7CC340CD01A0FC3FC86F9F" box="[1380,1436,1505,1530]" pageId="7" pageNumber="456" type="male">5♂♂</specimenCount>
<specimenCount id="9D34BFD71C7CC340C8F2A31B3A9A6C7B" box="[151,206,1542,1566]" pageId="7" pageNumber="456" type="female">3♀♀</specimenCount>
<collectingCountry id="F32534CE1C7CC340C8B8A31B3B076C7B" box="[221,339,1542,1566]" name="Nigeria" pageId="7" pageNumber="456">NIGERIA</collectingCountry>
,
<location id="8EED22851C7CC340C900A31838446C78" LSID="urn:lsid:plazi:treatment:039BC5481C7CC34DC8F2A1673E1F6E8A:8EED22851C7CC340C900A31838446C78" box="[357,528,1541,1566]" country="Nigeria" name="Ologbo Forest" pageId="7" pageNumber="456">Ologbo Forest</location>
</materialsCitation>
,
<materialsCitation id="3B5A7E031C7CC340CA47A318394D6C7B" ID-GBIF-Occurrence="1323867391" box="[546,793,1541,1566]" country="Benin" location="South" pageId="7" pageNumber="456" specimenCount="1" stateProvince="South" typeStatus="paratype">
<collectingRegion id="49F6BABC1C7CC340CA47A31838306C7B" box="[546,612,1541,1566]" country="Tanzania" name="Kusini Unguja" pageId="7" pageNumber="456">South</collectingRegion>
of
<collectingCountry id="F32534CE1C7CC340CAF2A318388E6C7B" box="[663,730,1541,1566]" name="Benin" pageId="7" pageNumber="456">Benin</collectingCountry>
City
</materialsCitation>
,
<materialsCitation id="3B5A7E031C7CC340CB49A3183B906C24" ID-GBIF-Occurrence="1323867403" collectingDate="2010-11-19" collectingDateMax="2010-11-24" collectingDateMin="2010-11-19" country="Nigeria" location="Ologbo Forest" pageId="7" pageNumber="456" specimenCount="1" specimenCount-female="1" stateProvince="Edo State" typeStatus="paratype">
<collectingRegion id="49F6BABC1C7CC340CB49A31839CB6C7B" box="[812,927,1541,1566]" inferredBySuffix="true" pageId="7" pageNumber="456">
<collectingRegion id="49F6BABC1C7CC340CB49A318390D6C7B" box="[812,857,1541,1566]" country="Nigeria" name="Edo" pageId="7" pageNumber="456">Edo</collectingRegion>
State
</collectingRegion>
<date id="FF8C529E1C7CC340CBC8A31B3E056C7B" box="[941,1105,1542,1566]" pageId="7" pageNumber="456" value="2010-11-19" valueMax="2010-11-24" valueMin="2010-11-19">
<collectingDate id="EFC8AB761C7CC340CBC8A31B3E056C7B" box="[941,1105,1542,1566]" pageId="7" pageNumber="456" value="2010-11-19" valueMax="2010-11-24" valueMin="2010-11-19">19-24.XI.2010</collectingDate>
</date>
Leg.: Oskar Brattström;
<specimenCount id="9D34BFD71C7CC340CDE2A31B3FC86C78" box="[1415,1436,1542,1565]" pageId="7" pageNumber="456" type="female"></specimenCount>
<collectingCountry id="F32534CE1C7CC340C8F2A3373B596C27" box="[151,269,1578,1602]" name="Nigeria" pageId="7" pageNumber="456">NIGERIA</collectingCountry>
,
<location id="8EED22851C7CC340C979A3343B906C24" LSID="urn:lsid:plazi:treatment:039BC5481C7CC34DC8F2A1673E1F6E8A:8EED22851C7CC340C979A3343B906C24" box="[284,452,1577,1602]" country="Nigeria" name="Ologbo Forest" pageId="7" pageNumber="456" stateProvince="Edo State">Ologbo Forest</location>
</materialsCitation>
,
<materialsCitation id="3B5A7E031C7CC340C9B1A33438966C27" ID-GBIF-Occurrence="1323867400" box="[468,706,1577,1602]" country="Benin" location="South" pageId="7" pageNumber="456" specimenCount="1" stateProvince="South" typeStatus="paratype">
<collectingRegion id="49F6BABC1C7CC340C9B1A33438416C27" box="[468,533,1577,1602]" country="Tanzania" name="Kusini Unguja" pageId="7" pageNumber="456">South</collectingRegion>
of
<collectingCountry id="F32534CE1C7CC340CA26A33438D26C27" box="[579,646,1577,1602]" name="Benin" pageId="7" pageNumber="456">Benin</collectingCountry>
City
</materialsCitation>
,
<materialsCitation id="3B5A7E031C7CC340CAB4A3343FCC6C27" ID-GBIF-Occurrence="1323867408" box="[721,1432,1577,1602]" collectingDate="2009-03-26" country="Nigeria" location="Edo State" pageId="7" pageNumber="456" specimenCount="1" specimenCount-female="1" stateProvince="Edo State" typeStatus="paratype">
<collectingRegion id="49F6BABC1C7CC340CAB4A33439156C27" box="[721,833,1577,1602]" inferredBySuffix="true" pageId="7" pageNumber="456">
<collectingRegion id="49F6BABC1C7CC340CAB4A33438AA6C27" box="[721,766,1577,1602]" country="Nigeria" name="Edo" pageId="7" pageNumber="456">Edo</collectingRegion>
State
</collectingRegion>
<date id="FF8C529E1C7CC340CB29A334399E6C27" box="[844,970,1577,1602]" pageId="7" pageNumber="456" value="2009-03-26">
<collectingDate id="EFC8AB761C7CC340CB29A334399E6C27" box="[844,970,1577,1602]" pageId="7" pageNumber="456" value="2009-03-26">26.III.2009</collectingDate>
</date>
Leg.: Oskar Brattström;
<specimenCount id="9D34BFD71C7CC340CC91A3373F436C27" box="[1268,1303,1578,1602]" pageId="7" pageNumber="456" type="female">1♀</specimenCount>
<collectingCountry id="F32534CE1C7CC340CD47A3373FCC6C27" box="[1314,1432,1578,1602]" name="Nigeria" pageId="7" pageNumber="456">NIGERIA</collectingCountry>
</materialsCitation>
,
<materialsCitation id="3B5A7E031C7CC340C8F2A353398D6C00" ID-GBIF-Occurrence="1323867410" box="[151,985,1613,1638]" collectingDate="1954-08" country="Nigeria" location="Omo Forest" pageId="7" pageNumber="456" specimenCount="1" specimenCount-female="1" stateProvince="Lagos" typeStatus="paratype">
<collectingRegion id="49F6BABC1C7CC340C8F2A3533A8F6C03" box="[151,219,1614,1638]" country="Nigeria" name="Lagos" pageId="7" pageNumber="456">Lagos</collectingRegion>
<date id="FF8C529E1C7CC340C887A3533B086C03" box="[226,348,1613,1638]" pageId="7" pageNumber="456" value="1954-08">
<collectingDate id="EFC8AB761C7CC340C887A3533B086C03" box="[226,348,1613,1638]" pageId="7" pageNumber="456" value="1954-08">VIII. 1954</collectingDate>
</date>
. leg.: John Patrick Boorman,
<specimenCount id="9D34BFD71C7CC340CAC3A353389D6C03" box="[678,713,1614,1638]" pageId="7" pageNumber="456" type="female">1♀</specimenCount>
<collectingCountry id="F32534CE1C7CC340CAB5A35339126C03" box="[720,838,1614,1638]" name="Nigeria" pageId="7" pageNumber="456">NIGERIA</collectingCountry>
,
<location id="8EED22851C7CC340CB34A353398D6C00" LSID="urn:lsid:plazi:treatment:039BC5481C7CC34DC8F2A1673E1F6E8A:8EED22851C7CC340CB34A353398D6C00" box="[849,985,1614,1638]" country="Nigeria" name="Omo Forest" pageId="7" pageNumber="456" stateProvince="Lagos">Omo Forest</location>
</materialsCitation>
,
<materialsCitation id="3B5A7E031C7CC340CB81A35339956CEF" ID-GBIF-Occurrence="1323867409" collectingDate="2002-05" collectionCode="ABRI, MZUJ, BMNH, UMZC" country="Nigeria" location="Ogun State" pageId="7" pageNumber="456" specimenCount="1" stateProvince="Ogun State" typeStatus="paratype">
<collectingRegion id="49F6BABC1C7CC340CB81A3533E356C03" box="[996,1121,1614,1638]" country="Nigeria" name="Ogun" pageId="7" pageNumber="456">Ogun State</collectingRegion>
<date id="FF8C529E1C7CC340CC0DA3533E976C03" box="[1128,1219,1614,1638]" pageId="7" pageNumber="456" value="2002-05">
<collectingDate id="EFC8AB761C7CC340CC0DA3533E976C03" box="[1128,1219,1614,1638]" pageId="7" pageNumber="456" value="2002-05">V. 2002</collectingDate>
</date>
. leg.: Robert David Warren. Deposited in the
<collectionCode id="ED23EC9B1C7CC340C9D0A36F3BAE6CEF" box="[437,506,1650,1674]" pageId="7" pageNumber="456">ABRI</collectionCode>
,
<collectionCode id="ED23EC9B1C7CC340CA63A36F38066CEC" box="[518,594,1650,1673]" pageId="7" pageNumber="456">MZUJ</collectionCode>
,
<collectionCode id="ED23EC9B1C7CC340CA38A36F38E56CEC" box="[605,689,1650,1673]" pageId="7" pageNumber="456">BMNH</collectionCode>
and
<collectionCode id="ED23EC9B1C7CC340CA82A36F396C6CEF" box="[743,824,1650,1674]" pageId="7" pageNumber="456">UMZC</collectionCode>
collections.
</materialsCitation>
</paragraph>
<paragraph id="8B8D745E1C7CC340C8A2A3883F346D7F" blockId="7.[151,1437,1253,2037]" pageId="7" pageNumber="456">
<materialsCitation id="3B5A7E031C7CC340C8A2A38838F16CCB" ID-GBIF-Occurrence="1323867414" box="[199,677,1685,1710]" country="Nigeria" location="NIGERIA" pageId="7" pageNumber="456" specimenCount="1" specimenCount-male="1">
<emphasis id="B946A84C1C7CC340C8A2A3883BAA6CCB" bold="true" box="[199,510,1685,1710]" pageId="7" pageNumber="456">Other material examined:</emphasis>
<specimenCount id="9D34BFD71C7CC340CA60A38B387C6CCB" box="[517,552,1686,1710]" pageId="7" pageNumber="456" type="male">1♂</specimenCount>
<collectingCountry id="F32534CE1C7CC340CA4AA38B38F16CCB" box="[559,677,1686,1710]" name="Nigeria" pageId="7" pageNumber="456">NIGERIA</collectingCountry>
</materialsCitation>
,
<materialsCitation id="3B5A7E031C7CC340CAD5A38B3F426CCB" ID-GBIF-Occurrence="1323867405" box="[688,1302,1685,1710]" country="Nigeria" location="Eastern" pageId="7" pageNumber="456" specimenCount="1" stateProvince="Eastern">
<collectingRegion id="49F6BABC1C7CC340CAD5A38B39506CC8" box="[688,772,1686,1709]" country="Zambia" name="Eastern" pageId="7" pageNumber="456">Eastern</collectingRegion>
<collectingCountry id="F32534CE1C7CC340CB6EA388390B6CCB" box="[779,863,1685,1710]" name="Nigeria" pageId="7" pageNumber="456">Nigeria</collectingCountry>
Leg.: Frank Davey, could be Uzuakoli
</materialsCitation>
,
<materialsCitation id="3B5A7E031C7CC340CD44A3883E5E6CB7" ID-GBIF-Occurrence="1323867412" collectionCode="ABRI" country="Nigeria" location="Delta" pageId="7" pageNumber="456" specimenCount="1" specimenCount-female="1" stateProvince="Delta">
<collectingRegion id="49F6BABC1C7CC340CD44A3883F0A6CCB" box="[1313,1374,1685,1710]" country="Nigeria" name="Delta" pageId="7" pageNumber="456">Delta</collectingRegion>
State (Libert pers. comm.),
<specimenCount id="9D34BFD71C7CC340C9E8A3A73BE46CB7" box="[397,432,1722,1746]" pageId="7" pageNumber="456" type="female">1♀</specimenCount>
<collectingCountry id="F32534CE1C7CC340C9DDA3A7387F6CB7" box="[440,555,1722,1746]" name="Nigeria" pageId="7" pageNumber="456">NIGERIA</collectingCountry>
(sine locus et anno) both
<collectionCode id="ED23EC9B1C7CC340CB2BA3A739C56CB7" box="[846,913,1722,1746]" pageId="7" pageNumber="456">ABRI</collectionCode>
collection
</materialsCitation>
. 3♂♂,
<materialsCitation id="3B5A7E031C7CC340CC3EA3A73F3A6C93" ID-GBIF-Occurrence="1323867399" collectionCode="NNM" country="Nigeria" location="Cornes-Riley" pageId="7" pageNumber="456" specimenCount="1" specimenCount-female="1" stateProvince="Lagos">
<specimenCount id="9D34BFD71C7CC340CC3EA3A73E2A6CB7" box="[1115,1150,1722,1746]" pageId="7" pageNumber="456" type="female">1♀</specimenCount>
<collectingCountry id="F32534CE1C7CC340CCE0A3A73EAF6CB7" box="[1157,1275,1722,1746]" name="Nigeria" pageId="7" pageNumber="456">NIGERIA</collectingCountry>
,
<location id="8EED22851C7CC340CD62A3A43FC86CB7" LSID="urn:lsid:plazi:treatment:039BC5481C7CC34DC8F2A1673E1F6E8A:8EED22851C7CC340CD62A3A43FC86CB7" box="[1287,1436,1721,1746]" country="Nigeria" name="Cornes-Riley" pageId="7" pageNumber="456" stateProvince="Lagos">Cornes-Riley</location>
collection collected possibly in the vicinity of
<collectingRegion id="49F6BABC1C7CC340CAFCA3C338896C93" box="[665,733,1758,1782]" country="Nigeria" name="Lagos" pageId="7" pageNumber="456">Lagos</collectingRegion>
(deposited in
<collectionCode id="ED23EC9B1C7CC340CB1FA3C039E96C93" box="[890,957,1757,1782]" country="Netherlands" httpUri="http://grbio.org/cool/wkmc-b7hm" name="National Museum of Natural History, Naturalis" pageId="7" pageNumber="456">NNM</collectionCode>
)(from photographs by Robert Warren)
</materialsCitation>
,
<materialsCitation id="3B5A7E031C7CC340CD1CA3C33B0B6D7F" ID-GBIF-Occurrence="1323867417" country="Nigeria" location="Lagos" pageId="7" pageNumber="456" specimenCount="1" specimenCount-male="1" stateProvince="Lagos">
<specimenCount id="9D34BFD71C7CC340CD1CA3C33FC86C93" box="[1401,1436,1758,1782]" pageId="7" pageNumber="456" type="male">1♂</specimenCount>
<collectingCountry id="F32534CE1C7CC340C8F2A21F3B596D7F" box="[151,269,1794,1818]" name="Nigeria" pageId="7" pageNumber="456">NIGERIA</collectingCountry>
,
<collectingRegion id="49F6BABC1C7CC340C97DA21F3B0B6D7F" box="[280,351,1794,1818]" country="Nigeria" name="Lagos" pageId="7" pageNumber="456">Lagos</collectingRegion>
</materialsCitation>
,
<materialsCitation id="3B5A7E031C7CC340C90FA21F3F346D7F" ID-GBIF-Occurrence="1323867418" box="[362,1376,1793,1818]" collectionCode="BMNH" country="Nigeria" location="Olokemeji" pageId="7" pageNumber="456" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D34BFD71C7CC340C90FA21F3BD96D7F" box="[362,397,1794,1818]" pageId="7" pageNumber="456" type="female">1♀</specimenCount>
<collectingCountry id="F32534CE1C7CC340C9F1A21F385E6D7F" box="[404,522,1794,1818]" name="Nigeria" pageId="7" pageNumber="456">NIGERIA</collectingCountry>
,
<location id="8EED22851C7CC340CA70A21C38DF6D7F" LSID="urn:lsid:plazi:treatment:039BC5481C7CC34DC8F2A1673E1F6E8A:8EED22851C7CC340CA70A21C38DF6D7F" box="[533,651,1793,1818]" country="Nigeria" name="Olokemeji" pageId="7" pageNumber="456">Olokemeji</location>
(deposited in the
<collectionCode id="ED23EC9B1C7CC340CB31A21C39FD6D7F" box="[852,937,1793,1818]" pageId="7" pageNumber="456">BMNH</collectionCode>
)(from photographs by Torben Larsen).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32827D51C7CC34FC8A2A238394B6D1F" lastPageId="8" lastPageNumber="457" pageId="7" pageNumber="456" type="description">
<paragraph id="8B8D745E1C7CC34FC8A2A23838A36BCA" blockId="7.[151,1437,1253,2037]" lastBlockId="8.[151,1437,151,791]" lastPageId="8" lastPageNumber="457" pageId="7" pageNumber="456">
<emphasis id="B946A84C1C7CC340C8A2A23838406D5B" bold="true" box="[199,532,1829,1854]" pageId="7" pageNumber="456">
Description of the
<typeStatus id="5489CAFC1C7CC340C9C3A238385B6D5B" box="[422,527,1829,1854]" pageId="7" pageNumber="456" type="holotype">holotype</typeStatus>
.
</emphasis>
Forewing length:
<quantity id="4CCAD9BB1C7CC340CA83A23B396D6D58" box="[742,825,1830,1854]" metricMagnitude="-2" metricUnit="m" metricValue="3.7" pageId="7" pageNumber="456" unit="mm" value="37.0">37 mm</quantity>
. Wingspan:
<quantity id="4CCAD9BB1C7CC340CBA0A2383E7A6D58" box="[965,1070,1829,1854]" metricMagnitude="-2" metricUnit="m" metricValue="6.55" pageId="7" pageNumber="456" unit="mm" value="65.5">65.5 mm</quantity>
. The upperside ground colour is dark chestnut-red, but the apical half of the wing is black (covering just over half of the wing, as a triangle from the apex to the discoidal cell along the costa and the tornus. There are two rows of bluish-white spots along the outer margin in the black area from space 1b to the apex. Two well defined white rectangular spots are found in spaces 5 and 6 (respectively). The forewing apical black area has an intense iridescent blue reflection, best visible in direct sun or in strong artificial illumination. The hindwing is dominated by the chestnut-red ground colour, which is slightly darker in the marginal area, with a dark undulating sub-marginal line. The underside is light hazel-brown with the shades of the white spots in spaces 5 and 6 on the upperside showing through, and two blackish spots in spaces 4 and 5, as well as a small ill-defined black dot in space 8 near the apex. The usual “
<taxonomicName id="4C320FDD1C73C34FCCEDA5C23EBB6A9D" box="[1160,1263,223,248]" class="Insecta" family="Nymphalidae" genus="Bebearia" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="457" phylum="Arthropoda" rank="genus">
<emphasis id="B946A84C1C73C34FCCEDA5C23EBB6A9D" box="[1160,1263,223,248]" italics="true" pageId="8" pageNumber="457">Bebearia</emphasis>
</taxonomicName>
” pattern is also visible as a figure-of-eight spot and other amoeboid patches in the discoidal cell and irregular lines in the median and post-median area of both wings, as well as a tiny black ring in the hindwing cell. The body is strong and bold, black on the underside with a whitish band on the abdomen and a couple of whitish lateral spots. The body colour is light hazel-brown on the upperside, the same tone as the wings. The antennae are long (
<quantity id="4CCAD9BB1C73C34FCCC0A46E3EAE6BEE" box="[1189,1274,371,395]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" pageId="8" pageNumber="457" unit="mm" value="21.0">21 mm</quantity>
), with a black shaft and three-fourth of the club is light hazel-brown.
</paragraph>
<paragraph id="8B8D745E1C73C34FC8A2A4A73BF06972" blockId="8.[151,1437,151,791]" pageId="8" pageNumber="457">
<emphasis id="B946A84C1C73C34FC8A2A4A73B2F6BB6" bold="true" box="[199,379,442,467]" pageId="8" pageNumber="457">Male genitalia.</emphasis>
As normal for
<taxonomicName id="4C320FDD1C73C34FCA57A4A738CD6BB6" box="[562,665,442,467]" class="Insecta" family="Nymphalidae" genus="Bebearia" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="457" phylum="Arthropoda" rank="genus">
<emphasis id="B946A84C1C73C34FCA57A4A738CD6BB6" box="[562,665,442,467]" italics="true" pageId="8" pageNumber="457">Bebearia</emphasis>
</taxonomicName>
, the genitalia are rather large (
<quantity id="4CCAD9BB1C73C34FCC67A4A63E086BB7" box="[1026,1116,443,467]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="8" pageNumber="457" unit="mm" value="2.1">2.1 mm</quantity>
dorsoventrally and
<quantity id="4CCAD9BB1C73C34FCD26A4A63FC86BB7" box="[1347,1436,443,467]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="8" pageNumber="457" unit="mm" value="2.3">2.3 mm</quantity>
laterally) and strongly sclerotised. The tegumen is rather broad and T-shaped on the lateral view, curving evenly below to the anterior tip. The uncus is longer than the tegumen and gently curving downward, strongly resembling a renaissance carnival mask with a long, bird-bill shaped structure. The gnathos is twin-lobed, the lobes are slender, gently curved and almost as long as the uncus, unusually turning down parallel to the stem of the tegumen. The valvae are rather long, fan-shaped, and are without projections. They are slightly bent upwards and their tips are evenly rounded. The valvae are only sparsely covered with short setae. The saccus is small and gently curved upwards with a narrow and rounded tip. The aedeagus is rather short and broad, narrower at the anterior end as the posterior tip is broad and rectangular.
<collectingRegion id="49F6BABC1C73C34FCA27A7C638026896" box="[578,598,731,755]" country="Nigeria" name="Rivers" pageId="8" pageNumber="457">A</collectingRegion>
downward curving cornutus is visible in the vesica (not everted). It is half as long as the aedeagus.
</paragraph>
<caption id="DF4D24D61C73C34FC8F2A0803BEA6F94" httpUri="https://zenodo.org/record/263144/files/figure.png" pageId="8" pageNumber="457" targetBox="[177,1388,862,1400]" targetPageId="8">
<paragraph id="8B8D745E1C73C34FC8F2A0803BEA6F94" blockId="8.[151,1437,1437,1521]" pageId="8" pageNumber="457">
<emphasis id="B946A84C1C73C34FC8F2A0803B416FD6" bold="true" box="[151,277,1437,1459]" pageId="8" pageNumber="457">FIGURE 7.</emphasis>
Male genitalia with removed aedeagi in the
<taxonomicName id="4C320FDD1C73C34FCA8FA08239316FD6" box="[746,869,1437,1460]" class="Insecta" family="Nymphalidae" genus="Bebearia" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="457" phylum="Arthropoda" rank="species" species="plistonax">
<emphasis id="B946A84C1C73C34FCA8FA08239316FD6" box="[746,869,1437,1460]" italics="true" pageId="8" pageNumber="457">B. plistonax</emphasis>
</taxonomicName>
group: A—
<taxonomicName id="4C320FDD1C73C34FCB86A0803EE46FD6" box="[995,1200,1437,1459]" class="Insecta" family="Nymphalidae" genus="Bebearia" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="457" phylum="Arthropoda" rank="species" species="wojtusiaki" status="sp. nov.">
<emphasis id="B946A84C1C73C34FCB86A0803EE46FD6" box="[995,1200,1437,1459]" italics="true" pageId="8" pageNumber="457">Bebearia wojtusiaki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27515371C73C34FCCDCA0833EA66FD6" box="[1209,1266,1438,1459]" pageId="8" pageNumber="457" rank="species">sp. n.</taxonomicNameLabel>
(paratype), gen. prep.: MZUJ-JL-155; B—
<taxonomicName id="4C320FDD1C73C34FC9C6A0A0382F6FB7" authority="(Liberia)" baseAuthorityName="Liberia" box="[419,635,1468,1490]" class="Insecta" family="Nymphalidae" genus="Bebearia" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="457" phylum="Arthropoda" rank="species" species="arcadius">
<emphasis id="B946A84C1C73C34FC9C6A0A0384C6FB7" box="[419,536,1468,1490]" italics="true" pageId="8" pageNumber="457">B. arcadius</emphasis>
(Liberia)
</taxonomicName>
, gen. prep.: MZUJ-JL-153 and C—
<taxonomicName id="4C320FDD1C73C34FCB8DA0A03E366FB7" box="[1000,1122,1468,1490]" class="Insecta" family="Nymphalidae" genus="Bebearia" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="457" phylum="Arthropoda" rank="species" species="plistonax">
<emphasis id="B946A84C1C73C34FCB8DA0A03E366FB7" box="[1000,1122,1468,1490]" italics="true" pageId="8" pageNumber="457">B. plistonax</emphasis>
</taxonomicName>
(Bitje, Cameroon), gen. prep.: MZUJ-JL-152 (lateral view).
</paragraph>
</caption>
<paragraph id="8B8D745E1C73C34FC8A2A3003B3E6D6B" blockId="8.[151,1437,1565,2022]" pageId="8" pageNumber="457">
<emphasis id="B946A84C1C73C34FC8A2A3003B886C53" bold="true" box="[199,476,1565,1590]" pageId="8" pageNumber="457">Description of female.</emphasis>
Forewing length:
<quantity id="4CCAD9BB1C73C34FCAA5A303394E6C53" box="[704,794,1566,1590]" metricMagnitude="-2" metricUnit="m" metricValue="4.9" pageId="8" pageNumber="457" unit="mm" value="49.0">49 mm</quantity>
. Wingspan:
<quantity id="4CCAD9BB1C73C34FCBD6A3033E586C53" box="[947,1036,1566,1590]" metricMagnitude="-2" metricUnit="m" metricValue="8.4" pageId="8" pageNumber="457" unit="mm" value="84.0">84 mm</quantity>
. Antennal length:
<quantity id="4CCAD9BB1C73C34FCC95A3003F0B6C53" box="[1264,1375,1565,1590]" metricMagnitude="-2" metricUnit="m" metricValue="2.35" pageId="8" pageNumber="457" unit="mm" value="23.5">23.5 mm</quantity>
. The appearance of the female is very similar to that of the male, although the size difference between the sexes is substantial. The white spotting in the outer half of the forewing is conspicuous and the spots form a narrow, but obvious white band in the spaces between veins 4 and 7. The outer margin of the forewing is concave, making the forewing apex rather acute. The hindwing is evenly rounded from the apex to the tornus. There are two, rather illdefined whitish spots in the hindwing apex, both missing in the male. The underside colour and pattern is identical to that of the male.
</paragraph>
<paragraph id="8B8D745E1C73C34FC8A2A204394B6D1F" blockId="8.[151,1437,1565,2022]" pageId="8" pageNumber="457">
<emphasis id="B946A84C1C73C34FC8A2A2043BC16D57" bold="true" box="[199,405,1817,1842]" pageId="8" pageNumber="457">Female genitalia.</emphasis>
The papillae anales are prominent and are covered with rather dense hair. The apophyses posteriores are long and think. The von Siebolds organ is medium sized, weakly sclerotized. The ductus bursae is very narrow, and straight, the bursa copulatrix is rounded.
</paragraph>
</subSubSection>
<subSubSection id="C32827D51C73C34EC8A2A2983E366895" lastPageId="9" lastPageNumber="458" pageId="8" pageNumber="457" type="diagnosis">
<paragraph id="8B8D745E1C73C34EC8A2A2983E366895" blockId="8.[151,1437,1565,2022]" lastBlockId="9.[151,1437,151,1184]" lastPageId="9" lastPageNumber="458" pageId="8" pageNumber="457">
<emphasis id="B946A84C1C73C34FC8A2A2983B156DFB" bold="true" box="[199,321,1925,1950]" pageId="8" pageNumber="457">Diagnosis.</emphasis>
<taxonomicName id="4C320FDD1C73C34FC92BA29A3BB36DFB" box="[334,487,1925,1950]" class="Insecta" family="Nymphalidae" genus="Bebearia" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="457" phylum="Arthropoda" rank="species" species="wojtusiaki">
<emphasis id="B946A84C1C73C34FC92BA29A3BB36DFB" box="[334,487,1925,1950]" italics="true" pageId="8" pageNumber="457">B. wojtusiaki</emphasis>
</taxonomicName>
, at first impression, resembles a slightly darker and well spotted
<taxonomicName id="4C320FDD1C73C34FCD6DA29A3FC16DFB" box="[1288,1429,1925,1950]" class="Insecta" family="Nymphalidae" genus="Bebearia" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="457" phylum="Arthropoda" rank="species" species="plistonax">
<emphasis id="B946A84C1C73C34FCD6DA29A3FC16DFB" box="[1288,1429,1925,1950]" italics="true" pageId="8" pageNumber="457">B. plistonax</emphasis>
</taxonomicName>
. However, it is instead a morphologically intermediate species between
<taxonomicName id="4C320FDD1C73C34FCBB5A2B63E0D6DA7" box="[976,1113,1961,1986]" class="Insecta" family="Nymphalidae" genus="Bebearia" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="457" phylum="Arthropoda" rank="species" species="plistonax">
<emphasis id="B946A84C1C73C34FCBB5A2B63E0D6DA7" box="[976,1113,1961,1986]" italics="true" pageId="8" pageNumber="457">B. plistonax</emphasis>
</taxonomicName>
and
<taxonomicName id="4C320FDD1C73C34FCCF3A2B63F4D6DA7" box="[1174,1305,1961,1986]" class="Insecta" family="Nymphalidae" genus="Bebearia" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="457" phylum="Arthropoda" rank="species" species="arcadius">
<emphasis id="B946A84C1C73C34FCCF3A2B63F4D6DA7" box="[1174,1305,1961,1986]" italics="true" pageId="8" pageNumber="457">B. arcadius</emphasis>
</taxonomicName>
, as the red colour on the upperside is always slightly darker than on
<taxonomicName id="4C320FDD1C73C34FCB7EA2D239F56D83" box="[795,929,1997,2022]" class="Insecta" family="Nymphalidae" genus="Bebearia" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="457" phylum="Arthropoda" rank="species" species="plistonax">
<emphasis id="B946A84C1C73C34FCB7EA2D239F56D83" box="[795,929,1997,2022]" italics="true" pageId="8" pageNumber="457">B. plistonax</emphasis>
</taxonomicName>
and similarly dark to, or only slightly lighter than
<taxonomicName id="4C320FDD1C72C34EC8B1A5843B0D6AD5" box="[212,345,151,176]" class="Insecta" family="Nymphalidae" genus="Bebearia" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="458" phylum="Arthropoda" rank="species" species="arcadius">
<emphasis id="B946A84C1C72C34EC8B1A5843B0D6AD5" box="[212,345,151,176]" italics="true" pageId="9" pageNumber="458">B. arcadius</emphasis>
</taxonomicName>
. The bluish-white spotting on the forewing of
<taxonomicName id="4C320FDD1C72C34ECBF7A5843E7D6AD5" box="[914,1065,151,176]" class="Insecta" family="Nymphalidae" genus="Bebearia" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="458" phylum="Arthropoda" rank="species" species="wojtusiaki">
<emphasis id="B946A84C1C72C34ECBF7A5843E7D6AD5" box="[914,1065,151,176]" italics="true" pageId="9" pageNumber="458">B. wojtusiaki</emphasis>
</taxonomicName>
is always more bluish than
<taxonomicName id="4C320FDD1C72C34ECDE1A5843AAA6AB0" class="Insecta" family="Nymphalidae" genus="Bebearia" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="458" phylum="Arthropoda" rank="species" species="plistonax">
<emphasis id="B946A84C1C72C34ECDE1A5843AAA6AB0" italics="true" pageId="9" pageNumber="458">B. plistonax</emphasis>
</taxonomicName>
, as the blue iridescent sheen, similarly to
<emphasis id="B946A84C1C72C34ECA8AA5A0392D6AB0" box="[751,889,188,213]" italics="true" pageId="9" pageNumber="458">
<taxonomicName id="4C320FDD1C72C34ECA8AA5A039226AB0" box="[751,886,188,213]" class="Insecta" family="Nymphalidae" genus="Bebearia" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="458" phylum="Arthropoda" rank="species" species="arcadius">B. arcadius</taxonomicName>
,
</emphasis>
is more extensive on
<taxonomicName id="4C320FDD1C72C34ECCE4A5A03F426AB0" box="[1153,1302,188,213]" class="Insecta" family="Nymphalidae" genus="Bebearia" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="458" phylum="Arthropoda" rank="species" species="wojtusiaki">
<emphasis id="B946A84C1C72C34ECCE4A5A03F426AB0" box="[1153,1302,188,213]" italics="true" pageId="9" pageNumber="458">B. wojtusiaki</emphasis>
</taxonomicName>
, and is not restricted just to the close environment of the whitish spots, which is usually the case in
<taxonomicName id="4C320FDD1C72C34ECCE9A5FC3F476A9D" box="[1164,1299,223,248]" class="Insecta" family="Nymphalidae" genus="Bebearia" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="458" phylum="Arthropoda" rank="species" species="plistonax">
<emphasis id="B946A84C1C72C34ECCE9A5FC3F476A9D" box="[1164,1299,223,248]" italics="true" pageId="9" pageNumber="458">B. plistonax</emphasis>
</taxonomicName>
(apart from certain specimens from
<collectingCountry id="F32534CE1C72C34EC9CDA41938576B78" box="[424,515,260,285]" name="Zambia" pageId="9" pageNumber="458">Zambia</collectingCountry>
). The bluish-white spotting is also more extensive in
<taxonomicName id="4C320FDD1C72C34ECC00A4183EAD6B78" box="[1125,1273,260,285]" class="Insecta" family="Nymphalidae" genus="Bebearia" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="458" phylum="Arthropoda" rank="species" species="wojtusiaki">
<emphasis id="B946A84C1C72C34ECC00A4183EAD6B78" box="[1125,1273,260,285]" italics="true" pageId="9" pageNumber="458">B. wojtusiaki</emphasis>
</taxonomicName>
which always has two spots in the space between veins 1 and 2, three between 3 and 4, also three between 4 and 5, the inner ones being rather ill-defined. In females there is always a bluish-white, rather undefined spot at the end of the discoidal cell, visible as bluish-white shading on the males.
<taxonomicName id="4C320FDD1C72C34ECABFA46C39366BED" box="[730,866,367,392]" class="Insecta" family="Nymphalidae" genus="Bebearia" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="458" phylum="Arthropoda" rank="species" species="plistonax">
<emphasis id="B946A84C1C72C34ECABFA46C39366BED" box="[730,866,367,392]" italics="true" pageId="9" pageNumber="458">B. plistonax</emphasis>
</taxonomicName>
in most cases have a single whitish spot between veins 1 and 2, a very ill-defined bluish-white spot can occur in space between veins 1 and 2 but it was found only on females from
<collectingRegion id="49F6BABC1C72C34EC905A4AA3BE76BB5" box="[352,435,439,464]" country="Zambia" name="Central" pageId="9" pageNumber="458">Central</collectingRegion>
Africa,
<collectingCountry id="F32534CE1C72C34ECA73A4AA38396BB5" box="[534,621,439,464]" name="Uganda" pageId="9" pageNumber="458">Uganda</collectingCountry>
and
<collectingCountry id="F32534CE1C72C34ECACEA4AA395B6BB5" box="[683,783,439,464]" name="Tanzania" pageId="9" pageNumber="458">Tanzania</collectingCountry>
populations, and only a single male was found with an additional ill-defined bluish-white spot in the spaces between veins 1 and
<quantity id="4CCAD9BB1C72C34ECBB0A4C13E546B90" box="[981,1024,476,501]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="9" pageNumber="458" unit="in" value="2.0">2 in</quantity>
the series of examined specimens in the ABRI collection from
<collectingRegion id="49F6BABC1C72C34EC9DEA4E23874687D" box="[443,544,511,536]" country="Zambia" name="Northern" pageId="9" pageNumber="458">Northern</collectingRegion>
<collectingCountry id="F32534CE1C72C34ECA42A4E23828687D" box="[551,636,511,536]" name="Zambia" pageId="9" pageNumber="458">Zambia</collectingCountry>
(among
<specimenCount id="9D34BFD71C72C34ECABAA71D391C687D" box="[735,840,511,536]" pageId="9" pageNumber="458" type="male">17 males</specimenCount>
). The structure of the male genitalia is very similar in all three species, and dissection of multiple specimens from different localities revealed high individual variation and overlap of multiple features between them. We therefore consider male genitalia as lacking diagnostic value in discerning the three species. However, we found morphologically small, but significant differences in the structure of female genitalia with regards to the shape of ductus bursae, which is straight and considerably thinner in
<taxonomicName id="4C320FDD1C72C34ECDE1A78C3B5E68A8" class="Insecta" family="Nymphalidae" genus="Bebearia" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="458" phylum="Arthropoda" rank="species" species="wojtusiaki">
<emphasis id="B946A84C1C72C34ECDE1A78C3B5E68A8" italics="true" pageId="9" pageNumber="458">B. wojtusiaki</emphasis>
</taxonomicName>
, not gently bent before opening into corpus bursae as in
<taxonomicName id="4C320FDD1C72C34ECBA7A7A83E1968A8" box="[962,1101,692,717]" class="Insecta" family="Nymphalidae" genus="Bebearia" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="458" phylum="Arthropoda" rank="species" species="plistonax">
<emphasis id="B946A84C1C72C34ECBA7A7A83E1968A8" box="[962,1101,692,717]" italics="true" pageId="9" pageNumber="458">B. plistonax</emphasis>
</taxonomicName>
and
<taxonomicName id="4C320FDD1C72C34ECCEBA7A83F4068A8" box="[1166,1300,692,717]" class="Insecta" family="Nymphalidae" genus="Bebearia" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="458" phylum="Arthropoda" rank="species" species="arcadius">
<emphasis id="B946A84C1C72C34ECCEBA7A83F4068A8" box="[1166,1300,692,717]" italics="true" pageId="9" pageNumber="458">B. arcadius</emphasis>
</taxonomicName>
. The bursa copulatrix is sphere-shaped, while it is more oval or pear-shaped in the related species.
</paragraph>
</subSubSection>
<subSubSection id="C32827D51C72C34EC8A2A7E13EFA6EC5" pageId="9" pageNumber="458" type="discussion">
<paragraph id="8B8D745E1C72C34EC8A2A7E13EFA6EC5" blockId="9.[151,1437,151,1184]" pageId="9" pageNumber="458">
<emphasis id="B946A84C1C72C34EC8A2A7E13B186970" bold="true" box="[199,332,764,789]" pageId="9" pageNumber="458">Discussion.</emphasis>
The differences between the eastern Nigerian
<taxonomicName id="4C320FDD1C72C34ECB03A7E039BB6970" box="[870,1007,764,789]" class="Insecta" family="Nymphalidae" genus="Bebearia" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="458" phylum="Arthropoda" rank="species" species="plistonax">
<emphasis id="B946A84C1C72C34ECB03A7E039BB6970" box="[870,1007,764,789]" italics="true" pageId="9" pageNumber="458">B. plistonax</emphasis>
</taxonomicName>
and
<taxonomicName id="4C320FDD1C72C34ECC4FA7E03EEB6970" box="[1066,1215,764,789]" class="Insecta" family="Nymphalidae" genus="Bebearia" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="458" phylum="Arthropoda" rank="species" species="wojtusiaki">
<emphasis id="B946A84C1C72C34ECC4FA7E03EEB6970" box="[1066,1215,764,789]" italics="true" pageId="9" pageNumber="458">B. wojtusiaki</emphasis>
</taxonomicName>
in western
<collectingCountry id="F32534CE1C72C34ECD2DA7E13FC86970" box="[1352,1436,764,789]" name="Nigeria" pageId="9" pageNumber="458">Nigeria</collectingCountry>
were initially recognised over thirty years ago by the late Prof. Janusz Wojtusiak, who collected extensively in
<collectingCountry id="F32534CE1C72C34EC8F2A6593ABF6938" box="[151,235,836,861]" name="Nigeria" pageId="9" pageNumber="458">Nigeria</collectingCountry>
between
<date id="FF8C529E1C72C34EC933A65938546938" box="[342,512,836,861]" pageId="9" pageNumber="458" value="1982" valueMax="1986">1982 and 1986</date>
. However, he hesitated to describe the taxon from the limited material available at that time, maybe believing that the differences between the two populations would not prove sufficient enough to justify the separation of the western Nigerian populations from typical
<taxonomicName id="4C320FDD1C72C34ECB81A6903E3B69C0" box="[996,1135,908,933]" class="Insecta" family="Nymphalidae" genus="Bebearia" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="458" phylum="Arthropoda" rank="species" species="plistonax">
<emphasis id="B946A84C1C72C34ECB81A6903E3B69C0" box="[996,1135,908,933]" italics="true" pageId="9" pageNumber="458">B. plistonax</emphasis>
</taxonomicName>
. In the last two decades, knowledge about the fauna and biogeography of
<collectingRegion id="49F6BABC1C72C34ECAA6A6AD38AE69A2" box="[707,762,944,967]" country="Tanzania" name="Mjini Magharibi" pageId="9" pageNumber="458">West</collectingRegion>
African butterflies has improved significantly, and western
<collectingCountry id="F32534CE1C72C34EC8F2A6C93ABF6988" box="[151,235,980,1005]" name="Nigeria" pageId="9" pageNumber="458">Nigeria</collectingCountry>
is now beginning to be recognised as a distinct biogeographic sub-region, harbouring significant level of endemism and playing a role of a linkage between the more
<collectingRegion id="49F6BABC1C72C34ECB5DA6EA39DF6E75" box="[824,907,1015,1040]" country="Zambia" name="Central" pageId="9" pageNumber="458">Central</collectingRegion>
Africa-related
<collectingRegion id="49F6BABC1C72C34ECC50A6E53EEE6E75" box="[1077,1210,1015,1040]" country="Nigeria" name="Cross River" pageId="9" pageNumber="458">Cross River</collectingRegion>
loop and the forests west of the
<collectingCountry id="F32534CE1C72C34EC979A1013BDD6E50" box="[284,393,1052,1077]" name="Benin" pageId="9" pageNumber="458">Dahomey</collectingCountry>
Gap. We found that the morphological differences between
<taxonomicName id="4C320FDD1C72C34ECC51A1003EEF6E50" box="[1076,1211,1052,1077]" class="Insecta" family="Nymphalidae" genus="Bebearia" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="458" phylum="Arthropoda" rank="species" species="plistonax">
<emphasis id="B946A84C1C72C34ECC51A1003EEF6E50" box="[1076,1211,1052,1077]" italics="true" pageId="9" pageNumber="458">B. plistonax</emphasis>
</taxonomicName>
,
<taxonomicName id="4C320FDD1C72C34ECCAFA1003F1F6E50" box="[1226,1355,1052,1077]" class="Insecta" family="Nymphalidae" genus="Bebearia" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="458" phylum="Arthropoda" rank="species" species="arcadius">
<emphasis id="B946A84C1C72C34ECCAFA1003F1F6E50" box="[1226,1355,1052,1077]" italics="true" pageId="9" pageNumber="458">B. arcadius</emphasis>
</taxonomicName>
and
<taxonomicName id="4C320FDD1C72C34ECDE1A1003B5F6E3D" class="Insecta" family="Nymphalidae" genus="Bebearia" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="458" phylum="Arthropoda" rank="species" species="wojtusiaki">
<emphasis id="B946A84C1C72C34ECDE1A1003B5F6E3D" italics="true" pageId="9" pageNumber="458">B. wojtusiaki</emphasis>
</taxonomicName>
are stable on a geographical scale and there is no evidence for clinal patterns. Furthermore, morphological characters of
<taxonomicName id="4C320FDD1C72C34EC9B8A17838256E18" box="[477,625,1124,1149]" class="Insecta" family="Nymphalidae" genus="Bebearia" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="458" phylum="Arthropoda" rank="species" species="wojtusiaki">
<emphasis id="B946A84C1C72C34EC9B8A17838256E18" box="[477,625,1124,1149]" italics="true" pageId="9" pageNumber="458">B. wojtusiaki</emphasis>
</taxonomicName>
do not justify its placement as a subspecies of either. Thus, despite no apparent geographical overlap we consider that the three should be treated as distinct species.
</paragraph>
</subSubSection>
<caption id="DF4D24D61C72C34EC8F2A2A53EF76D88" httpUri="https://zenodo.org/record/263145/files/figure.png" pageId="9" pageNumber="458" targetBox="[171,1422,1241,1942]" targetPageId="9">
<paragraph id="8B8D745E1C72C34EC8F2A2A53EF76D88" blockId="9.[151,1436,1976,2029]" pageId="9" pageNumber="458">
<emphasis id="B946A84C1C72C34EC8F2A2A53B416DA8" bold="true" box="[151,277,1976,1998]" pageId="9" pageNumber="458">FIGURE 8.</emphasis>
<taxonomicName id="4C320FDD1C72C34EC97AA2A73BF26DAB" box="[287,422,1976,1999]" class="Insecta" family="Nymphalidae" genus="Bebearia" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="458" phylum="Arthropoda" rank="species" species="wojtusiaki">
<emphasis id="B946A84C1C72C34EC97AA2A73BF26DAB" box="[287,422,1976,1999]" italics="true" pageId="9" pageNumber="458">B. wojtusiaki</emphasis>
</taxonomicName>
female (paratype) A—upperside, D—underside;
<taxonomicName id="4C320FDD1C72C34ECBCFA2A73E756DAB" box="[938,1057,1976,1999]" class="Insecta" family="Nymphalidae" genus="Bebearia" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="458" phylum="Arthropoda" rank="species" species="arcadius">
<emphasis id="B946A84C1C72C34ECBCFA2A73E756DAB" box="[938,1057,1976,1999]" italics="true" pageId="9" pageNumber="458">B. arcadius</emphasis>
</taxonomicName>
female (Nimba Mountains, Liberia) B—upperside, E—underside;
<taxonomicName id="4C320FDD1C72C34EC9ACA2C538166D88" box="[457,578,2007,2029]" class="Insecta" family="Nymphalidae" genus="Bebearia" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="458" phylum="Arthropoda" rank="species" species="plistonax">
<emphasis id="B946A84C1C72C34EC9ACA2C538166D88" box="[457,578,2007,2029]" italics="true" pageId="9" pageNumber="458">B. plistonax</emphasis>
</taxonomicName>
female (Ngel Nyaki, Nigeria) C—upperside, F—underside.
</paragraph>
</caption>
<caption id="DF4D24D61C71C34DC8F2A11339136E07" httpUri="https://zenodo.org/record/263146/files/figure.png" pageId="10" pageNumber="459" targetBox="[163,1415,210,1006]" targetPageId="10">
<paragraph id="8B8D745E1C71C34DC8F2A11339136E07" blockId="10.[151,1436,1038,1122]" pageId="10" pageNumber="459">
<emphasis id="B946A84C1C71C34DC8F2A1133B406E46" bold="true" box="[151,276,1038,1060]" pageId="10" pageNumber="459">FIGURE 9.</emphasis>
Female genitalia (full armature and magnified papillae and antrum) in the
<taxonomicName id="4C320FDD1C71C34DCC78A1123ECC6E41" box="[1053,1176,1038,1060]" class="Insecta" family="Nymphalidae" genus="Bebearia" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="459" phylum="Arthropoda" rank="species" species="plistonax">
<emphasis id="B946A84C1C71C34DCC78A1123ECC6E41" box="[1053,1176,1038,1060]" italics="true" pageId="10" pageNumber="459">B. plistonax</emphasis>
</taxonomicName>
-group: A
<emphasis id="B946A84C1C71C34DCC98A1123F426E41" bold="true" box="[1277,1302,1039,1060]" pageId="10" pageNumber="459"></emphasis>
<taxonomicName id="4C320FDD1C71C34DCD73A1123FC86E41" box="[1302,1436,1038,1060]" class="Insecta" family="Nymphalidae" genus="Bebearia" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="459" phylum="Arthropoda" rank="species" species="wojtusiaki">
<emphasis id="B946A84C1C71C34DCD73A1123FC86E41" box="[1302,1436,1038,1060]" italics="true" pageId="10" pageNumber="459">B. wojtusiaki</emphasis>
</taxonomicName>
(paratype), gen. prep.: MZUJ-JL-01/09.05.2014; B
<emphasis id="B946A84C1C71C34DCAFBA13038E36E27" bold="true" box="[670,695,1069,1090]" pageId="10" pageNumber="459"></emphasis>
<taxonomicName id="4C320FDD1C71C34DCAD2A133397F6E26" box="[695,811,1069,1091]" class="Insecta" family="Nymphalidae" genus="Bebearia" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="459" phylum="Arthropoda" rank="species" species="arcadius">
<emphasis id="B946A84C1C71C34DCAD2A133397F6E26" box="[695,811,1069,1091]" italics="true" pageId="10" pageNumber="459">B. arcadius</emphasis>
</taxonomicName>
(Labé, Guinea), gen. prep.: MZUJ-JL-03/09.05.2014; C
<emphasis id="B946A84C1C71C34DCD08A1303FD26E27" bold="true" box="[1389,1414,1069,1090]" pageId="10" pageNumber="459"></emphasis>
<taxonomicName id="4C320FDD1C71C34DCDE3A1333AA06E07" class="Insecta" family="Nymphalidae" genus="Bebearia" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="459" phylum="Arthropoda" rank="species" species="plistonax">
<emphasis id="B946A84C1C71C34DCDE3A1333AA06E07" italics="true" pageId="10" pageNumber="459">B. plistonax</emphasis>
</taxonomicName>
(Ebogo, Cameroon), gen. prep.: MZUJ-JL-02/09.05.2014.
</paragraph>
</caption>
<subSubSection id="C32827D51C71C34DC8A2A1933E1F6E8A" pageId="10" pageNumber="459" type="etymology">
<paragraph id="8B8D745E1C71C34DC8A2A1933E1F6E8A" blockId="10.[151,1436,1166,1263]" pageId="10" pageNumber="459">
<emphasis id="B946A84C1C71C34DC8A2A1933B1A6EC2" bold="true" box="[199,334,1166,1191]" pageId="10" pageNumber="459">Etymology.</emphasis>
We gratefully pay tribute to the late Prof. Janusz Wojtusiak, who studied Nigerian butterflies between
<date id="FF8C529E1C71C34DC899A1AE3BF26EAE" box="[252,422,1202,1227]" pageId="10" pageNumber="459" value="1982" valueMax="1986">1982 and 1986</date>
, by naming this species in his honour. He was the first known person to recognize that the western Nigerian
<taxonomicName id="4C320FDD1C71C34DC902A1CA3BB96E8A" box="[359,493,1238,1263]" class="Insecta" family="Nymphalidae" genus="Bebearia" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="459" phylum="Arthropoda" rank="species" species="plistonax">
<emphasis id="B946A84C1C71C34DC902A1CA3BB96E8A" box="[359,493,1238,1263]" italics="true" pageId="10" pageNumber="459">B. plistonax</emphasis>
</taxonomicName>
differed from the ones found east of the
<collectingRegion id="49F6BABC1C71C34DCBA5A1CA3E136E8A" box="[960,1095,1238,1263]" country="Nigeria" name="Cross River" pageId="10" pageNumber="459">Cross River</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>